treatments-xml/data/03/DE/29/03DE2968FF85FF85FEDE45A06833F9EB.xml

<document id="49E99DAACD64315E480D53138B4073EF" ID-DOI="10.1515/aemnp-2017-0118" ID-ISSN="0374-1036" ID-Zenodo-Dep="5324218" ID-ZooBank="11CB4607-CB26-4A43-89BC-F9B5B72C6372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630148606604" checkinUser="marcus" docAuthor="Straka, Jakub, Batelka, Jan &amp; Pauly, Alain" docDate="2017" docId="03DE2968FF85FF85FEDE45A06833F9EB" docLanguage="en" docName="ActaEntMusNatPra.2017.57.suppl.183-219.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57" docSource="http://dx.doi.org/10.1515/aemnp-2017-0118" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Lasioglossum (Hemihalictus) boswelliae Pauly &amp; Straka 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="199" masterDocId="FFE75110FF8BFF95FFC846556C54FFB9" masterDocTitle="Bees of the Socotra Archipelago (Hymenoptera: Anthophila), their biogeography and association with parasites" masterLastPageNumber="219" masterPageNumber="183" pageNumber="197" updateTime="1699164397122" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3A0251BD53EB49BEE86DF905536CE13D">Bees of the Socotra Archipelago (Hymenoptera: Anthophila), their biogeography and association with parasites</mods:title>
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<mods:namePart id="69EEE314E12902093DEF57D2FDF6465D">Straka, Jakub</mods:namePart>
<mods:affiliation id="D55C2BF2D92FC0097373B8065E972DAD">) &amp;) &amp;) Department of Zoology, Charles University in Prague, Viničná 7, CZ- 12844 Praha 2, Czech Republic; e-mails: jakub. straka @ aculeataresearch. com; janbat @ centrum. cz</mods:affiliation>
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<mods:namePart id="DB8C89AFCAE669F986993BF64BE5DA6B">Batelka, Jan</mods:namePart>
<mods:affiliation id="B7D141BA97E201BF63A4598CCCA325AE">) &amp;) &amp;) Department of Zoology, Charles University in Prague, Viničná 7, CZ- 12844 Praha 2, Czech Republic; e-mails: jakub. straka @ aculeataresearch. com; janbat @ centrum. cz</mods:affiliation>
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<mods:namePart id="22DB9EDED7B807456AD5AC98EB379A4A">Pauly, Alain</mods:namePart>
<mods:affiliation id="B6528250EAEBF1A0BAE2EF8635FC335B">) &amp;) Royal Belgian Institute of Natural Sciences, Entomology, Rue Vautier 29, B- 1000 Brussels, Belgium; e-mail: alain. pauly @ brutele. be</mods:affiliation>
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<paragraph id="8BC8987EFF85FF9BFEDE45A06FBCFBB5" blockId="14.[278,1000,1013,1036]" box="[278,1000,1013,1036]" pageId="14" pageNumber="197">
<heading id="D0802F12FF85FF9BFEDE45A06FBCFBB5" bold="true" box="[278,1000,1013,1036]" centered="true" fontSize="8" level="2" pageId="14" pageNumber="197" reason="3">
<emphasis id="B903446CFF85FF9BFEDE45A06FBCFBB5" bold="true" box="[278,1000,1013,1036]" pageId="14" pageNumber="197">
<taxonomicName id="4C77E3FDFF85FF9BFEDE45A06FDFFBB5" authority="Pauly &amp; Straka" authorityName="Pauly &amp; Straka" authorityYear="2017" box="[278,907,1013,1036]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="197" phylum="Arthropoda" rank="species" species="boswelliae" status="sp. nov." subGenus="Hemihalictus">
<emphasis id="B903446CFF85FF9BFEDE45A06DF9FBB5" bold="true" box="[278,429,1013,1036]" italics="true" pageId="14" pageNumber="197">Lasioglossum</emphasis>
(
<emphasis id="B903446CFF85FF9BFE7545A06E05FBB5" bold="true" box="[445,593,1013,1036]" italics="true" pageId="14" pageNumber="197">Hemihalictus</emphasis>
) 
<emphasis id="B903446CFF85FF9BFDA945A06E87FBB5" bold="true" box="[609,723,1013,1036]" italics="true" pageId="14" pageNumber="197">boswelliae</emphasis>
Pauly &amp; Straka
</taxonomicName>
<taxonomicNameLabel id="A230F917FF85FF9BFC5A45A06FBCFBB5" box="[914,1000,1013,1036]" pageId="14" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36DCBF5FF85FF9BFD8042466EE2FB9F" box="[584,694,1043,1062]" pageId="14" pageNumber="197" type="description">
<paragraph id="8BC8987EFF85FF9BFD8042466EE2FB9F" blockId="14.[151,1128,1043,1272]" box="[584,694,1043,1062]" pageId="14" pageNumber="197">
(
<figureCitation id="134C84FBFF85FF9BFD8642466EE4FB9F" box="[590,688,1043,1062]" captionStart="Figs 51–59" captionStartId="16.[151,188,1333,1352]" captionTargetBox="[160,1115,219,1311]" captionTargetId="figure-10@16.[160,1117,219,1311]" captionTargetPageId="16" captionText="Figs 51–59. Lasioglossum boswelliae Pauly &amp; Straka sp. nov., female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="14" pageNumber="197">Figs 51–59</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36DCBF5FF85FF9BFF5F42606848FBF1" box="[151,1052,1077,1096]" pageId="14" pageNumber="197" type="reference_group">
<paragraph id="8BC8987EFF85FF9BFF5F42606848FBF1" blockId="14.[151,1128,1043,1272]" box="[151,1052,1077,1096]" pageId="14" pageNumber="197">
<treatmentCitationGroup id="AB67BF50FF85FF9BFF5F42606848FBF1" box="[151,1052,1077,1096]" pageId="14" pageNumber="197">
<taxonomicName id="4C77E3FDFF85FF9BFF5F42606E31FBF1" ID-CoL="6NZK8" authority="(Cockerell, 1937)" baseAuthorityName="Cockerell" baseAuthorityYear="1937" box="[151,613,1077,1096]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="197" phylum="Arthropoda" rank="species" species="kowitense" subGenus="Evylaeus">
<emphasis id="B903446CFF85FF9BFF5F42606D58FBF1" box="[151,268,1077,1096]" italics="true" pageId="14" pageNumber="197">Lasioglossum</emphasis>
(
<emphasis id="B903446CFF85FF9BFED142606D32FBF1" box="[281,358,1077,1096]" italics="true" pageId="14" pageNumber="197">Evylaeus</emphasis>
) 
<emphasis id="B903446CFF85FF9BFEBA42606D92FBF1" box="[370,454,1077,1096]" italics="true" pageId="14" pageNumber="197">kowitense</emphasis>
(
<bibRefCitation id="EFE6E58FFF85FF9BFE1A42606E0BFBF1" author="COCKERELL T. D. A." box="[466,607,1077,1096]" pageId="14" pageNumber="197" refId="ref15063" refString="COCKERELL T. D. A. 1937: African bees of the genera Ceratina, Halictus and Megachile. British Museum, London, xvi + 254." type="book" year="1937">Cockerell, 1937</bibRefCitation>
)
</taxonomicName>
: 
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<bibRefCitation id="EFE6E58FFF85FF9BFDA642606EB2FBF1" author="EBMER A. W." box="[622,742,1077,1096]" pageId="14" pageNumber="197" pagination="399 - 453" refId="ref15264" refString="EBMER A. W. 2000: Asiatische Halictidae - 9. Die Artengruppe des Lasioglossum pauperatum (Insecta: Hymenoptera: Apoidea: Halictidae: Halictinae). Linzer Biologische Beitrage 32: 399 - 453." type="journal article" year="2000">EBMER (2000)</bibRefCitation>
: 433
</treatmentCitation>
, figs 94, 95 (misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<emphasis id="B903446CFF85FF9BFF5F420A6D49FBCB" bold="true" box="[151,285,1119,1138]" pageId="14" pageNumber="197">Type material.</emphasis>
<typeStatus id="54CC26DCFF85FF9BFEEA420A6D22FBC9" box="[290,374,1119,1138]" pageId="14" pageNumber="197" type="holotype">HOLOTYPE</typeStatus>
: 
<specimenCount id="9D7153F7FF85FF9BFE4A420C6DC6FBCB" box="[386,402,1113,1138]" pageId="14" pageNumber="197" type="female">♀</specimenCount>
, ‘ 
<collectingCountry id="F360D8EEFF85FF9BFE68420A6DA6FBCB" box="[416,498,1119,1138]" name="Yemen" pageId="14" pageNumber="197">YEMEN</collectingCountry>
, 
<collectingCountry id="F360D8EEFF85FF9BFE33420A6E09FBCB" box="[507,605,1119,1138]" name="Yemen" pageId="14" pageNumber="197">SOCOTRA</collectingCountry>
isl. E / Kesa env., 
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/ 
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, 
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/ 
<date id="FFC9BEBEFF85FF9BFB8D420A6CBEFB34" pageId="14" pageNumber="197" value="2010-01-28" valueMax="2010-01-29" valueMin="2010-01-28">28.- 29.i.2010</date>
, 
<collectionCode id="ED6600BBFF85FF9BFF3A422F6D4FFB34" box="[242,283,1146,1165]" pageId="14" pageNumber="197">YPT</collectionCode>
, L. Purchart lgt.’ (
<collectionCode id="ED6600BBFF85FF9BFE70422F6DADFB34" box="[440,505,1146,1165]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="14" pageNumber="197">NMPC</collectionCode>
). 
<typeStatus id="54CC26DCFF85FF9BFDC0422F6E34FB32" box="[520,608,1146,1165]" pageId="14" pageNumber="197" type="paratype">PARATYPES</typeStatus>
: 
<collectingCountry id="F360D8EEFF85FF9BFDA4422F6EE5FB34" box="[620,689,1146,1165]" name="Yemen" pageId="14" pageNumber="197">Socotra</collectingCountry>
: Dixam plateau, Wadi Dirhor, open woodland with 
<taxonomicName id="4C77E3FDFF85FF9BFF5F42C06D64FB11" box="[151,304,1173,1192]" class="Magnoliopsida" family="Burseraceae" genus="Boswellia" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="197" phylum="Tracheophyta" rank="species" species="ameero">
<emphasis id="B903446CFF85FF9BFF5F42C06D64FB11" box="[151,304,1173,1192]" italics="true" pageId="14" pageNumber="197">Boswellia ameero</emphasis>
</taxonomicName>
trees, 
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, 
<geoCoordinate id="EE43FEB9FF85FF9BFE6242C06E56FB11" box="[426,514,1173,1192]" degrees="12" direction="north" minutes="28.0" orientation="latitude" pageId="14" pageNumber="197" precision="92" value="12.466666">12°28.0′N</geoCoordinate>
, 
<geoCoordinate id="EE43FEB9FF85FF9BFDC442C06E36FB11" box="[524,610,1173,1192]" degrees="54" direction="east" minutes="00.5" orientation="longitude" pageId="14" pageNumber="197" precision="92" value="54.008335">54°00.5′E</geoCoordinate>
, 
<specimenCount id="9D7153F7FF85FF9BFDA342C06EDFFB1E" box="[619,651,1166,1192]" pageId="14" pageNumber="197" type="female">1 ♀</specimenCount>
, 
<date id="FFC9BEBEFF85FF9BFD5E42C06EA5FB11" box="[662,753,1173,1192]" pageId="14" pageNumber="197" value="2012-06-15">15.vi.2012</date>
et 
<date id="FFC9BEBEFF85FF9BFCC442C06F3CFB11" box="[780,872,1173,1192]" pageId="14" pageNumber="197" value="2012-06-22">22.vi.2012</date>
, J. Bezděk, J. Hájek, 
<collectionCode id="ED6600BBFF85FF9BFBE842C0687BFB11" box="[1056,1071,1173,1192]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="197" type="Museum">V</collectionCode>
. Hula, P. Kment, I. Malenovský, J. Niedobová &amp; L. Purchart lgt.; Aloove area, Aloove vill. env., 
<taxonomicName id="4C77E3FDFF85FF9BFC7242FA6833FB7B" box="[954,1127,1199,1218]" class="Magnoliopsida" family="Euphorbiaceae" genus="Jatropha" kingdom="Plantae" order="Malpighiales" pageId="14" pageNumber="197" phylum="Tracheophyta" rank="species" species="unicostata">
<emphasis id="B903446CFF85FF9BFC7242FA6833FB7B" box="[954,1127,1199,1218]" italics="true" pageId="14" pageNumber="197">Jatropha unicostata</emphasis>
</taxonomicName>
shrubland, with 
<taxonomicName id="4C77E3FDFF85FF9BFEE8429F6D90FB64" box="[288,452,1226,1245]" class="Magnoliopsida" family="Burseraceae" genus="Boswellia" kingdom="Plantae" order="Sapindales" pageId="14" pageNumber="197" phylum="Tracheophyta" rank="species" species="elongata">
<emphasis id="B903446CFF85FF9BFEE8429F6D90FB64" box="[288,452,1226,1245]" italics="true" pageId="14" pageNumber="197">Boswellia elongata</emphasis>
</taxonomicName>
trees, 
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, 
<geoCoordinate id="EE43FEB9FF85FF9BFDF2429F6EC5FB64" box="[570,657,1226,1245]" degrees="12" direction="north" minutes="31.2" orientation="latitude" pageId="14" pageNumber="197" precision="92" value="12.52">12°31.2′N</geoCoordinate>
<geoCoordinate id="EE43FEB9FF85FF9BFD5E429F6EB9FB64" box="[662,749,1226,1245]" degrees="54" direction="east" minutes="07.4" orientation="longitude" pageId="14" pageNumber="197" precision="92" value="54.123333">54°07.4′E</geoCoordinate>
, 
<specimenCount id="9D7153F7FF85FF9BFD3D429F6F40FB65" box="[757,788,1219,1245]" pageId="14" pageNumber="197" type="female">1 ♀</specimenCount>
, 
<date id="FFC9BEBEFF85FF9BFCD6429F6FF5FB64" box="[798,929,1226,1245]" pageId="14" pageNumber="197" value="2012-06-19" valueMax="2012-06-20" valueMin="2012-06-19">19.–20.vi.2012</date>
, J. Bezděk, J. Hájek, 
<collectionCode id="ED6600BBFF85FF9BFB9C429F6837FB64" box="[1108,1123,1226,1245]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="197" type="Museum">V</collectionCode>
. Hula, P. Kment, I. Malenovský, J. Niedobová &amp; L. Purchart lgt. (all in 
<collectionCode id="ED6600BBFF85FF9BFD3C42B06F60FB41" box="[756,820,1253,1272]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="14" pageNumber="197">NMPC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36DCBF5FF85FF9AFF5F43456D6DFBCA" lastPageId="15" lastPageNumber="198" pageId="14" pageNumber="197" type="description">
<paragraph id="8BC8987EFF85FF9BFF5F43456DB7FADE" blockId="14.[151,1127,1296,1575]" pageId="14" pageNumber="197">
<emphasis id="B903446CFF85FF9BFF5F43456DD0FA9E" bold="true" box="[151,388,1296,1319]" pageId="14" pageNumber="197">
Description. 
<emphasis id="B903446CFF85FF9BFEE343456DD0FA9E" bold="true" box="[299,388,1296,1319]" italics="true" pageId="14" pageNumber="197">Female.</emphasis>
</emphasis>
Small species: total body length 6.0 mm, forewing length 
<quantity id="4C8F359BFF85FF9BFBC443456836FA9E" box="[1036,1122,1296,1319]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="197" unit="mm" value="4.5">4.5 mm</quantity>
, intertegular distance 
<quantity id="4C8F359BFF85FF9BFEB143656D89FAFE" box="[377,477,1328,1351]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="197" unit="mm" value="1.25">1.25 mm</quantity>
. Black body with beige pubescence, apical margins of terga brown to reddish (
<figureCitation id="134C84FBFF85FF9BFE9243056D82FADE" box="[346,470,1360,1383]" captionStart="Figs 51–59" captionStartId="16.[151,188,1333,1352]" captionTargetBox="[160,1115,219,1311]" captionTargetId="figure-10@16.[160,1117,219,1311]" captionTargetPageId="16" captionText="Figs 51–59. Lasioglossum boswelliae Pauly &amp; Straka sp. nov., female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="14" pageNumber="197">Figs 51, 52</figureCitation>
).
</paragraph>
<paragraph id="8BC8987EFF85FF9AFF7D43256DBDFE0A" blockId="14.[151,1127,1296,1575]" lastBlockId="15.[113,1089,220,1619]" lastPageId="15" lastPageNumber="198" pageId="14" pageNumber="197">
<emphasis id="B903446CFF85FF9BFF7D43256CBBFA3E" box="[181,239,1392,1415]" italics="true" pageId="14" pageNumber="197">Head</emphasis>
almost as long as wide (Length/width = 0.96) (
<figureCitation id="134C84FBFF85FF9BFD1443256F7EFA3E" box="[732,810,1392,1415]" captionStart="Figs 51–59" captionStartId="16.[151,188,1333,1352]" captionTargetBox="[160,1115,219,1311]" captionTargetId="figure-10@16.[160,1117,219,1311]" captionTargetPageId="16" captionText="Figs 51–59. Lasioglossum boswelliae Pauly &amp; Straka sp. nov., female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="14" pageNumber="197">Fig. 53</figureCitation>
). Measurements (mm): length of clypeus 0.35; width of clypeus apically 0.38; lower interocular distance 0.78; upper interocular distance 0.95; clypeo-antennal distance 0.28; length of eye 1.18; interantennal distance 0.20; interocellar distance 0.35; ocellocular distance 0.20; antennocellar distance 0.58; antennocular distance 0.28. Mandibles black, dark brown in middle. Labrum black. Clypeus shiny with large irregular punctures, punctures less than half diameter apart basally, but more than three diameters apart apically, longitudinal swellings among punctures ill-defined. Supraclypeal area dull, irregularly microsculptured, with fine punctures; punctures one to less than one diameter apart. Frons with fine and dense punctures, interspaces hardly visible (
<figureCitation id="134C84FBFF84FF9AFC2D47496860FE8A" box="[997,1076,284,307]" captionStart="Figs 51–59" captionStartId="16.[151,188,1333,1352]" captionTargetBox="[160,1115,219,1311]" captionTargetId="figure-10@16.[160,1117,219,1311]" captionTargetPageId="16" captionText="Figs 51–59. Lasioglossum boswelliae Pauly &amp; Straka sp. nov., female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="15" pageNumber="198">Fig. 53</figureCitation>
). Paraocular area variably punctate, punctures one and half to less than half diameter apart, interspaces slightly shiny. Ocelli of medium size (
<figureCitation id="134C84FBFF84FF9AFD5847096EB6FECA" box="[656,738,348,371]" captionStart="Figs 51–59" captionStartId="16.[151,188,1333,1352]" captionTargetBox="[160,1115,219,1311]" captionTargetId="figure-10@16.[160,1117,219,1311]" captionTargetPageId="16" captionText="Figs 51–59. Lasioglossum boswelliae Pauly &amp; Straka sp. nov., female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="15" pageNumber="198">Fig. 54</figureCitation>
). Vertex narrow, nearly missing. Gena thin, finely longitudinally microsculptured ventrally. Scape black, flagellum black dorsally, pale ochraceous ventrally.
</paragraph>
<paragraph id="8BC8987EFF84FF9AFF5847E96D33FD6A" blockId="15.[113,1089,220,1619]" pageId="15" pageNumber="198">
<emphasis id="B903446CFF84FF9AFF5847E96D5CFE6A" box="[144,264,444,467]" italics="true" pageId="15" pageNumber="198">
<taxonomicName id="4C77E3FDFF84FF9AFF5847E96D51FE6A" box="[144,261,444,467]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="198" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
.
</emphasis>
Scutum with fine and dense punctation, punctures one to less than one diameter apart, interspaces between punctures brightly shiny (
<figureCitation id="134C84FBFF84FF9AFD5F47896EB1FE4A" box="[663,741,476,499]" captionStart="Figs 51–59" captionStartId="16.[151,188,1333,1352]" captionTargetBox="[160,1115,219,1311]" captionTargetId="figure-10@16.[160,1117,219,1311]" captionTargetPageId="16" captionText="Figs 51–59. Lasioglossum boswelliae Pauly &amp; Straka sp. nov., female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="15" pageNumber="198">Fig. 55</figureCitation>
). Scutum laterally, anteriorly and posteriorly with tomentum, center with short sparse and thin setae and may appear glabrous. Scutellum punctate like scutum on margins and medially, but with two sparsely punctate areas in centre, with punctures up to five diameters apart, with thin erect setae and without tomentum. Metanotum covered with tomentum. Mesopleura irregularly punctate, punctures ill-defined and evanescent in coarse microsculpture, dull. Hypoepimeral area with fine and dense punctures, dull. Propodeum not carinate, dorsal propodeal area in form of crescent, finely plicate (
<figureCitation id="134C84FBFF84FF9AFEC244E96D0EFD6A" box="[266,346,700,723]" captionStart="Figs 51–59" captionStartId="16.[151,188,1333,1352]" captionTargetBox="[160,1115,219,1311]" captionTargetId="figure-10@16.[160,1117,219,1311]" captionTargetPageId="16" captionText="Figs 51–59. Lasioglossum boswelliae Pauly &amp; Straka sp. nov., female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="15" pageNumber="198">Fig. 56</figureCitation>
).
</paragraph>
<paragraph id="8BC8987EFF84FF9AFF5844896F3DFCAA" blockId="15.[113,1089,220,1619]" pageId="15" pageNumber="198">
<emphasis id="B903446CFF84FF9AFF5844896C9DFD4A" box="[144,201,732,755]" italics="true" pageId="15" pageNumber="198">Legs.</emphasis>
Black, base of pro- and mesotibiae with small yellow spot. Inner metatibial spur with five teeth of decreasing length (
<figureCitation id="134C84FBFF84FF9AFE3F44A96E13FCAA" box="[503,583,764,787]" captionStart="Figs 51–59" captionStartId="16.[151,188,1333,1352]" captionTargetBox="[160,1115,219,1311]" captionTargetId="figure-10@16.[160,1117,219,1311]" captionTargetPageId="16" captionText="Figs 51–59. Lasioglossum boswelliae Pauly &amp; Straka sp. nov., female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="15" pageNumber="198">Fig. 57</figureCitation>
). Scopa of hind legs white.
</paragraph>
<paragraph id="8BC8987EFF84FF9AFF5845496DF4FCEA" blockId="15.[113,1089,220,1619]" pageId="15" pageNumber="198">
<emphasis id="B903446CFF84FF9AFF5845496C85FC8A" box="[144,209,796,819]" italics="true" pageId="15" pageNumber="198">Wings</emphasis>
relatively long, well exceeding end of metasoma. Membranes hyaline. Tegulae, stigma and veins testaceous.
</paragraph>
<paragraph id="8BC8987EFF84FF9AFF5845096DFFFBEA" blockId="15.[113,1089,220,1619]" pageId="15" pageNumber="198">
<emphasis id="B903446CFF84FF9AFF5845096D52FCCA" box="[144,262,860,883]" italics="true" pageId="15" pageNumber="198">Metasoma.</emphasis>
Tergum I with sloping base and centre completely striate and dull, only posterolaterally with fine and dense punctation, punctures well defined, about one diameter apart or less, sides of tergal base with patch of tomentum. Apical margin of tergum I smooth, punctate as main part of tergum, but impunctated in middle (
<figureCitation id="134C84FBFF84FF9AFD3845E96F6BFC6A" box="[752,831,956,979]" captionStart="Figs 51–59" captionStartId="16.[151,188,1333,1352]" captionTargetBox="[160,1115,219,1311]" captionTargetId="figure-10@16.[160,1117,219,1311]" captionTargetPageId="16" captionText="Figs 51–59. Lasioglossum boswelliae Pauly &amp; Straka sp. nov., female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="15" pageNumber="198">Fig. 59</figureCitation>
). Apical depressions not developed.Apex and also base of terga II–VI light brown to orange-brown. All apical margins of tergites largely discoloured, translucent (
<figureCitation id="134C84FBFF84FF9AFD8145A96ECEFBAA" box="[585,666,1020,1043]" captionStart="Figs 60–68" captionStartId="18.[151,188,1346,1365]" captionTargetBox="[151,1126,219,1330]" captionTargetId="figure-10@18.[151,1126,219,1330]" captionTargetPageId="18" captionText="Figs 60–68.Lasioglossum dioscoridis Pauly &amp; Straka sp.nov., female. 60 – dorsal view; 61 – lateral view; 62 – head, frontal view; 63 – vertex; 64 – scutum and scutellum; 65 – propodeum; 66 – inner metatibial spur; 67 – tergum I; 68 – metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5324242" httpUri="https://zenodo.org/record/5324242/files/figure.png" pageId="15" pageNumber="198">Fig. 60</figureCitation>
). Terga II–III with basolateral patch of tomentum. Terga V–VI with light yellowish to white setae. Sterna black with half apical part pale brown, with white setae.
</paragraph>
<paragraph id="8BC8987EFF84FF9AFF5842096D6DFBCA" blockId="15.[113,1089,220,1619]" box="[144,313,1116,1139]" pageId="15" pageNumber="198">
<emphasis id="B903446CFF84FF9AFF5842096C9CFBCA" bold="true" box="[144,200,1116,1139]" italics="true" pageId="15" pageNumber="198">Male</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36DCBF5FF84FF9AFFB942296F9AF9AA" pageId="15" pageNumber="198" type="diagnosis">
<paragraph id="8BC8987EFF84FF9AFFB942296F9AF9AA" blockId="15.[113,1089,220,1619]" pageId="15" pageNumber="198">
<emphasis id="B903446CFF84FF9AFFB942296D39FB2A" bold="true" box="[113,365,1148,1171]" pageId="15" pageNumber="198">Differential diagnosis.</emphasis>
This species belongs to the Holarctic subgenus 
<taxonomicName id="4C77E3FDFF84FF9AFCBF42296CA4FB0A" authority="Cockerell, 1897" authorityName="Cockerell" authorityYear="1897" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="subGenus" subGenus="Hemihalictus">
<emphasis id="B903446CFF84FF9AFCBF42296851FB2A" box="[887,1029,1148,1171]" italics="true" pageId="15" pageNumber="198">Hemihalictus</emphasis>
Cockerell, 1897
</taxonomicName>
based on the weakened distal wing venation, the second submarginal crossvein narrower than the first, the posterior surface of propodeum without carina. It is close to 
<taxonomicName id="4C77E3FDFF84FF9AFBE342E96DDEFB4A" authority="(Bluthgen, 1934)" baseAuthorityName="Bluthgen" baseAuthorityYear="1934" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="ablenum">
<emphasis id="B903446CFF84FF9AFBE342E96C99FB4A" italics="true" pageId="15" pageNumber="198">L. ablenum</emphasis>
(Blüthgen, 1934)
</taxonomicName>
from the Sahara, 
<taxonomicName id="4C77E3FDFF84FF9AFD8042896FC4FB4A" authority="(Cockerell, 1937)" baseAuthorityName="Cockerell" baseAuthorityYear="1937" box="[584,912,1244,1267]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="kowitense">
<emphasis id="B903446CFF84FF9AFD8042896E99FB4A" box="[584,717,1244,1267]" italics="true" pageId="15" pageNumber="198">L. kowitense</emphasis>
(
<bibRefCitation id="EFE6E58FFF84FF9AFD1342896FDDFB4A" author="COCKERELL T. D. A." box="[731,905,1244,1267]" pageId="15" pageNumber="198" refId="ref15063" refString="COCKERELL T. D. A. 1937: African bees of the genera Ceratina, Halictus and Megachile. British Museum, London, xvi + 254." type="book" year="1937">Cockerell, 1937</bibRefCitation>
)
</taxonomicName>
from 
<collectingCountry id="F360D8EEFF84FF9AFC1942896840FB4A" box="[977,1044,1244,1267]" name="Sudan" pageId="15" pageNumber="198">Sudan</collectingCountry>
and an undescribed species found in northern 
<collectingCountry id="F360D8EEFF84FF9AFDE942A96E2AFAAA" box="[545,638,1276,1299]" name="Ethiopia" pageId="15" pageNumber="198">Ethiopia</collectingCountry>
. The head is longer in 
<taxonomicName id="4C77E3FDFF84FF9AFCAD42A96FA5FAAA" authorityName="Pauly &amp; Straka" authorityYear="2017" box="[869,1009,1276,1299]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="boswelliae">
<emphasis id="B903446CFF84FF9AFCAD42A96FA5FAAA" box="[869,1009,1276,1299]" italics="true" pageId="15" pageNumber="198">L. boswelliae</emphasis>
</taxonomicName>
(length / width = 0.96) than in 
<taxonomicName id="4C77E3FDFF84FF9AFEA643496DBCFA8A" baseAuthorityName="Bluthgen" baseAuthorityYear="1934" box="[366,488,1308,1331]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="ablenum">
<emphasis id="B903446CFF84FF9AFEA643496DBCFA8A" box="[366,488,1308,1331]" italics="true" pageId="15" pageNumber="198">L. ablenum</emphasis>
</taxonomicName>
(L / 1 = 0.86) and 
<taxonomicName id="4C77E3FDFF84FF9AFD7143496F6BFA8A" baseAuthorityName="Cockerell" baseAuthorityYear="1937" box="[697,831,1308,1331]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="kowitense">
<emphasis id="B903446CFF84FF9AFD7143496F6BFA8A" box="[697,831,1308,1331]" italics="true" pageId="15" pageNumber="198">L. kowitense</emphasis>
</taxonomicName>
(L / 1 = 0.90). From 
<taxonomicName id="4C77E3FDFF84FF9AFBE343496C99FAEA" baseAuthorityName="Bluthgen" baseAuthorityYear="1934" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="ablenum">
<emphasis id="B903446CFF84FF9AFBE343496C99FAEA" italics="true" pageId="15" pageNumber="198">L. ablenum</emphasis>
</taxonomicName>
it differs in the punctation of the lateral parts of tergum I reaching the apical margin, the basolateral parts of the tergum I with allied tomentum. From 
<taxonomicName id="4C77E3FDFF84FF9AFC9143096F8AFACA" baseAuthorityName="Cockerell" baseAuthorityYear="1937" box="[857,990,1372,1395]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="kowitense">
<emphasis id="B903446CFF84FF9AFC9143096F8AFACA" box="[857,990,1372,1395]" italics="true" pageId="15" pageNumber="198">L. kowitense</emphasis>
</taxonomicName>
it differs in the tergum I, with its dorsal base completely striated and without punctation (densely striato-punctuated in 
<taxonomicName id="4C77E3FDFF84FF9AFE9D43C96D8FFA0A" baseAuthorityName="Cockerell" baseAuthorityYear="1937" box="[341,475,1436,1459]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="kowitense">
<emphasis id="B903446CFF84FF9AFE9D43C96D8FFA0A" box="[341,475,1436,1459]" italics="true" pageId="15" pageNumber="198">L. kowitense</emphasis>
</taxonomicName>
), punctation limited to the sides, the middle of the apical margin is not punctated (in 
<taxonomicName id="4C77E3FDFF84FF9AFE5E43E96E4FFA6A" baseAuthorityName="Cockerell" baseAuthorityYear="1937" box="[406,539,1468,1491]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="kowitense">
<emphasis id="B903446CFF84FF9AFE5E43E96E4FFA6A" box="[406,539,1468,1491]" italics="true" pageId="15" pageNumber="198">L. kowitense</emphasis>
</taxonomicName>
it is striato-punctated). The plumose pubescence of the baso-lateral parts of tergum I is denser in 
<taxonomicName id="4C77E3FDFF84FF9AFD8743896E88FA4A" authorityName="Pauly &amp; Straka" authorityYear="2017" box="[591,732,1500,1523]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="boswelliae">
<emphasis id="B903446CFF84FF9AFD8743896E88FA4A" box="[591,732,1500,1523]" italics="true" pageId="15" pageNumber="198">L. boswelliae</emphasis>
</taxonomicName>
. The pubescence of the scutum is denser in the middle in 
<taxonomicName id="4C77E3FDFF84FF9AFEA443A96DA6F9AA" baseAuthorityName="Cockerell" baseAuthorityYear="1937" box="[364,498,1532,1555]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="kowitense">
<emphasis id="B903446CFF84FF9AFEA443A96DA6F9AA" box="[364,498,1532,1555]" italics="true" pageId="15" pageNumber="198">L. kowitense</emphasis>
</taxonomicName>
but denser in the periphery in 
<taxonomicName id="4C77E3FDFF84FF9AFCF043A96F92F9AA" authorityName="Pauly &amp; Straka" authorityYear="2017" box="[824,966,1532,1555]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="198" phylum="Arthropoda" rank="species" species="boswelliae">
<emphasis id="B903446CFF84FF9AFCF043A96F92F9AA" box="[824,966,1532,1555]" italics="true" pageId="15" pageNumber="198">L. boswelliae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DCBF5FF84FF9AFFB940496815F9EA" pageId="15" pageNumber="198" type="etymology">
<paragraph id="8BC8987EFF84FF9AFFB940496815F9EA" blockId="15.[113,1089,220,1619]" pageId="15" pageNumber="198">
<emphasis id="B903446CFF84FF9AFFB940496CA6F98A" bold="true" box="[113,242,1564,1587]" pageId="15" pageNumber="198">Etymology.</emphasis>
Named after the Frankincense Trees (
<taxonomicName id="4C77E3FDFF84FF9AFD4140496F73F98A" box="[649,807,1564,1587]" class="Magnoliopsida" family="Burseraceae" genus="Boswellia" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="198" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B903446CFF84FF9AFD4140496EA6F98A" box="[649,754,1564,1587]" italics="true" pageId="15" pageNumber="198">Boswellia</emphasis>
spp.
</taxonomicName>
), as this bee was collected in an area with 
<taxonomicName id="4C77E3FDFF84FF9AFEDD40696D87F9EA" box="[277,467,1596,1619]" class="Magnoliopsida" family="Burseraceae" genus="Boswellia" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="198" phylum="Tracheophyta" rank="species" species="ameero">
<emphasis id="B903446CFF84FF9AFEDD40696D87F9EA" box="[277,467,1596,1619]" italics="true" pageId="15" pageNumber="198">Boswellia ameero</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C77E3FDFF84FF9AFDCF40696ED5F9EA" box="[519,641,1596,1619]" class="Magnoliopsida" family="Burseraceae" genus="Boswellia" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="198" phylum="Tracheophyta" rank="species" species="elongata">
<emphasis id="B903446CFF84FF9AFDCF40696E43F9EA" box="[519,535,1596,1619]" italics="true" pageId="15" pageNumber="198">B</emphasis>
. 
<emphasis id="B903446CFF84FF9AFDEC40696ED5F9EA" box="[548,641,1596,1619]" italics="true" pageId="15" pageNumber="198">elongata</emphasis>
</taxonomicName>
trees; noun in apposition in genitive case.
</paragraph>
</subSubSection>
<caption id="DF08C8F6FF9BFF85FF5F43606DDFFAC4" ID-DOI="http://doi.org/10.5281/zenodo.5324240" ID-Zenodo-Dep="5324240" httpUri="https://zenodo.org/record/5324240/files/figure.png" pageId="16" pageNumber="199" startId="16.[151,188,1333,1352]" targetBox="[160,1115,219,1311]" targetPageId="16">
<paragraph id="8BC8987EFF9BFF85FF5F43606DDFFAC4" blockId="16.[151,1127,1333,1405]" pageId="16" pageNumber="199">
Figs 51–59. 
<taxonomicName id="4C77E3FDFF9BFF85FF3743606E0CFAF1" authority="Pauly &amp; Straka" authorityName="Pauly &amp; Straka" authorityYear="2017" box="[255,600,1333,1352]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="199" phylum="Arthropoda" rank="species" species="boswelliae" status="sp. nov.">
<emphasis id="B903446CFF9BFF85FF3743606D87FAF1" box="[255,467,1333,1352]" italics="true" pageId="16" pageNumber="199">Lasioglossum boswelliae</emphasis>
Pauly &amp; Straka
</taxonomicName>
<taxonomicNameLabel id="A230F917FF9BFF85FD9343606ECBFAF1" box="[603,671,1333,1352]" pageId="16" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
, female.51 – dorsal view; 52 – lateral view; 53 – head, frontal view; 54 – vertex; 55 – scutum and scutellum; 56 – propodeum; 57 – inner metatibial spur; 58 – tergum I; 59 – metasoma, dorsal view.
</paragraph>
</caption>
<subSubSection id="C36DCBF5FF9BFF85FF5F43EE6E6FF9AB" pageId="16" pageNumber="199" type="discussion">
<paragraph id="8BC8987EFF9BFF85FF5F43EE6E6FF9AB" blockId="16.[151,1127,1467,1618]" pageId="16" pageNumber="199">
<emphasis id="B903446CFF9BFF85FF5F43EE6D52FA6B" bold="true" box="[151,262,1467,1490]" pageId="16" pageNumber="199">Remarks.</emphasis>
The species resembles the Afrotropical species. We suspect its Ethiopian origin. It was previously reported from 
<collectingCountry id="F360D8EEFF9BFF85FE1B438E6E73FA4B" box="[467,551,1499,1522]" name="Yemen" pageId="16" pageNumber="199">Socotra</collectingCountry>
, based on 
<specimenCount id="9D7153F7FF9BFF85FD5B438E6F70FA4B" box="[659,804,1499,1522]" pageId="16" pageNumber="199" type="female">three females</specimenCount>
, under the name 
<taxonomicName id="4C77E3FDFF9BFF85FC1C438E6D2DF9AB" baseAuthorityName="Cockerell" baseAuthorityYear="1937" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="199" phylum="Arthropoda" rank="species" species="kowitense" subGenus="Evylaeus">
<emphasis id="B903446CFF9BFF85FC1C438E6833FA4B" box="[980,1127,1499,1522]" italics="true" pageId="16" pageNumber="199">Lasioglossum</emphasis>
(
<emphasis id="B903446CFF9BFF85FF6843AE6D54F9AB" box="[160,256,1531,1554]" italics="true" pageId="16" pageNumber="199">Evylaeus</emphasis>
) 
<emphasis id="B903446CFF9BFF85FED843AE6D2DF9AB" box="[272,377,1531,1554]" italics="true" pageId="16" pageNumber="199">kowitense</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFE6E58FFF9BFF85FE6943AE6E6CF9AB" author="EBMER A. W." box="[417,568,1531,1554]" pageId="16" pageNumber="199" pagination="399 - 453" refId="ref15264" refString="EBMER A. W. 2000: Asiatische Halictidae - 9. Die Artengruppe des Lasioglossum pauperatum (Insecta: Hymenoptera: Apoidea: Halictidae: Halictinae). Linzer Biologische Beitrage 32: 399 - 453." type="journal article" year="2000">EBMER (2000)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DCBF5FF9BFF85FF5F404E6833F9EB" pageId="16" pageNumber="199" type="distribution">
<paragraph id="8BC8987EFF9BFF85FF5F404E6833F9EB" blockId="16.[151,1127,1467,1618]" pageId="16" pageNumber="199">
<emphasis id="B903446CFF9BFF85FF5F404E6D7DF98B" bold="true" box="[151,297,1563,1586]" pageId="16" pageNumber="199">Distribution.</emphasis>
The species is endemic to 
<collectingCountry id="F360D8EEFF9BFF85FDF5404E6EC6F98B" box="[573,658,1563,1586]" name="Yemen" pageId="16" pageNumber="199">Socotra</collectingCountry>
: Adho Demalu [= Adho Dimello], Hamadara [= Hamadero] (
<bibRefCitation id="EFE6E58FFF9BFF85FEFC406E6DE2F9EB" author="EBMER A. W." box="[308,438,1595,1618]" pageId="16" pageNumber="199" pagination="399 - 453" refId="ref15264" refString="EBMER A. W. 2000: Asiatische Halictidae - 9. Die Artengruppe des Lasioglossum pauperatum (Insecta: Hymenoptera: Apoidea: Halictidae: Halictinae). Linzer Biologische Beitrage 32: 399 - 453." type="journal article" year="2000">EBMER 2000</bibRefCitation>
), Aloove area, Dixam plateau and environment of Kesa (this paper).
</paragraph>
</subSubSection>
</treatment>
</document>