treatments-xml/data/03/A7/87/03A787F7FFCDAA36FE420E61FBDE89E0.xml
2024-06-21 12:22:17 +02:00

567 lines
53 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="B31D44840E59BB28F790FD70799401F1" ID-DOI="10.24349/acarologia/20214432" ID-ISSN="2107-7207" ID-Zenodo-Dep="5392448" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630316612985" checkinUser="marcus" docAuthor="Ermilov, Sergey G. &amp; Liao, Jhih-Rong" docDate="2021" docId="03A787F7FFCDAA36FE420E61FBDE89E0" docLanguage="en" docName="Acarologia.2021.61.2.297-320.pdf" docOrigin="Acarologia 61 (2)" docSource="http://dx.doi.org/10.24349/acarologia/20214432" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.6:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="6" docTitle="Tyrphonothrus nivnu Ermilov &amp; Liao 2021, n. sp." docType="treatment" docVersion="3" lastPageNumber="303" masterDocId="FF9EFF8FFFCEAA31FFA90B07FFC68C33" masterDocTitle="New faunistical and taxonomic data on oribatid mites (Acari: Oribatida) of Taiwan" masterLastPageNumber="320" masterPageNumber="297" pageNumber="299" updateTime="1699224043532" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="170D9DCFE43932F61C5635F9B0A3EBB8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CFE53700C807DA2799C4F97D352F24CC">
<mods:title id="78B0ACED311C92F20B84857F2CB1049B">New faunistical and taxonomic data on oribatid mites (Acari: Oribatida) of Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="08180BF67875954ED697D840B1E474A1" type="personal">
<mods:role id="1831D442547CEF8F7DD8D685B85CA551">
<mods:roleTerm id="BE27F56170D774C3925D37C9AF3E4BA9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="305C2AA9D986D293A4609DD17D8E961E">Ermilov, Sergey G.</mods:namePart>
<mods:affiliation id="07B332F3C8F11BF7CB8621E28EC88E86">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
</mods:name>
<mods:name id="FF02578408A3D10B97B8EF0D0AAFCE25" type="personal">
<mods:role id="B1FF974C7CDB4FBC7DDC364DFF8B4019">
<mods:roleTerm id="031694C3A336A4371E1EE6713A442DB4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7DFD160EC506E34313A71711C40CF78C">Liao, Jhih-Rong</mods:namePart>
<mods:affiliation id="42F5803D5E2F93E957E202F88190AD4B">Department of Entomology, National Taiwan University, Taipei, Taiwan. &amp; Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C50937C0C18F311EA5B4C3DA3BD62D8F">text</mods:typeOfResource>
<mods:relatedItem id="EBCB396554557757B345BE2B6F439655" type="host">
<mods:titleInfo id="F741908AD1C705288B41D173997F95CE">
<mods:title id="DB208847E959D164C8B0EE81631D9A92">Acarologia</mods:title>
</mods:titleInfo>
<mods:part id="A48096BF31111C66FAC9AA49EE3CF993">
<mods:date id="E38B0AE0D6ADCD5C005AC155C7A3DF3A">2021</mods:date>
<mods:detail id="0105763CB1F49FDFBCBA451273EC48A2" type="pubDate">
<mods:number id="DD81A5C95B22211F224B596B56D150CF">2021-04-29</mods:number>
</mods:detail>
<mods:detail id="16B7BDA4B0783F64310CFE74BEB1D54A" type="volume">
<mods:number id="D550B539C63E19C931C45F0E32D2D4A5">61</mods:number>
</mods:detail>
<mods:detail id="140E026194CBD14F28BA82D5BF08F0A3" type="issue">
<mods:number id="E73ABB53D86857C7350E0EBEDBB92D23">2</mods:number>
</mods:detail>
<mods:extent id="69A4D8847F209C2F5422F52DBD695C07" unit="page">
<mods:start id="9AB788CEB0C8FF65CAAA93754B46A5AC">297</mods:start>
<mods:end id="A763BB77F5DCAF744749BFFBB86544B9">320</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="CDD71CF341F662C8ACE6B980BFA3DB4A">
<mods:url id="4A5DC47FAF00F0055502BD3FCE50BECA">http://dx.doi.org/10.24349/acarologia/20214432</mods:url>
</mods:location>
<mods:classification id="81CAB9E886656A6684D746E6FE303702">journal article</mods:classification>
<mods:identifier id="EC4A4C19CDDBBDCDC1F7A2969B66F424" type="DOI">10.24349/acarologia/20214432</mods:identifier>
<mods:identifier id="198FEA54ED9740ECADB58670174FEEE1" type="ISSN">2107-7207</mods:identifier>
<mods:identifier id="859ED3212DFC573D109ADE61B57B7CD4" type="Zenodo-Dep">5392448</mods:identifier>
</mods:mods>
<treatment id="03A787F7FFCDAA36FE420E61FBDE89E0" LSID="urn:lsid:plazi:treatment:03A787F7FFCDAA36FE420E61FBDE89E0" httpUri="http://treatment.plazi.org/id/03A787F7FFCDAA36FE420E61FBDE89E0" lastPageId="7" lastPageNumber="303" pageId="3" pageNumber="299">
<subSubSection id="C314656AFFCDAA32FE420E61FC238982" pageId="3" pageNumber="299" type="nomenclature">
<paragraph id="8BB136E1FFCDAA32FE420E61FCF189B3" blockId="3.[491,823,1382,1408]" box="[491,823,1382,1408]" pageId="3" pageNumber="299">
<heading id="D0F9818DFFCDAA32FE420E61FCF189B3" bold="true" box="[491,823,1382,1408]" fontSize="11" level="2" pageId="3" pageNumber="299" reason="3">
<emphasis id="B97AEAF3FFCDAA32FE420E61FCF189B3" bold="true" box="[491,823,1382,1408]" pageId="3" pageNumber="299">
<taxonomicName id="4C0E4D62FFCDAA32FE420E61FD2B89B3" ID-CoL="359bfe3b-be73-4f6a-9269-9a6b136b1535" authority="Ermilov &amp; Liao, 2021" authorityName="Ermilov &amp; Liao" authorityYear="2021" box="[491,749,1382,1408]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="nivnu" status="sp. nov.">
<emphasis id="B97AEAF3FFCDAA32FE420E61FD2B89B3" bold="true" box="[491,749,1382,1408]" italics="true" pageId="3" pageNumber="299">Tyrphonothrus nivnu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2495788FFCDAA32FD5D0E61FCF189B3" box="[756,823,1382,1408]" pageId="3" pageNumber="299" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136E1FFCDAA32FE420E99FC238982" blockId="3.[491,1534,1438,2045]" box="[491,997,1438,1457]" pageId="3" pageNumber="299">
Zoobank:
<uuid id="FFA80C34FFCDAA32FDED0E99FC238982" box="[580,997,1438,1457]" pageId="3" pageNumber="299">F9F7AF0D-F80C-4E3D-B023-96E25AE36B02</uuid>
</paragraph>
</subSubSection>
<subSubSection id="C314656AFFCDAA32FDBA0EBAFD6389E6" box="[531,677,1469,1493]" pageId="3" pageNumber="299" type="description">
<paragraph id="8BB136E1FFCDAA32FDBA0EBAFD6389E6" blockId="3.[491,1534,1438,2045]" box="[531,677,1469,1493]" pageId="3" pageNumber="299">(Figures 24)</paragraph>
</subSubSection>
<subSubSection id="C314656AFFCDAA32FDBA0EE7FB758ADA" pageId="3" pageNumber="299" type="diagnosis">
<paragraph id="8BB136E1FFCDAA32FDBA0EE7FB758ADA" blockId="3.[491,1534,1438,2045]" pageId="3" pageNumber="299">
<emphasis id="B97AEAF3FFCDAA32FDBA0EE7FD4F89CB" bold="true" box="[531,649,1504,1528]" pageId="3" pageNumber="299">Diagnosis</emphasis>
— Adult. Body size: 614630 × 348357. Body with sparse tuberculate cerotegument; ornamentation and sculpturing absent. Rostral, lamellar and interlamellar setae and exobothridial seta
<emphasis id="B97AEAF3FFCDAA32FD480D21FD3F8A0E" box="[737,761,1574,1597]" italics="true" pageId="3" pageNumber="299">ex</emphasis>
<subScript id="178A34A4FFCDAA32FD510D36FCC78A71" attach="left" box="[760,769,1585,1602]" fontSize="7" pageId="3" pageNumber="299">1</subScript>
setiform, roughened;
<emphasis id="B97AEAF3FFCDAA32FC5A0D21FBCF8A0E" box="[1011,1033,1574,1597]" italics="true" pageId="3" pageNumber="299">in</emphasis>
longest,
<emphasis id="B97AEAF3FFCDAA32FBC20D21FB448A0E" box="[1131,1154,1574,1597]" italics="true" pageId="3" pageNumber="299">ro</emphasis>
and
<emphasis id="B97AEAF3FFCDAA32FB110D22FB0D8A0E" box="[1208,1227,1573,1597]" italics="true" pageId="3" pageNumber="299">le</emphasis>
of medium length,
<emphasis id="B97AEAF3FFCDAA32FA0B0D21FA7C8A0E" box="[1442,1466,1574,1597]" italics="true" pageId="3" pageNumber="299">ex</emphasis>
<subScript id="178A34A4FFCDAA32FA100D36FA048A71" attach="left" box="[1465,1474,1585,1602]" fontSize="7" pageId="3" pageNumber="299">1</subScript>
shortest.
<taxonomicName id="4C0E4D62FFCDAA32FDFD0D4FFD088A53" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[596,718,1608,1632]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="307" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
with four longitudinal ridges. Notogastral setae
<emphasis id="B97AEAF3FFCDAA32FB7F0D4FFB248A6C" box="[1238,1250,1608,1631]" italics="true" pageId="3" pageNumber="299">e</emphasis>
<subScript id="178A34A4FFCDAA32FB4B0D54FB2E8A57" attach="left" box="[1250,1256,1619,1636]" fontSize="7" pageId="3" pageNumber="299">2</subScript>
,
<emphasis id="B97AEAF3FFCDAA32FB520D40FACE8A6C" box="[1275,1288,1607,1631]" italics="true" pageId="3" pageNumber="299">h</emphasis>
<subScript id="178A34A4FFCDAA32FAA00D54FAD48A57" attach="left" box="[1289,1298,1619,1636]" fontSize="7" pageId="3" pageNumber="299">1</subScript>
,
<emphasis id="B97AEAF3FFCDAA32FA8B0D40FAE98A6C" box="[1314,1327,1607,1631]" italics="true" pageId="3" pageNumber="299">h</emphasis>
<subScript id="178A34A4FFCDAA32FA860D54FAF38A57" attach="left" box="[1327,1333,1619,1636]" fontSize="7" pageId="3" pageNumber="299">2</subScript>
, and
<emphasis id="B97AEAF3FFCDAA32FADE0D4FFA428A6C" box="[1399,1412,1608,1631]" italics="true" pageId="3" pageNumber="299">p</emphasis>
<subScript id="178A34A4FFCDAA32FA2D0D54FA4C8A57" attach="left" box="[1412,1418,1619,1636]" fontSize="7" pageId="3" pageNumber="299">2</subScript>
very long, setiform, with attenuate tip, barbed, other setae of medium length, setiform, barbed. Cheliceral seta
<emphasis id="B97AEAF3FFCDAA32FD3B0D8BFD068A97" box="[658,704,1676,1700]" italics="true" pageId="3" pageNumber="299">cha</emphasis>
spiniform, barbed apically,
<emphasis id="B97AEAF3FFCDAA32FC4E0D8BFBD38A97" box="[999,1045,1676,1700]" italics="true" pageId="3" pageNumber="299">chb</emphasis>
falciform. Epimeral setae
<emphasis id="B97AEAF3FFCDAA32FA980D8BFA8A8A97" box="[1329,1356,1676,1700]" italics="true" pageId="3" pageNumber="299">1a</emphasis>
,
<emphasis id="B97AEAF3FFCDAA32FAF20D8BFABD8A97" box="[1371,1403,1676,1700]" italics="true" pageId="3" pageNumber="299">2a</emphasis>
and
<emphasis id="B97AEAF3FFCDAA32FA020D8BFA0D8A97" box="[1451,1483,1676,1700]" italics="true" pageId="3" pageNumber="299">3a</emphasis>
spiniform, barbed apically, others setiform, slightly barbed. Genital plate with nine or 10 pairs of setiform, roughened genital setae. Leg tarsi with three claws.
</paragraph>
</subSubSection>
<subSubSection id="C314656AFFCDAA36FDBA0DF3FD838F98" lastPageId="7" lastPageNumber="303" pageId="3" pageNumber="299" type="description">
<paragraph id="8BB136E1FFCDAA32FDBA0DF3FAA28B1D" blockId="3.[491,1534,1438,2045]" pageId="3" pageNumber="299">
<emphasis id="B97AEAF3FFCDAA32FDBA0DF3FD388B3F" bold="true" box="[531,766,1780,1804]" pageId="3" pageNumber="299">Description of adult</emphasis>
<emphasis id="B97AEAF3FFCDAA32FC890DF2FC078B3F" box="[800,961,1781,1804]" italics="true" pageId="3" pageNumber="299">Measurements</emphasis>
Body length: 614 (
<typeStatus id="54B58843FFCDAA32FB1A0DF3FAD18B3F" box="[1203,1303,1780,1804]" pageId="3" pageNumber="299" type="holotype">holotype</typeStatus>
, female), 614630 (
<specimenCount id="9D08FD68FFCDAA32FE5B0C11FD498B1C" box="[498,655,1814,1839]" pageId="3" pageNumber="299" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
, all females); body width: 348 (
<typeStatus id="54B58843FFCDAA32FC450C11FB978B1D" box="[1004,1105,1814,1838]" pageId="3" pageNumber="299" type="holotype">holotype</typeStatus>
), 348357 (
<specimenCount id="9D08FD68FFCDAA32FB130C11FA9F8B1D" box="[1210,1369,1814,1838]" pageId="3" pageNumber="299" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
).
</paragraph>
<paragraph id="8BB136E1FFCDAA32FDBA0C3DFA6D8B47" blockId="3.[491,1534,1438,2045]" pageId="3" pageNumber="299">
<emphasis id="B97AEAF3FFCDAA32FDBA0C3DFD498B62" box="[531,655,1850,1873]" italics="true" pageId="3" pageNumber="299">Integument</emphasis>
(
<figureCitation id="13352A64FFCDAA32FD350C3EFCDC8B62" box="[668,794,1849,1873]" captionStart="Figure 4" captionStartId="6.[119,190,1905,1926]" captionTargetBox="[95,1501,221,1848]" captionTargetId="figure-2@6.[95,1501,221,1851]" captionTargetPageId="6" captionText="Figure 4 Tyrphonothrus nivnun. sp., adult, SEM micrographs (a-d) and microscope images (e-i): a dorsal view; b lateral view; c tuberculate notogastral cerotegument; d claws of leg tarsus I; e epimeral setae3aand3b; f epimeral integument (deep focus); g, h genital plates (superficial and deep focus, respectively); i claws of leg tarsus III. Scale bar 200 μm (a, b), 10 μm (c), 50 μm (d); images without scale bar." figureDoi="http://doi.org/10.5281/zenodo.5392454" httpUri="https://zenodo.org/record/5392454/files/figure.png" pageId="3" pageNumber="299">Figs 4c, 4f</figureCitation>
) Body color light brown. Body surface densely porose and covered by gel-like and sparse tuberculate (diameter of tubercle up to 2) cerotegument.
</paragraph>
<paragraph id="8BB136E1FFCDAA32FDBA0C79FB0E8BE8" blockId="3.[491,1534,1438,2045]" pageId="3" pageNumber="299">
<emphasis id="B97AEAF3FFCDAA32FDBA0C79FD4B8BA5" box="[531,653,1918,1942]" italics="true" pageId="3" pageNumber="299">Prodorsum</emphasis>
(Figs 2a, 4a, 4b) Rostrum rounded. Lateral carina well-developed. Rostral (5357), lamellar (5357) and interlamellar (110118) setae, and exobothridial seta
<emphasis id="B97AEAF3FFCDAA32FADB0CA6FA4C8B8B" box="[1394,1418,1953,1976]" italics="true" pageId="3" pageNumber="299">ex</emphasis>
<subScript id="178A34A4FFCDAA32FA230CABFA558B8E" attach="left" box="[1418,1427,1964,1981]" fontSize="7" pageId="3" pageNumber="299">1</subScript>
(20) setiform, roughened; exobothridial seta
<emphasis id="B97AEAF3FFCDAA32FC3D0CC3FC6A8BE8" box="[916,940,1988,2011]" italics="true" pageId="3" pageNumber="299">ex</emphasis>
<subScript id="178A34A4FFCDAA32FC020CC8FC778BD3" attach="left" box="[939,945,1999,2016]" fontSize="7" pageId="3" pageNumber="299">2</subScript>
represented by alveolus.
</paragraph>
<paragraph id="8BB136E1FFCDAA34FDBA0CE1FBB28437" blockId="3.[491,1534,1438,2045]" lastBlockId="5.[119,1525,1985,2052]" lastPageId="5" lastPageNumber="301" pageId="3" pageNumber="299">
<taxonomicName id="4C0E4D62FFCDAA32FDBA0CE1FD568BCE" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[531,656,2022,2045]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEAF3FFCDAA32FDBA0CE1FD568BCE" box="[531,656,2022,2045]" italics="true" pageId="3" pageNumber="299">Notogaster</emphasis>
</taxonomicName>
(Figs 2a, 2b, 4a, 4b) Four longitudinal ridges; medial ridges bifurcate d leg I, without trochanter, right, ventroantiaxial view; e leg II, without trochanter and tarsus, right, ventroantiaxial view; f leg III, without tarsus, left, antiaxial view; g leg IV, left, antiaxial view. Scale bar 20 μm (a, c), 10 μm (b), 50 μm (d-g).
</paragraph>
<paragraph id="8BB136E1FFCAAA35FFDE0EBAFA268BCE" pageId="4" pageNumber="300">
<table id="F90EC441FFCA55CEFFDE0EBAFA258BCE" box="[119,1507,1469,2045]" colsContinueIn="8.[119,1516,1619,2025]" gridcols="1" gridrows="12" pageId="4" pageNumber="300">
<tr id="353E34A3FFCA55CEFFDE0EBAFA2589E0" box="[119,1507,1469,1491]" gridrow="0" pageId="4" pageNumber="300">
<th id="76EF5DDFFFCA55CEFFDE0EBAFA2589E0" box="[119,1507,1469,1491]" gridcol="0" gridrow="0" pageId="4" pageNumber="300">
<emphasis id="B97AEAF3FFCAAA35FFDE0EBAFF1689E1" bold="true" box="[119,208,1469,1490]" pageId="4" pageNumber="300">Figure 2</emphasis>
<taxonomicName id="4C0E4D62FFCAAA35FF710EBAFE6089E0" authority="Ermilov &amp; Liao, 2021" authorityName="Ermilov &amp; Liao" authorityYear="2021" box="[216,422,1469,1491]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="300" phylum="Arthropoda" rank="species" species="nivnu" status="sp. nov.">
<emphasis id="B97AEAF3FFCAAA35FF710EBAFE6089E0" box="[216,422,1469,1491]" italics="true" pageId="4" pageNumber="300">Tyrphonothrus nivnu</emphasis>
</taxonomicName>
<emphasis id="B97AEAF3FFCAAA35FE0F0EBAFE1A89E1" bold="true" box="[422,476,1469,1490]" pageId="4" pageNumber="300">
<taxonomicNameLabel id="A2495788FFCAAA35FE0F0EBAFE1A89E1" box="[422,476,1469,1490]" pageId="4" pageNumber="300" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult: a dorsal view (legs omitted); b ventral view (legs omitted). Scale bar 100 μm.
</th>
</tr>
<tr id="353E34A3FFCA55CEFFDE0D4FFA258A96" box="[119,1507,1608,1701]" gridrow="1" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0D4FFA258A96" box="[119,1507,1608,1701]" gridcol="0" gridrow="1" pageId="4" pageNumber="300">
posteriorly. Notogastral setae
<emphasis id="B97AEAF3FFCAAA35FC990D4EFCFA8A53" box="[816,828,1609,1632]" italics="true" pageId="4" pageNumber="300">e</emphasis>
2,
<emphasis id="B97AEAF3FFCAAA35FCFC0D4FFCA48A53" box="[853,866,1608,1632]" italics="true" pageId="4" pageNumber="300">h</emphasis>
1,
<emphasis id="B97AEAF3FFCAAA35FCD50D4FFC4F8A53" box="[892,905,1608,1632]" italics="true" pageId="4" pageNumber="300">h</emphasis>
2, and
<emphasis id="B97AEAF3FFCAAA35FC790D4EFC1B8A53" box="[976,989,1609,1632]" italics="true" pageId="4" pageNumber="300">p</emphasis>
2 (8694) setiform, with attenuate tip, barbed, other setae (4953) setiform, barbed. Setae
<emphasis id="B97AEAF3FFCAAA35FC680D6CFC088AB1" box="[961,974,1643,1666]" italics="true" pageId="4" pageNumber="300">p</emphasis>
2 and
<emphasis id="B97AEAF3FFCAAA35FBA40D6CFBDC8AB1" box="[1037,1050,1643,1666]" italics="true" pageId="4" pageNumber="300">p</emphasis>
3 distant from each other. All notogastral lyrifissures (
<emphasis id="B97AEAF3FFCAAA35FDD80D89FD408A96" box="[625,646,1678,1701]" italics="true" pageId="4" pageNumber="300">ia</emphasis>
,
<emphasis id="B97AEAF3FFCAAA35FD3A0D89FD688A96" box="[659,686,1678,1701]" italics="true" pageId="4" pageNumber="300">im</emphasis>
,
<emphasis id="B97AEAF3FFCAAA35FD120D8AFD168A96" box="[699,720,1677,1701]" italics="true" pageId="4" pageNumber="300">ih</emphasis>
, and
<emphasis id="B97AEAF3FFCAAA35FCA30D89FCD98A96" box="[778,799,1678,1701]" italics="true" pageId="4" pageNumber="300">ip</emphasis>
) distinct.
</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0DA8FA258AF4" box="[119,1507,1711,1735]" gridrow="2" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0DA8FA258AF4" box="[119,1507,1711,1735]" gridcol="0" gridrow="2" pageId="4" pageNumber="300">
<taxonomicName id="4C0E4D62FFCAAA35FDBA0DA8FD658AF4" authorityName="Krefft" authorityYear="1871" box="[531,675,1711,1735]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="300" phylum="Nematoda" rank="genus">
<emphasis id="B97AEAF3FFCAAA35FDBA0DA8FD658AF4" box="[531,675,1711,1735]" italics="true" pageId="4" pageNumber="300">Gnathosoma</emphasis>
</taxonomicName>
(Figs 3a-c) Subcapitulum size: 106114 × 8286. Subcapitular setae (
<emphasis id="B97AEAF3FFCAAA35FA1A0DB7FA068AF4" box="[1459,1472,1712,1735]" italics="true" pageId="4" pageNumber="300">a</emphasis>
,
</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0DD5FA258AD9" box="[119,1507,1746,1770]" gridrow="3" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0DD5FA258AD9" box="[119,1507,1746,1770]" gridcol="0" gridrow="3" pageId="4" pageNumber="300">
2628;
<emphasis id="B97AEAF3FFCAAA35FD940DD4FD968AD9" box="[573,592,1747,1770]" italics="true" pageId="4" pageNumber="300">m</emphasis>
, 1214;
<emphasis id="B97AEAF3FFCAAA35FD190DD5FD7B8AD9" box="[688,701,1746,1770]" italics="true" pageId="4" pageNumber="300">h</emphasis>
, 1416) setiform, slightly barbed;
<emphasis id="B97AEAF3FFCAAA35FB9B0DD4FB838AD9" box="[1074,1093,1747,1770]" italics="true" pageId="4" pageNumber="300">m</emphasis>
thinnest. Three pairs of adoral setae
</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0DF3FA258B3F" box="[119,1507,1780,1804]" gridrow="4" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0DF3FA258B3F" box="[119,1507,1780,1804]" gridcol="0" gridrow="4" pageId="4" pageNumber="300">(8) setiform, thin, smooth. Palp (4145) with setation 0-0-1-3-9(+ω); solenidion thickened,</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0C10FA258B1C" box="[119,1507,1815,1839]" gridrow="5" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0C10FA258B1C" box="[119,1507,1815,1839]" gridcol="0" gridrow="5" pageId="4" pageNumber="300">rounded apically. Postpalpal seta (10) rod-like, slightly barbed. Chelicera (98102) with</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0C3EFA258B62" box="[119,1507,1849,1873]" gridrow="6" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0C3EFA258B62" box="[119,1507,1849,1873]" gridcol="0" gridrow="6" pageId="4" pageNumber="300">
two setae;
<emphasis id="B97AEAF3FFCAAA35FDCB0C3EFD578B62" box="[610,657,1849,1873]" italics="true" pageId="4" pageNumber="300">cha</emphasis>
(4) spiniform, barbed apically,
<emphasis id="B97AEAF3FFCAAA35FC4B0C3EFBD78B62" box="[994,1041,1849,1873]" italics="true" pageId="4" pageNumber="300">chb</emphasis>
(16) falciform, dorsally slightly barbed.
</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0C5BFA258B47" box="[119,1507,1884,1908]" gridrow="7" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0C5BFA258B47" box="[119,1507,1884,1908]" gridcol="0" gridrow="7" pageId="4" pageNumber="300">Trägårdhs organ not observed.</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0C79FA258BA5" box="[119,1507,1918,1942]" gridrow="8" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0C79FA258BA5" box="[119,1507,1918,1942]" gridcol="0" gridrow="8" pageId="4" pageNumber="300">
<emphasis id="B97AEAF3FFCAAA35FDBA0C79FC0B8BA5" box="[531,973,1918,1942]" italics="true" pageId="4" pageNumber="300">Epimeral and lateral podosomal regions</emphasis>
(Figs 2b, 4e) Epimeral setal formula: 3-1-3-3.
</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0CA7FA258B8B" box="[119,1507,1952,1976]" gridrow="9" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0CA7FA258B8B" box="[119,1507,1952,1976]" gridcol="0" gridrow="9" pageId="4" pageNumber="300">
Epimeral setae
<emphasis id="B97AEAF3FFCAAA35FD390CA7FD6C8B8B" box="[656,682,1952,1976]" italics="true" pageId="4" pageNumber="300">1a</emphasis>
,
<emphasis id="B97AEAF3FFCAAA35FD110CA7FD1F8B8B" box="[696,729,1952,1976]" italics="true" pageId="4" pageNumber="300">2a</emphasis>
and
<emphasis id="B97AEAF3FFCAAA35FCAE0CA7FCE18B8B" box="[775,807,1952,1976]" italics="true" pageId="4" pageNumber="300">3a</emphasis>
(4) spiniform, barbed apically;
<emphasis id="B97AEAF3FFCAAA35FBDD0CA7FB4B8B8B" box="[1140,1165,1952,1976]" italics="true" pageId="4" pageNumber="300">1c</emphasis>
,
<emphasis id="B97AEAF3FFCAAA35FB320CA7FB7E8B8B" box="[1179,1208,1952,1976]" italics="true" pageId="4" pageNumber="300">4b</emphasis>
(20) and
<emphasis id="B97AEAF3FFCAAA35FAB40CA7FAFE8B8B" box="[1309,1336,1952,1976]" italics="true" pageId="4" pageNumber="300">1b</emphasis>
,
<emphasis id="B97AEAF3FFCAAA35FAEC0CA7FA998B8B" box="[1349,1375,1952,1976]" italics="true" pageId="4" pageNumber="300">3b</emphasis>
,
<emphasis id="B97AEAF3FFCAAA35FAC40CA7FA408B8B" box="[1389,1414,1952,1976]" italics="true" pageId="4" pageNumber="300">3c</emphasis>
,
<emphasis id="B97AEAF3FFCAAA35FA3D0CA7FA688B8B" box="[1428,1454,1952,1976]" italics="true" pageId="4" pageNumber="300">4a</emphasis>
,
<emphasis id="B97AEAF3FFCAAA35FA150CA7FA138B8B" box="[1468,1493,1952,1976]" italics="true" pageId="4" pageNumber="300">4c</emphasis>
</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0CC4FA258BE8" box="[119,1507,1987,2011]" gridrow="10" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0CC4FA258BE8" box="[119,1507,1987,2011]" gridcol="0" gridrow="10" pageId="4" pageNumber="300">(2832) setiform, slightly barbed.</td>
</tr>
<tr id="353E34A3FFCA55CEFFDE0CE2FA258BCE" box="[119,1507,2021,2045]" gridrow="11" pageId="4" pageNumber="300">
<td id="76EF5DDFFFCA55CEFFDE0CE2FA258BCE" box="[119,1507,2021,2045]" gridcol="0" gridrow="11" pageId="4" pageNumber="300">
<emphasis id="B97AEAF3FFCAAA35FDBA0CE2FD1D8BCE" box="[531,731,2021,2045]" italics="true" pageId="4" pageNumber="300">Anogenital region</emphasis>
(Figs 2b, 4g, 4h) Nine or 10 pairs of genital (5761) and one pair of
</td>
</tr>
</table>
</paragraph>
<caption id="DF716669FFCBAA34FFDE0CC6FA1B8BE4" ID-DOI="http://doi.org/10.5281/zenodo.5392452" ID-Zenodo-Dep="5392452" box="[119,1501,1985,2007]" httpUri="https://zenodo.org/record/5392452/files/figure.png" pageId="5" pageNumber="301" startId="5.[119,190,1985,2006]" targetBox="[213,1389,224,1853]" targetPageId="5">
<paragraph id="8BB136E1FFCBAA34FFDE0CC6FA1B8BE4" blockId="5.[119,1525,1985,2052]" box="[119,1501,1985,2007]" pageId="5" pageNumber="301">
<emphasis id="B97AEAF3FFCBAA34FFDE0CC6FF168BE5" bold="true" box="[119,208,1985,2006]" pageId="5" pageNumber="301">Figure 3</emphasis>
<taxonomicName id="4C0E4D62FFCBAA34FF710CC6FE638BE4" authority="Ermilov &amp; Liao, 2021" authorityName="Ermilov &amp; Liao" authorityYear="2021" box="[216,421,1985,2007]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="301" phylum="Arthropoda" rank="species" species="nivnu" status="sp. nov.">
<emphasis id="B97AEAF3FFCBAA34FF710CC6FE638BE4" box="[216,421,1985,2007]" italics="true" pageId="5" pageNumber="301">Tyrphonothrus nivnu</emphasis>
</taxonomicName>
<emphasis id="B97AEAF3FFCBAA34FE0C0CC6FE1D8BE5" bold="true" box="[421,475,1985,2006]" pageId="5" pageNumber="301">
<taxonomicNameLabel id="A2495788FFCBAA34FE0C0CC6FE1D8BE5" box="[421,475,1985,2006]" pageId="5" pageNumber="301" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult: a subcapitulum, ventral view; b palp, right, antiaxial view; c chelicera, right, antiaxial view;
</paragraph>
</caption>
<caption id="DF716669FFC8AA37FFDE0C76FEE28BF9" ID-DOI="http://doi.org/10.5281/zenodo.5392454" ID-Zenodo-Dep="5392454" httpUri="https://zenodo.org/record/5392454/files/figure.png" pageId="6" pageNumber="302" startId="6.[119,190,1905,1926]" targetBox="[95,1501,221,1848]" targetPageId="6">
<paragraph id="8BB136E1FFC8AA37FFDE0C76FEE28BF9" blockId="6.[118,1517,1905,1994]" pageId="6" pageNumber="302">
<emphasis id="B97AEAF3FFC8AA37FFDE0C76FF158BB5" bold="true" box="[119,211,1905,1926]" pageId="6" pageNumber="302">Figure 4</emphasis>
<taxonomicName id="4C0E4D62FFC8AA37FF490C76FE748BB4" authority="Ermilov &amp; Liao, 2021" authorityName="Ermilov &amp; Liao" authorityYear="2021" box="[224,434,1905,1927]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="302" phylum="Arthropoda" rank="species" species="nivnu" status="sp. nov.">
<emphasis id="B97AEAF3FFC8AA37FF490C76FE748BB4" box="[224,434,1905,1927]" italics="true" pageId="6" pageNumber="302">Tyrphonothrus nivnu</emphasis>
</taxonomicName>
<emphasis id="B97AEAF3FFC8AA37FE1B0C76FE2D8BB5" bold="true" box="[434,491,1905,1926]" pageId="6" pageNumber="302">
<taxonomicNameLabel id="A2495788FFC8AA37FE1B0C76FE2D8BB5" box="[434,491,1905,1926]" pageId="6" pageNumber="302" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult, SEM micrographs (a-d) and microscope images (e-i): a dorsal view; b lateral view; c tuberculate notogastral cerotegument; d claws of leg tarsus I; e epimeral setae
<emphasis id="B97AEAF3FFC8AA37FC0B0C80FC068BAE" box="[930,960,1927,1949]" italics="true" pageId="6" pageNumber="302">3a</emphasis>
and
<emphasis id="B97AEAF3FFC8AA37FC450C80FBC28BAE" box="[1004,1028,1927,1949]" italics="true" pageId="6" pageNumber="302">3b</emphasis>
; f epimeral integument (deep focus); g, h genital plates (superficial and deep focus, respectively); i claws of leg tarsus III. Scale bar 200 μm (a, b), 10 μm (c), 50 μm (d); images without scale bar.
</paragraph>
</caption>
<caption id="DF716669FFC9AA36FDAA0BFFFB6B8D3E" box="[515,1197,247,270]" pageId="7" pageNumber="303" startId="7.[515,578,248,269]">
<paragraph id="8BB136E1FFC9AA36FDAA0BFFFB6B8D3E" blockId="7.[515,1197,247,270]" box="[515,1197,247,270]" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDAA0BFFFD928D3E" bold="true" box="[515,596,248,269]" pageId="7" pageNumber="303">Table 1</emphasis>
Leg setation and solenidia of adult
<taxonomicName id="4C0E4D62FFC9AA36FC030BF0FBB18D3E" authority="Ermilov &amp; Liao, 2021" authorityName="Ermilov &amp; Liao" authorityYear="2021" box="[938,1143,247,269]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="303" phylum="Arthropoda" rank="species" species="nivnu" status="sp. nov.">
<emphasis id="B97AEAF3FFC9AA36FC030BF0FBB18D3E" box="[938,1143,247,269]" italics="true" pageId="7" pageNumber="303">Tyrphonothrus nivnu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2495788FFC9AA36FBDE0BFFFB6B8D3E" box="[1143,1197,248,269]" pageId="7" pageNumber="303" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BB136E1FFC9AA36FE5A0A51FAEC8E35" pageId="7" pageNumber="303">
<table id="F90EC441FFC955CEFE5A0A51FA5C8E35" box="[499,1434,342,518]" colsContinueIn="12.[103,1443,343,535]" gridcols="6" gridrows="5" pageId="7" pageNumber="303">
<tr id="353E34A3FFC955CEFE5A0A51FA5C8D5F" box="[499,1434,342,364]" gridrow="0" pageId="7" pageNumber="303">
<th id="76EF5DDFFFC955CEFE5A0A51FDDE8D5F" box="[499,536,342,364]" gridcol="0" gridrow="0" pageId="7" pageNumber="303">Leg</th>
<th id="76EF5DDFFFC955CEFDE70A51FDBD8D5F" box="[590,635,342,364]" gridcol="1" gridrow="0" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDE70A50FDA38D5F" box="[590,613,343,364]" italics="true" pageId="7" pageNumber="303">Tr</emphasis>
</th>
<th id="76EF5DDFFFC955CEFD3C0A51FCD18D5F" box="[661,791,342,364]" gridcol="2" gridrow="0" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FD3C0A50FD688D5F" box="[661,686,343,364]" italics="true" pageId="7" pageNumber="303">Fe</emphasis>
</th>
<th id="76EF5DDFFFC955CEFCEB0A51FC598D5F" box="[834,927,342,364]" gridcol="3" gridrow="0" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FCEB0A51FC988D5F" box="[834,862,342,364]" italics="true" pageId="7" pageNumber="303">Ge</emphasis>
</th>
<th id="76EF5DDFFFC955CEFC680A51FBE68D5F" box="[961,1056,342,364]" gridcol="4" gridrow="0" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FC680A50FC138D5F" box="[961,981,343,364]" italics="true" pageId="7" pageNumber="303">Ti</emphasis>
</th>
<th id="76EF5DDFFFC955CEFBE10A51FA5C8D5F" box="[1096,1434,342,364]" gridcol="5" gridrow="0" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FBE10A50FBA78D5F" box="[1096,1121,343,364]" italics="true" pageId="7" pageNumber="303">Ta</emphasis>
</th>
</tr>
<tr id="353E34A3FFC955CEFE5A0A7CFA5C8DA7" box="[499,1434,379,404]" gridrow="1" pageId="7" pageNumber="303">
<th id="76EF5DDFFFC955CEFE5A0A7CFDDE8DA7" box="[499,536,379,404]" gridcol="0" gridrow="1" pageId="7" pageNumber="303">I</th>
<td id="76EF5DDFFFC955CEFDE70A7CFDBD8DA7" box="[590,635,379,404]" gridcol="1" gridrow="1" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDE70A7AFDA68DA0" box="[590,608,381,403]" italics="true" pageId="7" pageNumber="303">v</emphasis>
</td>
<td id="76EF5DDFFFC955CEFD3C0A7CFCD18DA7" box="[661,791,379,404]" gridcol="2" gridrow="1" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FD3C0A7AFCD18DA0" box="[661,791,381,403]" italics="true" pageId="7" pageNumber="303">d, (l), bv”, v”</emphasis>
</td>
<td id="76EF5DDFFFC955CEFCEB0A7CFC598DA7" box="[834,927,379,404]" gridcol="3" gridrow="1" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FCEB0A7AFC488DA0" box="[834,910,381,403]" italics="true" pageId="7" pageNumber="303">(l), v, d</emphasis>
σ
</td>
<td id="76EF5DDFFFC955CEFC680A7CFBE68DA7" box="[961,1056,379,404]" gridcol="4" gridrow="1" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FC680A7AFBC88DA0" box="[961,1038,381,403]" italics="true" pageId="7" pageNumber="303">(l), v, d</emphasis>
φ
</td>
<td id="76EF5DDFFFC955CEFBE10A7CFA5C8DA7" box="[1096,1434,379,404]" gridcol="5" gridrow="1" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FBE10A7CFAC88DA2" box="[1096,1294,379,401]" italics="true" pageId="7" pageNumber="303">(ft), (tc), (p), (u), (a),</emphasis>
ɛ, ω1, ω2, ω3
</td>
</tr>
<tr id="353E34A3FFC955CEFE5A0AA4FA5C8D8A" box="[499,1434,419,441]" gridrow="2" pageId="7" pageNumber="303">
<th id="76EF5DDFFFC955CEFE5A0AA4FDDE8D8A" box="[499,536,419,441]" gridcol="0" gridrow="2" pageId="7" pageNumber="303">II</th>
<td id="76EF5DDFFFC955CEFDE70AA4FDBD8D8A" box="[590,635,419,441]" gridcol="1" gridrow="2" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDE70AA4FDA68D8A" box="[590,608,419,441]" italics="true" pageId="7" pageNumber="303">v</emphasis>
</td>
<td id="76EF5DDFFFC955CEFD3C0AA4FCD18D8A" box="[661,791,419,441]" gridcol="2" gridrow="2" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FD3C0AA4FCD18D8A" box="[661,791,419,441]" italics="true" pageId="7" pageNumber="303">d, (l), bv”, v”</emphasis>
</td>
<td id="76EF5DDFFFC955CEFCEB0AA4FC598D8A" box="[834,927,419,441]" gridcol="3" gridrow="2" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FCEB0AA4FCB68D8A" box="[834,880,419,441]" italics="true" pageId="7" pageNumber="303">(l), d</emphasis>
σ
</td>
<td id="76EF5DDFFFC955CEFC680AA4FBE68D8A" box="[961,1056,419,441]" gridcol="4" gridrow="2" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FC680AA4FBC88D8A" box="[961,1038,419,441]" italics="true" pageId="7" pageNumber="303">(l), v, d</emphasis>
φ
</td>
<td id="76EF5DDFFFC955CEFBE10AA4FA5C8D8A" box="[1096,1434,419,441]" gridcol="5" gridrow="2" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FBE10AA4FAC88D8A" box="[1096,1294,419,441]" italics="true" pageId="7" pageNumber="303">(ft), (tc), (p), (u), (a),</emphasis>
ω
</td>
</tr>
<tr id="353E34A3FFC955CEFE5A0ACEFA5C8DEC" box="[499,1434,457,479]" gridrow="3" pageId="7" pageNumber="303">
<th id="76EF5DDFFFC955CEFE5A0ACEFDDE8DEC" box="[499,536,457,479]" gridcol="0" gridrow="3" pageId="7" pageNumber="303">III</th>
<td id="76EF5DDFFFC955CEFDE70ACEFDBD8DEC" box="[590,635,457,479]" gridcol="1" gridrow="3" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDE70ACEFDBD8DEC" box="[590,635,457,479]" italics="true" pageId="7" pageNumber="303">l, v</emphasis>
</td>
<td id="76EF5DDFFFC955CEFD3C0ACEFCD18DEC" box="[661,791,457,479]" gridcol="2" gridrow="3" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FD3C0ACEFD238DEC" box="[661,741,457,479]" italics="true" pageId="7" pageNumber="303">d, l, ev</emphasis>
</td>
<td id="76EF5DDFFFC955CEFCEB0ACEFC598DEC" box="[834,927,457,479]" gridcol="3" gridrow="3" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FCEB0ACEFC888DEC" box="[834,846,457,479]" italics="true" pageId="7" pageNumber="303">d</emphasis>
</td>
<td id="76EF5DDFFFC955CEFC680ACEFBE68DEC" box="[961,1056,457,479]" gridcol="4" gridrow="3" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FC680ACEFC2A8DEC" box="[961,1004,457,479]" italics="true" pageId="7" pageNumber="303">v, d</emphasis>
φ
</td>
<td id="76EF5DDFFFC955CEFBE10ACEFA5C8DEC" box="[1096,1434,457,479]" gridcol="5" gridrow="3" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FBE10ACEFAC18DEC" box="[1096,1287,457,479]" italics="true" pageId="7" pageNumber="303">(ft), (tc), (p), (u), (a)</emphasis>
</td>
</tr>
<tr id="353E34A3FFC955CEFE5A0AF7FA5C8E35" box="[499,1434,496,518]" gridrow="4" pageId="7" pageNumber="303">
<th id="76EF5DDFFFC955CEFE5A0AF7FDDE8E35" box="[499,536,496,518]" gridcol="0" gridrow="4" pageId="7" pageNumber="303">IV</th>
<td id="76EF5DDFFFC955CEFDE70AF7FDBD8E35" box="[590,635,496,518]" gridcol="1" gridrow="4" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDE70AF7FDA68E35" box="[590,608,496,518]" italics="true" pageId="7" pageNumber="303">v</emphasis>
</td>
<td id="76EF5DDFFFC955CEFD3C0AF7FCD18E35" box="[661,791,496,518]" gridcol="2" gridrow="4" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FD3C0AF7FD0C8E35" box="[661,714,496,518]" italics="true" pageId="7" pageNumber="303">d, ev</emphasis>
</td>
<td id="76EF5DDFFFC955CEFCEB0AF7FC598E35" box="[834,927,496,518]" gridcol="3" gridrow="4" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FCEB0AF7FC888E35" box="[834,846,496,518]" italics="true" pageId="7" pageNumber="303">d</emphasis>
</td>
<td id="76EF5DDFFFC955CEFC680AF7FBE68E35" box="[961,1056,496,518]" gridcol="4" gridrow="4" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FC680AF7FC2D8E35" box="[961,1003,496,518]" italics="true" pageId="7" pageNumber="303">d, v</emphasis>
</td>
<td id="76EF5DDFFFC955CEFBE10AF7FA5C8E35" box="[1096,1434,496,518]" gridcol="5" gridrow="4" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FBE10AF7FB358E35" box="[1096,1267,496,518]" italics="true" pageId="7" pageNumber="303">ft”, (tc), (p), (u), s</emphasis>
, (pv)
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB136E1FFC9AA36FE5B0915FB1E8E6F" blockId="7.[498,1447,530,604]" pageId="7" pageNumber="303">
<tableNote id="76E8376FFFC9AA36FE5B0915FB1E8E6F" pageId="7" pageNumber="303" targetBox="[499,1434,342,518]" targetPageId="7">
Note: Roman letters refer to normal setae (except ɛ = famulus); Greek letters refer to solenidia. Single quotation mark (
<emphasis id="B97AEAF3FFC9AA36FD82092AFDF48E73" box="[555,562,557,576]" italics="true" pageId="7" pageNumber="303"></emphasis>
) marks setae on the anterior and double quotation mark (
<emphasis id="B97AEAF3FFC9AA36FBB7092AFBEC8E73" box="[1054,1066,557,576]" italics="true" pageId="7" pageNumber="303"></emphasis>
) setae on the posterior side of a given leg segment;
<emphasis id="B97AEAF3FFC9AA36FDED094EFD898E6F" box="[580,591,585,604]" italics="true" pageId="7" pageNumber="303">d</emphasis>
φ and
<emphasis id="B97AEAF3FFC9AA36FD20094EFD528E6F" box="[649,660,585,604]" italics="true" pageId="7" pageNumber="303">d</emphasis>
σ seta and solenidion coupled; parentheses refer to a pair of setae.
</tableNote>
</paragraph>
<paragraph id="8BB136E1FFC9AA36FE4209A6FD9D8ECD" blockId="7.[491,1559,673,1497]" pageId="7" pageNumber="303">anal (20) setae developed, setiform, roughened; all genital seta directed backwards. Three pairs of adanal setae (4953) setiform, barbed. Adanal lyrifissure distinct, anal lyrifissure not observed.</paragraph>
<paragraph id="8BB136E1FFC9AA36FDBA080DFD838F98" blockId="7.[491,1559,673,1497]" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDBA080DFD818F12" box="[531,583,778,801]" italics="true" pageId="7" pageNumber="303">Legs</emphasis>
(
<figureCitation id="13352A64FFC9AA36FDFF080EFD668F12" box="[598,672,777,801]" captionStart="Figure 3" captionStartId="5.[119,190,1985,2006]" captionTargetBox="[213,1389,224,1853]" captionTargetId="figure-2@5.[193,1402,221,1931]" captionTargetPageId="5" captionText="Figure 3 Tyrphonothrus nivnun. sp., adult: a subcapitulum, ventral view; b palp, right, antiaxial view; c chelicera, right, antiaxial view;" figureDoi="http://doi.org/10.5281/zenodo.5392452" httpUri="https://zenodo.org/record/5392452/files/figure.png" pageId="7" pageNumber="303">Figs 3</figureCitation>
d-g, 4b, 4d, 4i) Heterotridactylous; claws mediodistally with slight longitudinal stria, dorsally slightly and rarely barbed. Formulas of leg setation and solenidia: I (154411) [113], II (153410) [111], III (231210) [010], IV (121210) [000]; homology of setae and solenidia indicated in
<tableCitation id="C68C035AFFC9AA36FBEF0877FB518FBB" box="[1094,1175,880,904]" captionStart="Table 1" captionStartId="7.[515,578,248,269]" captionText="Table 1 Leg setation and solenidia of adultTyrphonothrus nivnun. sp." pageId="7" pageNumber="303">Table 1</tableCitation>
. All solenidia thickened, rounded apically.
</paragraph>
</subSubSection>
<subSubSection id="C314656AFFC9AA36FDBA08B1FC2E886B" pageId="7" pageNumber="303" type="materials_examined">
<paragraph id="8BB136E1FFC9AA36FDBA08B1FA728FFE" blockId="7.[491,1559,673,1497]" box="[531,1460,949,974]" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDBA08B1FD328FFD" bold="true" box="[531,756,950,974]" pageId="7" pageNumber="303">Material examined</emphasis>
<typeStatus id="54B58843FFC9AA36FCBE08B2FCBC8FFE" box="[791,890,949,973]" pageId="7" pageNumber="303" type="holotype">Holotype</typeStatus>
(female) and
<specimenCount id="9D08FD68FFC9AA36FBB008B2FB758FFD" box="[1049,1203,949,974]" pageId="7" pageNumber="303" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
(all females):
<date id="FFB01021FFC9AA36FAFF08B2FA768FFE" box="[1366,1456,949,973]" pageId="7" pageNumber="303" value="2018-07-21">2018.7.21</date>
,
</paragraph>
<paragraph id="8BB136E1FFC9AA36FE4208DFFD2B8820" blockId="7.[491,1559,673,1497]" pageId="7" pageNumber="303">
<collectingRegion id="49CAF803FFC9AA36FE4208DFFD4B8FC3" box="[491,653,984,1008]" country="Taiwan" name="Chiayi" pageId="7" pageNumber="303">Chiayi County</collectingRegion>
, Zhuqi Township, Shizhuo,
<quantity id="4CF69B04FFC9AA36FC6308DFFBD58FC3" box="[970,1043,984,1008]" metricMagnitude="3" metricUnit="m" metricValue="1.528" pageId="7" pageNumber="303" unit="m" value="1528.0">1528 m</quantity>
a.s.l., soil,
<geoCoordinate id="EE3A5026FFC9AA36FB3B08DFFADC8FC3" box="[1170,1306,979,1008]" degrees="23" direction="north" minutes="29.232" orientation="latitude" pageId="7" pageNumber="303" precision="1" value="23.4872">
23°29.232
<emphasis id="B97AEAF3FFC9AA36FB5208D4FAC58FD6" box="[1275,1283,979,997]" italics="true" pageId="7" pageNumber="303">
<superScript id="7C7B9BA9FFC9AA36FB5208D4FAC58FD6" attach="right" box="[1275,1283,979,997]" fontSize="7" pageId="7" pageNumber="303"></superScript>
</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE3A5026FFC9AA36FA8F08DFFA7B8FC3" box="[1318,1469,979,1008]" degrees="120" direction="east" minutes="42.221" orientation="longitude" pageId="7" pageNumber="303" precision="1" value="120.70368">
120°42.221
<emphasis id="B97AEAF3FFC9AA36FA0B08D4FA6C8FD6" box="[1442,1450,979,997]" italics="true" pageId="7" pageNumber="303">
<superScript id="7C7B9BA9FFC9AA36FA0B08D4FA6C8FD6" attach="right" box="[1442,1450,979,997]" fontSize="7" pageId="7" pageNumber="303"></superScript>
</emphasis>
E
</geoCoordinate>
, collected by H.C. Liao.
</paragraph>
<paragraph id="8BB136E1FFC9AA36FDBA0F1AFC2E886B" blockId="7.[491,1559,673,1497]" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDBA0F1AFD018806" bold="true" box="[531,711,1053,1077]" pageId="7" pageNumber="303">Type deposition</emphasis>
— The
<typeStatus id="54B58843FFC9AA36FC880F1AFC478806" box="[801,897,1053,1077]" pageId="7" pageNumber="303" type="holotype">holotype</typeStatus>
is deposited in the collection of the NTU;
<specimenCount id="9D08FD68FFC9AA36FAF10F1AFA378805" box="[1368,1521,1053,1078]" pageId="7" pageNumber="303" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
are deposited in the collection of the
<collectionCode id="ED1FAE24FFC9AA36FC2F0F47FC22886B" box="[902,996,1088,1112]" pageId="7" pageNumber="303">TSUMZ</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C314656AFFC9AA36FDBA0F64FA5D88AE" pageId="7" pageNumber="303" type="etymology">
<paragraph id="8BB136E1FFC9AA36FDBA0F64FA5D88AE" blockId="7.[491,1559,673,1497]" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDBA0F64FD548848" bold="true" box="[531,658,1123,1147]" pageId="7" pageNumber="303">Etymology</emphasis>
— The species name
<taxonomicName id="4C0E4D62FFC9AA36FC290F64FC788849" authorityName="Ermilov &amp; Liao" authorityYear="2021" box="[896,958,1123,1146]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="303" phylum="Arthropoda" rank="species" species="nivnu">
<emphasis id="B97AEAF3FFC9AA36FC290F64FC788849" box="[896,958,1123,1146]" italics="true" pageId="7" pageNumber="303">nivnu</emphasis>
</taxonomicName>
refers to Nivnu, Goddess of
<collectingCountry id="F3197671FFC9AA36FAAA0F65FA978849" box="[1283,1361,1122,1146]" name="Taiwan" pageId="7" pageNumber="303">Taiwan</collectingCountry>
indigenous people Cou, who is the creator of the universe, heaven and earth, in the Cous legend.
</paragraph>
</subSubSection>
<subSubSection id="C314656AFFC9AA36FDBA0FAFFBDE89E0" pageId="7" pageNumber="303" type="discussion">
<paragraph id="8BB136E1FFC9AA36FDBA0FAFFA7B897A" blockId="7.[491,1559,673,1497]" pageId="7" pageNumber="303">
<emphasis id="B97AEAF3FFC9AA36FDBA0FAFFD4488F3" bold="true" box="[531,642,1192,1216]" pageId="7" pageNumber="303">Remarks</emphasis>
—In general morphological traits (notogastral ridges developed; tridactylous legs; notogastral setae
<emphasis id="B97AEAF3FFC9AA36FD4C0FCCFD3788D1" box="[741,753,1227,1250]" italics="true" pageId="7" pageNumber="303">e</emphasis>
<subScript id="178A34A4FFC9AA36FD590FD1FD3088D4" attach="left" box="[752,758,1238,1255]" fontSize="7" pageId="7" pageNumber="303">2</subScript>
,
<emphasis id="B97AEAF3FFC9AA36FCA20FCDFCDE88D1" box="[779,792,1226,1250]" italics="true" pageId="7" pageNumber="303">h</emphasis>
<subScript id="178A34A4FFC9AA36FCB00FD1FCE488D4" attach="left" box="[793,802,1238,1255]" fontSize="7" pageId="7" pageNumber="303">1</subScript>
,
<emphasis id="B97AEAF3FFC9AA36FC9A0FCDFC8688D1" box="[819,832,1226,1250]" italics="true" pageId="7" pageNumber="303">h</emphasis>
<subScript id="178A34A4FFC9AA36FCE80FD1FC8188D4" attach="left" box="[833,839,1238,1255]" fontSize="7" pageId="7" pageNumber="303">2</subScript>
, and
<emphasis id="B97AEAF3FFC9AA36FC220FCCFC5E88D1" box="[907,920,1227,1250]" italics="true" pageId="7" pageNumber="303">p</emphasis>
<subScript id="178A34A4FFC9AA36FC310FD1FC5888D4" attach="left" box="[920,926,1238,1255]" fontSize="7" pageId="7" pageNumber="303">2</subScript>
long, setiform, with attenuate tip, other setae of medium length, setiform; high number of genital setae; absence of notogastral ornamentation; and comparatively large body size), the new species is morphologically most similar to
<taxonomicName id="4C0E4D62FFC9AA36FE420E36FCBE897A" authority="(Piffl, 1971)" baseAuthorityName="Piffl" baseAuthorityYear="1971" box="[491,888,1329,1353]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="303" phylum="Arthropoda" rank="species" species="altissimus">
<emphasis id="B97AEAF3FFC9AA36FE420E36FCC7897A" box="[491,769,1329,1353]" italics="true" pageId="7" pageNumber="303">Tyrphonothrus altissimus</emphasis>
(
<bibRefCitation id="EF9F4B10FFC9AA36FCA10E36FCB5897A" author="Piffl E." box="[776,883,1329,1353]" pageId="7" pageNumber="303" pagination="23 - 54" refId="ref6315" refString="Piffl E. 1971. Neue Oribatiden (Acari) aus dem Himalaya. Khumbu Himal, 4 (1): 23 - 54." type="journal article" year="1971">Piffl, 1971</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F3197671FFC9AA36FC150E36FC38897A" box="[956,1022,1329,1353]" name="Nepal" pageId="7" pageNumber="303">Nepal</collectingCountry>
,
<taxonomicName id="4C0E4D62FFC9AA36FBB10E35FA47897A" authority="(Hammer, 1961)" baseAuthorityName="Hammer" baseAuthorityYear="1961" box="[1048,1409,1329,1353]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="303" phylum="Arthropoda" rank="species" species="cajamarcensis">
<emphasis id="B97AEAF3FFC9AA36FBB10E35FB13897A" box="[1048,1237,1330,1353]" italics="true" pageId="7" pageNumber="303">T. cajamarcensis</emphasis>
(
<bibRefCitation id="EF9F4B10FFC9AA36FB720E36FABC897A" author="Hammer M." box="[1243,1402,1329,1353]" pageId="7" pageNumber="303" pagination="1 - 157" refId="ref5989" refString="Hammer M. 1961. Investigations on the oribatid fauna of the Andes Mountains. II. Peru. Det Kong. Dansk. Vidensk. Selsk. Biol. Skr., 13 (1): 1 - 157." type="journal article" year="1961">Hammer, 1961</bibRefCitation>
)
</taxonomicName>
from
</paragraph>
<paragraph id="8BB136E1FFC9AA36FE420E52FBDE89E0" blockId="7.[491,1559,673,1497]" pageId="7" pageNumber="303">
<collectingCountry id="F3197671FFC9AA36FE420E52FDDA895F" box="[491,540,1365,1388]" name="Peru" pageId="7" pageNumber="303">Peru</collectingCountry>
and North
<collectingCountry id="F3197671FFC9AA36FD350E53FD1E895F" box="[668,728,1364,1388]" name="India" pageId="7" pageNumber="303">India</collectingCountry>
and
<taxonomicName id="4C0E4D62FFC9AA36FD550E52FBBB895F" authority="(Stary and Block, 1995)" baseAuthorityName="Stary and Block" baseAuthorityYear="1995" box="[764,1149,1364,1388]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="303" phylum="Arthropoda" rank="species" species="wallworki">
<emphasis id="B97AEAF3FFC9AA36FD550E52FC40895F" box="[764,902,1364,1388]" italics="true" pageId="7" pageNumber="303">T. wallworki</emphasis>
(
<bibRefCitation id="EF9F4B10FFC9AA36FC270E53FBB1895F" author="Stary J. &amp; Block W." box="[910,1143,1364,1388]" pageId="7" pageNumber="303" refId="ref6438" refString="Stary J., Block W. 1995. Oribatid mites (Acari: Oribatida) of South Georgia, South Atlantic. J. Nat. Hist.," type="book" year="1995">Starý and Block, 1995</bibRefCitation>
)
</taxonomicName>
from the
<collectingCountry id="F3197671FFC9AA36FB4F0E53FA4E895F" box="[1254,1416,1364,1388]" name="South Georgia and the South Sandwich Islands" pageId="7" pageNumber="303">South Georgia</collectingCountry>
Islands, however it differs from these listed species by the barbed notogastral setae (versus smooth) and comparatively long (
<emphasis id="B97AEAF3FFC9AA36FD630E9DFD108982" box="[714,726,1434,1457]" italics="true" pageId="7" pageNumber="303">c</emphasis>
<subScript id="178A34A4FFC9AA36FD7F0EA2FD198985" attach="left" box="[726,735,1445,1462]" fontSize="7" pageId="7" pageNumber="303">1</subScript>
about 4/5 length of distance
<emphasis id="B97AEAF3FFC9AA36FBB10E9DFBE28982" box="[1048,1060,1434,1457]" italics="true" pageId="7" pageNumber="303">c</emphasis>
<subScript id="178A34A4FFC9AA36FB8D0EA2FBEB8985" attach="left" box="[1060,1069,1445,1462]" fontSize="7" pageId="7" pageNumber="303">1</subScript>
<emphasis id="B97AEAF3FFC9AA36FB950E9EFB8C8982" box="[1084,1098,1433,1457]" italics="true" pageId="7" pageNumber="303">d</emphasis>
<subScript id="178A34A4FFC9AA36FBE30EA2FB958985" attach="left" box="[1098,1107,1445,1462]" fontSize="7" pageId="7" pageNumber="303">1</subScript>
) notogastral setae
<emphasis id="B97AEAF3FFC9AA36FAB60E9DFAED8982" box="[1311,1323,1434,1457]" italics="true" pageId="7" pageNumber="303">c</emphasis>
<subScript id="178A34A4FFC9AA36FA830EA2FAF58985" attach="left" box="[1322,1331,1445,1462]" fontSize="7" pageId="7" pageNumber="303">1</subScript>
,
<emphasis id="B97AEAF3FFC9AA36FAEA0E9DFA898982" box="[1347,1359,1434,1457]" italics="true" pageId="7" pageNumber="303">c</emphasis>
<subScript id="178A34A4FFC9AA36FAE60EA2FA938985" attach="left" box="[1359,1365,1445,1462]" fontSize="7" pageId="7" pageNumber="303">2</subScript>
,
<emphasis id="B97AEAF3FFC9AA36FACE0E9DFAB58982" box="[1383,1395,1434,1457]" italics="true" pageId="7" pageNumber="303">c</emphasis>
<subScript id="178A34A4FFC9AA36FADA0EA2FABF8985" attach="left" box="[1395,1401,1445,1462]" fontSize="7" pageId="7" pageNumber="303">3</subScript>
, and
<emphasis id="B97AEAF3FFC9AA36FA100E9DFA038982" box="[1465,1477,1434,1457]" italics="true" pageId="7" pageNumber="303">e</emphasis>
<subScript id="178A34A4FFC9AA36FA6C0EA2FA088985" attach="left" box="[1477,1486,1445,1462]" fontSize="7" pageId="7" pageNumber="303">1</subScript>
(versus short,
<emphasis id="B97AEAF3FFC9AA36FD290EBBFD4A89E0" box="[640,652,1468,1491]" italics="true" pageId="7" pageNumber="303">c</emphasis>
<subScript id="178A34A4FFC9AA36FD250ECFFD5389EA" attach="left" box="[652,661,1480,1497]" fontSize="7" pageId="7" pageNumber="303">1</subScript>
about 1/2 length of distance
<emphasis id="B97AEAF3FFC9AA36FC620EBBFC1189E0" box="[971,983,1468,1491]" italics="true" pageId="7" pageNumber="303">c</emphasis>
<subScript id="178A34A4FFC9AA36FC7E0ECFFC2689EA" attach="left" box="[983,992,1480,1497]" fontSize="7" pageId="7" pageNumber="303">1</subScript>
<emphasis id="B97AEAF3FFC9AA36FC460EBCFC3B89E0" box="[1007,1021,1467,1491]" italics="true" pageId="7" pageNumber="303">d</emphasis>
<subScript id="178A34A4FFC9AA36FC540ECFFBC089EA" attach="left" box="[1021,1030,1480,1497]" fontSize="7" pageId="7" pageNumber="303">1</subScript>
).
</paragraph>
</subSubSection>
</treatment>
</document>