209 lines
34 KiB
XML
209 lines
34 KiB
XML
<document id="97E38DFA558E3CFDD81539A90E7DF548" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFF1E50250C685BB6677FAF5" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) nigrihirtus Assing 2017, spec. nov." docType="treatment" docUuid="4BCA38DF-D00B-4833-81D4-05557C34488E" docUuidSource="ZooBank" docVersion="10" lastPageNumber="165" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="165" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="93E5A6659D183F7F5DA9D8A51BF7EA0F">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="2245228D21BF016BD5189E26A124C712">Assing, Volker</mods:namePart>
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<mods:title id="5CFAFCBB19CEE34FBD90B740A46D1A1B">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="1948712B8F672401DED5B4D1717C0ABE">
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<mods:date id="6D1A3F99771B71BA8297DCCCDEF7D18D">2017</mods:date>
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<mods:number id="7CEAB3DA579B10A0D25FCDFBAEF758E3">2017-06-30</mods:number>
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<mods:number id="D51454D59336161977F7F8A87BA401F7">67</mods:number>
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<treatment id="03A787BAFFF1E50250C685BB6677FAF5" ID-DOI="http://doi.org/10.5281/zenodo.5888442" ID-GBIF-Taxon="192498203" ID-Zenodo-Dep="5888442" LSID="urn:lsid:zoobank.org:act:4BCA38DF-D00B-4833-81D4-05557C34488E" httpUri="http://treatment.plazi.org/id/03A787BAFFF1E50250C685BB6677FAF5" lastPageNumber="165" pageId="48" pageNumber="165">
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<subSubSection id="C3146527FFF1E50250C685BB61E0FD5B" pageId="48" pageNumber="165" type="nomenclature">
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<paragraph id="8BB136ACFFF1E50250C685BB6149FD7A" blockId="48.[227,747,691,783]" box="[227,578,691,718]" pageId="48" pageNumber="165">
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<heading id="D0F981C0FFF1E50250C685BB6149FD7A" box="[227,578,691,718]" fontSize="12" level="2" pageId="48" pageNumber="165" reason="2">
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<taxonomicName id="4C0E4D2FFFF1E50250C685BB62D9FD7A" ID-CoL="7GHYQ" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[227,466,691,718]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="nigrihirtus" status="sp. nov." subGenus="Zyras">
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<emphasis id="B97AEABEFFF1E50250C685BB6212FD7A" box="[227,281,691,718]" italics="true" pageId="48" pageNumber="165">Zyras</emphasis>
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(
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<emphasis id="B97AEABEFFF1E502510285BB6256FD7A" box="[295,349,691,718]" italics="true" pageId="48" pageNumber="165">Zyras</emphasis>
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)
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<emphasis id="B97AEABEFFF1E502514F85BB62D9FD7A" box="[362,466,691,718]" italics="true" pageId="48" pageNumber="165">nigrihirtus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24957C5FFF1E50251FC85BB6149FD7A" box="[473,578,691,718]" pageId="48" pageNumber="165" rank="species">spec. nov.</taxonomicNameLabel>
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<paragraph id="8BB136ACFFF1E50250C685D361E0FD5B" blockId="48.[227,747,691,783]" box="[227,747,731,751]" pageId="48" pageNumber="165">
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urn: lsid:zoobank.org:act:
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<uuid id="FFA80C79FFF1E50251B485D361E0FD5B" box="[401,747,731,751]" pageId="48" pageNumber="165">4BCA38DF-D00B-4833-81D4-05557C34488E</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146527FFF1E50250C685F362F8FCBB" box="[227,499,763,783]" pageId="48" pageNumber="165" type="description">
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<paragraph id="8BB136ACFFF1E50250C685F362F8FCBB" blockId="48.[227,747,691,783]" box="[227,499,763,783]" pageId="48" pageNumber="165">
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(
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<figureCitation id="13352A29FFF1E50250C285F3622BFCBB" box="[231,288,763,783]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="48" pageNumber="165">Figs 38</figureCitation>
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,
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<figureCitation id="13352A29FFF1E502510285F36237FCBB" box="[295,316,763,783]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="48" pageNumber="165">78</figureCitation>
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,
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<figureCitation id="13352A29FFF1E502516685F3626AFCBB" box="[323,353,763,783]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="48" pageNumber="165">116</figureCitation>
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,
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<figureCitation id="13352A29FFF1E502514D85F362A6FCBB" box="[360,429,763,783]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="48" pageNumber="165">255–259</figureCitation>
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,
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<figureCitation id="13352A29FFF1E502519185F362E4FCBB" box="[436,495,763,783]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="48" pageNumber="165">Map 10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3146527FFF1E502509884216261FC55" pageId="48" pageNumber="165" type="materials_examined">
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<paragraph id="8BB136ACFFF1E502509884216261FC55" blockId="48.[189,794,809,993]" pageId="48" pageNumber="165">
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<emphasis id="B97AEABEFFF1E50250988421625EFCF5" bold="true" box="[189,341,809,833]" pageId="48" pageNumber="165">Type material</emphasis>
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||
:
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<materialsCitation id="3B663CF1FFF1E502514784216261FC55" ID-GBIF-Occurrence="3450162394" collectingDate="1985-02" collectingDateMax="1985-04-31" collectingDateMin="1985-02" collectingMethod="Pit fall trap" collectionCode="BMNH" collectorName="R. Ent. Soc. Lond." determinerName="V. Assing" elevation="400" location="Dumoga-Bone N. P." pageId="48" pageNumber="165" specimenCount="1" stateProvince="SULAWESI UTARA" typeStatus="holotype">
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<typeStatus id="54B5880EFFF1E5025147842162CEFCF5" box="[354,453,809,833]" pageId="48" pageNumber="165" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D08FD25FFF1E50251E8842262E9FCF5" box="[461,482,810,833]" pageId="48" pageNumber="165">♂</specimenCount>
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[elytra missing]: “INDONESIA:
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<collectingRegion id="49CAF84EFFF1E50250CA844162C0FCD5" box="[239,459,841,865]" country="Indonesia" name="Sulawesi Utara" pageId="48" pageNumber="165">SULAWESI UTARA</collectingRegion>
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,
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<location id="8ED16077FFF1E50251F1844261AEFCD5" LSID="urn:lsid:plazi:treatment:03A787BAFFF1E50250C685BB6677FAF5:8ED16077FFF1E50251F1844261AEFCD5" box="[468,677,842,866]" name="Dumoga-Bone N. P." pageId="48" pageNumber="165" stateProvince="SULAWESI UTARA">Dumoga-Bone N.P.</location>
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,
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<date id="FFB0106CFFF1E502528B844163F8FC35" pageId="48" pageNumber="165" value="1985-02" valueMax="1985-04-31" valueMin="1985-02-00">
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<collectingDate id="EFF4E984FFF1E502528B844163F8FC35" pageId="48" pageNumber="165" value="1985-02" valueMax="1985-04-31" valueMin="1985-02-00">Feb–April 1985</collectingDate>
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</date>
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. / Plot C, ca
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<quantity id="4CF69B49FFF1E50251A7846162CCFC35" box="[386,455,873,897]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="165" unit="m" value="400.0">
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<elevation id="0023D19FFFF1E50251A7846162CCFC35" box="[386,455,873,897]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="165" unit="m" value="400.0">400 m</elevation>
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</quantity>
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,
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<location id="8ED16077FFF1E50251F68461617EFC35" LSID="urn:lsid:plazi:treatment:03A787BAFFF1E50250C685BB6677FAF5:8ED16077FFF1E50251F68461617EFC35" box="[467,629,873,897]" name="Lowland forest" pageId="48" pageNumber="165" stateProvince="SULAWESI UTARA">Lowland forest</location>
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/
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<collectingMethod id="524F4EBBFFF1E50252AB84616003FC35" box="[654,776,873,897]" pageId="48" pageNumber="165">Pit fall trap</collectingMethod>
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/ R.Ent.Soc. Lond., Project Wallace, B.M. 1985-10 / 29.123 / Holotypus ♂
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<taxonomicName id="4C0E4D2FFFF1E502515584A2612EFC75" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[368,549,937,961]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="nigrihirtus" status="sp. nov.">
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<emphasis id="B97AEABEFFF1E502515584A2612EFC75" box="[368,549,937,961]" italics="true" pageId="48" pageNumber="165">Zyras nigrihirtus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24957C5FFF1E502521784A2617AFC75" box="[562,625,938,961]" pageId="48" pageNumber="165" rank="species">sp. n.</taxonomicNameLabel>
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, det.
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<determinerName id="7E6DDA68FFF1E502529284A26012FC75" box="[695,793,937,961]" pageId="48" pageNumber="165">V. Assing</determinerName>
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2016” (
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||
<collectionCode id="ED1FAE69FFF1E502512D84C16255FC55" box="[264,350,969,993]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="48" pageNumber="165">BMNH</collectionCode>
|
||
).
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||
</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146527FFF1E5025098830161B9FB35" pageId="48" pageNumber="165" type="etymology">
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<paragraph id="8BB136ACFFF1E5025098830161B9FB35" blockId="48.[189,794,1033,1153]" pageId="48" pageNumber="165">
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<emphasis id="B97AEABEFFF1E50250988301623FFB95" bold="true" box="[189,308,1033,1057]" pageId="48" pageNumber="165">Etymology</emphasis>
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||
: The specific epithet (adjective) is composed of the Latin adjectives niger (black) and hirtus (pubescent). It alludes to the black coloration of the body and the hypothesized close relationship to
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<taxonomicName id="4C0E4D2FFFF1E5025276836261A6FB35" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[595,685,1129,1153]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="hirtus">
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<emphasis id="B97AEABEFFF1E5025276836261A6FB35" box="[595,685,1129,1153]" italics="true" pageId="48" pageNumber="165">Z. hirtus</emphasis>
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||
</taxonomicName>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFF1E502509883A167F1FB15" pageId="48" pageNumber="165" type="description">
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<paragraph id="8BB136ACFFF1E502509883A16275FA35" blockId="48.[189,795,1193,2081]" pageId="48" pageNumber="165">
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<emphasis id="B97AEABEFFF1E502509883A16234FB75" bold="true" box="[189,319,1193,1217]" pageId="48" pageNumber="165">Description</emphasis>
|
||
: Body length
|
||
<quantity id="4CF69B49FFF1E50251FA83A16138FB75" box="[479,563,1193,1217]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="48" pageNumber="165" unit="mm" value="6.8">6.8 mm</quantity>
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||
. Coloration (
|
||
<figureCitation id="13352A29FFF1E50252E383A1601EFB75" box="[710,789,1193,1217]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="48" pageNumber="165">Figs 38</figureCitation>
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,
|
||
<figureCitation id="13352A29FFF1E502509883C163D2FB55" box="[189,217,1225,1249]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="48" pageNumber="165">78</figureCitation>
|
||
,
|
||
<figureCitation id="13352A29FFF1E50250C183C16205FB55" box="[228,270,1225,1249]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="48" pageNumber="165">116</figureCitation>
|
||
): body black (note that the elytra of the
|
||
<typeStatus id="54B5880EFFF1E502529983C16011FB55" box="[700,794,1225,1249]" pageId="48" pageNumber="165" type="holotype">holotype</typeStatus>
|
||
are missing); legs pale-yellowish; antennae with antennomeres I–II pale-brown, III–VII blackish-brown, and VIII–XI gradually becoming paler, XI yellowish-brown; maxillary palpi blackish-brown, with the terminal palpomere yellow.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFF1E502509882816121F815" blockId="48.[189,795,1193,2081]" pageId="48" pageNumber="165">
|
||
Head (
|
||
<figureCitation id="13352A29FFF1E50251338281626FFA15" box="[278,356,1417,1441]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="48" pageNumber="165">Fig. 78</figureCitation>
|
||
) moderately transverse, extensively impunctate along middle; lateral dorsal portions with moderately dense and coarse punctation. Eyes large and bulging, much longer than postocular region in dorsal view. Antenna (
|
||
<figureCitation id="13352A29FFF1E5025152810162CFF995" box="[375,452,1545,1569]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="48" pageNumber="165">Fig. 38</figureCitation>
|
||
)
|
||
<quantity id="4CF69B49FFF1E50251FF81016126F995" box="[474,557,1545,1569]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="48" pageNumber="165" unit="mm" value="1.9">1.9 mm</quantity>
|
||
long; antennomeres IV approximately as long as broad, VI–X of gradually increasing width and increasingly transverse, X approximately twice as broad as long, and XI of conical shape, approximately as long as the combined length of VIII–X. Pronotum (
|
||
<figureCitation id="13352A29FFF1E502511E81A1628EF975" box="[315,389,1705,1729]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="48" pageNumber="165">Fig. 78</figureCitation>
|
||
) rather weakly transverse, 1.08 times as broad as long and 1.3 times as broad as head, broadest in anterior half; lateral margins straight in posterior twothirds (dorsal view); punctation coarse, dense, and nearly regularly distributed; midline very narrowly impunctate; lateral margins each with seven long and erect black setae; anterior and posterior margins each with one long and erect black seta on either side.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFF1E502509880A161CCF855" blockId="48.[189,795,1193,2081]" pageId="48" pageNumber="165">Hind wings fully developed. Legs very slender; metatarsomere I as long as the combined length of II–IV.</paragraph>
|
||
<paragraph id="8BB136ACFFF1E502509880E160E8FD95" blockId="48.[189,795,1193,2081]" lastBlockId="48.[831,1437,233,834]" pageId="48" pageNumber="165">
|
||
Abdomen (
|
||
<figureCitation id="13352A29FFF1E502516480E16292F7B5" box="[321,409,2025,2049]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="48" pageNumber="165">Fig. 116</figureCitation>
|
||
) with moderately deep anterior impressions on tergites III–V; anterior impression of tergite III with a transverse row of sparse, anterior impressions of tergites IV–V with a transverse row of denser non-setiferous punctures; discs of tergites III–V with fine non-setiferous punctation on whole surface and with interspersed setiferous punctures; tergites VI–VII with an anterior transverse band of dense non-setiferous punctures, on remainder of disc with less dense nonsetiferous punctures everywhere and with interspersed setiferous punctures; posterior margin of tergite VII with palisade fringe.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFF1E502531A85226690FCF5" blockId="48.[831,1437,233,834]" pageId="48" pageNumber="165">
|
||
♂: tergite VIII (
|
||
<figureCitation id="13352A29FFF1E50253CB8521674FFDF5" box="[1006,1092,553,577]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="48" pageNumber="165">Fig. 258</figureCitation>
|
||
) with numerous long and erect black setae in posterior third, posterior margin convex, without median concavity; sternite VIII (
|
||
<figureCitation id="13352A29FFF1E502552085616650FD35" box="[1285,1371,617,641]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="48" pageNumber="165">Fig. 259</figureCitation>
|
||
) with numerous long black setae in posterior half, posterior margin strongly convex; median lobe of aedeagus (
|
||
<figureCitation id="13352A29FFF1E502536285C160DFFD55" box="[839,980,713,737]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="48" pageNumber="165">Figs 255–256</figureCitation>
|
||
)
|
||
<quantity id="4CF69B49FFF1E50253C785C1673AFD55" box="[994,1073,713,737]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="48" pageNumber="165" unit="mm" value="0.9">0.9 mm</quantity>
|
||
long; ventral process of sub-triangular shape and apically acute in ventral view, at base with median carina (lateral view); paramere (
|
||
<figureCitation id="13352A29FFF1E50254FC84016624FC95" box="[1241,1327,777,801]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="48" pageNumber="165">Fig. 257</figureCitation>
|
||
) as long as median lobe and with very small and flattened apical lobe.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFF1E502531A846167F1FB15" blockId="48.[831,1436,873,1185]" pageId="48" pageNumber="165">
|
||
<emphasis id="B97AEABEFFF1E502531A84616705FC35" bold="true" box="[831,1038,873,897]" pageId="48" pageNumber="165">Comparative notes</emphasis>
|
||
: As can be inferred from the subconical shape of antennomere XI, the shape of the ventral process of the aedeagus, and particularly from the short and flattened apical lobe of the paramere,
|
||
<taxonomicName id="4C0E4D2FFFF1E502550C84C26055FBB5" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="nigrihirtus">
|
||
<emphasis id="B97AEABEFFF1E502550C84C26055FBB5" italics="true" pageId="48" pageNumber="165">Z. nigrihirtus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C0E4D2FFFF1E502542384E2676BFBB5" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[1030,1120,1001,1025]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="hirtus">
|
||
<emphasis id="B97AEABEFFF1E502542384E2676BFBB5" box="[1030,1120,1001,1025]" italics="true" pageId="48" pageNumber="165">Z. hirtus</emphasis>
|
||
</taxonomicName>
|
||
group. Among the species of this group, it is characterized particularly by the coloration, the dense and coarse punctation of the pronotum, the conspicuous punctation pattern of the abdomen, the rather sparse long setation of the forebody, and by the shape of the median lobe of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFF1E502531A83C16677FAF5" pageId="48" pageNumber="165" type="distribution">
|
||
<paragraph id="8BB136ACFFF1E502531A83C16677FAF5" blockId="48.[831,1437,1225,1345]" pageId="48" pageNumber="165">
|
||
<emphasis id="B97AEABEFFF1E502531A83C167B5FB55" bold="true" box="[831,1214,1225,1249]" pageId="48" pageNumber="165">Distribution and natural history</emphasis>
|
||
: The type locality (
|
||
<figureCitation id="13352A29FFF1E502536283E16093FAB5" box="[839,920,1257,1281]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="48" pageNumber="165">Map 10</figureCitation>
|
||
) is identical to that of
|
||
<taxonomicName id="4C0E4D2FFFF1E502545A83E26619FAB5" authorityName="Assing" authorityYear="2017" box="[1151,1298,1257,1281]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="densissimus">
|
||
<emphasis id="B97AEABEFFF1E502545A83E26619FAB5" box="[1151,1298,1257,1281]" italics="true" pageId="48" pageNumber="165">Z. densissimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0E4D2FFFF1E502553B83E26662FAB5" authorityName="Assing" authorityYear="2017" box="[1310,1385,1258,1281]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="titan">
|
||
<emphasis id="B97AEABEFFF1E502553B83E26662FAB5" box="[1310,1385,1258,1281]" italics="true" pageId="48" pageNumber="165">Z. titan</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0E4D2FFFF1E502531A820260C7FA95" authorityName="Assing" authorityYear="2017" box="[831,972,1289,1313]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="densihirtus">
|
||
<emphasis id="B97AEABEFFF1E502531A820260C7FA95" box="[831,972,1289,1313]" italics="true" pageId="48" pageNumber="165">Z. densihirtus</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="54B5880EFFF1E502542682016754FA95" box="[1027,1119,1289,1313]" pageId="48" pageNumber="165" type="holotype">holotype</typeStatus>
|
||
was collected with a pitfall trap in lowland forest at an altitude of approximately
|
||
<specimenCount id="9D08FD25FFF1E502551182216677FAF5" box="[1332,1404,1321,1345]" pageId="48" pageNumber="165" type="generic">
|
||
<quantity id="4CF69B49FFF1E502551182216673FAF5" box="[1332,1400,1321,1345]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="165" unit="m" value="400.0">400 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |