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<document id="860BFD2DCB1EF0B0F16AE534871482AB" ID-DOI="10.5281/zenodo.5460634" ID-ISSN="2345-7600" ID-Zenodo-Dep="5460634" ID-ZooBank="1E7214C5-24E8-4860-BF13-203FD93A2C19" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630239335276" checkinUser="valdenar" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2018" docId="03A587FA7827A033FCABB86D301E51A3" docLanguage="en" docName="66rbz763-781.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Macrochlamys caverna Pholyotha &amp; Sutcharit &amp; Panha 2018, new species" docType="treatment" docUuid="FFDB9E0E-1972-401D-B685-AC5466D5CAF6" docUuidSource="ZooBank" docVersion="3" lastPageNumber="771" masterDocId="FF9CFF827821A03BFFC7BF1034285476" masterDocTitle="The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)" masterLastPageNumber="781" masterPageNumber="763" pageNumber="769" updateTime="1699195389588" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="C0CB7B39B8516D35988771FA41C21D45" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CB2089BC4D3C4703C9B90910F1704DFB">
<mods:title id="1285BFD70F297B9A511EC6EF19493685">The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)</mods:title>
</mods:titleInfo>
<mods:name id="5AAA3FE1F107988007B1233F78539441" type="personal">
<mods:role id="350756C665ED36898CF4755E653FF03E">
<mods:roleTerm id="17A0D7AAB662A8F2FEABF3E0A46823B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C8EC4B48FFF5F20A8A2B0BA2CE155CC4">Pholyotha, Arthit</mods:namePart>
</mods:name>
<mods:name id="DC2FC6C6EDE2C57CCC2F2B36366EA9BA" type="personal">
<mods:role id="66AC41EB88E69E67F54027CA4BD0F306">
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<mods:namePart id="B8CFBA1986C8F467A827E0027F4CCA54">Sutcharit, Chirasak</mods:namePart>
</mods:name>
<mods:name id="975E3CA80D0DCFD391925383B44E6217" type="personal">
<mods:role id="B8A6009EC2D3642DA134684D39849F14">
<mods:roleTerm id="4784160562F4FDC225941B1A0617EFD4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DAA51C38BDEFB7B9CEDBCE7A95C5A14D">Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource id="4E36274345B7C952D729B4D32759185F">text</mods:typeOfResource>
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<mods:title id="FD8578DD80597400F2CEB86D1F1FD53F">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="866F2D366EEDA1A305049D23321BCA03">
<mods:date id="BBF6DA6900414F98480E20498FA4CAE9">2018</mods:date>
<mods:detail id="F80C1DAAC3638C7E3B6CFC7F7B20C6CA" type="pubDate">
<mods:number id="4D601144BB5DB3334921520532232854">2018-12-03</mods:number>
</mods:detail>
<mods:detail id="DCEC3CF840A093923D0CF3D8ABF2D641" type="volume">
<mods:number id="75FA46E27427A52D0C70814E5401DA8E">66</mods:number>
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<mods:classification id="582651137D5FC88D1122FD243A393F77">journal article</mods:classification>
<mods:identifier id="11D1D217795C664B39C8AA68E46F03A0" type="DOI">10.5281/zenodo.5460634</mods:identifier>
<mods:identifier id="1C5FF373D2737520769420AEC3C33B61" type="ISSN">2345-7600</mods:identifier>
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</mods:mods>
<treatment id="03A587FA7827A033FCABB86D301E51A3" LSID="urn:lsid:zoobank.org:act:FFDB9E0E-1972-401D-B685-AC5466D5CAF6" httpUri="http://treatment.plazi.org/id/03A587FA7827A033FCABB86D301E51A3" lastPageId="8" lastPageNumber="771" pageId="6" pageNumber="769">
<subSubSection id="C31665677827A03DFCABB86D308253C3" pageId="6" pageNumber="769" type="nomenclature">
<paragraph id="8BB336EC7827A03DFCABB86D308253C3" blockId="6.[876,1435,1917,2005]" pageId="6" pageNumber="769">
<heading id="D0FB81807827A03DFCABB86D308253C3" bold="true" centered="true" fontSize="10" level="2" pageId="6" pageNumber="769" reason="2">
<emphasis id="B978EAFE7827A03DFCABB86D308253C3" bold="true" pageId="6" pageNumber="769">
<taxonomicName id="4C0C4D6F7827A03DFCABB86D317753E3" ID-CoL="3X42X" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[876,1375,1917,1941]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="caverna" status="sp. nov.">
<emphasis id="B978EAFE7827A03DFCABB86D304753E3" bold="true" box="[876,1135,1917,1941]" italics="true" pageId="6" pageNumber="769">Macrochlamys caverna</emphasis>
Pholyotha &amp; Panha
</taxonomicName>
,
<taxonomicNameLabel id="A24B57857827A03DFAAAB86D308253C3" pageId="6" pageNumber="769" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31665677827A03DFC1DB8AD310353A3" box="[986,1323,1981,2005]" pageId="6" pageNumber="769" type="description">
<paragraph id="8BB336EC7827A03DFC1DB8AD310353A3" blockId="6.[876,1435,1917,2005]" box="[986,1323,1981,2005]" pageId="6" pageNumber="769">
(
<figureCitation id="13372A697827A03DFC25B8AD306A53A3" box="[994,1090,1981,2005]" captionStart="Fig" captionStartId="2.[135,170,1546,1568]" captionTargetBox="[136,1474,192,1529]" captionTargetId="figure-7@2.[136,1474,189,1531]" captionTargetPageId="2" captionText="Fig. 2. Living snails of Macrochlamys spp. from Thailand. A, Macrochlamys aurantia new species, paratype (CUMZ 7103), Thong Pha Phum, Kanchanaburi, shell width about 15 mm; B, M. coleus new species, paratype (CUMZ 7106), Sai Yok, Kanchanaburi, shell width about 15 mm; C,M. caverna new species, paratype (CUMZ 7109), Tha Wung, Lopburi, shell width about 17 mm; D, M. lemma new species, paratype (CUMZ 7116), Ban Mi, Lopburi, shell width about 15 mm; E, M. tanymentula new species, paratype (CUMZ 7119), Thong Pha Phum, Kanchanaburi, shell width about 10 mm; F, M. kelantanensis (CUMZ 7125), Lam Kaen, Phang-Nga, shell width about 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5460640" httpUri="https://zenodo.org/record/5460640/files/figure.png" pageId="6" pageNumber="769">Figs. 2C</figureCitation>
,
<figureCitation id="13372A697827A03DFB88B8AD30A553A3" box="[1103,1165,1981,2005]" captionStart="Fig" captionStartId="3.[113,148,1896,1918]" captionTargetBox="[113,1451,328,1881]" captionTargetId="figure-9@3.[113,1451,328,1881]" captionTargetPageId="3" captionText="Fig. 3. Shells of Macrochlamys aurantia new species (A, B), M. coleus new species (C, D), and M. caverna new species (E, F). A, Macrochlamys aurantia holotype (CUMZ 7101); B, paratype (CUMZ 7102); C, M. coleus holotype (CUMZ 7104); D, paratype (CUMZ 7105); E, M. caverna holotype (CUMZ 7107); F, paratype (CUMZ 7108)." figureDoi="http://doi.org/10.5281/zenodo.5460642" httpUri="https://zenodo.org/record/5460642/files/figure.png" pageId="6" pageNumber="769">3E, F</figureCitation>
,
<figureCitation id="13372A697827A03DFB5DB8AD30F653A3" box="[1178,1246,1981,2005]" captionStart="Fig" captionStartId="7.[113,148,1853,1875]" captionTargetBox="[113,1451,285,1838]" captionTargetId="figure-9@7.[113,1451,285,1838]" captionTargetPageId="7" captionText="Fig. 5. Genitalia of Macrochlamys caverna new species (A, B), and M. lemma new species (C, D). A, B, Macrochlamys caverna paratype (CUMZ 7109): A, whole reproductive system; B, internal structure of penis with penial verge.C, D, Macrochlamys lemma paratype (CUMZ 7116): C, whole reproductive system; D, internal structure of penis with penial verge. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460646" httpUri="https://zenodo.org/record/5460646/files/figure.png" pageId="6" pageNumber="769">5A, B</figureCitation>
,
<figureCitation id="13372A697827A03DFB2CB8AD310C53A3" box="[1259,1316,1981,2005]" captionStart="Fig" captionStartId="10.[136,171,1215,1237]" captionTargetBox="[136,1474,189,1200]" captionTargetId="figure-7@10.[136,1474,189,1200]" captionTargetPageId="10" captionText="Fig. 6. SEM images of the radula of Macrochlamys aurantia new species (A–C) (paratype; CUMZ 7103), M. coleus new species (D–F) (paratype; CUMZ 7106), and M. caverna new species (G–I) (paratype; CUMZ 7109). Central tooth (indicated by ‘C’) with lateral teeth (A, D, G); lateral teeth with tricuspid marginal teeth transition (B, E, H); bicuspid marginal teeth (C, F, I)." figureDoi="http://doi.org/10.5281/zenodo.5460650" httpUri="https://zenodo.org/record/5460650/files/figure.png" pageId="6" pageNumber="769">6G–I</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31665677827A033FC84B8ED36665603" lastPageId="8" lastPageNumber="771" pageId="6" pageNumber="769" type="materials_examined">
<paragraph id="8BB336EC7827A033FC84B8ED36665603" blockId="6.[835,1474,2045,2101]" lastBlockId="8.[134,777,188,629]" lastPageId="8" lastPageNumber="771" pageId="6" pageNumber="769">
<emphasis id="B978EAFE7827A03DFC84B8ED30025C63" bold="true" box="[835,1066,2045,2069]" pageId="6" pageNumber="769">Material examined.</emphasis>
<materialsCitation id="3B643CB17827A033FBF3B8ED372C5563" collectionCode="CUMZ" country="Thailand" lastPageId="8" lastPageNumber="771" latitude="14.901638" location="Wat Khao Samo Khon" longLatPrecision="1" longitude="100.508194" municipality="Tha Wung District" pageId="6" pageNumber="769" specimenCode="CUMZ 7107" specimenCount="1" stateProvince="Lop Buri" typeStatus="holotype">
<typeStatus id="54B7884E7827A03DFBF3B8ED30B15C63" box="[1076,1177,2045,2069]" pageId="6" pageNumber="769" type="holotype">Holotype</typeStatus>
(
<figureCitation id="13372A697827A03DFB6BB8ED312C5C63" box="[1196,1284,2045,2069]" captionStart="Fig" captionStartId="3.[113,148,1896,1918]" captionTargetBox="[113,1451,328,1881]" captionTargetId="figure-9@3.[113,1451,328,1881]" captionTargetPageId="3" captionText="Fig. 3. Shells of Macrochlamys aurantia new species (A, B), M. coleus new species (C, D), and M. caverna new species (E, F). A, Macrochlamys aurantia holotype (CUMZ 7101); B, paratype (CUMZ 7102); C, M. coleus holotype (CUMZ 7104); D, paratype (CUMZ 7105); E, M. caverna holotype (CUMZ 7107); F, paratype (CUMZ 7108)." figureDoi="http://doi.org/10.5281/zenodo.5460642" httpUri="https://zenodo.org/record/5460642/files/figure.png" pageId="6" pageNumber="769">Fig. 3E</figureCitation>
; shell height
<quantity id="4CF49B097827A03DFA67B8ED37475C43" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="769" unit="mm" value="8.6">8.6 mm</quantity>
, shell width
<quantity id="4CF49B097827A03DFC30B70D30715C43" box="[1015,1113,2077,2101]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="6" pageNumber="769" unit="mm" value="17.2">17.2 mm</quantity>
, aperture height
<quantity id="4CF49B097827A03DFACBB70D31485C43" box="[1292,1376,2077,2101]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="6" pageNumber="769" unit="mm" value="5.6">5.6 mm</quantity>
, aperture width 8.0 mm, 6¼ whorls) (
<specimenCode id="DBAA9E977829A033FE62BFAD361854A3" box="[421,560,189,213]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="8" pageNumber="771">CUMZ 7107</specimenCode>
), limestone outcrops at
<location id="8ED360377829A033FF62BFCE358F5483" LSID="urn:lsid:plazi:treatment:03A587FA7827A033FCABB86D301E51A3:8ED360377829A033FF62BFCE358F5483" box="[165,423,221,245]" country="Thailand" latitude="14.901638" longLatPrecision="1" longitude="100.508194" municipality="Tha Wung District" name="Wat Khao Samo Khon" pageId="8" pageNumber="771" stateProvince="Lop Buri">Wat Khao Samo Khon</location>
(Temple),
<collectingMunicipality id="6BD7AC967829A033FDEDBFCD37295483" box="[554,769,221,246]" pageId="8" pageNumber="771">Tha Wung District</collectingMunicipality>
,
<collectingRegion id="49C8F80E7829A033FF4FBFED356D5563" box="[136,325,253,277]" country="Thailand" name="Lop Buri" pageId="8" pageNumber="771">Lopburi Province</collectingRegion>
,
<collectingCountry id="F31B767C7829A033FE89BFED35985563" box="[334,432,253,277]" name="Thailand" pageId="8" pageNumber="771">Thailand</collectingCountry>
,
<geoCoordinate id="EE38502B7829A033FE7EBFED367C5563" box="[441,596,252,277]" degrees="14" direction="north" minutes="54" orientation="latitude" pageId="8" pageNumber="771" precision="1" seconds="05.9" value="14.901638">14° 54′05.9″N</geoCoordinate>
,
<geoCoordinate id="EE38502B7829A033FD9ABFED372C5563" box="[605,772,252,277]" degrees="100" direction="east" minutes="30" orientation="longitude" pageId="8" pageNumber="771" precision="1" seconds="29.5" value="100.508194">100° 30′29.5″E</geoCoordinate>
</materialsCitation>
.
<materialsCitation id="3B643CB17829A033FF4FBE0E35CF5663" collectionCode="NHMUK, ZRC" country="Thailand" county="Tha Wung District" latitude="14.902416" location="Non" longLatPrecision="1" longitude="100.50239" municipality="Wat Tham Chang Pueak" pageId="8" pageNumber="771" specimenCode="CUMZ 7108, CUMZ 7109, CUMZ 7110, CUMZ 7111" specimenCount="2" stateProvince="Lop Buri" typeStatus="paratype">
<typeStatus id="54B7884E7829A033FF4FBE0E34DB5540" box="[136,243,286,310]" pageId="8" pageNumber="771" type="paratype">Paratypes</typeStatus>
: 55 shells (
<figureCitation id="13372A697829A033FEA4BE0D359A5543" box="[355,434,285,309]" captionStart="Fig" captionStartId="3.[113,148,1896,1918]" captionTargetBox="[113,1451,328,1881]" captionTargetId="figure-9@3.[113,1451,328,1881]" captionTargetPageId="3" captionText="Fig. 3. Shells of Macrochlamys aurantia new species (A, B), M. coleus new species (C, D), and M. caverna new species (E, F). A, Macrochlamys aurantia holotype (CUMZ 7101); B, paratype (CUMZ 7102); C, M. coleus holotype (CUMZ 7104); D, paratype (CUMZ 7105); E, M. caverna holotype (CUMZ 7107); F, paratype (CUMZ 7108)." figureDoi="http://doi.org/10.5281/zenodo.5460642" httpUri="https://zenodo.org/record/5460642/files/figure.png" pageId="8" pageNumber="771">Fig. 3F</figureCitation>
; shell height
<quantity id="4CF49B097829A033FDFEBE0D36A35543" box="[569,651,285,309]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="8" pageNumber="771" unit="mm" value="8.1">8.1 mm</quantity>
, shell width 15.0 mm, aperture height
<quantity id="4CF49B097829A033FE65BE2D35D15523" box="[418,505,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="771" unit="mm" value="5.4">5.4 mm</quantity>
, aperture width
<quantity id="4CF49B097829A033FD6CBE2D37295523" box="[683,769,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="8" pageNumber="771" unit="mm" value="7.2">7.2 mm</quantity>
, 6⅛ whorls) (
<specimenCode id="DBAA9E977829A033FED7BE4D35B65503" box="[272,414,349,373]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="8" pageNumber="771">CUMZ 7108</specimenCode>
),
<specimenCount id="9D0AFD657829A033FE77BE4E364A5503" box="[432,610,349,373]" pageId="8" pageNumber="771" type="generic">seven specimens</specimenCount>
(
<specimenCode id="DBAA9E977829A033FDA8BE4D36D55503" box="[623,765,349,373]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="8" pageNumber="771">CUMZ 7109</specimenCode>
), five shells (
<collectionCode id="ED1DAE297829A033FEC5BE6D354555E3" box="[258,365,381,405]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="771" type="Museum">NHMUK</collectionCode>
), five shells (
<collectionCode id="ED1DAE297829A033FE3EBE6D361955E3" box="[505,561,381,405]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="771">ZRC</collectionCode>
.MOL.13772), same data as holotype.
<location id="8ED360377829A033FEFABE8E354355C3" LSID="urn:lsid:plazi:treatment:03A587FA7827A033FCABB86D301E51A3:8ED360377829A033FEFABE8E354355C3" box="[317,363,414,437]" country="Thailand" county="Tha Wung District" latitude="14.902416" longLatPrecision="1" longitude="100.50239" municipality="Wat Tham Chang Pueak" name="Non" pageId="8" pageNumber="771" stateProvince="Lop Buri">Non</location>
type material: 11 shells (
<specimenCode id="DBAA9E977829A033FDA8BE8D36D355C3" box="[623,763,413,437]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="8" pageNumber="771">CUMZ 7110</specimenCode>
),
<specimenCount id="9D0AFD657829A033FF40BEAD351F55A3" box="[135,311,445,469]" pageId="8" pageNumber="771" type="generic">three specimens</specimenCount>
(
<specimenCode id="DBAA9E977829A033FE8EBEAD35F455A3" box="[329,476,445,469]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="8" pageNumber="771">CUMZ 7111</specimenCode>
),
<collectingMunicipality id="6BD7AC967829A033FE35BEAE372D55A3" box="[498,773,445,469]" pageId="8" pageNumber="771">Wat Tham Chang Pueak</collectingMunicipality>
(Temple),
<collectingCounty id="62D24E607829A033FF3DBECD35E15583" box="[250,457,477,502]" pageId="8" pageNumber="771">Tha Wung District</collectingCounty>
,
<collectingRegion id="49C8F80E7829A033FE12BECD36BD5583" box="[469,661,477,501]" country="Thailand" name="Lop Buri" pageId="8" pageNumber="771">Lopburi Province</collectingRegion>
,
<collectingCountry id="F31B767C7829A033FD66BECD372B5583" box="[673,771,477,501]" name="Thailand" pageId="8" pageNumber="771">Thailand</collectingCountry>
,
<geoCoordinate id="EE38502B7829A033FF4DBEED35035663" box="[138,299,508,533]" degrees="14" direction="north" minutes="54" orientation="latitude" pageId="8" pageNumber="771" precision="1" seconds="08.7" value="14.902416">14° 54′08.7″N</geoCoordinate>
,
<geoCoordinate id="EE38502B7829A033FEFDBEED35CF5663" box="[314,487,508,533]" degrees="100" direction="east" minutes="30" orientation="longitude" pageId="8" pageNumber="771" precision="1" seconds="08.6" value="100.50239">100° 30′08.6″E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="3B643CB17829A033FE32BEED36625603" collectionCode="CUMZ" country="Thailand" county="Ban Mi District" latitude="14.924723" location="Khao Mon Ing Dharma Practice" longLatPrecision="1" longitude="100.516754" municipality="Ban Chi" pageId="8" pageNumber="771" specimenCode="CUMZ 7112, CUMZ 7113" specimenCount="1" stateProvince="Lop Buri" typeStatus="paratype">
20 shells (
<specimenCode id="DBAA9E977829A033FDAEBEED36D55663" box="[617,765,509,533]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="8" pageNumber="771">CUMZ 7112</specimenCode>
),
<specimenCount id="9D0AFD657829A033FF4FBD0D35155643" box="[136,317,541,565]" pageId="8" pageNumber="771" type="generic">eight specimens</specimenCount>
(
<specimenCode id="DBAA9E977829A033FE88BD0D35CD5643" box="[335,485,541,565]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="8" pageNumber="771">CUMZ 7113</specimenCode>
),
<location id="8ED360377829A033FE3BBD0D34F65623" LSID="urn:lsid:plazi:treatment:03A587FA7827A033FCABB86D301E51A3:8ED360377829A033FE3BBD0D34F65623" country="Thailand" county="Ban Mi District" latitude="14.924723" longLatPrecision="1" longitude="100.516754" municipality="Ban Chi" name="Khao Mon Ing Dharma Practice" pageId="8" pageNumber="771" stateProvince="Lop Buri">
Khao
<location id="8ED360377829A033FD85BD0E34F65623" LSID="urn:lsid:plazi:treatment:03A587FA7827A033FCABB86D301E51A3:8ED360377829A033FD85BD0E34F65623" country="Thailand" county="Ban Mi District" latitude="14.924723" longLatPrecision="1" longitude="100.516754" municipality="Ban Chi" name="Mon Ing Dharma Practice Place" pageId="8" pageNumber="771" stateProvince="Lop Buri">Mon Ing Dharma Practice</location>
</location>
Place,
<collectingMunicipality id="6BD7AC967829A033FEECBD2E35AE5623" box="[299,390,573,597]" pageId="8" pageNumber="771">Ban Chi</collectingMunicipality>
,
<collectingCounty id="62D24E607829A033FE56BD2E36135623" box="[401,571,573,597]" pageId="8" pageNumber="771">Ban Mi District</collectingCounty>
,
<collectingRegion id="49C8F80E7829A033FD82BD2D372B5623" box="[581,771,573,597]" country="Thailand" name="Lop Buri" pageId="8" pageNumber="771">Lopburi Province</collectingRegion>
,
<collectingCountry id="F31B767C7829A033FF4FBD4D34C25603" box="[136,234,605,629]" name="Thailand" pageId="8" pageNumber="771">Thailand</collectingCountry>
,
<geoCoordinate id="EE38502B7829A033FF31BD4D35BC5603" box="[246,404,604,629]" degrees="14" direction="north" minutes="55" orientation="latitude" pageId="8" pageNumber="771" precision="1" seconds="29.0" value="14.924723">14° 55′29.0″N</geoCoordinate>
,
<geoCoordinate id="EE38502B7829A033FE66BD4D36625603" box="[417,586,604,629]" degrees="100" direction="east" minutes="31" orientation="longitude" pageId="8" pageNumber="771" precision="1" seconds="00.3" value="100.516754">100° 31′00.3″E</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF7366647826A03CFFB6B82D30AA5395" ID-DOI="http://doi.org/10.5281/zenodo.5460646" ID-Zenodo-Dep="5460646" httpUri="https://zenodo.org/record/5460646/files/figure.png" pageId="7" pageNumber="770" startId="7.[113,148,1853,1875]" targetBox="[113,1451,285,1838]" targetPageId="7">
<paragraph id="8BB336EC7826A03CFFB6B82D30AA5395" blockId="7.[113,1453,1853,2019]" pageId="7" pageNumber="770">
Fig. 5. Genitalia of
<taxonomicName id="4C0C4D6F7826A03CFEF6B82D363B5325" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[305,531,1853,1875]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="770" phylum="Mollusca" rank="species" species="caverna" status="sp. nov.">
<emphasis id="B978EAFE7826A03CFEF6B82D363B5325" box="[305,531,1853,1875]" italics="true" pageId="7" pageNumber="770">Macrochlamys caverna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857826A03CFDDDB82D36A65325" box="[538,654,1853,1875]" pageId="7" pageNumber="770" rank="species">new species</taxonomicNameLabel>
(A, B), and
<taxonomicName id="4C0C4D6F7826A03CFCCEB82E37415324" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[777,873,1853,1875]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="770" phylum="Mollusca" rank="species" species="lemma" status="sp. nov.">
<emphasis id="B978EAFE7826A03CFCCEB82E37415324" box="[777,873,1853,1875]" italics="true" pageId="7" pageNumber="770">M. lemma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857826A03CFCB7B82D37CC5325" box="[880,996,1853,1875]" pageId="7" pageNumber="770" rank="species">new species</taxonomicNameLabel>
(C, D). A, B,
<taxonomicName id="4C0C4D6F7826A03CFBB7B82D317A5325" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1136,1362,1853,1875]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="770" phylum="Mollusca" rank="species" species="caverna">
<emphasis id="B978EAFE7826A03CFBB7B82D317A5325" box="[1136,1362,1853,1875]" italics="true" pageId="7" pageNumber="770">Macrochlamys caverna</emphasis>
</taxonomicName>
paratype (CUMZ 7109): A, whole reproductive system; B, internal structure of penis with penial verge. C, D,
<taxonomicName id="4C0C4D6F7826A03CFBEAB84A31295319" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1069,1281,1882,1903]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="770" phylum="Mollusca" rank="species" species="lemma">
<emphasis id="B978EAFE7826A03CFBEAB84A31295319" box="[1069,1281,1882,1903]" italics="true" pageId="7" pageNumber="770">Macrochlamys lemma</emphasis>
</taxonomicName>
paratype (CUMZ 7116): C, whole reproductive system; D, internal structure of penis with penial verge. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting.
</paragraph>
</caption>
<subSubSection id="C31665677829A033FF4FBD8D35CE5683" pageId="8" pageNumber="771" type="etymology">
<paragraph id="8BB336EC7829A033FF4FBD8D35CE5683" blockId="8.[136,775,669,757]" pageId="8" pageNumber="771">
<emphasis id="B978EAFE7829A033FF4FBD8D352256C3" bold="true" box="[136,266,669,693]" pageId="8" pageNumber="771">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="4C0C4D6F7829A033FE3FBD8E366656C3" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[504,590,670,693]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="caverna">
<emphasis id="B978EAFE7829A033FE3FBD8E366656C3" box="[504,590,670,693]" italics="true" pageId="8" pageNumber="771">caverna</emphasis>
</taxonomicName>
” is derived from the Latin “
<emphasis id="B978EAFE7829A033FEC6BDAE351656A3" box="[257,318,702,725]" italics="true" pageId="8" pageNumber="771">cavus</emphasis>
” meaning “hole or hollow”, in reference to the open and deep umbilicus.
</paragraph>
</subSubSection>
<subSubSection id="C31665677829A033FF4FBC0D308A5643" pageId="8" pageNumber="771" type="description">
<paragraph id="8BB336EC7829A033FF4FBC0D35055003" blockId="8.[135,777,797,1141]" pageId="8" pageNumber="771">
<emphasis id="B978EAFE7829A033FF4FBC0D353B5743" bold="true" box="[136,275,797,821]" pageId="8" pageNumber="771">Description.</emphasis>
Shell medium-sized (
<figureCitation id="13372A697829A033FE30BC0D364B5743" box="[503,611,797,821]" captionStart="Fig" captionStartId="3.[113,148,1896,1918]" captionTargetBox="[113,1451,328,1881]" captionTargetId="figure-9@3.[113,1451,328,1881]" captionTargetPageId="3" captionText="Fig. 3. Shells of Macrochlamys aurantia new species (A, B), M. coleus new species (C, D), and M. caverna new species (E, F). A, Macrochlamys aurantia holotype (CUMZ 7101); B, paratype (CUMZ 7102); C, M. coleus holotype (CUMZ 7104); D, paratype (CUMZ 7105); E, M. caverna holotype (CUMZ 7107); F, paratype (CUMZ 7108)." figureDoi="http://doi.org/10.5281/zenodo.5460642" httpUri="https://zenodo.org/record/5460642/files/figure.png" pageId="8" pageNumber="771">Fig. 3E, F</figureCitation>
; shell height up to
<quantity id="4CF49B097829A033FF64BC2D34D25723" box="[163,250,829,853]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="8" pageNumber="771" unit="mm" value="8.6">8.6 mm</quantity>
; shell width up to
<quantity id="4CF49B097829A033FE02BC2D36015723" box="[453,553,829,853]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="8" pageNumber="771" unit="mm" value="17.2">17.2 mm</quantity>
), dextral, depressed, somewhat thin, translucent, shiny, and pale reddish-brown. Shell surface smooth with obvious microscopic growth lines. Embryonic shell surface smooth, small with 2½ whorls. Whorls 5–6, regularly increasing. Suture wide and shallow. Spire convex, apex raised. Spire angle about 134–142°. Last whorl narrow with rounded periphery. Aperture ovate-lunate, obliquely open, lip simple. Columellar margin slightly thickened and reflected close to umbilicus. Umbilicus widely open and deep.
</paragraph>
<paragraph id="8BB336EC7829A033FF4FBB8D355751A3" blockId="8.[135,776,1181,1493]" pageId="8" pageNumber="771">
<emphasis id="B978EAFE7829A033FF4FBB8D350750C3" box="[136,303,1181,1205]" italics="true" pageId="8" pageNumber="771">Genital organs</emphasis>
. Atrium (at) short. Penis prolonged tubeshaped. Distal end with short and swollen penial caecum (pc) that corresponds to penial verge inside. Epiphallus (e) cylindrical, about two times penis length. Epiphallic caecum (ec) coiled about one circle and located proximally about one-third of epiphallic length. Penial retractor muscle (prm) short and thick. Flagellum (fl) enlarged and short, about one-third of epiphalic length. Vas deference (vd) a long, thin tube, connected between distal end of epiphallus and free oviduct (
<figureCitation id="13372A697829A033FEDCBAAD355B51A3" box="[283,371,1469,1493]" captionStart="Fig" captionStartId="7.[113,148,1853,1875]" captionTargetBox="[113,1451,285,1838]" captionTargetId="figure-9@7.[113,1451,285,1838]" captionTargetPageId="7" captionText="Fig. 5. Genitalia of Macrochlamys caverna new species (A, B), and M. lemma new species (C, D). A, B, Macrochlamys caverna paratype (CUMZ 7109): A, whole reproductive system; B, internal structure of penis with penial verge.C, D, Macrochlamys lemma paratype (CUMZ 7116): C, whole reproductive system; D, internal structure of penis with penial verge. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460646" httpUri="https://zenodo.org/record/5460646/files/figure.png" pageId="8" pageNumber="771">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC7829A033FF4FBAED35AA52E3" blockId="8.[136,776,1533,1685]" pageId="8" pageNumber="771">
Internal wall of penis proximally with oblique trapezoid penial pilasters (pp) and distally around penial caecum with long and narrowed pilasters surrounding penial verge (pv). Penial verge short, triangular-shaped and located at distal end of penis (
<figureCitation id="13372A697829A033FED8B96D355D52E3" box="[287,373,1661,1685]" captionStart="Fig" captionStartId="7.[113,148,1853,1875]" captionTargetBox="[113,1451,285,1838]" captionTargetId="figure-9@7.[113,1451,285,1838]" captionTargetPageId="7" captionText="Fig. 5. Genitalia of Macrochlamys caverna new species (A, B), and M. lemma new species (C, D). A, B, Macrochlamys caverna paratype (CUMZ 7109): A, whole reproductive system; B, internal structure of penis with penial verge.C, D, Macrochlamys lemma paratype (CUMZ 7116): C, whole reproductive system; D, internal structure of penis with penial verge. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460646" httpUri="https://zenodo.org/record/5460646/files/figure.png" pageId="8" pageNumber="771">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC7829A033FF4FB9AD356553A3" blockId="8.[136,776,1725,2005]" pageId="8" pageNumber="771">
Vagina (v) about same length as penis and cylindrical. Dart apparatus (da) large and cylindrical, located at proximal end of vagina. Gametolytic sac (gs) prolonged and bulbous, gametolytic duct (gd) long and cylindrical. Free oviduct (fo) about same length as penis, proximally encircled with thick brownish tissue close to gametolytic duct and free oviduct junction at about one-fourth of its length. Oviduct (ov) large lobules; prostate gland (pg) running alongside oviduct (
<figureCitation id="13372A697829A033FF2EB8AD356953A3" box="[233,321,1981,2005]" captionStart="Fig" captionStartId="7.[113,148,1853,1875]" captionTargetBox="[113,1451,285,1838]" captionTargetId="figure-9@7.[113,1451,285,1838]" captionTargetPageId="7" captionText="Fig. 5. Genitalia of Macrochlamys caverna new species (A, B), and M. lemma new species (C, D). A, B, Macrochlamys caverna paratype (CUMZ 7109): A, whole reproductive system; B, internal structure of penis with penial verge.C, D, Macrochlamys lemma paratype (CUMZ 7116): C, whole reproductive system; D, internal structure of penis with penial verge. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460646" httpUri="https://zenodo.org/record/5460646/files/figure.png" pageId="8" pageNumber="771">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC7829A033FF4FB8ED31EA5543" blockId="8.[136,776,2045,2101]" lastBlockId="8.[835,1476,189,309]" pageId="8" pageNumber="771">
<emphasis id="B978EAFE7829A033FF4FB8ED34FE5C63" box="[136,214,2045,2069]" italics="true" pageId="8" pageNumber="771">Radula</emphasis>
. Each row consists of 93 teeth with formula (47- (13-12)-1-(12-13)-45). Central tooth symmetrical tricuspid. Mesocone large and lanceolate. Lateral teeth asymmetrical tricuspid with small endocone, large mesocone. Ectocone larger than endocone. Elongate and obliquely bicuspid marginal teeth start from tooth number 12–13 (
<figureCitation id="13372A697829A033FA80BE0D319E5543" box="[1351,1462,285,309]" captionStart="Fig" captionStartId="10.[136,171,1215,1237]" captionTargetBox="[136,1474,189,1200]" captionTargetId="figure-7@10.[136,1474,189,1200]" captionTargetPageId="10" captionText="Fig. 6. SEM images of the radula of Macrochlamys aurantia new species (A–C) (paratype; CUMZ 7103), M. coleus new species (D–F) (paratype; CUMZ 7106), and M. caverna new species (G–I) (paratype; CUMZ 7109). Central tooth (indicated by ‘C’) with lateral teeth (A, D, G); lateral teeth with tricuspid marginal teeth transition (B, E, H); bicuspid marginal teeth (C, F, I)." figureDoi="http://doi.org/10.5281/zenodo.5460650" httpUri="https://zenodo.org/record/5460650/files/figure.png" pageId="8" pageNumber="771">Fig. 6G–I</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC7829A033FC84BE4D308A5643" blockId="8.[834,1475,349,565]" pageId="8" pageNumber="771">
<emphasis id="B978EAFE7829A033FC84BE4D37D35503" box="[835,1019,349,373]" italics="true" pageId="8" pageNumber="771">External features</emphasis>
. Animal with dark grey head and tentacles that gradually becomes paler posteriorly into the yellowish body and tail. Caudal foss (cf) and caudal horn (ch) large and slightly dark grey in colour. Mantle edge well developed, shell lobes and dorsal lobes grey, except right dorsal lobe (rdl) which has yellowish colouration (
<figureCitation id="13372A697829A033FB15BEED310C5663" box="[1234,1316,509,533]" captionStart="Fig" captionStartId="2.[135,170,1546,1568]" captionTargetBox="[136,1474,192,1529]" captionTargetId="figure-7@2.[136,1474,189,1531]" captionTargetPageId="2" captionText="Fig. 2. Living snails of Macrochlamys spp. from Thailand. A, Macrochlamys aurantia new species, paratype (CUMZ 7103), Thong Pha Phum, Kanchanaburi, shell width about 15 mm; B, M. coleus new species, paratype (CUMZ 7106), Sai Yok, Kanchanaburi, shell width about 15 mm; C,M. caverna new species, paratype (CUMZ 7109), Tha Wung, Lopburi, shell width about 17 mm; D, M. lemma new species, paratype (CUMZ 7116), Ban Mi, Lopburi, shell width about 15 mm; E, M. tanymentula new species, paratype (CUMZ 7119), Thong Pha Phum, Kanchanaburi, shell width about 10 mm; F, M. kelantanensis (CUMZ 7125), Lam Kaen, Phang-Nga, shell width about 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5460640" httpUri="https://zenodo.org/record/5460640/files/figure.png" pageId="8" pageNumber="771">Fig. 2C</figureCitation>
). Snails secrete yellowish slime when disturbed.
</paragraph>
</subSubSection>
<subSubSection id="C31665677829A033FC84BD4D30E556A3" pageId="8" pageNumber="771" type="distribution">
<paragraph id="8BB336EC7829A033FC84BD4D30E556A3" blockId="8.[835,1474,605,725]" pageId="8" pageNumber="771">
<emphasis id="B978EAFE7829A033FC84BD4D304E5603" bold="true" box="[835,1126,605,629]" pageId="8" pageNumber="771">Distribution and habitat.</emphasis>
Population density of this new species was generally high and abundant where found. They tend to live in small limestone hills in the Tha Wung and Ban Mi Districts,
<collectingRegion id="49C8F80E7829A033FBCFBDAD30E156A3" box="[1032,1225,701,725]" country="Thailand" name="Lop Buri" pageId="8" pageNumber="771">Lopburi Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C31665677829A033FC84BDED301E51A3" pageId="8" pageNumber="771" type="discussion">
<paragraph id="8BB336EC7829A033FC84BDED31475783" blockId="8.[835,1476,765,1014]" pageId="8" pageNumber="771">
<emphasis id="B978EAFE7829A033FC84BDED37905763" bold="true" box="[835,952,765,789]" pageId="8" pageNumber="771">Remarks.</emphasis>
<taxonomicName id="4C0C4D6F7829A033FC04BDED30FA5763" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[963,1234,765,789]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="caverna" status="sp. nov.">
<emphasis id="B978EAFE7829A033FC04BDED30FA5763" box="[963,1234,765,789]" italics="true" pageId="8" pageNumber="771">Macrochlamys caverna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857829A033FB19BDEE31425763" box="[1246,1386,765,789]" pageId="8" pageNumber="771" rank="species">new species</taxonomicNameLabel>
can be distinguished by its rather depressed shell with a wide and shallow suture and a widely open and deep umbilicus. Its genitalia have a short penial caecum, one circle of coiled epiphallic caecum, a rather long vagina and free oviduct, and very large dart apparatus that is located at the base of the vagina opposite the penis. Radula morphology of this species is very similar to
<taxonomicName id="4C0C4D6F7829A033FB99BCCE30F75783" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1118,1247,990,1013]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE7829A033FB99BCCE30F75783" box="[1118,1247,990,1013]" italics="true" pageId="8" pageNumber="771">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857829A033FB21BCCE31435783" box="[1254,1387,989,1013]" pageId="8" pageNumber="771" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
<paragraph id="8BB336EC7829A033FC84BB0D301E51A3" blockId="8.[834,1476,1053,1493]" pageId="8" pageNumber="771">
<taxonomicName id="4C0C4D6F7829A033FC84BB0D306D5043" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[835,1093,1053,1077]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="caverna" status="sp. nov.">
<emphasis id="B978EAFE7829A033FC84BB0D306D5043" box="[835,1093,1053,1077]" italics="true" pageId="8" pageNumber="771">Macrochlamys caverna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857829A033FB88BB0E30FC5043" box="[1103,1236,1053,1077]" pageId="8" pageNumber="771" rank="species">new species</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4C0C4D6F7829A033FC46BB2E302C5023" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[897,1028,1086,1109]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE7829A033FC46BB2E302C5023" box="[897,1028,1086,1109]" italics="true" pageId="8" pageNumber="771">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857829A033FBCABB2E30B85023" box="[1037,1168,1085,1109]" pageId="8" pageNumber="771" rank="species">new species</taxonomicNameLabel>
and
<taxonomicName id="4C0C4D6F7829A033FB0DBB2E311D5023" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1226,1333,1085,1109]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="coleus" status="sp. nov.">
<emphasis id="B978EAFE7829A033FB0DBB2E311D5023" box="[1226,1333,1085,1109]" italics="true" pageId="8" pageNumber="771">M. coleus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857829A033FAF8BB2E31EA5023" box="[1343,1474,1085,1109]" pageId="8" pageNumber="771" rank="species">new species</taxonomicNameLabel>
by its more elevated spire, shallow suture, widely open umbilicus and oblique trapezoid penial pilasters on the penial wall. In contrast, both
<taxonomicName id="4C0C4D6F7829A033FB4FBB8E312050C3" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1160,1288,1182,1205]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE7829A033FB4FBB8E312050C3" box="[1160,1288,1182,1205]" italics="true" pageId="8" pageNumber="771">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857829A033FAD7BB8E31BB50C3" box="[1296,1427,1181,1205]" pageId="8" pageNumber="771" rank="species">new species</taxonomicNameLabel>
and
<taxonomicName id="4C0C4D6F7829A033FC82BBAE379850A3" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[837,944,1213,1237]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="coleus" status="sp. nov.">
<emphasis id="B978EAFE7829A033FC82BBAE379850A3" box="[837,944,1213,1237]" italics="true" pageId="8" pageNumber="771">M. coleus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857829A033FC7FBBAE301350A3" box="[952,1083,1213,1237]" pageId="8" pageNumber="771" rank="species">new species</taxonomicNameLabel>
have a depressed spire, channelled suture, narrow umbilicus, and small wrinkled and longitudinal folds on the penial wall. The rounded last whorl and wide open umbilicus of this new species differs from
<taxonomicName id="4C0C4D6F7829A033FA93BA0E30675123" authority="(Crosse &amp; Fischer, 1863)" baseAuthorityName="Crosse &amp; Fischer" baseAuthorityYear="1863" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="benoiti">
<emphasis id="B978EAFE7829A033FA93BA0E31EC5143" box="[1364,1476,1309,1333]" italics="true" pageId="8" pageNumber="771">M. benoiti</emphasis>
(
<bibRefCitation id="EF9D4B1D7829A033FC8CBA2D306F5123" author="Crosse H &amp; Fischer P" box="[843,1095,1341,1365]" pageId="8" pageNumber="771" pagination="343 - 379" refId="ref12759" refString="Crosse H &amp; Fischer P (1863) Note sur la faune malacologique de Cochinchine, comprenant la description des especes nouvelles ou peu connues. Journal de Conchyliologie, 11: 343 - 379." type="journal article" year="1863">Crosse &amp; Fischer, 1863</bibRefCitation>
)
</taxonomicName>
from Indochina and
<taxonomicName id="4C0C4D6F7829A033FAEDBA2E37C55103" authority="(Pfeiffer, 1854)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1854" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="malaccana">
<emphasis id="B978EAFE7829A033FAEDBA2E31E95123" box="[1322,1473,1341,1365]" italics="true" pageId="8" pageNumber="771">M. malaccana</emphasis>
(
<bibRefCitation id="EF9D4B1D7829A033FC8EBA4D37CD5103" author="Pfeiffer L" box="[841,997,1373,1397]" pageId="8" pageNumber="771" pagination="145 - 150" refId="ref13877" refString="Pfeiffer L (1854) Descriptions of twenty-three species of Helicea, from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 22: 145 - 150." type="journal article" year="1854">Pfeiffer, 1854</bibRefCitation>
)
</taxonomicName>
from Peninsular
<collectingCountry id="F31B767C7829A033FB77BA4D313E5103" box="[1200,1302,1373,1397]" name="Malaysia" pageId="8" pageNumber="771">Malaysia</collectingCountry>
.
<taxonomicName id="4C0C4D6F7829A033FAE4BA4D37A551E3" baseAuthorityName="Crosse &amp; Fischer" baseAuthorityYear="1863" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="benoiti">
<emphasis id="B978EAFE7829A033FAE4BA4D37A551E3" italics="true" pageId="8" pageNumber="771">Macrochlamys benoiti</emphasis>
</taxonomicName>
has a shouldered last whorl and narrow umbilicus, while
<taxonomicName id="4C0C4D6F7829A033FC42BA8E303651C3" baseAuthorityName="Pfeiffer" baseAuthorityYear="1854" box="[901,1054,1437,1461]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="771" phylum="Mollusca" rank="species" species="malaccana">
<emphasis id="B978EAFE7829A033FC42BA8E303651C3" box="[901,1054,1437,1461]" italics="true" pageId="8" pageNumber="771">M. malaccana</emphasis>
</taxonomicName>
has a comparatively broader last whorl and minute umbilicus.
</paragraph>
</subSubSection>
</treatment>
</document>
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