treatments-xml/data/03/A5/87/03A5879A682EFFB32FE8FF4C95D0FDB0.xml

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<mods:title id="32590E1F99355DC73E1C5CCD9C17F293">Eviota rubriceps, a new goby from the Southwestern Pacific Ocean, with comments on E. mikiae and E. raja (Teleostei: Gobiidae)</mods:title>
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<taxonomicName id="4C0C4D0F682EFFB42FE8FF4C9479FF62" ID-CoL="3DHNJ" box="[151,350,151,177]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="rubriceps" status="sp. nov.">
<emphasis id="B978EA9E682EFFB42FE8FF4C9479FF62" bold="true" box="[151,350,151,177]" italics="true" pageId="1" pageNumber="54">Eviota rubriceps</emphasis>
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<paragraph id="8BB3368C682EFFB42FE8FF6594BBFF05" blockId="1.[151,423,151,214]" box="[151,412,190,214]" pageId="1" pageNumber="54">Red-headed warfgoby</paragraph>
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<taxonomicName id="4C0C4D0F682EFFB42FE8FEDA95FEFEC5" ID-CoL="4HNC" box="[151,217,257,278]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E682EFFB42FE8FEDA95FEFEC5" box="[151,217,257,278]" italics="true" pageId="1" pageNumber="54">Eviota</emphasis>
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sp. 3. 
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: 699
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.
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</paragraph>
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<taxonomicName id="4C0C4D0F682EFFB42FE8FEC495FEFEE7" ID-CoL="4HNC" box="[151,217,287,308]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E682EFFB42FE8FEC495FEFEE7" box="[151,217,287,308]" italics="true" pageId="1" pageNumber="54">Eviota</emphasis>
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sp. 1. 
<treatmentCitation id="0AAD109D682EFFB42E63FEC59760FEE7" author="Allen" box="[284,583,286,308]" page="617" pageId="1" pageNumber="54" year="2009">
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: 617
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.
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</paragraph>
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<taxonomicName id="4C0C4D0F682EFFB42FE8FEE5947EFE81" authority="Raja Ampat" authorityName="Raja Ampat" box="[151,345,317,339]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E682EFFB42FE8FEE595FEFE80" box="[151,217,318,339]" italics="true" pageId="1" pageNumber="54">Eviota</emphasis>
Raja Ampat
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sp. 1. 
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Dimara 
<emphasis id="B978EA9E682EFFB42E91FEE59706FE80" box="[494,545,317,339]" italics="true" pageId="1" pageNumber="54">et al.</emphasis>
, 2010
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(pg. 21, 
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Fig. 
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I).
</paragraph>
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<typeStatus id="54B7882E682EFFB42FE8FEA2942FFE42" box="[151,264,377,401]" pageId="1" pageNumber="54" type="holotype">Holotype</typeStatus>
:
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233447, 
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, male, 
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, 
<taxonomicName id="4C0C4D0F682EFFB42D9EFEA29633FE42" box="[737,788,377,401]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="61" phylum="Chordata" rank="species" species="raja">Raja</taxonomicName>
Ampat Islands, Waisilip, West Waigeo 
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, 0 
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, 
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, 
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, clove oil, M. V. Erdmann, 
<date id="FFB2104C682EFFB42DD8FE479679FE67" box="[679,862,412,437]" pageId="1" pageNumber="54" value="2010-08-28">28 August 2010</date>
.
</paragraph>
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:
</emphasis>
<collectingCountry id="F31B761C682EFFB42E31FE1B970EFE04" box="[334,553,447,471]" name="Papua New Guinea" pageId="1" pageNumber="54">Papua New Guinea</collectingCountry>
: 
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222548, 
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, male, Bismark Archipelago, Ninigo Islands, S.E. of Ami 
<collectingCountry id="F31B761C682EFFB42FB0FE389431FE28" box="[207,278,483,507]" name="Iceland" pageId="1" pageNumber="54">Island</collectingCountry>
, 0 
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, 
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, 
<quantity id="4CF49B69682EFFB42D4AFE3897B5FE28" box="[565,658,483,507]" metricMagnitude="0" metricUnit="m" metricValue="4.55" metricValueMax="9.1" metricValueMin="0.0" pageId="1" pageNumber="54" unit="m" value="4.55" valueMax="9.1" valueMin="0.0">0–9.1 m</quantity>
, patch reef, rotenone, field number VGS78-3, V.G. Springer, 
<emphasis id="B978EA9E682EFFB42A44FE3F9054FE28" box="[1339,1395,483,507]" italics="true" pageId="1" pageNumber="54">et al.</emphasis>
, 
<date id="FFB2104C682EFFB42AFFFE38941EFDCD" pageId="1" pageNumber="54" value="1978-10-23">23 October 1978</date>
. 
<collectingCountry id="F31B761C682EFFB42E35FDDD949CFDCD" box="[330,443,518,542]" name="Indonesia" pageId="1" pageNumber="54">Indonesia</collectingCountry>
: 
<collectionCode id="ED1DAE49682EFFB42EB4FDDD973BFDCD" LSID="urn:lsid:biocol.org:col:34871" box="[459,540,518,542]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="54">USNM</collectionCode>
222550, 10.0 &amp; 
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, males, Banda Islands, Gunung Api, 0 
<geoCoordinate id="EE38504B682EFFB42A51FDDD90B1FDCD" box="[1326,1430,518,542]" direction="south" orientation="latitude" pageId="1" pageNumber="54" precision="5" value="-4.5361">4.5361°S</geoCoordinate>
, 
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, 
<quantity id="4CF49B69682EFFB42E54FDF29451FD92" box="[299,374,553,577]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="1" pageNumber="54" unit="m" value="1.5" valueMax="3.0" valueMin="0.0">0–3 m</quantity>
, coral rubble, rotenone, field number VGS74-8, V.G. Springer &amp; M. Gomon, 
<date id="FFB2104C682EFFB42B8CFDF190B0FD92" box="[1267,1431,553,577]" pageId="1" pageNumber="54" value="1974-03-07">7 March 1974</date>
; 
<collectionCode id="ED1DAE49682EFFB42FE8FD9795CFFDB7" LSID="urn:lsid:biocol.org:col:34871" box="[151,232,588,612]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="54">USNM</collectionCode>
209978, 
<specimenCount id="9D0AFD05682EFFB42E30FD979480FDB7" box="[335,423,588,612]" pageId="1" pageNumber="54" type="male">4 males</specimenCount>
, 
<quantity id="4CF49B69682EFFB42ECDFD979771FDB7" box="[434,598,588,612]" metricMagnitude="-2" metricUnit="m" metricValue="1.085" metricValueMax="1.15" metricValueMin="1.02" pageId="1" pageNumber="54" unit="mm" value="10.85" valueMax="11.5" valueMin="10.2">10.2–11.5 mm</quantity>
, 
<quantity id="4CF49B69682EFFB42D1FFD9797E3FDB7" box="[608,708,588,612]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="54" unit="mm" value="12.7">12.7 mm</quantity>
female, Saparua, ca. 0 
<geoCoordinate id="EE38504B682EFFB42CC1FD979100FDB7" box="[958,1063,588,612]" direction="south" orientation="latitude" pageId="1" pageNumber="54" precision="5" value="-3.5528">3.5528°S</geoCoordinate>
, 
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, isolated coral head, rotenone, field number VGS73-14, V.G. Springer &amp; M. Gomon, 
<date id="FFB2104C682EFFB42C03FDB4911AFD54" box="[892,1085,623,648]" pageId="1" pageNumber="54" value="1973-01-18">18 January 1973</date>
; 
<collectionCode id="ED1DAE49682EFFB42B34FDB491ACFD54" LSID="urn:lsid:biocol.org:col:34778" box="[1099,1163,623,647]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="54">ROM</collectionCode>
85198, 
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, male, 
<collectingCountry id="F31B761C682EFFB42FE8FD489420FD78" box="[151,263,659,683]" name="Indonesia" pageId="1" pageNumber="54">Indonesia</collectingCountry>
, 
<taxonomicName id="4C0C4D0F682EFFB42E6CFD489461FD78" box="[275,326,659,683]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="61" phylum="Chordata" rank="species" species="raja">Raja</taxonomicName>
Ampat Islands, Waisilip, Waigeo 
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, 0 
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, 
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, 
<taxonomicName id="4C0C4D0F682EFFB42B4EFD4F91A5FD78" box="[1073,1154,660,683]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="1" pageNumber="54" phylum="Cnidaria" rank="genus">
<emphasis id="B978EA9E682EFFB42B4EFD4F91A5FD78" box="[1073,1154,660,683]" italics="true" pageId="1" pageNumber="54">Porites</emphasis>
</taxonomicName>
patch reef, 
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, clove oil, field number RW10-24A, M. V. Erdmann, 
<date id="FFB2104C682EFFB42DCFFD6D9656FD1D" box="[688,881,694,719]" pageId="1" pageNumber="54" value="2010-01-29">29 January 2010</date>
; 
<collectionCode id="ED1DAE49682EFFB42C01FD6D9693FD1D" LSID="urn:lsid:biocol.org:col:34700" box="[894,948,694,718]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="1" pageNumber="54">CAS</collectionCode>
233448, 
<quantity id="4CF49B69682EFFB42B5DFD6D91ACFD1D" box="[1058,1163,694,718]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="1" pageNumber="54" unit="mm" value="10.6">10.6 mm</quantity>
, male, 10.7 &amp; 
<quantity id="4CF49B69682EFFB42A48FD6D90BAFD1D" box="[1335,1437,694,718]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="1" pageNumber="54" unit="mm" value="10.8">10.8 mm</quantity>
females, taken with 
<typeStatus id="54B7882E682EFFB42E07FD0294FAFD22" box="[376,477,729,753]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
; 
<collectionCode id="ED1DAE49682EFFB42E96FD02970EFD22" LSID="urn:lsid:biocol.org:col:34778" box="[489,553,729,753]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="54">ROM</collectionCode>
88204, 10.8 &amp; 
<quantity id="4CF49B69682EFFB42DA7FD02961CFD22" box="[728,827,729,753]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="1" pageNumber="54" unit="mm" value="11.8">11.8 mm</quantity>
females, taken with 
<typeStatus id="54B7882E682EFFB42B5CFD0291AEFD22" box="[1059,1161,729,753]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
; 
<collectionCode id="ED1DAE49682EFFB42BECFD0191F0FD21" LSID="urn:lsid:biocol.org:col:34981" box="[1171,1239,730,754]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="1" pageNumber="54">WAM</collectionCode>
P.3353-001, 10.0 mm male, four females 
<quantity id="4CF49B69682EFFB42EDEFD27971FFCC7" box="[417,568,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="9.65" metricValueMax="10.8" metricValueMin="8.5" pageId="1" pageNumber="54" unit="mm" value="9.65" valueMax="10.8" valueMin="8.5">8.5–10.8 mm</quantity>
, 
<collectingCountry id="F31B761C682EFFB42D3BFD279793FCC7" box="[580,692,764,788]" name="Indonesia" pageId="1" pageNumber="54">Indonesia</collectingCountry>
, 
<collectingCountry id="F31B761C682EFFB42DC1FD27963FFCC7" box="[702,792,764,788]" name="Indonesia" pageId="1" pageNumber="54">Maluku</collectingCountry>
, Banda Islands, Banda Harbor, 0 
<geoCoordinate id="EE38504B682EFFB42BF9FD2791C8FCC7" box="[1158,1263,764,788]" direction="south" orientation="latitude" pageId="1" pageNumber="54" precision="5" value="-4.5075">4.5075°S</geoCoordinate>
, 
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, 
<quantity id="4CF49B69682EFFB42AF2FD279588FCE4" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="54" unit="m" value="5.0">5 m</quantity>
, clove oil, M.V. Erdmann &amp; G.R. Allen, 
<date id="FFB2104C682EFFB42D04FCC4961BFCE4" box="[635,828,799,823]" pageId="1" pageNumber="54" value="2010-10-28">28 October 2010</date>
; 
<collectionCode id="ED1DAE49682EFFB42C38FCFB96B2FCE4" LSID="urn:lsid:biocol.org:col:34697" box="[839,917,800,823]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="1" pageNumber="54">BPBM</collectionCode>
41076, five females 
<quantity id="4CF49B69682EFFB42BFCFCC4903DFCE4" box="[1155,1306,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="10.7" metricValueMin="8.9" pageId="1" pageNumber="54" unit="mm" value="9.8" valueMax="10.7" valueMin="8.9">8.9–10.7 mm</quantity>
, taken with 
<collectionCode id="ED1DAE49682EFFB42FE8FC9895FCFC88" LSID="urn:lsid:biocol.org:col:34981" box="[151,219,835,859]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="1" pageNumber="54">WAM</collectionCode>
P.3353-001; 
<collectionCode id="ED1DAE49682EFFB42E0FFC9F948AFC88" box="[368,429,836,859]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="54">MZB</collectionCode>
20024, four females 8.0– 
<quantity id="4CF49B69682EFFB42DB2FC989614FC88" box="[717,819,835,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="1" pageNumber="54" unit="mm" value="10.7">10.7 mm</quantity>
, taken with 
<collectionCode id="ED1DAE49682EFFB42CC5FC9896D9FC88" LSID="urn:lsid:biocol.org:col:34981" box="[954,1022,835,859]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="1" pageNumber="54">WAM</collectionCode>
P.3353-001.
</paragraph>
<paragraph id="8BB3368C682EFFB42FB8FCBD9003FC17" blockId="1.[151,1437,377,2026]" pageId="1" pageNumber="54">
<emphasis id="B978EA9E682EFFB42FB8FCBD946EFCAD" bold="true" box="[199,329,870,894]" pageId="1" pageNumber="54">Non-types:</emphasis>
<collectionCode id="ED1DAE49682EFFB42E2CFCBD9483FCAD" LSID="urn:lsid:biocol.org:col:34871" box="[339,420,870,894]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="54">USNM</collectionCode>
222549, 
<quantity id="4CF49B69682EFFB42D6CFCBD975CFCAD" box="[531,635,870,894]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="54" unit="mm" value="12.7">12.7 mm</quantity>
, male, 
<collectingCountry id="F31B761C682EFFB42DAEFCBD9687FCAD" box="[721,928,870,894]" name="Philippines" pageId="1" pageNumber="54">Philippine Islands</collectingCountry>
, Palawan Province, N. N-E. side of Bararin 
<collectingCountry id="F31B761C682EFFB42FE8FC5295FBFC72" box="[151,220,905,929]" name="Iceland" pageId="1" pageNumber="54">Island</collectingCountry>
(Cuyo 
<collectingCountry id="F31B761C682EFFB42E4AFC529459FC72" box="[309,382,905,929]" name="Iceland" pageId="1" pageNumber="54">Island</collectingCountry>
), 
<geoCoordinate id="EE38504B682EFFB42EECFC529728FC72" box="[403,527,905,929]" direction="north" orientation="latitude" pageId="1" pageNumber="54" precision="5" value="10.8783">10.8783°N</geoCoordinate>
, 
<geoCoordinate id="EE38504B682EFFB42D62FC529783FC72" box="[541,676,905,929]" direction="east" orientation="longitude" pageId="1" pageNumber="54" precision="5" value="120.9456">120.9456°E</geoCoordinate>
, 
<quantity id="4CF49B69682EFFB42DCDFC529607FC72" box="[690,800,905,929]" metricMagnitude="0" metricUnit="m" metricValue="8.7" metricValueMax="17.4" metricValueMin="0.0" pageId="1" pageNumber="54" unit="m" value="8.7" valueMax="17.4" valueMin="0.0">0–17.4 m</quantity>
, coral, rotenone,, field number SP78-21, L. Knapp, V. Springer, J. Libbey, 
<date id="FFB2104C682EFFB42E07FC7794F7FC16" box="[376,464,940,965]" pageId="1" pageNumber="54" value="1924-05">24 May</date>
, 1978; 
<collectionCode id="ED1DAE49682EFFB42D5EFC77977FFC17" LSID="urn:lsid:biocol.org:col:34700" box="[545,600,940,964]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="1" pageNumber="54">CAS</collectionCode>
233449, 
<specimenCount id="9D0AFD05682EFFB42DBEFC77961DFC17" box="[705,826,940,964]" pageId="1" pageNumber="54" type="female">11 females</specimenCount>
<quantity id="4CF49B69682EFFB42C3EFC7796ECFC17" box="[833,971,940,964]" metricMagnitude="-3" metricUnit="m" metricValue="8.35" metricValueMax="8.9" metricValueMin="7.8" pageId="1" pageNumber="54" unit="mm" value="8.35" valueMax="8.9" valueMin="7.8">7.8–8.9 mm</quantity>
, taken with 
<collectionCode id="ED1DAE49682EFFB42B2DFC7691B1FC16" LSID="urn:lsid:biocol.org:col:34981" box="[1106,1174,941,965]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="1" pageNumber="54">WAM</collectionCode>
P.3353-001.
</paragraph>
</subSubSection>
<subSubSection id="C3166507682EFFB42FB8FC1497C1FB4E" pageId="1" pageNumber="54" type="diagnosis">
<paragraph id="8BB3368C682EFFB42FB8FC1497C1FB4E" blockId="1.[151,1437,377,2026]" pageId="1" pageNumber="54">
<emphasis id="B978EA9E682EFFB42FB8FC14941DFC3B" bold="true" box="[199,314,975,1000]" pageId="1" pageNumber="54">Diagnosis</emphasis>
. The following combination of characters distinguish 
<taxonomicName id="4C0C4D0F682EFFB42CD3FC0A911EFC3B" box="[940,1081,976,1000]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682EFFB42CD3FC0A911EFC3B" box="[940,1081,976,1000]" italics="true" pageId="1" pageNumber="54">E. rubriceps</emphasis>
</taxonomicName>
from congeners: cephalic sensory-pore system pattern 2 (only IT pore missing); dorsal/anal fin-ray formula almost always 8/7; pectoral-fin rays unbranched; first three dorsal spines may be filamentous in males; 5th pelvic-fin ray absent or rudimentary to 20% maximum of 4th ray; strong black pigment along scale pockets on ventral two-thirds of sides and peppering of dark chromatophores on nape and upper back when preserved; snout, top and sides of head red in life; dark area of abdomen not crossed by a curved yellow-gold line in life.
</paragraph>
</subSubSection>
<subSubSection id="C3166507682EFFB42FB8FB7390BBF997" pageId="1" pageNumber="54" type="description">
<paragraph id="8BB3368C682EFFB42FB8FB739752FA47" blockId="1.[151,1437,377,2026]" pageId="1" pageNumber="54">
<emphasis id="B978EA9E682EFFB42FB8FB73947EFB12" bold="true" box="[199,345,1192,1217]" pageId="1" pageNumber="54">Description.</emphasis>
Dorsal-fin rays VI-I,8; anal-fin rays I,7 (26), I,8 (3); pectoral-fin rays 15 (14–16, usually 15), rays not branched; pelvic fin I,4 (I,4+ a rudiment to 20%); branches on 4th pelvic-fin ray 5 (3–5), 2 (2–6) segments between branches; 11 branched caudal-fin rays; segmented caudal-fin rays 17; lateral scale rows 24 (21–24); transverse scale rows 6; breast and pectoral-fin base scaleless; first three dorsal spines may be filamentous in males, the second spine longest, reaching to end of second dorsal-fin base; depressed pelvic fin always extends beyond analfin origin; cephalic sensory-pore system pattern 2 (only IT missing); male genital papilla non-fimbriate; precaudal vertebrae 10, caudal vertebrae 15, total 25.
</paragraph>
<paragraph id="8BB3368C682EFFB42FB8FA4490BBF997" blockId="1.[151,1437,377,2026]" pageId="1" pageNumber="54">
<emphasis id="B978EA9E682EFFB42FB8FA449451FA64" bold="true" box="[199,374,1439,1463]" pageId="1" pageNumber="54">Measurements</emphasis>
(based on 
<typeStatus id="54B7882E682EFFB42E8EFA449774FA64" box="[497,595,1439,1463]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
and 14 
<typeStatus id="54B7882E682EFFB42DD2FA7B963CFA6B" box="[685,795,1440,1464]" pageId="1" pageNumber="54" type="paratype">paratypes</typeStatus>
, 
<quantity id="4CF49B69682EFFB42C59FA4496E9FA64" box="[806,974,1439,1463]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" metricValueMax="1.35" metricValueMin="1.01" pageId="1" pageNumber="54" unit="mm" value="11.8" valueMax="13.5" valueMin="10.1">10.1–13.5 mm</quantity>
). head length 31.8 (29.5–33.3, 31.2); origin of first dorsal fin 33.7 (32.4–39.0, 35.9); origin of second dorsal fin 55.5 (54.8–59.5, 56.7); origin of anal fin 58.5 (55.2–62.7, 59.5); caudal-peduncle length 24.4 (16.9–29.0, 23.8); caudal-peduncle depth 11.8 (9.1–12.7, 10.9); body depth 22.2 (17.8–23.8, 21.7); eye diameter 8.5 (8.5–11.3, 10.3); snout length 4.4 (2.8–4.7, 3.8); upperjaw length 10.7 (7.6–13.7, 9.9); pectoral-fin length 22.2 (22.2–33.6, 29.9); pelvic-fin length 31.1 (23.3–40.6, 33.2).
</paragraph>
</subSubSection>
<subSubSection id="C3166507682EFFB62FB8F9959797FDE5" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54" type="materials_examined">
<paragraph id="8BB3368C682EFFB42FB8F99596B0F839" blockId="1.[151,1437,377,2026]" pageId="1" pageNumber="54">
<emphasis id="B978EA9E682EFFB42FB8F99597AEF9B4" bold="true" box="[199,649,1614,1639]" pageId="1" pageNumber="54">
Color of preserved specimens (
<figureCitation id="13372A09682EFFB42D45F9959758F9B4" box="[570,639,1614,1639]" captionStart="FIGURE 1" captionStartId="2.[151,250,608,631]" captionTargetBox="[265,1321,195,583]" captionTargetId="figure@2.[265,1321,195,583]" captionTargetPageId="2" captionText="FIGURE 1. Eviota rubriceps, holotype, CAS 223447, 13.5 mm male, photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/205708/files/figure.png" pageId="1" pageNumber="54">Fig. 1</figureCitation>
)
</emphasis>
. The characteristic color pattern consists of dark pigmentation on the lower two-thirds of the body, pale coloration on the dorsal portion of nape and upper one-third of body but with a peppering of dark chromatophores, and obliquely arranged dark subcutaneous bars on lower caudal peduncle. The coloration of the 
<typeStatus id="54B7882E682EFFB42E21F96394E4F903" box="[350,451,1720,1744]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
, as shown in 
<figureCitation id="13372A09682EFFB42D1CF96397B8F903" box="[611,671,1720,1744]" captionStart="FIGURE 1" captionStartId="2.[151,250,608,631]" captionTargetBox="[265,1321,195,583]" captionTargetId="figure@2.[265,1321,195,583]" captionTargetPageId="2" captionText="FIGURE 1. Eviota rubriceps, holotype, CAS 223447, 13.5 mm male, photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/205708/files/figure.png" pageId="1" pageNumber="54">Fig.1</figureCitation>
, is typical of the species except for the subcutaneous bars that are indistinct on this specimen. The salient color characters of the species are the fine, dense brown chromatophores on snout, laterally on head, on pectoral-fin base, and present on lower two-thirds of body. The scale pockets are typically pronounced, pigmented with a narrow to wide margin of dark chromatophores, the pattern becoming less intense posteriorly; lower two-thirds of body also with dusky subcutaneous shading; upper third of body and dorsalmost portion of nape pale, but with a peppering of dark chromatophores. About five dark oblique subcutaneous bars directed upward and forward from faint spots along lower margin of caudal peduncle to just above midline of body and may or may not be in line with, but separated from, small dark subcutaneous spots dorsally along caudal peduncle and beneath second dorsal fin, spots often obscure (
<figureCitation id="13372A09682EFFB42C3AF80996ADF839" box="[837,906,2002,2026]" captionStart="FIGURE 2" captionStartId="2.[151,250,1154,1177]" captionTargetBox="[299,1286,693,1081]" captionTargetId="figure@2.[265,1321,680,1133]" captionTargetPageId="2" captionText="FIGURE 2. Eviota rubriceps, paratype, USNM 209978, 11.5 mm male. Drawn by G. (Trudy) Nicholson, Smithsonian Institution, NMNH, Division of Fishes." httpUri="https://zenodo.org/record/205709/files/figure.png" pageId="1" pageNumber="54">Fig. 2</figureCitation>
).
</paragraph>
<caption id="DF736604682DFFB72FE8FDBB918DFDA5" box="[151,1194,608,631]" httpUri="https://zenodo.org/record/205708/files/figure.png" pageId="2" pageNumber="55" targetBox="[265,1321,195,583]" targetPageId="2">
<paragraph id="8BB3368C682DFFB72FE8FDBB918DFDA5" blockId="2.[151,1194,608,631]" box="[151,1194,608,631]" pageId="2" pageNumber="55">
<emphasis id="B978EA9E682DFFB72FE8FDBB9434FDA6" bold="true" box="[151,275,608,631]" pageId="2" pageNumber="55">FIGURE 1.</emphasis>
<taxonomicName id="4C0C4D0F682DFFB72E66FDBA94E5FDA5" box="[281,450,608,630]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682DFFB72E66FDBA94E5FDA5" box="[281,450,608,630]" italics="true" pageId="2" pageNumber="55">Eviota rubriceps</emphasis>
</taxonomicName>
, holotype, CAS 223447, 13.5 mm male, photograph by D.W. Greenfield.
</paragraph>
</caption>
<caption id="DF736604682DFFB72FE8FB5994C1FB64" httpUri="https://zenodo.org/record/205709/files/figure.png" pageId="2" pageNumber="55" targetBox="[299,1286,693,1081]" targetPageId="2">
<paragraph id="8BB3368C682DFFB72FE8FB5994C1FB64" blockId="2.[151,1436,1154,1207]" pageId="2" pageNumber="55">
<emphasis id="B978EA9E682DFFB72FE8FB599434FB4B" bold="true" box="[151,275,1154,1177]" pageId="2" pageNumber="55">FIGURE 2.</emphasis>
<taxonomicName id="4C0C4D0F682DFFB72E65FB5F94E3FB4A" box="[282,452,1155,1177]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682DFFB72E65FB5F94E3FB4A" box="[282,452,1155,1177]" italics="true" pageId="2" pageNumber="55">Eviota rubriceps</emphasis>
</taxonomicName>
, paratype, USNM 209978, 11.5 mm male. Drawn by G. (Trudy) Nicholson, Smithsonian Institution, NMNH, Division of Fishes.
</paragraph>
</caption>
<caption id="DF736604682DFFB72FE8F894979AF857" httpUri="https://zenodo.org/record/205710/files/figure.png" pageId="2" pageNumber="55" targetBox="[265,1321,1257,1849]" targetPageId="2">
<paragraph id="8BB3368C682DFFB72FE8F894979AF857" blockId="2.[151,1436,1871,1924]" pageId="2" pageNumber="55">
<emphasis id="B978EA9E682DFFB72FE8F8949434F8B6" bold="true" box="[151,275,1871,1894]" pageId="2" pageNumber="55">FIGURE 3.</emphasis>
<taxonomicName id="4C0C4D0F682DFFB72E64F88A94E2F8B6" box="[283,453,1871,1894]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682DFFB72E64F88A94E2F8B6" box="[283,453,1871,1894]" italics="true" pageId="2" pageNumber="55">Eviota rubriceps</emphasis>
</taxonomicName>
, underwater photograph, Halmahera, Pulau Gifao, Buli Bay, 00°47.447 N, 128°19.298 E, 10 m, Maluku Islands, Indonesia. Photograph by G.R. Allen.
</paragraph>
</caption>
<paragraph id="8BB3368C682DFFB62FB8F87791B5FF02" blockId="2.[151,1436,1963,2021]" lastBlockId="3.[151,1436,151,566]" lastPageId="3" lastPageNumber="56" pageId="2" pageNumber="55">Lower portion of first dorsal fin with narrow to wide dusky pigmentation, outer portion pale, the filamentous spines with tiny spots formed by fine dark chromatophores. Second dorsal fin dusky throughout, more intense basally. Anal fin similar to second dorsal fin. Caudal fin dusky but paler than dorsal and anal fins, some specimens with heavier irregular pigmentation on fin, especially basally. Pectoral and pelvic fins pale.</paragraph>
<paragraph id="8BB3368C682CFFB62FB8FF019797FDE5" blockId="3.[151,1436,151,566]" pageId="3" pageNumber="56">
<emphasis id="B978EA9E682CFFB62FB8FF019470FF20" bold="true" box="[199,343,218,243]" pageId="3" pageNumber="56">Color in life</emphasis>
(
<figureCitation id="13372A09682CFFB62E17FF0194F1FF21" box="[360,470,218,242]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="2.[151,250,1871,1894]" captionStartId-1="3.[151,250,1419,1442]" captionTargetBox-0="[265,1321,1257,1849]" captionTargetBox-1="[265,1321,620,1398]" captionTargetId-0="figure@2.[265,1321,1257,1850]" captionTargetId-1="figure@3.[265,1321,620,1399]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 3. Eviota rubriceps, underwater photograph, Halmahera, Pulau Gifao, Buli Bay, 00 ° 47.447 N, 128 ° 19.298 E, 10 m, Maluku Islands, Indonesia. Photograph by G. R. Allen." captionText-1="FIGURE 4. Eviota rubriceps, underwater photograph, Pulau Mommon, 03 ° 56.377 S, 132 ° 48.208 E, 12 m, Fakfak Peninsula, West Papua. Photograph by G. R. Allen." httpUri-0="https://zenodo.org/record/205710/files/figure.png" httpUri-1="https://zenodo.org/record/205711/files/figure.png" pageId="3" pageNumber="56">Figs. 3–4</figureCitation>
). Body translucent white. A black band running from behind head to caudal-fin base. Band widest over abdomen, slightly above midline down to lower portion of belly, narrowing posteriorly to vertebral column with six obvious black lines extending ventrally along posterior half of body. Black band over abdomen sprinkled with tiny gold flecks. Brown scale edges over white background extending above band to dorsal-fin bases. A peppering of dark chromatophores on nape. Head and snout from level of ventral eye margin dorsally reddish. Upper and tip of lower lip reddish. Lower part of head below eye white, extending posteriorly along belly to caudal-fin base. Top of head with a narrow white line from between eyes posteriorly to nape. Fins translucent white with redish bands across first dorsal fin. Two rows of small red dots running along bases of second dorsal and anal fins. Membranes with a peppering of fine dark chromatophores. Pupil and iris of eye black, often with a narrow white line running across upper third onto head.
</paragraph>
</subSubSection>
<caption id="DF736604682CFFB62FE8FA509700FA13" httpUri="https://zenodo.org/record/205711/files/figure.png" pageId="3" pageNumber="56" targetBox="[265,1321,620,1398]" targetPageId="3">
<paragraph id="8BB3368C682CFFB62FE8FA509700FA13" blockId="3.[151,1436,1419,1473]" pageId="3" pageNumber="56">
<emphasis id="B978EA9E682CFFB62FE8FA509434FA72" bold="true" box="[151,275,1419,1442]" pageId="3" pageNumber="56">FIGURE 4.</emphasis>
<taxonomicName id="4C0C4D0F682CFFB62E64FA5694E2FA71" box="[283,453,1420,1442]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682CFFB62E64FA5694E2FA71" box="[283,453,1420,1442]" italics="true" pageId="3" pageNumber="56">Eviota rubriceps</emphasis>
</taxonomicName>
, underwater photograph, Pulau Mommon, 03°56.377 S, 132°48.208 E, 12 m, Fakfak Peninsula, West Papua. Photograph by G.R. Allen.
</paragraph>
</caption>
<subSubSection id="C3166507682CFFB62FB8FA3D912BF9F1" pageId="3" pageNumber="56" type="distribution">
<paragraph id="8BB3368C682CFFB62FB8FA3D912BF9F1" blockId="3.[151,1436,1510,2008]" pageId="3" pageNumber="56">
<emphasis id="B978EA9E682CFFB62FB8FA3D9447FA2C" bold="true" box="[199,352,1510,1535]" pageId="3" pageNumber="56">Distribution.</emphasis>
Known from the 
<taxonomicName id="4C0C4D0F682CFFB62D55FA3C977AFA2C" box="[554,605,1511,1535]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="61" phylum="Chordata" rank="species" species="raja">Raja</taxonomicName>
Ampat Islands, Banda Islands, and Maluka Islands in 
<collectingCountry id="F31B761C682CFFB62BB8FA3C901FFA2C" box="[1223,1336,1511,1535]" name="Indonesia" pageId="3" pageNumber="56">Indonesia</collectingCountry>
; the Bismark Archipelago, 
<collectingCountry id="F31B761C682CFFB62E0FF9D0976CF9F1" box="[368,587,1546,1570]" name="Papua New Guinea" pageId="3" pageNumber="56">Papua New Guinea</collectingCountry>
; and the Palawan Province, 
<collectingCountry id="F31B761C682CFFB62CF8F9D1912EF9F1" box="[903,1033,1546,1570]" name="Philippines" pageId="3" pageNumber="56">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3166507682CFFB12FB8F9F49047FEB7" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56" type="etymology">
<paragraph id="8BB3368C682CFFB62FB8F9F4972EF9B9" blockId="3.[151,1436,1510,2008]" pageId="3" pageNumber="56">
<emphasis id="B978EA9E682CFFB62FB8F9F49469F994" bold="true" box="[199,334,1583,1607]" pageId="3" pageNumber="56">Etymology.</emphasis>
The specific epithet is an adjective combining the Latin 
<emphasis id="B978EA9E682CFFB62CACF9F49100F994" box="[979,1063,1583,1607]" italics="true" pageId="3" pageNumber="56">rubrum</emphasis>
(red) and 
<emphasis id="B978EA9E682CFFB62BE2F9EB91FCF994" box="[1181,1243,1584,1607]" italics="true" pageId="3" pageNumber="56">caput</emphasis>
(head), referring to the distinctive red head in life.
</paragraph>
<paragraph id="8BB3368C682CFFB12FB8F9AD9047FEB7" blockId="3.[151,1436,1510,2008]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
<emphasis id="B978EA9E682CFFB62FB8F9AD9445F95C" bold="true" box="[199,354,1654,1679]" pageId="3" pageNumber="56">Comparison.</emphasis>
<taxonomicName id="4C0C4D0F682CFFB62E17F9A39704F95C" box="[360,547,1655,1679]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682CFFB62E17F9A39704F95C" box="[360,547,1655,1679]" italics="true" pageId="3" pageNumber="56">Eviota rubriceps</emphasis>
</taxonomicName>
belongs to the cephalic sensory-pore pattern 2 of 
<bibRefCitation id="EF9D4B7D682CFFB62B2EF9AC90B0F95C" author="Lachner" box="[1105,1431,1655,1679]" pageId="3" pageNumber="56" refString="Lachner, E. A. &amp; Karnella, S. J. (1980) Fishes of the Indo-Pacific genus Eviota with descriptions of eight new species (Teleostei: Gobiidae). Smithsonian Contributions to Zoology, No. 315, 1 - 127." type="book chapter" year="1980">Lachner and Karnella (1980)</bibRefCitation>
, lacking only the IT pore, and differs from species in that group as listed in 
<bibRefCitation id="EF9D4B7D682CFFB62C9DF94191F8F961" author="Gill" box="[994,1247,1690,1714]" pageId="3" pageNumber="56" refString="Gill, A. C., &amp; Jewett, S. L. (2004) Eviota hoesei and E. readerae, new species of fish from the southwest Pacific, with comments on the identity of E. corneliae Fricke (Perciformes: Gobiidae). Records of the Australian Museum, 56, 235 - 240." type="journal article" year="2004">Gill and Jewett (2004)</bibRefCitation>
, 
<bibRefCitation id="EF9D4B7D682CFFB62B8BF9419464F904" author="Greenfield" pageId="3" pageNumber="56" refString="Greenfield, D. W., &amp; Randall, J. E. (2008) Eviota tigrina, a new goby from Tonga (Teleostei: Gobiidae). Proceedings of the California Academy of Sciences, Series 4, 59 (11), 497 - 501." type="journal article" year="2008">Greenfield and Randall (2008)</bibRefCitation>
, and 
<bibRefCitation id="EF9D4B7D682CFFB62EFFF9649799F904" author="Greenfield" box="[384,702,1727,1751]" pageId="3" pageNumber="56" refString="Greenfield, D. W. &amp; Suzuki, T. (2010) Eviota nigrispina, a new goby from the Ryukyu Islands, Japan (Teleostei: Gobiidae). Zootaxa, 2655, 57 - 62." type="journal article" year="2010">Greenfield &amp; Suzuki (2010)</bibRefCitation>
as follows. Only eight described species in pattern group 2 have simple unbranched pectoral-fin rays as in 
<taxonomicName id="4C0C4D0F682CFFB62D13F93F97D1F928" box="[620,758,1763,1787]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682CFFB62D13F93F97D1F928" box="[620,758,1763,1787]" italics="true" pageId="3" pageNumber="56">E. rubriceps</emphasis>
</taxonomicName>
: 
<taxonomicName id="4C0C4D0F682CFFB62C7AF93F918EF929" authority="Jewett and Lachner (1983)" authorityName="Jewett and Lachner" authorityYear="1983" box="[773,1193,1762,1787]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="cometa">
<emphasis id="B978EA9E682CFFB62C7AF93F9652F928" box="[773,885,1764,1787]" italics="true" pageId="3" pageNumber="56">E. cometa</emphasis>
<bibRefCitation id="EF9D4B7D682CFFB62C03F938918EF929" author="Jewett" box="[892,1193,1762,1786]" pageId="3" pageNumber="56" refString="Jewett, S. L. &amp; Lachner, E. A. (1983) Seven new species of the Indo-Pacific genus Eviota (Pisces: Gobiidae). Proceedings of the Biological Society of Washington, 96 (4), 780 - 806." type="journal article" year="1983">Jewett and Lachner (1983)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0C4D0F682CFFB62BCAF93F948FF8CC" authority="Greenfield and Suzuki (2010)" authorityName="Greenfield and Suzuki" authorityYear="2010" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682CFFB62BCAF93F9060F928" box="[1205,1351,1764,1787]" italics="true" pageId="3" pageNumber="56">E. nigrispina</emphasis>
<bibRefCitation id="EF9D4B7D682CFFB62A31F939948FF8CC" author="Greenfield" pageId="3" pageNumber="56" refString="Greenfield, D. W. &amp; Suzuki, T. (2010) Eviota nigrispina, a new goby from the Ryukyu Islands, Japan (Teleostei: Gobiidae). Zootaxa, 2655, 57 - 62." type="journal article" year="2010">Greenfield and Suzuki (2010)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0C4D0F682CFFB62ECAF8D397CDF8CC" authority="Larson (1976)" authorityName="Larson" authorityYear="1976" box="[437,746,1799,1823]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="pellucida">
<emphasis id="B978EA9E682CFFB62ECAF8D39719F8CC" box="[437,574,1799,1823]" italics="true" pageId="3" pageNumber="56">E. pellucida</emphasis>
Larson (1976)
</taxonomicName>
, 
<taxonomicName id="4C0C4D0F682CFFB62D88F8D39656F8CC" box="[759,881,1800,1823]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="prasites">
<emphasis id="B978EA9E682CFFB62D88F8D39656F8CC" box="[759,881,1800,1823]" italics="true" pageId="3" pageNumber="56">E. prasites</emphasis>
</taxonomicName>
<collectingCountry id="F31B761C682CFFB62C05F8DC96E2F8CC" box="[890,965,1799,1823]" name="Jordan" pageId="3" pageNumber="56">Jordan</collectingCountry>
and Seale (1906), 
<taxonomicName id="4C0C4D0F682CFFB62BE1F8D3946FF891" authority="Jewett and Lachner (1983)" authorityName="Jewett and Lachner" authorityYear="1983" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="sigillata">
<emphasis id="B978EA9E682CFFB62BE1F8D3903CF8CC" box="[1182,1307,1799,1823]" italics="true" pageId="3" pageNumber="56">E. sigillata</emphasis>
<bibRefCitation id="EF9D4B7D682CFFB62A5CF8D3946FF891" author="Jewett" pageId="3" pageNumber="56" refString="Jewett, S. L. &amp; Lachner, E. A. (1983) Seven new species of the Indo-Pacific genus Eviota (Pisces: Gobiidae). Proceedings of the Biological Society of Washington, 96 (4), 780 - 806." type="journal article" year="1983">Jewett and Lachner (1983)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0C4D0F682CFFB62E29F8F7963FF891" authority="Lachner and Karnella (1980)" authorityName="Lachner and Karnella" authorityYear="1980" box="[342,792,1834,1859]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="spilota">
<emphasis id="B978EA9E682CFFB62E29F8F794E4F890" box="[342,451,1835,1859]" italics="true" pageId="3" pageNumber="56">E. spilota</emphasis>
<bibRefCitation id="EF9D4B7D682CFFB62EB3F8F1963FF891" author="Lachner" box="[460,792,1834,1858]" pageId="3" pageNumber="56" refString="Lachner, E. A. &amp; Karnella, S. J. (1980) Fishes of the Indo-Pacific genus Eviota with descriptions of eight new species (Teleostei: Gobiidae). Smithsonian Contributions to Zoology, No. 315, 1 - 127." type="book chapter" year="1980">Lachner and Karnella (1980)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0C4D0F682CFFB62C59F8F79175F891" authority="Rofen (1959)" authorityName="Rofen" authorityYear="1959" box="[806,1106,1834,1859]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="storthynx">
<emphasis id="B978EA9E682CFFB62C59F8F79688F890" box="[806,943,1835,1859]" italics="true" pageId="3" pageNumber="56">E. storthynx</emphasis>
Rofen (1959)
</taxonomicName>
, and 
<taxonomicName id="4C0C4D0F682CFFB62BEEF8F7946BF8B4" authority="Lachner and Karnella (1978)" authorityName="Lachner and Karnella" authorityYear="1978" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="zebrina">
<emphasis id="B978EA9E682CFFB62BEEF8F79021F890" box="[1169,1286,1835,1859]" italics="true" pageId="3" pageNumber="56">E. zebrina</emphasis>
<bibRefCitation id="EF9D4B7D682CFFB62A71F8F1946BF8B4" author="Lachner" pageId="3" pageNumber="56" refString="Lachner, E. A. &amp; Karnella, S. J. (1978) Fishes of the genus Eviota of the Red Sea with descriptions of three new species (Teleostei: Gobiidae). Smithsonian Contributions to Zoology, No. 286, 1 - 23." type="book chapter" year="1978">Lachner and Karnella (1978)</bibRefCitation>
</taxonomicName>
. Of these, only 
<taxonomicName id="4C0C4D0F682CFFB62E84F88B97AAF8B4" box="[507,653,1872,1895]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682CFFB62E84F88B97AAF8B4" box="[507,653,1872,1895]" italics="true" pageId="3" pageNumber="56">E. nigrispina</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0C4D0F682CFFB62DE4F88B9635F8B4" box="[667,786,1872,1895]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="prasites">
<emphasis id="B978EA9E682CFFB62DE4F88B9635F8B4" box="[667,786,1872,1895]" italics="true" pageId="3" pageNumber="56">E. prasites</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0C4D0F682CFFB62C5FF88B9680F8B4" box="[800,935,1871,1895]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="pellucida">
<emphasis id="B978EA9E682CFFB62C5FF88B9616F8B4" box="[800,817,1872,1895]" italics="true" pageId="3" pageNumber="56">E</emphasis>
. 
<emphasis id="B978EA9E682CFFB62C40F8949680F8B4" box="[831,935,1871,1895]" italics="true" pageId="3" pageNumber="56">pellucida</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0C4D0F682CFFB62C9BF88B914CF8B4" box="[996,1131,1871,1895]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="storthynx">
<emphasis id="B978EA9E682CFFB62C9BF88B914CF8B4" box="[996,1131,1871,1895]" italics="true" pageId="3" pageNumber="56">E. storthynx</emphasis>
</taxonomicName>
have a dorsal/anal formula of 8/7 as in 
<taxonomicName id="4C0C4D0F682CFFB62E66F8AC9484F85E" box="[281,419,1909,1934]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682CFFB62E66F8AC9484F85E" box="[281,419,1909,1934]" italics="true" pageId="3" pageNumber="56">E. rubriceps</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C0C4D0F682CFFB62ECEF8AC974CF85E" box="[433,619,1909,1934]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682CFFB62ECEF8AC974CF85E" box="[433,619,1909,1934]" italics="true" pageId="3" pageNumber="56">Eviota rubriceps</emphasis>
</taxonomicName>
differs from both 
<taxonomicName id="4C0C4D0F682CFFB62C49F8AC9689F85D" box="[822,942,1911,1934]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="prasites">
<emphasis id="B978EA9E682CFFB62C49F8AC9689F85D" box="[822,942,1911,1934]" italics="true" pageId="3" pageNumber="56">E. prasites</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0C4D0F682CFFB62C9CF8AC914DF85E" box="[995,1130,1909,1934]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="pellucida">
<emphasis id="B978EA9E682CFFB62C9CF8AC914DF85E" box="[995,1130,1909,1934]" italics="true" pageId="3" pageNumber="56">E. pellucida</emphasis>
</taxonomicName>
by having the 5th pelvic-fin ray about 20% or less of the 4th ray, versus about 40%. 
<taxonomicName id="4C0C4D0F682CFFB62C6DF84696E8F867" box="[786,975,1948,1972]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682CFFB62C6DF84696E8F867" box="[786,975,1948,1972]" italics="true" pageId="3" pageNumber="56">Eviota rubriceps</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C0C4D0F682CFFB62B17F84691D6F867" box="[1128,1265,1948,1972]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="storthynx">
<emphasis id="B978EA9E682CFFB62B17F84691D6F867" box="[1128,1265,1948,1972]" italics="true" pageId="3" pageNumber="56">E. storthynx</emphasis>
</taxonomicName>
by lacking the dark postocular spot and vertical bars or dark reticulated markings on the caudal fin present in that species. 
<taxonomicName id="4C0C4D0F682CFFB12A2CF81A9424FF63" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" lastPageId="4" lastPageNumber="57" order="Perciformes" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682CFFB12A2CF81A9424FF63" italics="true" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">Eviota rubriceps</emphasis>
</taxonomicName>
shares the most characters with 
<taxonomicName id="4C0C4D0F682BFFB12D10FF429626FF63" box="[623,769,153,176]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682BFFB12D10FF429626FF63" box="[623,769,153,176]" italics="true" pageId="4" pageNumber="57">E. nigrispina</emphasis>
</taxonomicName>
, but differs by lacking a curved yellow-gold line across the abdomen (
<figureCitation id="13372A09682BFFB12E70FF679474FF07" box="[271,339,188,212]" captionStart="FIGURE 5" captionStartId="4.[151,250,1112,1135]" captionTargetBox="[265,1321,409,1089]" captionTargetId="figure@4.[265,1321,409,1091]" captionTargetPageId="4" captionText="FIGURE 5. Eviota nigrispina. Figure 4 from Greenfield &amp; Suzuki (2010)." httpUri="https://zenodo.org/record/205712/files/figure.png" pageId="4" pageNumber="57">Fig. 5</figureCitation>
) in life and in having a peppering of small dark spots on the nape and upper back lacking in 
<taxonomicName id="4C0C4D0F682BFFB12AFBFF66942DFF2B" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682BFFB12AFBFF6690B2FF07" box="[1412,1429,189,212]" italics="true" pageId="4" pageNumber="57">E</emphasis>
. 
<emphasis id="B978EA9E682BFFB12FE8FF3A942DFF2B" box="[151,266,225,248]" italics="true" pageId="4" pageNumber="57">nigrispina</emphasis>
</taxonomicName>
(
<figureCitation id="13372A09682BFFB12E66FF049479FF24" box="[281,350,223,247]" captionStart="FIGURE 6" captionStartId="4.[151,250,1592,1615]" captionTargetBox="[265,1321,1182,1570]" captionTargetId="figure@4.[265,1321,1182,1570]" captionTargetPageId="4" captionText="FIGURE 6. Eviota nigrispina. Figure 5 from Greenfield &amp; Suzuki (2010)." httpUri="https://zenodo.org/record/205713/files/figure.png" pageId="4" pageNumber="57">Fig. 6</figureCitation>
). The small dark spots on the nape and upper back are very obvious in fresh material, but tend to fade in older specimens (30 plus years) and high magnification is needed to see them. The species also differ in caudal-peduncle depth, with 
<taxonomicName id="4C0C4D0F682BFFB12E9BFEF2975DFE93" box="[484,634,297,320]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682BFFB12E9BFEF2975DFE93" box="[484,634,297,320]" italics="true" pageId="4" pageNumber="57">E. nigrispina</emphasis>
</taxonomicName>
having a slightly deeper caudal peduncle (
<figureCitation id="13372A09682BFFB12B0FFEFC919FFEEC" box="[1136,1208,295,319]" captionStart="FIGURE 7" captionStartId="5.[151,250,1050,1073]" captionTargetBox="[346,1237,431,1015]" captionTargetId="figure@5.[330,1257,409,1029]" captionTargetPageId="5" captionText="FIGURE 7. Caudal-peduncle depth as percentage of SL. Blue diamonds = E. rubriceps; Magenta squares = E. nigrispina." httpUri="https://zenodo.org/record/205714/files/figure.png" pageId="4" pageNumber="57">Fig. 7</figureCitation>
). The 
<taxonomicName id="4C0C4D0F682BFFB12A79FEF290BBFE93" box="[1286,1436,297,320]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682BFFB12A79FEF290BBFE93" box="[1286,1436,297,320]" italics="true" pageId="4" pageNumber="57">E. nigrispina</emphasis>
</taxonomicName>
specimens in 
<figureCitation id="13372A09682BFFB12E50FE979456FEB7" box="[303,369,332,356]" captionStart="FIGURE 7" captionStartId="5.[151,250,1050,1073]" captionTargetBox="[346,1237,431,1015]" captionTargetId="figure@5.[330,1257,409,1029]" captionTargetPageId="5" captionText="FIGURE 7. Caudal-peduncle depth as percentage of SL. Blue diamonds = E. rubriceps; Magenta squares = E. nigrispina." httpUri="https://zenodo.org/record/205714/files/figure.png" pageId="4" pageNumber="57">Fig. 7</figureCitation>
are 
<typeStatus id="54B7882E682BFFB12EDEFE9694FBFEB6" box="[417,476,333,357]" pageId="4" pageNumber="57">types</typeStatus>
from 
<collectingCountry id="F31B761C682BFFB12D5FFE969747FEB7" box="[544,608,333,356]" name="Japan" pageId="4" pageNumber="57">Japan</collectingCountry>
and two specimens from 
<collectingCountry id="F31B761C682BFFB12CFFFE9796CAFEB7" box="[896,1005,332,356]" name="Indonesia" pageId="4" pageNumber="57">Indonesia</collectingCountry>
(ROM 85168, 11.4 &amp; 
<quantity id="4CF49B69682BFFB12B93FE979073FEB7" box="[1260,1364,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="4" pageNumber="57" unit="mm" value="12.1">12.1 mm</quantity>
).
</paragraph>
</subSubSection>
<caption id="DF736604682BFFB12FE8FB8396AAFBBC" box="[151,909,1112,1135]" httpUri="https://zenodo.org/record/205712/files/figure.png" pageId="4" pageNumber="57" targetBox="[265,1321,409,1089]" targetPageId="4">
<paragraph id="8BB3368C682BFFB12FE8FB8396AAFBBC" blockId="4.[151,909,1112,1135]" box="[151,909,1112,1135]" pageId="4" pageNumber="57">
<emphasis id="B978EA9E682BFFB12FE8FB839435FBBD" bold="true" box="[151,274,1112,1135]" pageId="4" pageNumber="57">FIGURE 5.</emphasis>
<taxonomicName id="4C0C4D0F682BFFB12E66FB8294EEFBBD" box="[281,457,1113,1134]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682BFFB12E66FB8294EEFBBD" box="[281,457,1113,1134]" italics="true" pageId="4" pageNumber="57">Eviota nigrispina</emphasis>
</taxonomicName>
. Figure 4 from Greenfield &amp; Suzuki (2010).
</paragraph>
</caption>
<caption id="DF736604682BFFB12FE8F9E396AAF99C" box="[151,909,1592,1615]" httpUri="https://zenodo.org/record/205713/files/figure.png" pageId="4" pageNumber="57" targetBox="[265,1321,1182,1570]" targetPageId="4">
<paragraph id="8BB3368C682BFFB12FE8F9E396AAF99C" blockId="4.[151,909,1592,1615]" box="[151,909,1592,1615]" pageId="4" pageNumber="57">
<emphasis id="B978EA9E682BFFB12FE8F9E39435F99D" bold="true" box="[151,274,1592,1615]" pageId="4" pageNumber="57">FIGURE 6.</emphasis>
<taxonomicName id="4C0C4D0F682BFFB12E66F9E194EEF99C" box="[281,457,1594,1615]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682BFFB12E66F9E194EEF99C" box="[281,457,1594,1615]" italics="true" pageId="4" pageNumber="57">Eviota nigrispina</emphasis>
</taxonomicName>
. Figure 5 from Greenfield &amp; Suzuki (2010).
</paragraph>
</caption>
<subSubSection id="C3166507682BFFB12FB8F9AF91F8F907" pageId="4" pageNumber="57" type="materials_examined">
<paragraph id="8BB3368C682BFFB12FB8F9AF91F8F907" blockId="4.[151,1436,1652,2037]" pageId="4" pageNumber="57">
<emphasis id="B978EA9E682BFFB12FB8F9AF972CF95E" bold="true" box="[199,523,1652,1677]" pageId="4" pageNumber="57">Other material examined.</emphasis>
<taxonomicName id="4C0C4D0F682BFFB12D65F9AD97C2F95E" box="[538,741,1654,1677]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682BFFB12D65F9AD97C2F95E" box="[538,741,1654,1677]" italics="true" pageId="4" pageNumber="57">Eviota nigrispina</emphasis>
</taxonomicName>
—OMNH-P35615, 
<typeStatus id="54B7882E682BFFB12CB4F9AF9117F95F" box="[971,1072,1652,1676]" pageId="4" pageNumber="57" type="holotype">holotype</typeStatus>
, Ryukyu Islands, 
<collectingCountry id="F31B761C682BFFB12A70F9AE9075F95F" box="[1295,1362,1653,1676]" name="Japan" pageId="4" pageNumber="57">Japan</collectingCountry>
; 
<collectionCode id="ED1DAE49682BFFB12A1AF9AF90BCF95F" LSID="urn:lsid:biocol.org:col:34700" box="[1381,1435,1652,1676]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="4" pageNumber="57">CAS</collectionCode>
229934, 
<typeStatus id="54B7882E682BFFB12F81F9429446F962" box="[254,353,1689,1713]" pageId="4" pageNumber="57" type="paratype">paratype</typeStatus>
, Ryukyu Islands, 
<collectingCountry id="F31B761C682BFFB12D47F942975BF963" box="[568,636,1689,1712]" name="Japan" pageId="4" pageNumber="57">Japan</collectingCountry>
; OMNH-P35614, 
<typeStatus id="54B7882E682BFFB12C2CF9429691F962" box="[851,950,1689,1713]" pageId="4" pageNumber="57" type="paratype">paratype</typeStatus>
, Ryukyu Islands, 
<collectingCountry id="F31B761C682BFFB12BF2F94291F7F963" box="[1165,1232,1689,1712]" name="Japan" pageId="4" pageNumber="57">Japan</collectingCountry>
; OMNH-P35640, 
<typeStatus id="54B7882E682BFFB12FE8F96695DDF906" box="[151,250,1725,1749]" pageId="4" pageNumber="57" type="paratype">paratype</typeStatus>
, Ryukyu Islands, 
<collectingCountry id="F31B761C682BFFB12EBEF9669722F907" box="[449,517,1725,1748]" name="Japan" pageId="4" pageNumber="57">Japan</collectingCountry>
; 
<collectionCode id="ED1DAE49682BFFB12D6FF9679777F907" LSID="urn:lsid:biocol.org:col:34778" box="[528,592,1724,1748]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="4" pageNumber="57">ROM</collectionCode>
85168 (2), 
<collectingCountry id="F31B761C682BFFB12DADF9679665F907" box="[722,834,1724,1748]" name="Indonesia" pageId="4" pageNumber="57">Indonesia</collectingCountry>
, 
<taxonomicName id="4C0C4D0F682BFFB12C32F96796A7F907" box="[845,896,1724,1748]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="61" phylum="Chordata" rank="species" species="raja">Raja</taxonomicName>
Ampat Islands, Waigeo 
<collectingCountry id="F31B761C682BFFB12BECF96791FCF907" box="[1171,1243,1724,1748]" name="Iceland" pageId="4" pageNumber="57">Island</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3166507682BFFB32FB8F93A95D0FDB0" lastPageId="6" lastPageNumber="59" pageId="4" pageNumber="57" type="discussion">
<paragraph id="8BB3368C682BFFB12FB8F93A96D4F8B6" blockId="4.[151,1436,1652,2037]" pageId="4" pageNumber="57">
<emphasis id="B978EA9E682BFFB12FB8F93A941CF92A" bold="true" box="[199,315,1761,1785]" pageId="4" pageNumber="57">Remarks.</emphasis>
While conducting a survey of the fishes at 
<taxonomicName id="4C0C4D0F682BFFB12C4AF93B964EF92B" box="[821,873,1760,1784]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="61" phylum="Chordata" rank="species" species="raja">Raja</taxonomicName>
Ampat, 
<collectingCountry id="F31B761C682BFFB12CB0F93B9118F92B" box="[975,1087,1760,1784]" name="Indonesia" pageId="4" pageNumber="57">Indonesia</collectingCountry>
, Gerald R. Allen and Mark V. Erdmann found a species of 
<taxonomicName id="4C0C4D0F682BFFB12EA3F8DD9702F8CE" box="[476,549,1798,1821]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E682BFFB12EA3F8DD9702F8CE" box="[476,549,1798,1821]" italics="true" pageId="4" pageNumber="57">Eviota</emphasis>
</taxonomicName>
with a distinctive red head and a body that was dark on the lower half, which they referred to as 
<taxonomicName id="4C0C4D0F682BFFB12E16F8F394BBF893" box="[361,412,1832,1856]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="61" phylum="Chordata" rank="species" species="raja">Raja</taxonomicName>
Ampat sp. 1. Subsequently they sent photographs to us and we recognized the similarity to the specimens previously given a manuscript name and awaiting description.
</paragraph>
<paragraph id="8BB3368C682BFFB02FB8F8AB97B4FEB7" blockId="4.[151,1436,1652,2037]" lastBlockId="5.[151,1437,151,356]" lastPageId="5" lastPageNumber="58" pageId="4" pageNumber="57">
Prior to this, 
<bibRefCitation id="EF9D4B7D682BFFB12E27F8AB9782F85B" author="Greenfield" box="[344,677,1904,1928]" pageId="4" pageNumber="57" refString="Greenfield, D. W. &amp; Suzuki, T. (2010) Eviota nigrispina, a new goby from the Ryukyu Islands, Japan (Teleostei: Gobiidae). Zootaxa, 2655, 57 - 62." type="journal article" year="2010">Greenfield and Suzuki (2010)</bibRefCitation>
described an 
<taxonomicName id="4C0C4D0F682BFFB12C3FF8A996AEF85A" box="[832,905,1906,1929]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E682BFFB12C3FF8A996AEF85A" box="[832,905,1906,1929]" italics="true" pageId="4" pageNumber="57">Eviota</emphasis>
</taxonomicName>
species from 
<collectingCountry id="F31B761C682BFFB12B5BF8AA9140F85B" box="[1060,1127,1905,1928]" name="Japan" pageId="4" pageNumber="57">Japan</collectingCountry>
, 
<taxonomicName id="4C0C4D0F682BFFB12B0DF8A99023F85A" box="[1138,1284,1906,1929]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682BFFB12B0DF8A99023F85A" box="[1138,1284,1906,1929]" italics="true" pageId="4" pageNumber="57">E. nigrispina</emphasis>
</taxonomicName>
, that also has a dark band on the ventral portion of the body when preserved. Comparison of photographs of live individuals of 
<taxonomicName id="4C0C4D0F682BFFB12AFBF84D942DF802" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682BFFB12AFBF84D942DF802" italics="true" pageId="4" pageNumber="57">E. nigrispina</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0C4D0F682BFFB12E39F86194F2F802" box="[326,469,1977,2001]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682BFFB12E39F86194F2F802" box="[326,469,1977,2001]" italics="true" pageId="4" pageNumber="57">E. rubriceps</emphasis>
</taxonomicName>
showed that they were quite different in coloration, however, they are very similar when preserved and have the same counts. 
<collectingCountry id="F31B761C682BFFB12DFDF8069786F826" box="[642,673,2013,2037]" name="American Samoa" pageId="4" pageNumber="57">As</collectingCountry>
discussed in the Comparisons section, in addition to the live coloration, there also are differences in preserved coloration and caudal-peduncle depth. There also are habitat differences between the two at the 
<taxonomicName id="4C0C4D0F682AFFB02EA2FF679736FF07" box="[477,529,188,212]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="61" phylum="Chordata" rank="species" species="raja">Raja</taxonomicName>
Ampat Islands. Erdmann (pers. comm., 
<date id="FFB2104C682AFFB02CAAFF679141FF07" box="[981,1126,188,212]" pageId="5" pageNumber="58" value="2011-07-20">20 July 2011</date>
) reports that 
<taxonomicName id="4C0C4D0F682AFFB02B87FF6690A5FF07" box="[1272,1410,188,212]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682AFFB02B87FF6690A5FF07" box="[1272,1410,188,212]" italics="true" pageId="5" pageNumber="58">E. rubriceps</emphasis>
</taxonomicName>
is a dominant species in shallow waters (mostly 
<quantity id="4CF49B69682AFFB02DCCFF049627FF24" box="[691,768,223,247]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="5" pageNumber="58" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">2–4 m</quantity>
, maximum 
<quantity id="4CF49B69682AFFB02CF3FF049699FF24" box="[908,958,223,247]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="58" unit="m" value="6.0">6 m</quantity>
). Two specimens of 
<taxonomicName id="4C0C4D0F682AFFB02BCDFF3A906FFF2B" box="[1202,1352,225,248]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682AFFB02BCDFF3A906FFF2B" box="[1202,1352,225,248]" italics="true" pageId="5" pageNumber="58">E. nigrispina</emphasis>
</taxonomicName>
(ROM 85168) were taken at a depth of 
<quantity id="4CF49B69682AFFB02D74FEDF9762FECF" box="[523,581,260,284]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="58" unit="m" value="22.0">22 m</quantity>
at Waigeo 
<collectingCountry id="F31B761C682AFFB02DB8FEDF9629FECF" box="[711,782,260,284]" name="Iceland" pageId="5" pageNumber="58">Island</collectingCountry>
, 
<taxonomicName id="4C0C4D0F682AFFB02C64FEDF9669FECF" box="[795,846,260,284]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="61" phylum="Chordata" rank="species" species="raja">Raja</taxonomicName>
Ampat Islands (
<figureCitation id="13372A09682AFFB02B73FEDF9176FECF" box="[1036,1105,260,284]" captionStart="FIGURE 8" captionStartId="5.[151,250,1527,1550]" captionTargetBox="[265,1321,1122,1506]" captionTargetId="figure@5.[265,1321,1122,1506]" captionTargetPageId="5" captionText="FIGURE 8. Eviota nigrispina, ROM 85168, Raja Ampat Islands." httpUri="https://zenodo.org/record/205715/files/figure.png" pageId="5" pageNumber="58">Fig. 8</figureCitation>
). Although 
<taxonomicName id="4C0C4D0F682AFFB02BA5FEDE9049FECF" box="[1242,1390,261,284]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682AFFB02BA5FEDE9049FECF" box="[1242,1390,261,284]" italics="true" pageId="5" pageNumber="58">E. nigrispina</emphasis>
</taxonomicName>
has been seen in shallow water at the Ryukyu Islands, 
<collectingCountry id="F31B761C682AFFB02DB5FEF3962DFEEC" box="[714,778,296,319]" name="Japan" pageId="5" pageNumber="58">Japan</collectingCountry>
where the water is cooler (pers. comm., T. Suzuki, 
<date id="FFB2104C682AFFB02A35FEFC95F5FEB7" pageId="5" pageNumber="58" value="2011-07-21">21 July 2011</date>
), the 
<typeStatus id="54B7882E682AFFB02E71FE979457FEB7" box="[270,368,332,356]" pageId="5" pageNumber="58" type="holotype">holotype</typeStatus>
was collected from 
<specimenCount id="9D0AFD05682AFFB02D2CFE9797B4FEB7" box="[595,659,332,356]" pageId="5" pageNumber="58" type="generic">
<quantity id="4CF49B69682AFFB02D2CFE9797A8FEB7" box="[595,655,332,356]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="58" unit="m" value="20.0">20 m</quantity>
.
</specimenCount>
</paragraph>
<caption id="DF736604682AFFB02FE8FBC1904BFBE2" box="[151,1388,1050,1073]" httpUri="https://zenodo.org/record/205714/files/figure.png" pageId="5" pageNumber="58" targetBox="[346,1237,431,1015]" targetPageId="5">
<paragraph id="8BB3368C682AFFB02FE8FBC1904BFBE2" blockId="5.[151,1388,1050,1073]" box="[151,1388,1050,1073]" pageId="5" pageNumber="58">
<emphasis id="B978EA9E682AFFB02FE8FBC19434FBE3" bold="true" box="[151,275,1050,1073]" pageId="5" pageNumber="58">FIGURE 7.</emphasis>
Caudal-peduncle depth as percentage of SL. Blue diamonds = 
<taxonomicName id="4C0C4D0F682AFFB02CEDFBC79128FBE2" box="[914,1039,1051,1073]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="rubriceps">
<emphasis id="B978EA9E682AFFB02CEDFBC79128FBE2" box="[914,1039,1051,1073]" italics="true" pageId="5" pageNumber="58">E. rubriceps</emphasis>
</taxonomicName>
; Magenta squares = 
<taxonomicName id="4C0C4D0F682AFFB02B9DFBC79041FBE2" box="[1250,1382,1052,1073]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682AFFB02B9DFBC79041FBE2" box="[1250,1382,1052,1073]" italics="true" pageId="5" pageNumber="58">E. nigrispina</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF736604682AFFB02FE8FA2C9616F9DD" box="[151,817,1527,1550]" httpUri="https://zenodo.org/record/205715/files/figure.png" pageId="5" pageNumber="58" targetBox="[265,1321,1122,1506]" targetPageId="5">
<paragraph id="8BB3368C682AFFB02FE8FA2C9616F9DD" blockId="5.[151,817,1527,1550]" box="[151,817,1527,1550]" pageId="5" pageNumber="58">
<emphasis id="B978EA9E682AFFB02FE8FA2C9435F9DE" bold="true" box="[151,274,1527,1550]" pageId="5" pageNumber="58">FIGURE 8.</emphasis>
<taxonomicName id="4C0C4D0F682AFFB02E66FA2294EEF9DD" box="[281,457,1529,1550]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="nigrispina">
<emphasis id="B978EA9E682AFFB02E66FA2294EEF9DD" box="[281,457,1529,1550]" italics="true" pageId="5" pageNumber="58">Eviota nigrispina</emphasis>
</taxonomicName>
, ROM 85168, 
<taxonomicName id="4C0C4D0F682AFFB02D1DFA2397B7F9DD" box="[610,656,1528,1550]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="61" phylum="Chordata" rank="species" species="raja">Raja</taxonomicName>
Ampat Islands.
</paragraph>
</caption>
<paragraph id="8BB3368C682AFFB02FB8F9EE9450F89E" blockId="5.[151,1437,1588,1979]" pageId="5" pageNumber="58">
Except for the close difference in caudal-peduncle depth, only color separates these two species. Color alone has been found to separate some other coral-reef fish species. 
<bibRefCitation id="EF9D4B7D682AFFB02C24F9839102F9A3" author="Greenfield" box="[859,1061,1624,1648]" pageId="5" pageNumber="58" refString="Greenfield, D. W. (1979) A review of the western Atlantic Starksia ocellata - complex (Pisces: Clinidae) with the description of two new species and proposal of superspecies status. Fieldiana Zoology, 73, 9 - 48." type="journal article" year="1979">Greenfield (1979)</bibRefCitation>
found that several species of the labrisomid genus 
<taxonomicName id="4C0C4D0F682AFFB02E1FF9A7949DF947" box="[352,442,1660,1684]" class="Actinopterygii" family="Labrisomidae" genus="Starksia" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E682AFFB02E1FF9A7949DF947" box="[352,442,1660,1684]" italics="true" pageId="5" pageNumber="58">Starksia</emphasis>
</taxonomicName>
could only be distinguished on the basis of color. 
<bibRefCitation id="EF9D4B7D682AFFB02C88F9A791CEF947" box="[1015,1257,1660,1684]" pageId="5" pageNumber="58" refString="Baldwin, C. C., Castillo, C. I., Weigt, L. A. &amp; Victor, B. C. (2011) Seven new species within western Atlantic Starksia atlantica, S. lepicoelia, and S. sluiteri (Teleostei, Labrisomidae), with comments on congruence of DNA barcodes and species. ZooKeys, 79, 21 - 72." type="journal article">
Baldwin 
<emphasis id="B978EA9E682AFFB02B21F9A691B0F947" box="[1118,1175,1660,1684]" italics="true" pageId="5" pageNumber="58">et al.</emphasis>
(2011)
</bibRefCitation>
reevaluated the species using cytochrome c oxidase (CO1) sequences and concluded that the data “support his decision to recognize the species almost entirely on the basis of minor differences in pigment Species in the gobiid genus 
<taxonomicName id="4C0C4D0F682AFFB02A36F91395E1F8D7" class="Actinopterygii" family="Gobiidae" genus="Elacatinus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E682AFFB02A36F91395E1F8D7" italics="true" pageId="5" pageNumber="58">Elacatinus</emphasis>
</taxonomicName>
provide another example of species that are separated only by color but are genetically distinct. 
<bibRefCitation id="EF9D4B7D682AFFB02A56F9369477F8F4" author="Tayor" pageId="5" pageNumber="58" refString="Taylor, M. S. &amp; Hellberg, M. E. (2005) Marine radiations at small geographic scales: speciation in neotropical reef gobies (Elacatinus). Evolution, 59 (2), 374 - 385." type="journal article" year="2005">Tayor and Hellberg (2005)</bibRefCitation>
suggest that color-based mate choice and ecological differences may have contributed to diversification in this genus.
</paragraph>
<paragraph id="8BB3368C682AFFB02FB8F88C9739F868" blockId="5.[151,1437,1588,1979]" pageId="5" pageNumber="58">
<bibRefCitation id="EF9D4B7D682AFFB02FB8F88C94BCF8BC" box="[199,411,1879,1904]" pageId="5" pageNumber="58" refString="Craig, M. T., Hastings, P. A., Pondella II, D. J., Robertson, D. R. &amp; Rosales-Casian, J. A. (2006) Phylogeography of the flag cabrilla Epinephelus labriformis (Serranidae): implications for the biogeography of the Tropical Eastern Pacific and the early stages of speciation in a marine shore fish. Journal of Biogeography, 33 (6), 969 - 979." type="journal article">
Craig 
<emphasis id="B978EA9E682AFFB02E72F8829461F8A3" box="[269,326,1880,1904]" italics="true" pageId="5" pageNumber="58">et al.</emphasis>
(2006)
</bibRefCitation>
, in discussing two genetically distinct grouper species that differ in coloration but are very similar morphologically, suggest that “the evolution of external colours precedes or occurs more rapidly than changes in molecular markers……
</paragraph>
<paragraph id="8BB3368C6829FFB32FB8FF4C9437FF24" blockId="6.[151,1436,151,611]" pageId="6" pageNumber="59">
<bibRefCitation id="EF9D4B7D6829FFB32FB8FF4C94FEFF7C" author="Victor" box="[199,473,151,175]" pageId="6" pageNumber="59" refString="Victor, B. C. &amp; Randall, J. E. (2010) Gramma dejongi, a new basslet (Perciformes: Grammatidae) from Cuba, a sympatric sibling species of G. loreto. Zoological Studies 49 (6), 865 - 871." type="journal article" year="2010">Victor &amp; Randall (2010)</bibRefCitation>
described 
<taxonomicName id="4C0C4D0F6829FFB32D2DFF439636FF63" box="[594,785,152,176]" class="Actinopterygii" family="Grammatidae" genus="Gramma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="59" phylum="Chordata" rank="species" species="dejongi">
<emphasis id="B978EA9E6829FFB32D2DFF439636FF63" box="[594,785,152,176]" italics="true" pageId="6" pageNumber="59">Gramma dejongi</emphasis>
</taxonomicName>
based solely on color, adult size, behavior, and ecological differences. In this case the barcode CO1 mtDNA was the same as a sympatric congener and they suggested recent speciation.
</paragraph>
<paragraph id="8BB3368C6829FFB32FB8FEDE95F2FE7C" blockId="6.[151,1436,151,611]" pageId="6" pageNumber="59">
<bibRefCitation id="EF9D4B7D6829FFB32FB8FEDE9492FECF" box="[199,437,260,284]" pageId="6" pageNumber="59" refString="Bowen, B. W., Muss, A., Rocha, L. A. &amp; Grant, W. S. (2006) Shallow mtDNA coalescence in Atlantic pygmy angelfishes (Genus Centropyge) indicates a recent invasion from the Indian Ocean. Journal of Heredity, 97 (1), 1 - 12." type="journal article">
Bowen 
<emphasis id="B978EA9E6829FFB32E5EFEDE947AFECF" box="[289,349,260,284]" italics="true" pageId="6" pageNumber="59">et al.</emphasis>
(2006)
</bibRefCitation>
, however, in discussing Atlantic pygmy angelfish species (
<emphasis id="B978EA9E6829FFB32B16FEDE91F9FECF" box="[1129,1246,261,284]" italics="true" pageId="6" pageNumber="59">Centropyg</emphasis>
e) could find no genetic differences between differently colored species and favored the conclusion that the species were color morphs of the same species. But they also stated that “color morphs subject to sexual selection or ecological differentiation (and lacking morphological or molecular diagnostics) are the raw edge of emerging species on coral reefs.
</paragraph>
<paragraph id="8BB3368C6829FFB32FB8FE6095D0FDB0" blockId="6.[151,1436,151,611]" pageId="6" pageNumber="59">
The small size of the dwarfgobies in the genus 
<taxonomicName id="4C0C4D0F6829FFB32DA4FE669603FE07" box="[731,804,445,468]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E6829FFB32DA4FE669603FE07" box="[731,804,445,468]" italics="true" pageId="6" pageNumber="59">Eviota</emphasis>
</taxonomicName>
, as well as some other small gobies such as those in the genus 
<taxonomicName id="4C0C4D0F6829FFB32FA0FE3B9412FE24" box="[223,309,480,503]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E6829FFB32FA0FE3B9412FE24" box="[223,309,480,503]" italics="true" pageId="6" pageNumber="59">Trimma</emphasis>
</taxonomicName>
, poses problems because this reduction in size often results in the reduction of the number of scales, fin rays and pores of the laterosensory canal system (
<bibRefCitation id="EF9D4B7D6829FFB32D9CFDD896D6FDC8" author="Weitzman" box="[739,1009,515,539]" pageId="6" pageNumber="59" refString="Weitzman, S. H. &amp; Vari, R. P. (1988) Miniaturization in South American freshwater fishes; an overview and discussion. Proceedings of the Biological Society of Washington, 101, 444 - 465." type="journal article" year="1988">Weitzman &amp; Vari, 1988</bibRefCitation>
). 
<collectingCountry id="F31B761C6829FFB32B7BFDDF9104FDCF" box="[1028,1059,516,540]" name="American Samoa" pageId="6" pageNumber="59">As</collectingCountry>
a result the potential for variation in morphological characters is reduced and many of the species of 
<taxonomicName id="4C0C4D0F6829FFB32CFEFDF396EDFDEC" box="[897,970,552,575]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="B978EA9E6829FFB32CFEFDF396EDFDEC" box="[897,970,552,575]" italics="true" pageId="6" pageNumber="59">Eviota</emphasis>
</taxonomicName>
are primarily distinguished by their color patterns.
</paragraph>
</subSubSection>
</treatment>
</document>