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<document id="9A0C4938329B16A084BFEC388C012561" ID-DOI="10.5281/zenodo.155299" ID-GBIF-Dataset="22750794-cbfc-4d09-a4ae-bcdb17a35113" ID-Zenodo-Dep="155299" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477431652181" checkinUser="plazi" docAuthor="Volker Mahnert &amp; Yun-chun Li" docDate="2016" docId="039980242836273CFC3D9C29FA9ACDF8" docLanguage="en" docName="7_Mahnert_Li.pdf" docOrigin="Revue suisse de Zoologie 123" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Bisetocreagris chuanensis Mahnert &amp; Li, 2016, n. sp." docType="treatment" docUuid="CE7E2922-9075-4577-B8B0-90FB5066D4B4" docUuidSource="ZooBank" docVersion="11" lastPageNumber="261" masterDocId="FFA0F85C2837273EFF819F3AFFF7CB6F" masterDocTitle="Cave-inhabiting Neobisiidae (Arachnida: Pseudoscorpiones) from China, with description of four new species of Bisetocreagris Ćurčić" masterLastPageNumber="268" masterPageNumber="259" pageNumber="260" updateTime="1698390683281" updateUser="plazi">
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<mods:title id="FA57485E40283E3F3CA9BE7EB8CA2787">Cave-inhabiting Neobisiidae (Arachnida: Pseudoscorpiones) from China, with description of four new species of Bisetocreagris Ćurčić</mods:title>
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<mods:namePart id="D7534CDB22F7F32596F1EFEDDC349C61">Volker Mahnert</mods:namePart>
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<mods:namePart id="E2E8BF3EB3E3F9D5CD027382AB962061">Yun-chun Li</mods:namePart>
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<mods:typeOfResource id="0DFCDF43F030DE8CB0F23756D653EFE2">text</mods:typeOfResource>
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<mods:title id="A027C915C02FDF5DDBA77AE19DE2E8C3">Revue suisse de Zoologie</mods:title>
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<mods:part id="5FCCD7F1F871B8558F7ECD8DE4BB6D81">
<mods:date id="D7985729C1D75F1A0698F2B9BDA3264F">2016</mods:date>
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<mods:number id="4AA6D1667E4765B91A39C28C65808539">123</mods:number>
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<treatment id="039980242836273CFC3D9C29FA9ACDF8" ID-DOI="http://doi.org/10.5281/zenodo.6080948" ID-GBIF-Taxon="124558068" ID-Zenodo-Dep="6080948" LSID="urn:lsid:zoobank.org:act:CE7E2922-9075-4577-B8B0-90FB5066D4B4" httpUri="http://treatment.plazi.org/id/039980242836273CFC3D9C29FA9ACDF8" lastPageId="2" lastPageNumber="261" pageId="1" pageNumber="260">
<subSubSection id="C32A62B92836273FFC3D9C29FAE8C842" box="[956,1311,787,813]" pageId="1" pageNumber="260" type="nomenclature">
<paragraph id="8B8F31322836273FFC3D9C29FAE8C842" blockId="1.[956,1311,787,843]" box="[956,1311,787,813]" pageId="1" pageNumber="260">
<heading id="D0C7865E2836273FFC3D9C29FAE8C842" bold="true" box="[956,1311,787,813]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="260" reason="4">
<emphasis id="B944ED202836273FFC3D9C29FAE8C842" bold="true" box="[956,1311,787,813]" pageId="1" pageNumber="260">
<taxonomicName id="4C304AB12836273FFC3D9C29FB2BC844" ID-CoL="LWS7" box="[956,1244,787,811]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="260" phylum="Arthropoda" rank="species" species="chuanensis" status="sp. nov.">
<emphasis id="B944ED202836273FFC3D9C29FB2BC844" bold="true" box="[956,1244,787,811]" italics="true" pageId="1" pageNumber="260">Bisetocreagris chuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277505B2836273FFB629C2FFAE8C842" box="[1251,1311,789,813]" pageId="1" pageNumber="260" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92836273FFBC39C0EFB6EC824" box="[1090,1177,820,843]" pageId="1" pageNumber="260" type="description">
<paragraph id="8B8F31322836273FFBC39C0EFB6EC824" blockId="1.[956,1311,787,843]" box="[1090,1177,820,843]" pageId="1" pageNumber="260">
<figureCitation id="130B2DB72836273FFBC39C0EFB6EC824" box="[1090,1177,820,843]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="1" pageNumber="260">Figs 1-5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92836273FFCBE9C5EFBCFCFA2" pageId="1" pageNumber="260" type="materials_examined">
<paragraph id="8B8F31322836273FFCBE9C5EFADCCF74" blockId="1.[831,1437,867,1052]" pageId="1" pageNumber="260">
<materialsCitation id="3B583B6F2836273FFCBE9C5EFC17C8B4" ID-GBIF-Occurrence="1324024076" collectingDate="2010-04-30" collectionCode="MSWU" country="China" elevation="1387" latitude="27.396616" location="Jin Sha" longLatPrecision="1" longitude="106.10173" municipality="Xin Hua Town" pageId="1" pageNumber="260" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou Province" typeStatus="holotype">
<emphasis id="B944ED202836273FFCBE9C5EFC44C813" bold="true" box="[831,947,868,892]" pageId="1" pageNumber="260">
<typeStatus id="548B8F902836273FFCBE9C5EFC5AC813" box="[831,941,868,892]" pageId="1" pageNumber="260" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED21A9F72836273FFC469C5EFBE8C813" box="[967,1055,868,892]" pageId="1" pageNumber="260">MSWU</collectionCode>
;
<specimenCount id="9D36FABB2836273FFBB69C5EFBBCC814" box="[1079,1099,868,891]" pageId="1" pageNumber="260" type="male"></specimenCount>
;
<collectingCountry id="F32771A22836273FFBE69C59FB5CC815" box="[1127,1195,867,890]" name="China" pageId="1" pageNumber="260">China</collectingCountry>
,
<collectingRegion id="49F4FFD02836273FFB429C59FA6EC815" box="[1219,1433,867,890]" country="China" name="Guizhou" pageId="1" pageNumber="260">Guizhou Province</collectingRegion>
,
<collectingMunicipality id="6BEBAB482836273FFCBE9CB9FC07C8F3" box="[831,1008,899,924]" pageId="1" pageNumber="260">Xin Hua Town</collectingMunicipality>
,
<location id="8EEF67E92836273FFB839CB9FBADC8F5" LSID="urn:lsid:plazi:treatment:039980242836273CFC3D9C29FA9ACDF8:8EEF67E92836273FFB839CB9FBADC8F5" box="[1026,1114,899,922]" country="China" latitude="27.396616" longLatPrecision="1" longitude="106.10173" municipality="Xin Hua Town" name="Jin Sha" pageId="1" pageNumber="260" stateProvince="Guizhou Province">Jin Sha</location>
,
<location id="8EEF67E92836273FFBEC9CB9FB0CC8F3" LSID="urn:lsid:plazi:treatment:039980242836273CFC3D9C29FA9ACDF8:8EEF67E92836273FFBEC9CB9FB0CC8F3" box="[1133,1275,899,924]" country="China" latitude="27.396616" longLatPrecision="1" longitude="106.10173" municipality="Xin Hua Town" name="Chuan Cave" pageId="1" pageNumber="260" stateProvince="Guizhou Province">Chuan Cave</location>
, alt.
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<elevation id="001DD6012836273FFABC9CBEFA6FC8F3" box="[1341,1432,900,924]" metricMagnitude="3" metricUnit="m" metricValue="1.387" pageId="1" pageNumber="260" unit="m" value="1387.0">1387 m</elevation>
</quantity>
,
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<geoCoordinate id="EE0457F52836273FFC629C9EFB77C8D4" box="[995,1152,932,955]" direction="east" orientation="longitude" pageId="1" pageNumber="260" precision="1" value="106.10173">106°06.104E</geoCoordinate>
;
<date id="FF8E17F22836273FFB1B9C9EFAA5C8D3" box="[1178,1362,932,956]" pageId="1" pageNumber="260" value="2010-04-30">
<collectingDate id="EFCAEE1A2836273FFB1B9C9EFAA5C8D3" box="[1178,1362,932,956]" pageId="1" pageNumber="260" value="2010-04-30">30 April 2010</collectingDate>
</date>
, coll.
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</materialsCitation>
. This large cave has a length of
<quantity id="4CC89CD72836273FFAD49CFEFA6BC8B3" box="[1365,1436,964,988]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="260" unit="m" value="700.0">700 m</quantity>
and is
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high; the pseudoscorpion was collected by hand under rocks, together with some spiders.
</paragraph>
<paragraph id="8B8F31322836273FFCBE9B0EFBCFCFA2" blockId="1.[831,1437,1075,1229]" pageId="1" pageNumber="260">
<emphasis id="B944ED202836273FFCBE9B0EFC49CF23" bold="true" box="[831,958,1076,1100]" pageId="1" pageNumber="260">
<typeStatus id="548B8F902836273FFCBE9B0EFC4FCF23" box="[831,952,1076,1100]" pageId="1" pageNumber="260" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D36FABB2836273FFC549B0EFC01CF24" box="[981,1014,1076,1100]" pageId="1" pageNumber="260" type="male">1♂</specimenCount>
(in
<collectionCode id="ED21A9F72836273FFBC29B0EFB5FCF23" box="[1091,1192,1076,1100]" pageId="1" pageNumber="260">MCSNV</collectionCode>
),
<specimenCount id="9D36FABB2836273FFB4B9B0EFB1BCF24" box="[1226,1260,1076,1100]" pageId="1" pageNumber="260" type="female">1♀</specimenCount>
(in
<collectionCode id="ED21A9F72836273FFAB89B0EFA66CF23" box="[1337,1425,1076,1100]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="260">MHNG</collectionCode>
);
<collectingCountry id="F32771A22836273FFCBE9B69FC75CF05" box="[831,898,1107,1130]" name="China" pageId="1" pageNumber="260">China</collectingCountry>
,
<collectingRegion id="49F4FFD02836273FFC129B69FB97CF05" box="[915,1120,1107,1130]" country="China" name="Guizhou" pageId="1" pageNumber="260">Guizhou Province</collectingRegion>
, Qianxi, Hong Lin Village, Wang Tian Dong, alt.
<quantity id="4CC89CD72836273FFBCA9B4EFB51CFE3" box="[1099,1190,1140,1164]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="260" unit="m" value="1500.0">1500 m</quantity>
, cave length
<quantity id="4CC89CD72836273FFACB9B4EFA6FCFE3" box="[1354,1432,1140,1164]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="260" unit="m" value="770.0">770 m</quantity>
,
<geoCoordinate id="EE0457F52836273FFCBE9BAEFC3DCFC4" box="[831,970,1172,1195]" direction="north" orientation="latitude" pageId="1" pageNumber="260" precision="15" value="27.116388">27°0659N</geoCoordinate>
,
<geoCoordinate id="EE0457F52836273FFC5B9BAEFB98CFC4" box="[986,1135,1172,1195]" direction="east" orientation="longitude" pageId="1" pageNumber="260" precision="15" value="105.89361">105°5337E</geoCoordinate>
;
<date id="FF8E17F22836273FFB019BAFFAAACFC2" box="[1152,1373,1172,1197]" pageId="1" pageNumber="260" value="2001-11-13">13 November 2001</date>
; coll. L. Latello &amp; G. Gozzo.
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92836273FFCBE9BD9FB91CD13" pageId="1" pageNumber="260" type="diagnosis">
<paragraph id="8B8F31322836273FFCBE9BD9FB91CD13" blockId="1.[831,1437,1251,1661]" pageId="1" pageNumber="260">
<emphasis id="B944ED202836273FFCBE9BD9FC4FCF95" bold="true" box="[831,952,1251,1274]" pageId="1" pageNumber="260">Diagnosis:</emphasis>
Troglobiomorphic habitus; carapace with 2 small eyes with indistinct lenses, 8 setae on posterior margin; anterior tergites with 10-11 marginal setae; pedipalp smooth excepted for hand with finely granular dorsodistal part; pedipalp slender, femur 4.4-4.8 times (length 1.56-1.70), patella 3.1-3.5 times (length 1.39- 1.53), club of patella 1.16-1.34 times longer than pedicel and about half the length of entire patella, hand with pedicel 1.7-2.0 times (length 1.22-1.42), chela with pedicel 3.7-4.0 times (length 2.57-2.83) longer than broad, finger as long as or longer than hand with pedicel. Chelicera: rallum with 8-9 pinnate setae, distal one with an expanded base.
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92836273FFCBE99AFFB3ECDA2" pageId="1" pageNumber="260" type="etymology">
<paragraph id="8B8F31322836273FFCBE99AFFB3ECDA2" blockId="1.[831,1437,1685,1741]" pageId="1" pageNumber="260">
<emphasis id="B944ED202836273FFCBE99AFFC33CDC2" bold="true" box="[831,964,1685,1709]" pageId="1" pageNumber="260">Etymology:</emphasis>
Latinized adjective, derived from the name of the locus typicus, the Chuan cave.
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92836273CFCBE99DEFAFEC8D6" lastPageId="2" lastPageNumber="261" pageId="1" pageNumber="260" type="description">
<paragraph id="8B8F31322836273CFCBE99DEFD7BC875" blockId="1.[831,1437,1764,2078]" lastBlockId="2.[151,757,258,2078]" lastPageId="2" lastPageNumber="261" pageId="1" pageNumber="260">
<emphasis id="B944ED202836273FFCBE99DEFA6ACD94" bold="true" box="[831,1437,1764,1789]" pageId="1" pageNumber="260">Description of males (female in parentheses):</emphasis>
Carapace in anterior half reddish or yellow, basal part lighter and with several unsclerotized areas; pedipalp and chelicera reddish to light brown; abdominal segments yellowish. Carapace smooth, with 2 (anterior) eyes with indistinct lenses, about 2 diameters from the anterior margin; anterior margin with a small rounded knob, without a distinct epistome; a shallow transversal furrow present; chaetotaxy: 6/8-9/6/7-8: 27-29 (6/6/4/7: 23). Tergites undivided, chaetotaxy: 10-11/10/9-
<date id="FF8E17F22836273CFAE2973CFF26CA76" lastPageId="2" lastPageNumber="261" pageId="1" pageNumber="260" value="2011-09-10" valueMax="2011-11-10" valueMin="2011-09-10">10/9- 11/11</date>
/10/10-11/
<date id="FF8E17F22835273CFEC39E38FE62CA76" box="[322,405,258,281]" pageId="2" pageNumber="261" value="2010-08-10">10/10/8</date>
/6-8 (7/7/7/8/7/7/9/
<date id="FF8E17F22835273CFDED9E39FD4ACA74" box="[620,701,259,283]" pageId="2" pageNumber="261" value="2011-10-09">9/10/11</date>
/9/7), X-XI with numerous pores. Anal cone 2+2. Manducatory process with 5 (4) marginal setae; pedipalpal coxa with 6-8 (4) setae, two round lyrifissures, coxa
<date id="FF8E17F22835273CFED89EB9FE78CAF4" box="[345,399,387,411]" pageId="2" pageNumber="261">I 8-9</date>
(5),
<date id="FF8E17F22835273CFE499EB9FDF0CAF6" box="[456,519,386,410]" pageId="2" pageNumber="261">II 6-8</date>
(5-6),
<date id="FF8E17F22835273CFDD69EB9FD68CAF6" box="[599,671,386,410]" pageId="2" pageNumber="261">III 4-6</date>
(3),
<date id="FF8E17F22835273CFD599EB9FF3ECAD4" pageId="2" pageNumber="261" value="2010-04-08">IV 8-10</date>
(6); genital operculum with 31 setae (posterior ones longer than anterior ones)(
<specimenCount id="9D36FABB2835273CFE799EF9FDFBCAB5" box="[504,524,451,474]" pageId="2" pageNumber="261"></specimenCount>
: 10 medial marginal and discal setae), genital opening with 4/4 simple interior setae, median genital sac undivided, apically enlarged, short (reaching anterior half of sternite IV); lateral genital sacs short, caudally bent and apically inflated. Sternite III with anteromedian groove flanked by one small seta on each side, with 18 long setae (7 medial discal setae)+ 5/6 (4) suprastigmal setae, IV-XI: 11+
<date id="FF8E17F22835273CFF3F9DF9FF2EC9B5" box="[190,217,707,730]" pageId="2" pageNumber="261">2x</date>
<date id="FF8E17F22835273CFF589DF9FEDEC9B5" box="[217,297,707,730]" pageId="2" pageNumber="261" value="2015-05-17">5/17/15</date>
/14/
<date id="FF8E17F22835273CFED59DF9FE44C9B5" box="[340,435,707,730]" pageId="2" pageNumber="261" value="2013-12-14">13/14/12</date>
/4 (2 lateral tactile setae) (
<specimenCount id="9D36FABB2835273CFD599DFEFD1BC9B4" box="[728,748,708,731]" pageId="2" pageNumber="261"></specimenCount>
: 11+
<date id="FF8E17F22835273CFF3F9DD9FF2DC995" box="[190,218,739,762]" pageId="2" pageNumber="261">2x</date>
4/16/
<date id="FF8E17F22835273CFE959DD9FE84C995" box="[276,371,739,762]" pageId="2" pageNumber="261" value="2014-12-13">14/13/12</date>
/
<date id="FF8E17F22835273CFEFC9DD9FE38C995" box="[381,463,739,762]" pageId="2" pageNumber="261" value="2012-08-10">12/10/8</date>
), VI-X with 2 submedial, submarginal setae. Pleural membrane granular.
</paragraph>
<paragraph id="8B8F31322835273CFF169C19FDBFCFF4" blockId="2.[151,757,258,2078]" pageId="2" pageNumber="261">
<emphasis id="B944ED202835273CFF169C19FEF7C855" box="[151,256,803,826]" italics="true" pageId="2" pageNumber="261">Chelicera</emphasis>
(
<figureCitation id="130B2DB72835273CFE949C19FEADC853" box="[277,346,803,828]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="2" pageNumber="261">Fig. 1</figureCitation>
): Hand with 6-7 setae, fixed finger with 13-15 small, pointed teeth, movable finger with 12-14 pointed teeth (medial ones on a slightly thickened lamella), subgaleal seta short, not reaching finger tip; galea (
<figureCitation id="130B2DB72835273CFF619C99FED5C8D3" box="[224,290,931,956]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="2" pageNumber="261">Fig. 2</figureCitation>
a, b): one male with a transparent rounded spinneret, the other one with a rounded spinneret (left) and a short transparent stump (right), indicating the presence of a broken galea; female with a short galea with two tiny apical branchlets (
<figureCitation id="130B2DB72835273CFED99B1EFE6FCF54" box="[344,408,1060,1083]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="2" pageNumber="261">Fig. 2</figureCitation>
b); serrula exterior 36-40, serrula interior 30-34 blades (distal one deeply forked). Rallum (
<figureCitation id="130B2DB72835273CFF1E9B5EFF29CF13" box="[159,222,1124,1148]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="2" pageNumber="261">Fig. 3</figureCitation>
) with 8 (9) pinnate setae, distal one separated and with expanded base, proximal one short.
</paragraph>
<paragraph id="8B8F31322835273CFF169B9EFEC4CDB1" blockId="2.[151,757,258,2078]" pageId="2" pageNumber="261">
<emphasis id="B944ED202835273CFF169B9EFF00CFD4" box="[151,247,1188,1211]" italics="true" pageId="2" pageNumber="261">Pedipalp</emphasis>
(
<figureCitation id="130B2DB72835273CFE8B9B9EFE83CFD4" box="[266,372,1188,1213]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="2" pageNumber="261">Figs 4, 5</figureCitation>
): Smooth excepted for dorsodistal part of hand with fine granulation; trochanter with a small rounded ventral hump, 2.3 (2.5) times, femur distally enlarged, 4.4-4.6 (4.8) times, patella 3.1 (3.5) times longer than broad, club 1.6-1.7 (2.0) times longer than broad and 1.16-1.24 (1.34) times longer than pedicel, hand with pedicel 1.7-2.0 (1.9) times, chela with pedicel 3.7-4.0 (4.0) times, without pedicel 3.4-3.7 (3.7) times longer than broad, finger as long as or 1.14-1.21 (
<specimenCount id="9D36FABB2835273CFD4D9A9FFD03CED3" box="[716,756,1445,1468]" pageId="2" pageNumber="261">♀♂</specimenCount>
<typeStatus id="548B8F902835273CFF169AFFFF08CEB2" box="[151,255,1477,1501]" pageId="2" pageNumber="261" type="paratype">paratypes</typeStatus>
) times longer than hand with pedicel. Fixed finger with 104-109 (91) cusped teeth, row shorter than on movable finger, movable finger with 103-112 (85) teeth, about 20 distal ones with tiny cusps. Trichobothria (
<figureCitation id="130B2DB72835273CFF1E997FFF28CD33" box="[159,223,1605,1628]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="2" pageNumber="261">Fig. 5</figureCitation>
): 12 (8+4) (
<specimenCount id="9D36FABB2835273CFEDD997CFE87CD32" box="[348,368,1606,1629]" pageId="2" pageNumber="261"></specimenCount>
: 8+3 on left finger:
<emphasis id="B944ED202835273CFDBF997CFDA7CD32" box="[574,592,1606,1629]" italics="true" pageId="2" pageNumber="261">st</emphasis>
absent),
<emphasis id="B944ED202835273CFD2F997FFD03CD33" box="[686,756,1605,1628]" italics="true" pageId="2" pageNumber="261">eb-esb</emphasis>
on hand,
<emphasis id="B944ED202835273CFF7C995FFEABCD13" box="[253,348,1637,1660]" italics="true" pageId="2" pageNumber="261">ib-isb-ist</emphasis>
grouped at fixed finger base,
<emphasis id="B944ED202835273CFD22995CFD45CD12" box="[675,690,1638,1661]" italics="true" pageId="2" pageNumber="261">it</emphasis>
distal of
<emphasis id="B944ED202835273CFF3599BCFF30CDF2" box="[180,199,1670,1693]" italics="true" pageId="2" pageNumber="261">et</emphasis>
,
<emphasis id="B944ED202835273CFF5499BCFF04CDF2" box="[213,243,1670,1693]" italics="true" pageId="2" pageNumber="261">est</emphasis>
distinctly nearer to
<emphasis id="B944ED202835273CFE4699BCFE2DCDF2" box="[455,474,1670,1693]" italics="true" pageId="2" pageNumber="261">et</emphasis>
than to
<emphasis id="B944ED202835273CFDB099BCFDBDCDF2" box="[561,586,1670,1693]" italics="true" pageId="2" pageNumber="261">ist</emphasis>
;
<emphasis id="B944ED202835273CFDD999BFFD87CDF3" box="[600,624,1669,1692]" italics="true" pageId="2" pageNumber="261">sb</emphasis>
on movable finger halfway between
<emphasis id="B944ED202835273CFE1D999FFE5ECDD3" box="[412,425,1701,1724]" italics="true" pageId="2" pageNumber="261">b</emphasis>
and
<emphasis id="B944ED202835273CFE61999CFE05CDD2" box="[480,498,1702,1725]" italics="true" pageId="2" pageNumber="261">st</emphasis>
; very short venom duct in fixed finger.
</paragraph>
<paragraph id="8B8F31322835273CFF1699DCFDA3CC13" blockId="2.[151,757,258,2078]" pageId="2" pageNumber="261">
<emphasis id="B944ED202835273CFF1699DCFF24CD92" box="[151,211,1766,1789]" italics="true" pageId="2" pageNumber="261">Leg I</emphasis>
: Femur 4.1-4.4 (4.5) times, patella 3.1-3.4 (3.1) times, tibia 5.8-6.1 (6.65) times, basitarsus 2.9-3.0 (3.0) times, telotarsus 4.6-5.0 (4.8) times longer than deep, femur 1.5-1.6 (1.6) times longer than patella, telotarsus 1.5-1.7 (1.7) times longer than basitarsus.
</paragraph>
<paragraph id="8B8F31322835273CFF1698BDFAE6CA14" blockId="2.[151,757,258,2078]" lastBlockId="2.[831,1437,258,379]" pageId="2" pageNumber="261">
<emphasis id="B944ED202835273CFF1698BDFF14CCF1" box="[151,227,1927,1950]" italics="true" pageId="2" pageNumber="261">Leg IV</emphasis>
: Femur+patella 4.5-4.8 (5.1) times longer than deep, femur shorter than patella, tibia 7.7-7.9 (7.3) times, basitarsus 3.0-3.4 (3.2) times, telotarsus 5.1 (5.2) times longer than deep; tibia with one tactile seta in middle (TS=0.47-0.55), basitarsus with two tactile setae (basally, distally) (TS= 0.13, TS= 0.83-0.86), telotarsus with a tactile seta near middle (TS=0.46-0.47); subterminal seta forked and dentate, claws slender and smooth, arolia undivided and distinctly shorter than claws.
</paragraph>
<paragraph id="8B8F31322835273CFCBE9EAFFAFEC8D6" blockId="2.[831,1437,404,953]" pageId="2" pageNumber="261">
<emphasis id="B944ED202835273CFCBE9EAFFC2FCAA3" bold="true" pageId="2" pageNumber="261">
Measurements of
<typeStatus id="548B8F902835273CFB959EAFFB80CAC2" box="[1044,1143,405,429]" pageId="2" pageNumber="261" type="holotype">holotype</typeStatus>
<specimenCount id="9D36FABB2835273CFB039EAEFB61CAC4" box="[1154,1174,404,427]" pageId="2" pageNumber="261"></specimenCount>
(of
<typeStatus id="548B8F902835273CFB4D9EAFFAC4CAC2" box="[1228,1331,405,429]" pageId="2" pageNumber="261" type="paratype">paratype</typeStatus>
<specimenCount id="9D36FABB2835273CFABF9EAEFAA5CAC4" box="[1342,1362,404,427]" pageId="2" pageNumber="261"></specimenCount>
;
<specimenCount id="9D36FABB2835273CFAE69EAEFA8CCAC4" box="[1383,1403,404,427]" pageId="2" pageNumber="261"></specimenCount>
in parentheses):
</emphasis>
Body length 4.43 (3.50; 3.45); carapace 1.24/1.12 (1.28/1.12; 1.12/0.96). Pedipalp: trochanter 0.92/0.39 (-; 0.89/0.34), femur 1.70/0.38 (1.61/0.35; 1.56/0.33), patella 1.53/0.50 (1.39/0.45; 1.44/0.41), length of pedicel 0.69 (0.64; 0.62), hand with pedicel 1.42/0.70 (1.22/0.72; 1.26/0.65), length of pedicel 0.25 (0.21; 0.21), length of finger 1.41 (1.49; 1.43), length of chela with pedicel 2.83 (2.69; 2.57). Leg I: femur 0.96/0.22 (0.94/0.23; 0.87/0.19), patella 0.59/0.19 (0.64/0.19; 0.55/0.18), tibia 0.87/0.15 (0.86/0.14); 0.78/0.12), basitarsus 0.38/0.13 (0.37/0.12; 0.35/0.12), telotarsus 0.58/0.13 (0.62/0.1; 0.58/0.12). Leg IV: femur+patella 1.59/0.35 (1.60/0.33; 1.48/0.29), length of femur 0.71 (0.70; 0.67), tibia 1.62/0.21 (1.48/0.19; 1.33/0.18), basitarsus 0.52/0.17 (0.51/0.15; 0.45/0.14), telotarsus 0.73/0.14 (0.73/0.14; 0.72/0.14).
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92835273CFCBE9CEAFA9ACDF8" pageId="2" pageNumber="261" type="discussion">
<paragraph id="8B8F31322835273CFCBE9CEAFA9ACDF8" blockId="2.[831,1437,976,1688]" pageId="2" pageNumber="261">
<emphasis id="B944ED202835273CFCBE9CEAFC46C887" bold="true" box="[831,945,976,1000]" pageId="2" pageNumber="261">Remarks:</emphasis>
This new species is similar, in pedipalpal measurements and proportions, to
<taxonomicName id="4C304AB12835273CFB499CC8FAD9CF66" box="[1224,1326,1010,1033]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="martii" status="comb. nov.">
<emphasis id="B944ED202835273CFB499CC8FAD9CF66" box="[1224,1326,1010,1033]" italics="true" pageId="2" pageNumber="261">B. martii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277505B2835273CFABB9CC8FA6ACF68" box="[1338,1437,1008,1033]" pageId="2" pageNumber="261">n. comb.</taxonomicNameLabel>
from the Guo Quan Dong Cave (
<collectingRegion id="49F4FFD02835273CFB519B28FA62CF47" box="[1232,1429,1041,1066]" country="China" name="Yunnan" pageId="2" pageNumber="261">Yunnan Province</collectingRegion>
) and to
<taxonomicName id="4C304AB12835273CFC129B09FB1FCF24" authority="Chamberlin, 1931" authorityName="Chamberlin" authorityYear="1931" box="[915,1256,1074,1099]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="lampra">
<emphasis id="B944ED202835273CFC129B09FBF4CF26" box="[915,1027,1074,1098]" italics="true" pageId="2" pageNumber="261">B. lampra</emphasis>
(
<bibRefCitation id="EFA14CC32835273CFB979B08FB17CF24" author="Chamberlin J. C." box="[1046,1248,1074,1099]" journalOrPublisher="Stanford University Publications, Biological Sciences" pageId="2" pageNumber="261" pagination="1 - 284" part="7" refString="Chamberlin J. C. 1931. The arachnid order Chelonethida. Stanford University Publications, Biological Sciences 7 (1): 1 - 284." title="The arachnid order Chelonethida" type="journal article" year="1931">Chamberlin, 1931</bibRefCitation>
)
</taxonomicName>
from Gu Shan, Fukien Province. All three belong to a group of species in which the pedicel of the patella is about half the length of the patella. Compared to
<taxonomicName id="4C304AB12835273CFB499BAFFADBCFC3" box="[1224,1324,1173,1196]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="martii" status="comb. nov.">
<emphasis id="B944ED202835273CFB499BAFFADBCFC3" box="[1224,1324,1173,1196]" italics="true" pageId="2" pageNumber="261">B. martii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277505B2835273CFAB49BAFFA6ECFC4" box="[1333,1433,1172,1196]" pageId="2" pageNumber="261">n. comb.</taxonomicNameLabel>
, the new species has only two tiny eyes (four distinct ones in
<taxonomicName id="4C304AB12835273CFC129BEDFC05CF81" box="[915,1010,1239,1262]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="martii" status="comb. nov.">
<emphasis id="B944ED202835273CFC129BEDFC05CF81" box="[915,1010,1239,1262]" italics="true" pageId="2" pageNumber="261">B. martii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277505B2835273CFC7B9BEDFBACCF82" box="[1018,1115,1238,1262]" pageId="2" pageNumber="261">n. comb.</taxonomicNameLabel>
), smooth pedipalpal segments except for the chelal hand, and more numerous teeth on the pedipalpal fingers (fixed/movable finger: 109/122 vs 83/81 vs 81).
<taxonomicName id="4C304AB12835273CFBE09A03FA89CE20" box="[1121,1406,1336,1360]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="chuanensis" status="sp. nov.">
<emphasis id="B944ED202835273CFBE09A03FA89CE20" box="[1121,1406,1336,1360]" italics="true" pageId="2" pageNumber="261">Bisetocreagris chuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277505B2835273CFA099A03FCA9CE1E" pageId="2" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C304AB12835273CFC739A60FBA8CE1F" box="[1010,1119,1369,1393]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="lampra">
<emphasis id="B944ED202835273CFC739A60FBA8CE1F" box="[1010,1119,1369,1393]" italics="true" pageId="2" pageNumber="261">B. lampra</emphasis>
</taxonomicName>
by the presence of only two eyes on the carapace (vs four in
<taxonomicName id="4C304AB12835273CFB449A41FAC0CEFF" box="[1221,1335,1401,1426]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="lampra">
<emphasis id="B944ED202835273CFB449A41FAC0CEFF" box="[1221,1335,1401,1426]" italics="true" pageId="2" pageNumber="261">B. lampra</emphasis>
</taxonomicName>
), by the smooth pedipalpal femur and patella, by a more slender pedipalpal femur (4.4 times vs 3.56 times) and a more slender patella (3.1 times vs 2.3 times), and by a stouter chela with pedicel 4.0 times (vs 4.7 times) longer than broad. Furthermore, trichobothria
<emphasis id="B944ED202835273CFB4C9924FB00CD5A" box="[1229,1271,1566,1589]" italics="true" pageId="2" pageNumber="261">et/it</emphasis>
are distinctly distal to
<emphasis id="B944ED202835273CFC1E9905FC51CD39" box="[927,934,1599,1622]" italics="true" pageId="2" pageNumber="261">t</emphasis>
in
<taxonomicName id="4C304AB12835273CFC4A9905FB95CD3A" box="[971,1122,1598,1622]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="chuanensis" status="sp. nov.">
<emphasis id="B944ED202835273CFC4A9905FB95CD3A" box="[971,1122,1598,1622]" italics="true" pageId="2" pageNumber="261">B. chuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277505B2835273CFBE89905FB51CD39" box="[1129,1190,1599,1622]" pageId="2" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
, but on the same level of
<emphasis id="B944ED202835273CFCE1995AFC90CD18" box="[864,871,1632,1655]" italics="true" pageId="2" pageNumber="261">t</emphasis>
in
<taxonomicName id="4C304AB12835273CFC11995AFC09CD1A" box="[912,1022,1630,1655]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="lampra">
<emphasis id="B944ED202835273CFC11995AFC09CD1A" box="[912,1022,1630,1655]" italics="true" pageId="2" pageNumber="261">B. lampra</emphasis>
</taxonomicName>
,
<emphasis id="B944ED202835273CFB8E995AFBD6CD18" box="[1039,1057,1632,1655]" italics="true" pageId="2" pageNumber="261">st</emphasis>
is on the same level of
<emphasis id="B944ED202835273CFAB4995AFAA4CD18" box="[1333,1363,1632,1655]" italics="true" pageId="2" pageNumber="261">est</emphasis>
in the new species, but distinctly distal to
<emphasis id="B944ED202835273CFB3A99BBFB2ECDF7" box="[1211,1241,1665,1688]" italics="true" pageId="2" pageNumber="261">est</emphasis>
in
<taxonomicName id="4C304AB12835273CFB7A99BBFA91CDF9" box="[1275,1382,1663,1688]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="lampra">
<emphasis id="B944ED202835273CFB7A99BBFA91CDF9" box="[1275,1382,1663,1688]" italics="true" pageId="2" pageNumber="261">B. lampra</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>