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<document id="07D741E8EEE635370E39C366E8D075AD" ID-CLB-Dataset="22365" ID-DOI="10.5852/ejt.2018.396" ID-GBIF-Dataset="2b95a851-e1bf-4bdd-9cce-441ddc4135d2" ID-Zenodo-Dep="3787049" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1588696404769" checkinUser="valdenar" docAuthor="Camfield, Rebecca &amp; Hughes, Mark" docDate="2018" docId="0397A96F2F79FFC103A0FD50E251290D" docLanguage="en" docName="ejt-396_camfield_hughes.pdf.imf" docOrigin="European Journal of Taxonomy 396" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia beddomei Hook." docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="FFAED1172F6DFFD7025FFFB1E00A2E5C" masterDocTitle="A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India" masterLastPageNumber="116" masterPageNumber="1" pageNumber="21" updateTime="1737026855421" updateUser="diego">
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<mods:title id="98F70F77558E70FF2681F647BBEE34E4">A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India</mods:title>
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<mods:namePart id="66DFFCA822E58CE819CAC052DEA004E0">Camfield, Rebecca</mods:namePart>
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<mods:namePart id="1B01A7CCA3C26C32E23F0A8578B37F9C">Hughes, Mark</mods:namePart>
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<mods:title id="83816463B65C781D98936F4637D02AC6">European Journal of Taxonomy</mods:title>
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<mods:date id="050C75894C7CB4C7A7B1AA16170B29DD">2018</mods:date>
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<treatment id="0397A96F2F79FFC103A0FD50E251290D" ID-DOI="http://doi.org/10.5281/zenodo.3794339" ID-GBIF-Taxon="163963841" ID-Zenodo-Dep="3794339" LSID="urn:lsid:plazi:treatment:0397A96F2F79FFC103A0FD50E251290D" httpUri="http://treatment.plazi.org/id/0397A96F2F79FFC103A0FD50E251290D" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" scope_family="Begoniaceae" scope_order="Cucurbitales">
<subSubSection id="C3244BF22F79FFC303A0FD50E43C2CA0" box="[511,1078,737,764]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8B8118792F79FFC303A0FD50E43C2CA0" blockId="20.[511,1078,737,799]" box="[511,1078,737,764]" pageId="20" pageNumber="21">
<heading id="D0C9AF152F79FFC303A0FD50E43C2CA0" box="[511,1078,737,764]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
<taxonomicName id="4C3E63FA2F79FFC303A0FD50E33B2CA7" ID-CoL="7SDVG" authority="Hook." authorityName="Hook." box="[511,817,737,764]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="beddomei">
<emphasis id="B94AC46B2F79FFC303A0FD50E2EA2CA0" bold="true" box="[511,736,737,764]" italics="true" pageId="20" pageNumber="21">Begonia beddomei</emphasis>
Hook.
</taxonomicName>
f. [sect.
<taxonomicName id="4C3E63FA2F79FFC301D1FD50E4262CA7" box="[910,1068,737,763]" class="Magnoliopsida" family="Begoniaceae" genus="Platycentrum" higherTaxonomySource="CoL" kingdom="Plantae" order="Cucurbitales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AC46B2F79FFC301D1FD50E4262CA7" box="[910,1068,737,763]" italics="true" pageId="20" pageNumber="21">Platycentrum</emphasis>
</taxonomicName>
]
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F79FFC30088FCB4E3042D3C" pageId="20" pageNumber="21" type="description">
<paragraph id="8B8118792F79FFC30088FCB4E3572D43" blockId="20.[511,1078,737,799]" box="[727,861,773,799]" pageId="20" pageNumber="21">
<figureCitation id="130504FC2F79FFC30088FCB4E3572D43" box="[727,861,773,799]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="20.[190,233,1992,2018]" captionStartId-1="21.[189,232,1957,1983]" captionTargetBox-1="[301,1287,265,1921]" captionTargetId-0="figure@20.[416,1172,1210,1962]" captionTargetId-1="figure@21.[301,1287,265,1921]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="Fig. 12. Map showing the location of B. beddomei Hook. specimens." captionText-1="Fig. 13. Illustration of B. beddomei Hook.f. by M. Smith (Hooker 1884). 1. Stamen. 2. Styles. 3. Fruit cross-section. Image from the Biodiversity Heritage Library, digitized by the Peter H. Raven Library." figureDoi-0="http://doi.org/10.5281/zenodo.14616263" figureDoi-1="http://doi.org/10.5281/zenodo.3787069" httpUri-0="https://zenodo.org/record/14616263/files/figure.png" httpUri-1="https://zenodo.org/record/3787069/files/figure.png" pageId="20" pageNumber="21">Figs 12–13</figureCitation>
</paragraph>
<paragraph id="8B8118792F79FFC302E1FCF7E3042D3C" blockId="20.[190,1398,838,935]" box="[190,782,838,864]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="AB2E3F572F79FFC302E1FCF7E3042D3C" box="[190,782,838,864]" pageId="20" pageNumber="21">
<emphasis id="B94AC46B2F79FFC302E1FCF7E1A02D3C" box="[190,426,838,864]" italics="true" pageId="20" pageNumber="21">Botanical Magazine</emphasis>
<treatmentCitation id="0A9F3E682F79FFC303EFFCF7E3002D3C" box="[432,778,838,864]" pageId="20" pageNumber="21">
110: tab. 6767 (
<bibRefCitation id="EFAF65882F79FFC3003DFCF7E30B2D3C" author="Hooker J. D." box="[610,769,838,864]" pageId="20" pageNumber="21" pagination="6767" refId="ref54769" refString="Hooker J. D. 1884. Begonia beddomei. Curtis's Botanical Magazine 110: tab. 6767." type="journal article" year="1884">Hooker 1884</bibRefCitation>
)
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F79FFC3014CFCF6E3722A4B" pageId="20" pageNumber="21" type="description">
<paragraph id="8B8118792F79FFC3014CFCF6E1832DFB" blockId="20.[190,1398,838,935]" pageId="20" pageNumber="21">
–
<materialsCitation id="3B5612242F79FFC30178FCF6E18F2DFB" collectingDate="1883-12-04" collectionCode="K" collectorName="Gustav Mann" country="India" location="Assam" pageId="20" pageNumber="21" specimenCode="K000761469" specimenCount="1" stateProvince="Assam" typeStatus="holotype">
Type: Cultivated collection,
<date id="FF803EB92F79FFC3062FFCF7E4F42D3C" box="[1136,1278,838,865]" pageId="20" pageNumber="21" value="1883-12-04">
<collectingDate id="EFC4C7512F79FFC3062FFCF7E4F42D3C" box="[1136,1278,838,865]" pageId="20" pageNumber="21" value="1883-12-04">4 Dec. 1883</collectingDate>
</date>
,
<emphasis id="B94AC46B2F79FFC30756FCF7E57C2D3C" box="[1289,1398,838,864]" italics="true" pageId="20" pageNumber="21">Beddome</emphasis>
s.n. (
<typeStatus id="5485A6DB2F79FFC302A8FCD8E1312DDF" box="[247,315,873,899]" pageId="20" pageNumber="21">holo-</typeStatus>
:
<specimenCode id="DB98B0022F79FFC30319FCD8E1EA2DDF" box="[326,480,873,899]" pageId="20" pageNumber="21">K000761469</specimenCode>
). Cultivated from vegetative material collected in the wild from
<collectingRegion id="49FAD69B2F79FFC30684FCDBE5242DD8" box="[1243,1326,874,900]" country="India" name="Assam" pageId="20" pageNumber="21">Assam</collectingRegion>
,
<collectingCountry id="F32958E92F79FFC30765FCD8E57C2DDF" box="[1338,1398,873,899]" name="India" pageId="20" pageNumber="21">India</collectingCountry>
by
<collectorName id="26CB7DAF2F79FFC302BCFC3CE18F2DFB" box="[227,389,909,935]" pageId="20" pageNumber="21">Gustav Mann</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8118792F79FFC302E1FC7FE2302DB4" blockId="20.[190,570,974,1000]" box="[190,570,974,1000]" pageId="20" pageNumber="21">
<heading id="D0C9AF152F79FFC302E1FC7FE2302DB4" bold="true" box="[190,570,974,1000]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B94AC46B2F79FFC302E1FC7FE2302DB4" bold="true" box="[190,570,974,1000]" pageId="20" pageNumber="21">Citations in other publications</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F79FFC302E1FC4DE3722A4B" blockId="20.[190,888,1020,1047]" box="[190,888,1020,1047]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="AB2E3F572F79FFC302E1FC4DE37F2A4B" box="[190,885,1020,1047]" pageId="20" pageNumber="21">
<treatmentCitation id="0A9F3E682F79FFC302E1FC4DE1932A4B" box="[190,409,1020,1047]" pageId="20" pageNumber="21">
<bibRefCitation id="EFAF65882F79FFC302E1FC4DE1932A4B" author="Fischer C. E. C." box="[190,409,1020,1047]" pageId="20" pageNumber="21" pagination="75 - 162" refId="ref53759" refString="Fischer C. E. C. 1938. The Flora of the Lushai Hills. Records of the Botanical Survey of India 12: 75 - 162." type="journal article" year="1938">Fischer (1938: 98)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F79FFC303FAFC4CE2812A4B" box="[421,651,1020,1047]" pageId="20" pageNumber="21">
<bibRefCitation id="EFAF65882F79FFC303FAFC4CE2812A4B" author="Kumar K. D." box="[421,651,1020,1047]" pageId="20" pageNumber="21" pagination="641 - 655" refId="ref55672" refString="Kumar K. D. 2002. Begoniaceae. In: Singh N. P., Singh K. P. &amp; Singh D. K. (eds) Flora of Mizoram 1: 641 - 655. Botanical Survey of India, Calcutta." type="journal article" year="2002">Kumar (2002: 643)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F79FFC300C8FC4DE37F2A4B" box="[663,885,1020,1047]" pageId="20" pageNumber="21">
<bibRefCitation id="EFAF65882F79FFC300C8FC4DE37F2A4B" author="Uddin A." box="[663,885,1020,1047]" pageId="20" pageNumber="21" pagination="591 - 597" refId="ref57753" refString="Uddin A. 2007. Distribution and status of India Begonia species. Journal of Economic and Taxonomic Botany 31 (3): 591 - 597." type="journal article" year="2007">Uddin (2007: 593)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
.
</paragraph>
</subSubSection>
<caption id="DF4148F12F79FFC302E1F879E3EF29BF" ID-DOI="http://doi.org/10.5281/zenodo.14616263" ID-Zenodo-Dep="14616263" box="[190,997,1992,2019]" httpUri="https://zenodo.org/record/14616263/files/figure.png" pageId="20" pageNumber="21" startId="20.[190,233,1992,2018]" targetBox="[427,1160,1220,1950]" targetPageId="20">
<paragraph id="8B8118792F79FFC302E1F879E3EF29BF" blockId="20.[190,997,1992,2019]" box="[190,997,1992,2019]" pageId="20" pageNumber="21">
<emphasis id="B94AC46B2F79FFC302E1F879E11029BF" bold="true" box="[190,282,1992,2019]" pageId="20" pageNumber="21">Fig. 12.</emphasis>
Map showing the location of
<taxonomicName id="4C3E63FA2F79FFC30025F87BE35729BF" authority="Hook." authorityName="Hook." box="[634,861,1993,2019]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="beddomei">
<emphasis id="B94AC46B2F79FFC30025F87BE30629BF" box="[634,780,1993,2019]" italics="true" pageId="20" pageNumber="21">B. beddomei</emphasis>
Hook.
</taxonomicName>
specimens.
</paragraph>
</caption>
<caption id="DF4148F12F78FFC202E2F814E56A29BE" ID-DOI="http://doi.org/10.5281/zenodo.3787069" ID-Zenodo-Dep="3787069" httpUri="https://zenodo.org/record/3787069/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1957,1983]" targetBox="[301,1287,265,1921]" targetPageId="21">
<paragraph id="8B8118792F78FFC202E2F814E56A29BE" blockId="21.[189,1398,1957,2019]" pageId="21" pageNumber="22">
<emphasis id="B94AC46B2F78FFC202E2F814E11029E3" bold="true" box="[189,282,1957,1983]" pageId="21" pageNumber="22">Fig. 13.</emphasis>
Illustration of
<taxonomicName id="4C3E63FA2F78FFC20393F817E2B829E3" authority="Hook." authorityName="Hook." box="[460,690,1957,1983]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="beddomei">
<emphasis id="B94AC46B2F78FFC20393F817E25529E3" box="[460,607,1957,1983]" italics="true" pageId="21" pageNumber="22">B. beddomei</emphasis>
Hook.
</taxonomicName>
f. by M. Smith (
<bibRefCitation id="EFAF65882F78FFC20131F814E40529E3" author="Hooker J. D." box="[878,1039,1957,1983]" pageId="21" pageNumber="22" pagination="6767" refId="ref54769" refString="Hooker J. D. 1884. Begonia beddomei. Curtis's Botanical Magazine 110: tab. 6767." type="journal article" year="1884">Hooker 1884</bibRefCitation>
). 1. Stamen. 2. Styles. 3. Fruit cross-section. Image from the Biodiversity Heritage Library, digitized by the Peter H. Raven Library.
</paragraph>
</caption>
<subSubSection id="C3244BF22F7BFFC102E2FEBAE4642FE2" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8B8118792F7BFFC102E2FEBAE1F32F79" blockId="22.[189,505,267,293]" box="[189,505,267,293]" pageId="22" pageNumber="23">
<heading id="D0C9AF152F7BFFC102E2FEBAE1F32F79" bold="true" box="[189,505,267,293]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B94AC46B2F7BFFC102E2FEBAE1F32F79" bold="true" box="[189,505,267,293]" pageId="22" pageNumber="23">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F7BFFC102E2FE8BE4642FE2" blockId="22.[189,1399,313,446]" pageId="22" pageNumber="23">
<materialsCitation id="3B5612242F7BFFC102E2FE8BE2852F2B" ID-GBIF-Occurrence="2610421320" pageId="22" pageNumber="23">
<collectingCountry id="F32958E92F7BFFC102E2FE8BE11F2F08" box="[189,277,314,340]" name="India" pageId="22" pageNumber="23">INDIA</collectingCountry>
:
<emphasis id="B94AC46B2F7BFFC1037BFE88E17D2F0F" bold="true" box="[292,375,313,339]" pageId="22" pageNumber="23">
<collectingRegion id="49FAD69B2F7BFFC1037BFE88E17D2F0F" box="[292,375,313,339]" country="India" name="Assam" pageId="22" pageNumber="23">Assam</collectingRegion>
</emphasis>
: Cultivated collection,
<date id="FF803EB92F7BFFC100C9FE8BE3132F08" box="[662,793,314,340]" pageId="22" pageNumber="23" value="1883-08">Aug. 1883</date>
,
<emphasis id="B94AC46B2F7BFFC10176FE88E39C2F0F" box="[809,918,313,339]" italics="true" pageId="22" pageNumber="23">Beddome</emphasis>
s.n. (
<collectionCode id="ED2F80BC2F7BFFC101BFFE8BE3F32F08" box="[992,1017,314,340]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K</collectionCode>
), cultivated from wild material collected by Gustav Mann from Assam
</materialsCitation>
;
<materialsCitation id="3B5612242F7BFFC100C4FEEDE2012FC6" ID-GBIF-Occurrence="2610421320" pageId="22" pageNumber="23">
Jorhat, Hoolong ‘Melling’ Forest Reserve,
<date id="FF803EB92F7BFFC106C8FEECE5182F2B" box="[1175,1298,349,375]" pageId="22" pageNumber="23" value="1972-12">Dec. 1972</date>
,
<emphasis id="B94AC46B2F7BFFC10740FEECE0E32FC6" italics="true" pageId="22" pageNumber="23">Yandell 213</emphasis>
(
<collectionCode id="ED2F80BC2F7BFFC102A2FE31E11E2FC6" box="[253,276,384,410]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K</collectionCode>
) [aff.
<taxonomicName id="4C3E63FA2F7BFFC1033BFE30E1F02FC6" box="[356,506,384,410]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="beddomei">
<emphasis id="B94AC46B2F7BFFC1033BFE30E1F02FC6" box="[356,506,384,410]" italics="true" pageId="22" pageNumber="23">B. beddomei</emphasis>
</taxonomicName>
].
</materialsCitation>
<materialsCitation id="3B5612242F7BFFC10049FE31E4C12FC6" ID-GBIF-Occurrence="2610421320" box="[534,1227,384,410]" pageId="22" pageNumber="23">
<emphasis id="B94AC46B2F7BFFC10049FE31E2802FC6" bold="true" box="[534,650,384,410]" pageId="22" pageNumber="23">
<collectingRegion id="49FAD69B2F7BFFC10049FE31E2802FC6" box="[534,650,384,410]" country="India" name="Mizoram" pageId="22" pageNumber="23">Mizoram</collectingRegion>
</emphasis>
: Lushai Hills, Aijal,
<date id="FF803EB92F7BFFC101D1FE31E4032FC6" box="[910,1033,384,410]" pageId="22" pageNumber="23" value="1929-09">Sep. 1929</date>
,
<emphasis id="B94AC46B2F7BFFC10646FE30E49F2FC6" box="[1049,1173,384,410]" italics="true" pageId="22" pageNumber="23">Parry 252</emphasis>
(
<collectionCode id="ED2F80BC2F7BFFC106F6FE31E4C82FC6" box="[1193,1218,384,410]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5612242F7BFFC10684FE31E2792FE1" ID-GBIF-Occurrence="2610421320" pageId="22" pageNumber="23">
Lushai Hills, Ramlaitui,
<date id="FF803EB92F7BFFC10360FE15E1B62FE1" box="[319,444,419,446]" pageId="22" pageNumber="23" value="1927-11">Nov. 1927</date>
,
<emphasis id="B94AC46B2F7BFFC10397FE15E24A2FE1" box="[456,576,419,445]" italics="true" pageId="22" pageNumber="23">Parry 306</emphasis>
(
<collectionCode id="ED2F80BC2F7BFFC1000FFE12E2632FE1" box="[592,617,419,445]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5612242F7BFFC10020FE12E4642FE2" ID-GBIF-Occurrence="2610421320" box="[639,1134,419,446]" pageId="22" pageNumber="23">
ibid.,
<date id="FF803EB92F7BFFC1009CFE12E3332FE1" box="[707,825,419,445]" pageId="22" pageNumber="23" value="1927-10">Oct. 1927</date>
,
<emphasis id="B94AC46B2F7BFFC1011AFE15E3B72FE1" box="[837,957,419,445]" italics="true" pageId="22" pageNumber="23">Parry 306</emphasis>
(
<collectionCode id="ED2F80BC2F7BFFC10192FE12E3EF2FE1" box="[973,997,419,445]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="22" pageNumber="23" type="Herbarium">K</collectionCode>
,
<collectionCode id="ED2F80BC2F7BFFC101ADFE12E4262FE1" box="[1010,1068,419,445]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="22" pageNumber="23" type="Herbarium">CAL</collectionCode>
n.v.).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F7BFFC102E2FE55E1AF2A3D" pageId="22" pageNumber="23" type="description">
<paragraph id="8B8118792F7BFFC102E2FE55E1442FA2" blockId="22.[189,334,484,510]" box="[189,334,484,510]" pageId="22" pageNumber="23">
<heading id="D0C9AF152F7BFFC102E2FE55E1442FA2" bold="true" box="[189,334,484,510]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B94AC46B2F7BFFC102E2FE55E1442FA2" bold="true" box="[189,334,484,510]" pageId="22" pageNumber="23">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F7BFFC102E2FDA2E1AF2A3D" blockId="22.[189,1399,531,1121]" pageId="22" pageNumber="23">
Tuberous, monoecious herb,
<quantity id="4CC6B59C2F7BFFC10053FDA2E28F2C71" box="[524,645,531,558]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="22" pageNumber="23" unit="cm" value="22.5" valueMax="30.0" valueMin="15.0">15–30 cm</quantity>
high. Stipules: lanceolate, 9–11 ×
<quantity id="4CC6B59C2F7BFFC1064BFDA2E4772C71" box="[1044,1149,531,557]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="22" pageNumber="23" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, glabrous, deciduous. Leaves: petiole
<quantity id="4CC6B59C2F7BFFC10328FD87E1D72C0D" box="[375,477,566,593]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageId="22" pageNumber="23" unit="cm" value="11.5" valueMax="16.0" valueMin="7.0">7–16 cm</quantity>
long, puberulous; lamina broadly ovate, basifixed, base cordate with lobes not overlapping, 10–22 ×
<quantity id="4CC6B59C2F7BFFC10392FDE8E2362C28" box="[461,572,601,628]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="22" pageNumber="23" unit="cm" value="11.5" valueMax="15.0" valueMin="8.0">8–15 cm</quantity>
, asymmetric, upper surface green with white/silvery spots, glabrous, underside red-purple, sparsely pubescent, denser on veins, venation palmate, midrib
<quantity id="4CC6B59C2F7BFFC1069CFDCCE5252CCB" box="[1219,1327,636,663]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="22" pageNumber="23" unit="cm" value="11.5" valueMax="15.0" valueMin="8.0">8–15 cm</quantity>
long; margin broadly dentate with fine serration, with sparse hairs; apex acute. Inflorescence: cymose, terminal, few; peduncle glabrous, branching 2–3 times, primary
<quantity id="4CC6B59C2F7BFFC1011EFD72E3B12C81" box="[833,955,707,733]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="22" pageNumber="23" unit="cm" value="14.0" valueMax="16.0" valueMin="12.0">12–16 cm</quantity>
, secondary
<quantity id="4CC6B59C2F7BFFC1061CFD72E4A92C81" box="[1091,1187,707,733]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="22" pageNumber="23" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 cm</quantity>
, tertiary
<quantity id="4CC6B59C2F7BFFC10756FD72E5782C81" box="[1289,1394,707,733]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="22" pageNumber="23" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, with 1–
<specimenCount id="9D38D3F02F7BFFC10346FD57E1892D5C" box="[281,387,742,768]" pageId="22" pageNumber="23" type="female">2 female</specimenCount>
and 4–
<specimenCount id="9D38D3F02F7BFFC103BBFD57E23F2D5C" box="[484,565,742,768]" pageId="22" pageNumber="23" type="male">8 male</specimenCount>
flowers; bracts lanceolate, 16–18 ×
<quantity id="4CC6B59C2F7BFFC101B1FD57E4502D5D" box="[1006,1114,742,769]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="22" pageNumber="23" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
, glabrous. Male flower: pedicel
<quantity id="4CC6B59C2F7BFFC10346FCB8E1902D78" box="[281,410,777,804]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.5" metricValueMin="1.1" pageId="22" pageNumber="23" unit="mm" value="18.0" valueMax="25.0" valueMin="11.0">11–25 mm</quantity>
long, glabrous; tepals 4; outer tepals ovate to broad ovate, 15–21 ×
<quantity id="4CC6B59C2F7BFFC1069CFCB8E53C2D78" box="[1219,1334,777,804]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="22" pageNumber="23" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
, pink to pale pink, sparsely pilose on reverse, margin entire; inner tepals elliptic, 10–15 ×
<quantity id="4CC6B59C2F7BFFC106F2FC9CE51C2D1B" box="[1197,1302,813,839]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="22" pageNumber="23" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
, pink to pale pink, glabrous; androecium with 80–100 stamens, symmetric; filaments
<quantity id="4CC6B59C2F7BFFC10602FCE1E4CE2D36" box="[1117,1220,848,874]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="22" pageNumber="23" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, unequal, fused at base into a long column; anther oblong elliptic,
<quantity id="4CC6B59C2F7BFFC1010DFCC2E39C2DD1" box="[850,918,883,909]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="mm" value="1.0">1 mm</quantity>
long, dehiscing through slits longer than half the length of the anther, not hooded, connective extended. Female flower: pedicel
<quantity id="4CC6B59C2F7BFFC106EFFC27E53E2DED" box="[1200,1332,918,945]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="22" pageNumber="23" unit="mm" value="18.5" valueMax="25.0" valueMin="12.0">12–25 mm</quantity>
long, glabrous; bracteoles absent; tepals 8, oblong elliptic, pink; ovary 2-locular, placentae bifid, capsule obovoid, glabrous, with one long oblong wing and two short oblong wings; styles 3, convoluted with twisted ends, caduceus. Fruit: recurved; capsule ovate-triangular, 9–11 ×
<quantity id="4CC6B59C2F7BFFC10664FBB1E4A32A46" box="[1083,1193,1024,1050]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="22" pageNumber="23" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
, glabrous; wings extending along the pedicel slightly, unequal; longest rounded oblong, 20–23 ×
<quantity id="4CC6B59C2F7BFFC106D6FB92E5062A61" box="[1161,1292,1059,1085]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="22" pageNumber="23" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
; shortest oblong, 2 ×
<quantity id="4CC6B59C2F7BFFC10314FBF7E1AB2A3D" box="[331,417,1094,1121]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="23" unit="mm" value="12.0">12 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F7BFFC102E2FB36E4102B91" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8B8118792F7BFFC102E2FB36E21F2AFE" blockId="22.[189,533,1159,1186]" box="[189,533,1159,1186]" pageId="22" pageNumber="23">
<heading id="D0C9AF152F7BFFC102E2FB36E21F2AFE" bold="true" box="[189,533,1159,1186]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B94AC46B2F7BFFC102E2FB36E21F2AFE" bold="true" box="[189,533,1159,1186]" pageId="22" pageNumber="23">Distribution and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F7BFFC102E2FB07E1832AAF" blockId="22.[189,1398,1206,1267]" pageId="22" pageNumber="23">
Possibly endemic to the Lushai Hills of
<collectingRegion id="49FAD69B2F7BFFC100EBFB07E3292A8C" box="[692,803,1206,1232]" country="India" name="Mizoram" pageId="22" pageNumber="23">Mizoram</collectingRegion>
;
<quantity id="4CC6B59C2F7BFFC1016BFB07E3D82A8D" box="[820,978,1206,1233]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="22" pageNumber="23" unit="m" value="1050.0" valueMax="1200.0" valueMin="900.0">900–1200 m</quantity>
. Flowering: August to September; fruiting: October.
</paragraph>
<paragraph id="8B8118792F7BFFC102E2FAAAE1BF2B69" blockId="22.[189,437,1307,1333]" box="[189,437,1307,1333]" pageId="22" pageNumber="23">
<heading id="D0C9AF152F7BFFC102E2FAAAE1BF2B69" bold="true" box="[189,437,1307,1333]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B94AC46B2F7BFFC102E2FAAAE1BF2B69" bold="true" box="[189,437,1307,1333]" pageId="22" pageNumber="23">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F7BFFC102E2FAFBE4102B91" blockId="22.[189,1399,1353,1485]" pageId="22" pageNumber="23">
Data Deficient. The species is known only from very few collections and none of them are recent. It is possibly Vulnerable to Endangered as the Khasi Hills have areas of extensive deforestation and without an exact locality for the
<typeStatus id="5485A6DB2F7BFFC1038AFA21E2022BF6" box="[469,520,1424,1450]" pageId="22" pageNumber="23">type</typeStatus>
and lack of recent collections it is difficult to assess accurately. The species was recorded at Hoollongapar Gibbon Sanctuary in
<collectingRegion id="49FAD69B2F7BFFC10142FA02E3672B91" box="[797,877,1459,1485]" country="India" name="Assam" pageId="22" pageNumber="23">Assam</collectingRegion>
(
<emphasis id="B94AC46B2F7BFFC10121FA02E4032B91" box="[894,1033,1459,1485]" italics="true" pageId="22" pageNumber="23">Yandell 213</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F7BFFC102E2FA44E251290D" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8B8118792F7BFFC102E2FA44E1252853" blockId="22.[189,303,1525,1551]" box="[189,303,1525,1551]" pageId="22" pageNumber="23">
<heading id="D0C9AF152F7BFFC102E2FA44E1252853" bold="true" box="[189,303,1525,1551]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B94AC46B2F7BFFC102E2FA44E1252853" bold="true" box="[189,303,1525,1551]" pageId="22" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F7BFFC102E2F995E1D828FA" blockId="22.[189,1399,1571,1702]" pageId="22" pageNumber="23">
<taxonomicName id="4C3E63FA2F7BFFC102E2F995E1922861" box="[189,408,1571,1597]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="beddomei">
<emphasis id="B94AC46B2F7BFFC102E2F995E1922861" box="[189,408,1571,1597]" italics="true" pageId="22" pageNumber="23">Begonia beddomei</emphasis>
</taxonomicName>
is known from only a few collections; the specimens seen lacked female flowers and the description for these is based on the protologue. This species is quite distinct amongst other members of
<taxonomicName id="4C3E63FA2F7BFFC10283F9DBE13428DF" box="[220,318,1642,1667]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AC46B2F7BFFC10283F9DBE13428DF" box="[220,318,1642,1667]" italics="true" pageId="22" pageNumber="23">Begonia</emphasis>
</taxonomicName>
sect.
<taxonomicName id="4C3E63FA2F7BFFC103DEF9D8E21528DF" box="[385,543,1641,1667]" class="Magnoliopsida" family="Begoniaceae" genus="Platycentrum" higherTaxonomySource="CoL" kingdom="Plantae" order="Cucurbitales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AC46B2F7BFFC103DEF9D8E21528DF" box="[385,543,1641,1667]" italics="true" pageId="22" pageNumber="23">Platycentrum</emphasis>
</taxonomicName>
in having female flowers with 8 tepals and 3 styles, and a tuberous rather than rhizomatous habit.
</paragraph>
<paragraph id="8B8118792F7BFFC102E2F97FE251290D" blockId="22.[189,1399,1742,1874]" pageId="22" pageNumber="23">
Living material was collected by Gustav Mann in
<collectingRegion id="49FAD69B2F7BFFC10150F97FE35528B4" box="[783,863,1742,1768]" country="India" name="Assam" pageId="22" pageNumber="23">Assam</collectingRegion>
and sent to Colonel Beddome, who grew the plants for the voucher specimens used by Hooker to describe the species. Mann did the majority of his collecting from the Khasi/Jaintia Hills of what is today
<collectingRegion id="49FAD69B2F7BFFC10136F8A5E3E12972" box="[873,1003,1812,1838]" country="India" name="Meghalaya" pageId="22" pageNumber="23">Meghalaya</collectingRegion>
into the Cachar Hills of
<collectingRegion id="49FAD69B2F7BFFC10740F8A4E5782973" box="[1311,1394,1813,1839]" country="India" name="Assam" pageId="22" pageNumber="23">Assam</collectingRegion>
, making this the likely
<typeStatus id="5485A6DB2F7BFFC1039CF889E1FC290E" box="[451,502,1848,1874]" pageId="22" pageNumber="23">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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