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<mods:title id="F2513B12771D1DEE4BE00EA4BC196932">An illustrated key to and diagnoses of the species of Staphylinidae (Coleoptera) associated with decaying carcasses in Argentina</mods:title>
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<mods:namePart id="9D3E3B9C67AE021996BB0B14393A5C80">Aballay, Fernando H.</mods:namePart>
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<mods:namePart id="DA78C91CED8E600FA5D6C2582B356515">Maldonado, María Belén</mods:namePart>
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<subSubSection id="C33611EEFFABB442FF6C7804BE7D058A" pageId="12" pageNumber="113" type="nomenclature">
<paragraph id="8B934265FFABB442FF6C7804BD6505AB" blockId="12.[151,565,364,423]" box="[151,565,364,390]" pageId="12" pageNumber="113">
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<taxonomicName id="4C2C39E6FFABB442FF6C7804BD6505AB" ID-CoL="ZD3Z" authority="Linnaeus" authorityName="Linnaeus" box="[151,565,364,390]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="maxillosus">
<emphasis id="B9589E77FFABB442FF6C7804BD6505AB" bold="true" box="[151,565,364,390]" pageId="12" pageNumber="113">
<emphasis id="B9589E77FFABB442FF6C7804BEF605AB" bold="true" box="[151,422,364,390]" italics="true" pageId="12" pageNumber="113">Creophilus maxillosus</emphasis>
(Linnaeus)
</emphasis>
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</heading>
</paragraph>
<paragraph id="8B934265FFABB442FF6C78E6BE7D058A" blockId="12.[151,565,364,423]" box="[151,301,398,423]" pageId="12" pageNumber="113">
(
<figureCitation id="13175EE0FFABB442FF6478E6BFAC058A" box="[159,252,398,423]" captionStart="FIGURE 1 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="12" pageNumber="113">Figs. 10</figureCitation>
,
<figureCitation id="13175EE0FFABB442FEF378E7BE75058A" box="[264,293,399,423]" captionStart="FIGURE 33 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="12" pageNumber="113">41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33611EEFFABB442FF6C78BEBD560723" pageId="12" pageNumber="113" type="diagnosis">
<paragraph id="8B934265FFABB442FF6C78BEBD560723" blockId="12.[151,1437,470,1179]" pageId="12" pageNumber="113">
<emphasis id="B9589E77FFABB442FF6C78BEBE5A05C2" bold="true" box="[151,266,470,495]" pageId="12" pageNumber="113">Diagnosis</emphasis>
(only referred to general habitus, modified from
<bibRefCitation id="EFBD3F94FFABB442FC7A78BEBB4805C2" author="Clarke" box="[897,1048,470,495]" pageId="12" pageNumber="113" refString="Clarke, D. J. (2011) Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae). Zoological Journal of the Linnean Society, 163 (3), 23 - 812. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00725. x" type="journal article" year="2011">Clarke 2011</bibRefCitation>
).
<taxonomicName id="4C2C39E6FFABB442FBCF78BEBA6905C3" box="[1076,1337,470,494]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="maxillosus">
<emphasis id="B9589E77FFABB442FBCF78BEBA6905C3" box="[1076,1337,470,494]" italics="true" pageId="12" pageNumber="113">Creophilus maxillosus</emphasis>
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can be distinguished by the following characters: antennal segments 16 brownish-black, 711 greyish-black; apex of antennal segment 11 slightly to moderately emarginate medially; pronotum moderately to distinctly narrowed posteriorly, hind angles indistinct; integument, including elytra, uniformly black; vestiture on hind margins of head, anterior pronotal declivities, and pterothorax mostly black; elytra with white vestiture arranged into welldemarcated (but variable) central transverse fascia; vestiture on dorsal surface of abdomen white, arranged into definite pattern, concentrated on terga 5 and 6, with 710 usually partly whitish.
<taxonomicName id="4C2C39E6FFABB442FBD77BC6BA7706EB" box="[1068,1319,686,710]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="maxillosus">
<emphasis id="B9589E77FFABB442FBD77BC6BA7706EB" box="[1068,1319,686,710]" italics="true" pageId="12" pageNumber="113">Creophilus maxillosus</emphasis>
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is distinct from all other
<taxonomicName id="4C2C39E6FFABB442FEB17BBBBE9606C6" box="[330,454,723,747]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFABB442FEB17BBBBE9606C6" box="[330,454,723,747]" italics="true" pageId="12" pageNumber="113">Creophilus</emphasis>
</taxonomicName>
species due to the definite whitish-grey elytral fascia, and dorsal pattern of body vestiture. Length 13.0
<quantity id="4CD4EF80FFABB442FE677B9EBD520723" box="[412,514,758,783]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="12" pageNumber="113" unit="mm" value="18.5">18.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33611EEFFABB442FF3C7A72BB3600B6" pageId="12" pageNumber="113" type="distribution">
<paragraph id="8B934265FFABB442FF3C7A72BC28077A" blockId="12.[151,1437,470,1179]" pageId="12" pageNumber="113">
<emphasis id="B9589E77FFABB442FF3C7A72BE30071E" bold="true" box="[199,352,794,819]" pageId="12" pageNumber="113">Distribution.</emphasis>
Widespread throughout the northern hemisphere; apparently absent from austral regions (except as adventive), South
<collectingCountry id="F33B02F5FFABB442FE797A57BEBF077A" box="[386,495,830,855]" name="China" pageId="12" pageNumber="113">East Asia</collectingCountry>
, and the East Indies (
<bibRefCitation id="EFBD3F94FFABB442FD267A56BC3C077A" author="Clarke" box="[733,876,830,855]" pageId="12" pageNumber="113" refString="Clarke, D. J. (2011) Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae). Zoological Journal of the Linnean Society, 163 (3), 23 - 812. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00725. x" type="journal article" year="2011">Clarke 2011</bibRefCitation>
).
</paragraph>
<paragraph id="8B934265FFABB442FF3C7A0ABB3600B6" blockId="12.[151,1437,470,1179]" pageId="12" pageNumber="113">
<emphasis id="B9589E77FFABB442FF3C7A0ABE1A0756" bold="true" box="[199,330,866,891]" pageId="12" pageNumber="113">Bionomics.</emphasis>
<taxonomicName id="4C2C39E6FFABB442FEAF7A0BBD010756" box="[340,593,867,891]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="maxillosus">
<emphasis id="B9589E77FFABB442FEAF7A0BBD010756" box="[340,593,867,891]" italics="true" pageId="12" pageNumber="113">Creophilus maxillosus</emphasis>
</taxonomicName>
is probably the best-studied rove beetle species (
<bibRefCitation id="EFBD3F94FFABB442FB727A0ABA4A0756" author="Clarke" box="[1161,1306,866,891]" pageId="12" pageNumber="113" refString="Clarke, D. J. (2011) Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae). Zoological Journal of the Linnean Society, 163 (3), 23 - 812. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00725. x" type="journal article" year="2011">Clarke 2011</bibRefCitation>
). The most complete list of literature on this species is given by
<bibRefCitation id="EFBD3F94FFABB442FD057AEFBCFC07B2" author="Herman" box="[766,940,903,927]" pageId="12" pageNumber="113" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman (2001)</bibRefCitation>
and updated by
<bibRefCitation id="EFBD3F94FFABB442FB947AEEBA4007B2" author="Clarke" box="[1135,1296,902,927]" pageId="12" pageNumber="113" refString="Clarke, D. J. (2011) Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae). Zoological Journal of the Linnean Society, 163 (3), 23 - 812. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00725. x" type="journal article" year="2011">Clarke (2011)</bibRefCitation>
. It has been found on carrion of all kinds where adults feed on maggots, in various kinds of both natural and synanthropic habitats (
<bibRefCitation id="EFBD3F94FFABB442FF057AA7BE8E07CA" author="Newton" box="[254,478,974,999]" pageId="12" pageNumber="113" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, pp. 272 - 418." type="book chapter" year="2000">
Newton
<emphasis id="B9589E77FFABB442FE9B7AA7BEC907CB" box="[352,409,974,998]" italics="true" pageId="12" pageNumber="113">et al.</emphasis>
2000
</bibRefCitation>
). In the “area of Cuyo (
<collectingCountry id="F33B02F5FFABB442FD117AA6BC0F07CA" box="[746,863,974,999]" name="Argentina" pageId="12" pageNumber="113">Argentina</collectingCountry>
)” it was mostly found in natural habitats (Aballay, pers. com.). According to
<bibRefCitation id="EFBD3F94FFABB442FE387A9ABD300026" author="Clarke" box="[451,608,1010,1035]" pageId="12" pageNumber="113" refString="Clarke, D. J. (2011) Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae). Zoological Journal of the Linnean Society, 163 (3), 23 - 812. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00725. x" type="journal article" year="2011">Clarke (2011)</bibRefCitation>
who has summarized the extensive information on this species from the forensic literature, adults of
<taxonomicName id="4C2C39E6FFABB442FE227D7EBD850003" box="[473,725,1046,1070]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="maxillosus">
<emphasis id="B9589E77FFABB442FE227D7EBD850003" box="[473,725,1046,1070]" italics="true" pageId="12" pageNumber="113">Creophilus maxillosus</emphasis>
</taxonomicName>
arrive early at a carcass during the bloat and decay stages, remaining there up to 13 days post-mortem (see also
<bibRefCitation id="EFBD3F94FFABB442FD0B7D52BB56007E" author="Matuszewski" box="[752,1030,1082,1107]" pageId="12" pageNumber="113" refString="Matuszewski, S., Bajerlein, D., Konwerski, S. &amp; Szpila, K. (2010) Insect succession and carrion decomposition in selected forests of Central Europe. Part 2: Composition and residency patterns of carrion fauna. Forensic Science International, 195, 42 - 51. http: // dx. doi. org / 10.1016 / j. forsciint. 2009.11.007" type="journal article" year="2010">
Matuszewski
<emphasis id="B9589E77FFABB442FC707D53BC94007E" box="[907,964,1083,1107]" italics="true" pageId="12" pageNumber="113">et al.</emphasis>
2010
</bibRefCitation>
;
<bibRefCitation id="EFBD3F94FFABB442FBE87D52BBB9007E" author="Matuszewski" box="[1043,1257,1082,1107]" pageId="12" pageNumber="113" refString="Matuszewski, S. (2012) Estimating the pre-appearance interval from temperature in Creophilus maxillosus L. (Coleoptera: Staphylinidae). Journal of Forensic Sciences, 57, 136 - 145. http: // dx. doi. org / 10.1111 / j. 1556 - 4029.2011.01958. x" type="journal article" year="2012">Matuszewski 2012</bibRefCitation>
;
<bibRefCitation id="EFBD3F94FFABB442FB0D7D52BE48005A" author="Prado" pageId="12" pageNumber="113" refString="Prado e Castro, C., Garcia, M. D. Martins da Silva, P. Faria e Silva, I. &amp; Serrano, A. (2013) Coleoptera of forensic interest: A study of seasonal community composition and succession in Lisbon, Portugal. Forensic Science International, 232, 73 - 83. http: // dx. doi. org / 10.1016 / j. forsciint. 2013.06.014" type="journal article" year="2013">
Prado e Castro
<emphasis id="B9589E77FFABB442FF6C7D37BF82005B" box="[151,210,1118,1142]" italics="true" pageId="12" pageNumber="113">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EFBD3F94FFABB442FEDD7D36BD30005A" author="Dekeirsschieter" box="[294,608,1118,1143]" pageId="12" pageNumber="113" refString="Dekeirsschieter, J., Frederick, C., Verheggen, F. J., Drugmand, D. &amp; Haubruge, E. (2013) Diversity of forensic rove beetles (Coleoptera, Staphylinidae) associated with decaying Pig carcass in a forest biotope. Journal of Forensic Science, 58 (4), 1032 - 1040. http: // dx. doi. org / 10.1111 / 1556 - 4029.12095" type="journal article" year="2013">
Dekeirsschieter
<emphasis id="B9589E77FFABB442FE257D37BD4A005B" box="[478,538,1118,1142]" italics="true" pageId="12" pageNumber="113">et al.</emphasis>
2013
</bibRefCitation>
).
<bibRefCitation id="EFBD3F94FFABB442FD8E7D36BC0E005A" author="Matuszewski" box="[629,862,1118,1143]" pageId="12" pageNumber="113" refString="Matuszewski, S. (2012) Estimating the pre-appearance interval from temperature in Creophilus maxillosus L. (Coleoptera: Staphylinidae). Journal of Forensic Sciences, 57, 136 - 145. http: // dx. doi. org / 10.1111 / j. 1556 - 4029.2011.01958. x" type="journal article" year="2012">Matuszewski (2012)</bibRefCitation>
has recently proposed this species to estimate an interval preceding its appearance on a cadaver, called the preappearance interval (PAI).
</paragraph>
</subSubSection>
</treatment>
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