121 lines
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121 lines
12 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1179.104037" ID-Pensoft-Pub="1313-2970-1179-243" ID-Pensoft-UUID="FD30B3684976524BBEFEED3F9C8679D1" ID-ZooBank="1C2AC35B27664077BA9B3EB4E8E8452A" ModsDocID="1313-2970-1179-243" checkinTime="1694565413728" checkinUser="pensoft" docAuthor="Jimenez-Ferbans, Larry, Maestre-Guerra, Ana, Villalba-Fuentes, Evelin, Barros-Barrios, Mayelis M. & Munoz-Montero, Jeison" docDate="2023" docId="110085C584D45F619B327B1DA0591F04" docLanguage="en" docName="ZooKeys 1179: 243-297" docOrigin="ZooKeys 1179" docPubDate="2023-09-12" docSource="http://dx.doi.org/10.3897/zookeys.1179.104037" docTitle="Passalus (Pertinax) rugosus Gravely 1918" docType="treatment" docVersion="1" id="FD30B3684976524BBEFEED3F9C8679D1" lastPageNumber="243" masterDocId="FD30B3684976524BBEFEED3F9C8679D1" masterDocTitle="Passalidae (Coleoptera, Scarabaeoidea) from the Caribbean coast of Colombia: synopsis, key, and new species description" masterLastPageNumber="297" masterPageNumber="243" pageNumber="243" updateTime="1694565413728" updateUser="pensoft">
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<mods:title id="73D224707E9918B0AAF5EE2DA9CC87F9">Passalidae (Coleoptera, Scarabaeoidea) from the Caribbean coast of Colombia: synopsis, key, and new species description</mods:title>
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<mods:namePart id="DBC2647AFEB0A270044B61DAC4E036B0">Jimenez-Ferbans, Larry</mods:namePart>
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<mods:affiliation id="AB30B69FB00664E04D313DAD8A3B975F">Facultad de Ciencias Basicas. Universidad del Magdalena, carrera 32 # 22 - 08, Santa Marta, Zip code 470004, Colombia</mods:affiliation>
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<mods:nameIdentifier id="173E52CB23FB23E80B00B970182E9FD4" type="email">larryjimenezferbans@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B726CD335309259B1CF01E02185E2035">Maestre-Guerra, Ana</mods:namePart>
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<mods:affiliation id="B6D64DFA4FDF01A15DBAC5E364AD3611">Facultad de Ciencias Basicas. Universidad del Magdalena, carrera 32 # 22 - 08, Santa Marta, Zip code 470004, Colombia</mods:affiliation>
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<mods:namePart id="AFCFDA39E297BA5D111CDA7E051334E6">Villalba-Fuentes, Evelin</mods:namePart>
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<mods:affiliation id="5F2CBF7E56D16A46FDC793C06C4404F3">Facultad de Ciencias Basicas. Universidad del Magdalena, carrera 32 # 22 - 08, Santa Marta, Zip code 470004, Colombia</mods:affiliation>
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<mods:namePart id="EAA73B0AF79B0DED061C5062DB9BFAE5">Barros-Barrios, Mayelis M.</mods:namePart>
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<mods:affiliation id="B023A735CCD9B0AF5A6A0C588CF976F9">Facultad de Ciencias Exactas y Naturales. Universidad de Caldas, Calle 65 # 26 - 10, Manizales, Zip code 170004, Colombia</mods:affiliation>
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<mods:namePart id="2A04C8E42C816492F0146B106F555CB1">Munoz-Montero, Jeison</mods:namePart>
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<mods:affiliation id="AB68EC2ACAE6B6DEC5A14EA03656BE26">Facultad de Ciencias Basicas. Universidad del Magdalena, carrera 32 # 22 - 08, Santa Marta, Zip code 470004, Colombia</mods:affiliation>
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<mods:date id="2F699345E975629293E83F66E6397092">2023</mods:date>
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<mods:number id="8156C75F1E28B1AA51F6986BDD49D29D">2023-09-12</mods:number>
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<mods:number id="D1753CFFB1C70EF6400A7D31A448946D">1179</mods:number>
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<mods:identifier id="E609DB20209D2C54CA84E1AA94035809" type="DOI">http://dx.doi.org/10.3897/zookeys.1179.104037</mods:identifier>
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<treatment id="110085C584D45F619B327B1DA0591F04" LSID="urn:lsid:plazi:treatment:110085C584D45F619B327B1DA0591F04" httpUri="http://treatment.plazi.org/id/110085C584D45F619B327B1DA0591F04" lastPageNumber="243" pageId="0" pageNumber="243">
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<subSubSection id="482B23CBE5390950A070D64654619112" pageId="0" pageNumber="243" type="nomenclature">
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<paragraph id="5F7848A1D5B4EB82945632C03DD7D24F" pageId="0" pageNumber="243">
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11.
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<taxonomicName id="C069A7230FE5EC11AE3CE166E22E62A5" ID-CoL="9G4SW" ID-ENA="2652932" LSID="110085C5-84D4-5F61-9B32-7B1DA0591F04" authority="Gravely, 1918" authorityName="Gravely" authorityYear="1918" class="Insecta" family="Passalidae" genus="Passalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passalus (Pertinax) rugosus" order="Coleoptera" pageId="0" pageNumber="243" phylum="Arthropoda" rank="species" species="rugosus" subGenus="Pertinax">Passalus (Pertinax) rugosus Gravely, 1918</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="FCCA87B8E0D10288114469D9DD9D355C" pageId="0" pageNumber="243" type="description">
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<paragraph id="08D9A27D305023119C7F1683F6ED13BB" pageId="0" pageNumber="243">
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<figureCitation id="A93AFE57B4FD63E4084C018E9D51E9C9" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Passalus (Pertinax) rugosus Gravely, 1918 A head and pronotum in dorsal view B habitus dorsal C habitus ventral D habitus lateral. Scale bars: 1.0 mm (A); 2.0 mm (B, C, D)." figureDoi="10.3897/zookeys.1179.104037.figure14" httpUri="https://binary.pensoft.net/fig/904578" pageId="0" pageNumber="243">Fig. 14</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1D213BBE76FF1AECE0F06818899417E8" pageId="0" pageNumber="243" type="diagnosis">
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<paragraph id="C1A33E86470FC68FBC42C4767C462AF3" pageId="0" pageNumber="243">Diagnosis.</paragraph>
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<paragraph id="0A4BBAFE589FF9DA209133E80F1B39E2" pageId="0" pageNumber="243">20.1-23.2 mm total length. Body subcylindrical. Anterior border of the frons with strong middle indentation, suggesting secondary mediofrontal tubercles. Mediofrontal and laterofrontal fused, large. Central tubercle with apex not free. Lateroposterior tubercles large, conspicuous, and conical. Eyes large. Antennal club tri-lamellate, with lamellae long. Lacinia with apex bidentate. Mediobasal area of mentum protruding and glabrous. Marginal groove narrow, occupying 2/3 of the anterior margin of the pronotum. Prosternellum rhomboidal, acute. Mesosternum glabrous; mesosternal scars distinct and elongated. Metasternum glabrous anterolaterally and in lateral groove; disc smooth and delimited by punctations posteriorly to laterally. Humeri pubescent and epipleura pubescent in basal third. Anterior ventral border of the profemur with conspicuous groove. Meso- and metatibiae with small spines or unarmed.</paragraph>
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<caption id="A9C7FC16AD2E58A6E691AC64B04C1053" doi="10.3897/zookeys.1179.104037.figure14" httpUri="https://binary.pensoft.net/fig/904578" pageId="0" pageNumber="243" start="Figure 14" startId="F14">
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<paragraph id="7906796EEFD9DCCD2CDB98DB0162630D" pageId="0" pageNumber="243">
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<emphasis id="F7F19727712BB8CB0A579EB033754BF3" bold="true" pageId="0" pageNumber="243">Figure 14.</emphasis>
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<taxonomicName id="C6CA6B2D8BCCF99673AEFD0B6C996289" authorityName="Gravely" authorityYear="1918" class="Insecta" family="Passalidae" genus="Passalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passalus (Pertinax) rugosus" order="Coleoptera" pageId="0" pageNumber="243" phylum="Arthropoda" rank="species" species="rugosus" subGenus="Pertinax">Passalus (Pertinax) rugosus</taxonomicName>
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Gravely, 1918
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<emphasis id="CE9A6E2244496DFC75E58FCB4EEB5CD9" bold="true" pageId="0" pageNumber="243">A</emphasis>
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head and pronotum in dorsal view
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<emphasis id="446FFCA084225DCF5C8E6F4FC80F770B" bold="true" pageId="0" pageNumber="243">B</emphasis>
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habitus dorsal
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<emphasis id="6759D3ADE95A6F4190D4841201DEDDFF" bold="true" pageId="0" pageNumber="243">C</emphasis>
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habitus ventral
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<emphasis id="7249511F3A51DB496444543B9CD8B85D" bold="true" pageId="0" pageNumber="243">D</emphasis>
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habitus lateral. Scale bars: 1.0 mm (
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<emphasis id="AC6B575F46858743935B0E516E6D886E" bold="true" pageId="0" pageNumber="243">A</emphasis>
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); 2.0 mm (
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<emphasis id="C33A443EB30CF57733B4EF2A11B869C2" bold="true" pageId="0" pageNumber="243">B, C, D</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="2349272110D83E722B0AC37F89FE3227" pageId="0" pageNumber="243" type="comments">
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<paragraph id="C8BDBBB945F54F4AEEA7DC883794212D" pageId="0" pageNumber="243">Comments.</paragraph>
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<paragraph id="57D654CF9C015923D6399947FD3C3DAF" pageId="0" pageNumber="243">
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<bibRefCitation id="4CF710DFF792D67D0030786179213D04" DOI="https://doi.org/10.1007/s13744-017-0584-1" author="Jimenez-Ferbans, L" journalOrPublisher="Neotropical Entomology" pageId="0" pageNumber="243" pagination="642 - 667" refId="B23" refString="Jimenez-Ferbans, L, Reyes-Castillo, P, Schuster, JC, 2018b. Passalidae (Coleoptera: Scarabaeoidea) of the Biogeographical Province of Choco and the West Andean Region of Colombia, with the Description of Two New Species. Neotropical Entomology 47 (5): 642 - 667, DOI: https://doi.org/10.1007/s13744-017-0584-1" title="Passalidae (Coleoptera: Scarabaeoidea) of the Biogeographical Province of Choco and the West Andean Region of Colombia, with the Description of Two New Species." url="https://doi.org/10.1007/s13744-017-0584-1" volume="47" year="2018 b">
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<normalizedToken id="729D0B70E42E5B8436BF6C46434AF0DA" originalValue="Jiménez-Ferbans">Jimenez-Ferbans</normalizedToken>
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et al. (2018b)
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</bibRefCitation>
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considered this species as endemic to the western Andes of Colombia. However, the present records extend its distribution to the eastern Andes.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |