treatments-xml/data/8C/5D/87/8C5D87D5BA57FFB7FD95950EFDE64D84.xml

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<mods:title id="1B113062E4E6381B628E3946CF3419F7">Six new species of Anthuridae (Peracarida: Isopoda) from the southern Mexican Pacific</mods:title>
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<mods:namePart id="F5B20B3199D41C3F515357D3775DDBBF">Jarquín-Martínez, Ubaldo</mods:namePart>
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<mods:affiliation id="AABD1DD85CF4C1959BE62BAE28644BB2">Universidad del Mar, campus Puerto Ángel, Laboratorio de Sistemática de Invertebrados Marinos (LABSIM), Ciudad Universitaria, Puerto Ángel, Oaxaca, 70902, México.</mods:affiliation>
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<mods:namePart id="A1961557AAC17488AD5A41DC8FA4D16C">García-Madrigal, María del Socorro</mods:namePart>
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<mods:affiliation id="AD7833A935ECBD2708D22EB841BD7189">Universidad del Mar, campus Puerto Ángel, Laboratorio de Sistemática de Invertebrados Marinos (LABSIM), Ciudad Universitaria, Puerto Ángel, Oaxaca, 70902, México.</mods:affiliation>
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<treatment id="8C5D87D5BA57FFB7FD95950EFDE64D84" ID-DOI="http://doi.org/10.5281/zenodo.5123270" ID-GBIF-Taxon="183727881" ID-Zenodo-Dep="5123270" LSID="urn:lsid:plazi:treatment:8C5D87D5BA57FFB7FD95950EFDE64D84" httpUri="http://treatment.plazi.org/id/8C5D87D5BA57FFB7FD95950EFDE64D84" lastPageId="30" lastPageNumber="91" pageId="24" pageNumber="85">
<subSubSection id="4CEE6548BA57FFB1FD95950EFC6F4E40" box="[590,997,330,357]" pageId="24" pageNumber="85" type="nomenclature">
<paragraph id="044B36C3BA57FFB1FD95950EFC6F4E40" blockId="24.[372,1215,330,427]" box="[590,997,330,357]" pageId="24" pageNumber="85">
<heading id="5F0381AFBA57FFB1FD95950EFC6F4E40" box="[590,997,330,357]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="85" reason="2">
<taxonomicName id="C3F44D40BA57FFB1FD95950EFC094E41" ID-CoL="8NCDC" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[590,899,330,357]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="85" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA57FFB1FD95950EFC094E41" bold="true" box="[590,899,330,357]" italics="true" pageId="24" pageNumber="85">Skuphonura oaxaquensis</emphasis>
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<taxonomicNameLabel id="2DB357AABA57FFB1FC50950FFC6F4E40" box="[907,997,331,357]" pageId="24" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act: 
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<paragraph id="044B36C3BA57FFB1FD5C95D5FC274E8E" blockId="24.[372,1215,330,427]" box="[647,941,401,427]" pageId="24" pageNumber="85">
<figureCitation id="9CCF2A46BA57FFB1FD5C95D5FC864E8E" box="[647,780,401,427]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="25.[189,232,1882,1908]" captionStartId-1="26.[189,232,1892,1918]" captionStartId-2="28.[189,232,1915,1941]" captionTargetBox-0="[211,1358,286,1821]" captionTargetBox-1="[205,1372,287,1839]" captionTargetBox-2="[236,1351,290,1848]" captionTargetId-0="figure-11@25.[193,1394,265,1841]" captionTargetId-1="figure-16@26.[189,1398,265,1853]" captionTargetId-2="figure-16@28.[189,1398,275,1863]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="28" captionText-0="Fig. 14. Skuphonura oaxaquensis sp. nov. A–B, D–H. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Habitus and detail of the pleotelson. B. Cephalon. C. Paratype, ♂, 7.3 mm (UMAR-PERA 549). Base of mouthparts. D. Antennule. E. Antenna and detail of flagellum. F. Maxilliped. G. Maxilla. H. Mandible and detail of the lacinia." captionText-1="Fig. 15. Skuphonura oaxaquensis sp. nov., holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Pereopod 1. B. Pereopod 2 and detail of the carpus (b) and sensory spine. C. Pereopod 3. D. Pereopod 4. E. Pereopod 5. F. Pereopod 6. G. Pereopod 7 and detail of the sensory spines.." captionText-2="Fig. 16. Skuphonura oaxaquensis sp. nov.A–D. Holotype, ♀, 7.9 mm, Oaxaca, PuertoÁngel, Estacahuite Beach (UMAR-PERA 548). A. Pleopod 1. B. Pleopod 2. C. Uropodal endopod. D. Uropodal exopod. E–G. Paratype, ♂, 7.3 mm (UMAR-PERA 549). E. Antennule. F. Pereopod 1. G. Pleopod 2." figureDoi-0="http://doi.org/10.5281/zenodo.5121796" figureDoi-1="http://doi.org/10.5281/zenodo.5121798" figureDoi-2="http://doi.org/10.5281/zenodo.5121800" httpUri-0="https://zenodo.org/record/5121796/files/figure.png" httpUri-1="https://zenodo.org/record/5121798/files/figure.png" httpUri-2="https://zenodo.org/record/5121800/files/figure.png" pageId="24" pageNumber="85">
Figs 14– 
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, 
<figureCitation id="9CCF2A46BA57FFB1FCC295D5FCC14E8E" box="[793,843,401,427]" captionStart="Fig" captionStartId="36.[189,232,1850,1876]" captionTargetBox="[205,1382,265,1809]" captionTargetId="figure-16@36.[205,1382,265,1809]" captionTargetPageId="36" captionText="Fig. 20. A. Amakusanthura guerrerensis sp. nov., holotype, ♀ (UMAR-PERA 522). B. Cortezura caeca sp. nov., holotype, ♀ (UMAR-PERA 534). C. Mesanthura antenniformis sp. nov., paratype, ♀ (ovigerous) (UMAR-PERA 540). D. M. estacahuitensis sp. nov., holotype, ♀ (UMAR-PERA 547). E. Skuphonura oaxaquensis sp. nov., holotype, ♀ (UMAR-PERA 548). F. Tinggianthura mexicana sp. nov., holotype, ♀ (UMAR-PERA 561). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5121808" httpUri="https://zenodo.org/record/5121808/files/figure.png" pageId="24" pageNumber="85">20E</figureCitation>
, 
<tableCitation id="49760378BA57FFB1FC8D95D5FC274E8E" box="[854,941,401,427]" captionStart="Table 2" captionStartId="29.[189,252,265,289]" captionTargetBox="[200,1366,348,1310]" captionTargetPageId="29" captionText="Table 2. Comparison between Skuphonura oaxaquensis sp. nov. and the closest species of genera." httpUri="http://table.plazi.org/id/508B664BBA52FFB4FF66954DFB594E04" pageId="24" pageNumber="85" tableUuid="508B664BBA52FFB4FF66954DFB594E04">Table 2</tableCitation>
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<emphasis id="3680EAD1BA57FFB1FF669596FECE4EC9" bold="true" box="[189,324,466,492]" pageId="24" pageNumber="85">Etymology</emphasis>
</heading>
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This species is named after 
<collectingRegion id="C630F821BA57FFB1FDD99645FDD74D3E" box="[514,605,513,539]" country="Mexico" name="Oaxaca" pageId="24" pageNumber="85">Oaxaca</collectingRegion>
, the Mexican State where the specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="4CEE6548BA57FFB1FF669606FC7E4C7F" pageId="24" pageNumber="85" type="materials_examined">
<paragraph id="044B36C3BA57FFB1FF669606FE204D79" blockId="24.[189,426,578,604]" box="[189,426,578,604]" pageId="24" pageNumber="85">
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<emphasis id="3680EAD1BA57FFB1FF669606FE204D79" bold="true" box="[189,426,578,604]" pageId="24" pageNumber="85">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="044B36C3BA57FFB1FF319635FB554DF7" blockId="24.[189,1399,625,722]" pageId="24" pageNumber="85">
<materialsCitation id="B49C3C9EBA57FFB1FF319635FB554DF7" ID-GBIF-Occurrence="3326572337" collectedFrom="coral rock" collectingDate="2005-09-10" collectionCode="R" collectorName="J. R. Bastida-Zavala et al." country="Mexico" elevation="3" latitude="15.666667" location="Estacahuite Beach" longLatPrecision="1284" longitude="-96.48333" municipality="Puerto Angel" pageId="24" pageNumber="85" specimenCode="UMAR-PERA 548" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca" typeStatus="holotype">
<emphasis id="3680EAD1BA57FFB1FF319635FED64DAE" bold="true" box="[234,348,625,651]" pageId="24" pageNumber="85">
<typeStatus id="DB4F8861BA57FFB1FF319635FED64DAE" box="[234,348,625,651]" pageId="24" pageNumber="85" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="7CE37653BA57FFB1FF6696D0FEBB4D8B" box="[189,305,660,686]" name="Mexico" pageId="24" pageNumber="85">MEXICO</collectingCountry>
• 
<specimenCount id="12F2FD4ABA57FFB1FE9696D1FEE94D8A" box="[333,355,661,687]" pageId="24" pageNumber="85" type="female">♀</specimenCount>
(body length 
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); 
<collectingRegion id="C630F821BA57FFB1FD5996D0FD574D8B" box="[642,733,660,686]" country="Mexico" name="Oaxaca" pageId="24" pageNumber="85">Oaxaca</collectingRegion>
, 
<collectingMunicipality id="E42FACB9BA57FFB1FD3096D1FC024D94" box="[747,904,661,689]" pageId="24" pageNumber="85">Puerto Ángel</collectingMunicipality>
, 
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; 
<geoCoordinate id="61C05004BA57FFB1FB5896D6FB7B4D8A" box="[1155,1265,658,687]" degrees="15" direction="north" minutes="40" orientation="latitude" pageId="24" pageNumber="85" precision="925" value="15.666667">15°40′ N</geoCoordinate>
, 
<geoCoordinate id="61C05004BA57FFB1FB2596D6FAF84D8A" box="[1278,1394,658,687]" degrees="96" direction="west" minutes="29" orientation="longitude" pageId="24" pageNumber="85" precision="925" value="-96.48333">96°29′ W</geoCoordinate>
; depth 
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</quantity>
; 
<date id="704A1003BA57FFB1FE8496F3FE764DF7" box="[351,508,695,722]" pageId="24" pageNumber="85" value="2005-09-10">
<collectingDate id="600EE9EBBA57FFB1FE8496F3FE764DF7" box="[351,508,695,722]" pageId="24" pageNumber="85" value="2005-09-10">10 Sep. 2005</collectingDate>
</date>
; 
<collectorName id="A9015315BA57FFB1FDD296FCFCBB4DF7" box="[521,817,695,722]" pageId="24" pageNumber="85">
J.R. Bastida-Zavala 
<emphasis id="3680EAD1BA57FFB1FD2D96FCFCA04DF4" box="[758,810,695,721]" italics="true" pageId="24" pageNumber="85">et al</emphasis>
.
</collectorName>
leg.; 
<collectedFrom id="B7E8753EBA57FFB1FCA896F3FC674DF4" box="[883,1005,695,721]" pageId="24" pageNumber="85">coral rock</collectedFrom>
; 
<specimenCode id="54529EB8BA57FFB1FC2296F3FB514DF7" box="[1017,1243,695,722]" pageId="24" pageNumber="85">UMAR-PERA 548</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="044B36C3BA57FFB1FF3196BDFC7E4C7F" blockId="24.[189,1398,761,858]" pageId="24" pageNumber="85">
<materialsCitation id="B49C3C9EBA57FFB1FF3196BDFAEB4C12" ID-GBIF-Occurrence="3326572314" collectedFrom="coral rock" collectingDate="2005-09-10" collectionCode="R" collectorName="J. R. Bastida-Zavala et al." country="Mexico" elevation="3" latitude="15.666667" location="MEXICO" longLatPrecision="1284" longitude="-96.48333" municipality="Puerto Angel" pageId="24" pageNumber="85" specimenCode="UMAR-PERA 549" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="paratype">
<emphasis id="3680EAD1BA57FFB1FF3196BDFEED4C36" bold="true" box="[234,359,761,787]" pageId="24" pageNumber="85">
<typeStatus id="DB4F8861BA57FFB1FF3196BDFEED4C36" box="[234,359,761,787]" pageId="24" pageNumber="85" type="paratype">Paratypes</typeStatus>
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<collectingCountry id="7CE37653BA57FFB1FF669758FEBB4C13" box="[189,305,796,822]" name="Mexico" pageId="24" pageNumber="85">MEXICO</collectingCountry>
• 
<specimenCount id="12F2FD4ABA57FFB1FE899758FE084C13" box="[338,386,796,822]" pageId="24" pageNumber="85" type="male">1 ♂</specimenCount>
(body length 
<quantity id="C30C9B26BA57FFB1FDE89758FD1C4C13" box="[563,662,796,822]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="24" pageNumber="85" unit="mm" value="7.3">7.3 mm</quantity>
); same collection data as for holotype; 
<specimenCode id="54529EB8BA57FFB1FBA59758FAEB4C12" box="[1150,1377,796,823]" pageId="24" pageNumber="85">UMAR-PERA 549</specimenCode>
</materialsCitation>
• 
<materialsCitation id="B49C3C9EBA57FFB1FF66977BFC7A4C7F" ID-GBIF-Occurrence="3326572302" box="[189,1008,831,858]" collectedFrom="coral rock" collectingDate="2005-09-10" collectionCode="R" collectorName="J. R. Bastida-Zavala et al." country="Mexico" elevation="3" latitude="15.666667" location="Estacahuite Beach" longLatPrecision="1284" longitude="-96.48333" municipality="Puerto Angel" pageId="24" pageNumber="85" specimenCode="UMAR-PERA 550" specimenCount="13" specimenCount-female="12" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="paratype">
<specimenCount id="12F2FD4ABA57FFB1FF66977BFE854C7F" box="[189,271,831,858]" pageId="24" pageNumber="85" type="female">12 ♀♀</specimenCount>
, 
<specimenCount id="12F2FD4ABA57FFB1FEC7977BFEC14C7F" box="[284,331,831,858]" pageId="24" pageNumber="85" type="male">1 ♂</specimenCount>
; same collection data as for holotype; 
<specimenCode id="54529EB8BA57FFB1FCD5977BFC7A4C7F" box="[782,1008,831,858]" pageId="24" pageNumber="85">UMAR-PERA 550</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4CEE6548BA57FFB2FF6697C4FAAF49D0" lastPageId="27" lastPageNumber="88" pageId="24" pageNumber="85" type="description">
<paragraph id="044B36C3BA57FFB1FF6697C4FEC44CBF" blockId="24.[189,334,896,922]" box="[189,334,896,922]" pageId="24" pageNumber="85">
<heading id="5F0381AFBA57FFB1FF6697C4FEC44CBF" bold="true" box="[189,334,896,922]" fontSize="11" level="3" pageId="24" pageNumber="85" reason="3">
<emphasis id="3680EAD1BA57FFB1FF6697C4FEC44CBF" bold="true" box="[189,334,896,922]" pageId="24" pageNumber="85">Description</emphasis>
</heading>
</paragraph>
<paragraph id="044B36C3BA57FFB1FF3197EBFDF04CEF" blockId="24.[188,1399,943,1181]" box="[234,634,943,970]" pageId="24" pageNumber="85">
<emphasis id="3680EAD1BA57FFB1FF3197EBFED64CEC" bold="true" box="[234,348,943,969]" pageId="24" pageNumber="85">
<typeStatus id="DB4F8861BA57FFB1FF3197EBFED64CEC" box="[234,348,943,969]" pageId="24" pageNumber="85" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="12F2FD4ABA57FFB1FEAB97EBFE0E4CEC" box="[368,388,943,969]" pageId="24" pageNumber="85" type="female">♀</specimenCount>
, UMAR-PERA 548)
</paragraph>
<paragraph id="044B36C3BA57FFB1FF669797FBDC4BB8" blockId="24.[188,1399,943,1181]" pageId="24" pageNumber="85">
BODY (
<figureCitation id="9CCF2A46BA57FFB1FEC99796FE224CC9" box="[274,424,978,1004]" captionStart="Fig" captionStartId="25.[189,232,1882,1908]" captionTargetBox="[211,1358,286,1821]" captionTargetId="figure-11@25.[193,1394,265,1841]" captionTargetPageId="25" captionText="Fig. 14. Skuphonura oaxaquensis sp. nov. A–B, D–H. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Habitus and detail of the pleotelson. B. Cephalon. C. Paratype, ♂, 7.3 mm (UMAR-PERA 549). Base of mouthparts. D. Antennule. E. Antenna and detail of flagellum. F. Maxilliped. G. Maxilla. H. Mandible and detail of the lacinia." figureDoi="http://doi.org/10.5281/zenodo.5121796" httpUri="https://zenodo.org/record/5121796/files/figure.png" pageId="24" pageNumber="85">Figs 14A–B</figureCitation>
, 
<figureCitation id="9CCF2A46BA57FFB1FE639796FE664CC9" box="[440,492,978,1004]" captionStart="Fig" captionStartId="36.[189,232,1850,1876]" captionTargetBox="[205,1382,265,1809]" captionTargetId="figure-16@36.[205,1382,265,1809]" captionTargetPageId="36" captionText="Fig. 20. A. Amakusanthura guerrerensis sp. nov., holotype, ♀ (UMAR-PERA 522). B. Cortezura caeca sp. nov., holotype, ♀ (UMAR-PERA 534). C. Mesanthura antenniformis sp. nov., paratype, ♀ (ovigerous) (UMAR-PERA 540). D. M. estacahuitensis sp. nov., holotype, ♀ (UMAR-PERA 547). E. Skuphonura oaxaquensis sp. nov., holotype, ♀ (UMAR-PERA 548). F. Tinggianthura mexicana sp. nov., holotype, ♀ (UMAR-PERA 561). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5121808" httpUri="https://zenodo.org/record/5121808/files/figure.png" pageId="24" pageNumber="85">20E</figureCitation>
). Length 
<quantity id="C30C9B26BA57FFB1FDB99796FD484CC8" box="[610,706,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="24" pageNumber="85" unit="mm" value="7.9">7.9 mm</quantity>
, 8.4 times as long as wide with pigment in the cephalon and margins of each somite; body proportion: C&lt;1&gt;2≥3&lt;4&gt;5&gt;6&gt;7. Cephalon 1.2 times as long as wide, rostrum as long as the lateral angles of cephalon, base of the mouthparts with two sharp spines. Pereonite 7 rectangular, 1.3 times as wide as long; pleonites 1–5 fused, broadened distally, slightly longer than pereonite 7; telson linguiform, narrow distal region, mid-distal region with a group of seven setae, apex slightly excavated with three pairs of setae; a pair of the statocysts.
</paragraph>
<paragraph id="044B36C3BA57FFB1FF669080FB5F4A6D" blockId="24.[188,1399,1220,1352]" pageId="24" pageNumber="85">
ANTENNULE (
<figureCitation id="9CCF2A46BA57FFB1FE8C9080FE434BFB" box="[343,457,1220,1246]" captionStart="Fig" captionStartId="25.[189,232,1882,1908]" captionTargetBox="[211,1358,286,1821]" captionTargetId="figure-11@25.[193,1394,265,1841]" captionTargetPageId="25" captionText="Fig. 14. Skuphonura oaxaquensis sp. nov. A–B, D–H. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Habitus and detail of the pleotelson. B. Cephalon. C. Paratype, ♂, 7.3 mm (UMAR-PERA 549). Base of mouthparts. D. Antennule. E. Antenna and detail of flagellum. F. Maxilliped. G. Maxilla. H. Mandible and detail of the lacinia." figureDoi="http://doi.org/10.5281/zenodo.5121796" httpUri="https://zenodo.org/record/5121796/files/figure.png" pageId="24" pageNumber="85">Fig. 14D</figureCitation>
). Peduncle with three articles. Articles 1 and 2 with plumose sensory setae; article 2 rectangular, slightly more than half the length of the first with six long distolateral setae; article 3 sub-equal in length to second with one long seta in the external distal angle and two in the internal one; flagellum with three articles, last tiny article with three aesthetascs and two smooth setae.
</paragraph>
<paragraph id="044B36C3BA57FFB1FF66912BFCBB4A89" blockId="24.[189,1398,1391,1452]" pageId="24" pageNumber="85">
ANTENNA (
<figureCitation id="9CCF2A46BA57FFB1FEE2912BFE294AAC" box="[313,419,1391,1417]" captionStart="Fig" captionStartId="25.[189,232,1882,1908]" captionTargetBox="[211,1358,286,1821]" captionTargetId="figure-11@25.[193,1394,265,1841]" captionTargetPageId="25" captionText="Fig. 14. Skuphonura oaxaquensis sp. nov. A–B, D–H. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Habitus and detail of the pleotelson. B. Cephalon. C. Paratype, ♂, 7.3 mm (UMAR-PERA 549). Base of mouthparts. D. Antennule. E. Antenna and detail of flagellum. F. Maxilliped. G. Maxilla. H. Mandible and detail of the lacinia." figureDoi="http://doi.org/10.5281/zenodo.5121796" httpUri="https://zenodo.org/record/5121796/files/figure.png" pageId="24" pageNumber="85">Fig. 14E</figureCitation>
). Peduncle with five articles. Article 2 with ventrodistal angle projected to article 3; flagellum with four articles with numerous thin setae.
</paragraph>
<paragraph id="044B36C3BA57FFB1FF669190FB6E4934" blockId="24.[188,1399,1491,1553]" pageId="24" pageNumber="85">
MAXILLIPED (
<figureCitation id="9CCF2A46BA57FFB1FE8F9197FE314AC8" box="[340,443,1491,1518]" captionStart="Fig" captionStartId="25.[189,232,1882,1908]" captionTargetBox="[211,1358,286,1821]" captionTargetId="figure-11@25.[193,1394,265,1841]" captionTargetPageId="25" captionText="Fig. 14. Skuphonura oaxaquensis sp. nov. A–B, D–H. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Habitus and detail of the pleotelson. B. Cephalon. C. Paratype, ♂, 7.3 mm (UMAR-PERA 549). Base of mouthparts. D. Antennule. E. Antenna and detail of flagellum. F. Maxilliped. G. Maxilla. H. Mandible and detail of the lacinia." figureDoi="http://doi.org/10.5281/zenodo.5121796" httpUri="https://zenodo.org/record/5121796/files/figure.png" pageId="24" pageNumber="85">Fig. 14F</figureCitation>
). Palp with three articles; article 1 with concave external margin; article 2 larger to the others with sinuous internal margin; distal article subcircular with a row of distal setae.
</paragraph>
<paragraph id="044B36C3BA57FFB1FF66927CFD4B4977" blockId="24.[189,705,1592,1618]" box="[189,705,1592,1618]" pageId="24" pageNumber="85">
MAXILLA (
<figureCitation id="9CCF2A46BA57FFB1FEED927CFE294977" box="[310,419,1592,1618]" captionStart="Fig" captionStartId="25.[189,232,1882,1908]" captionTargetBox="[211,1358,286,1821]" captionTargetId="figure-11@25.[193,1394,265,1841]" captionTargetPageId="25" captionText="Fig. 14. Skuphonura oaxaquensis sp. nov. A–B, D–H. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Habitus and detail of the pleotelson. B. Cephalon. C. Paratype, ♂, 7.3 mm (UMAR-PERA 549). Base of mouthparts. D. Antennule. E. Antenna and detail of flagellum. F. Maxilliped. G. Maxilla. H. Mandible and detail of the lacinia." figureDoi="http://doi.org/10.5281/zenodo.5121796" httpUri="https://zenodo.org/record/5121796/files/figure.png" pageId="24" pageNumber="85">Fig. 14G</figureCitation>
). With eight distal teeth.
</paragraph>
<paragraph id="044B36C3BA57FFB1FF66923EFCD649D8" blockId="24.[188,1399,1657,1789]" pageId="24" pageNumber="85">
MANDIBLE (
<figureCitation id="9CCF2A46BA57FFB1FE9F923DFE3849B6" box="[324,434,1657,1684]" captionStart="Fig" captionStartId="25.[189,232,1882,1908]" captionTargetBox="[211,1358,286,1821]" captionTargetId="figure-11@25.[193,1394,265,1841]" captionTargetPageId="25" captionText="Fig. 14. Skuphonura oaxaquensis sp. nov. A–B, D–H. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Habitus and detail of the pleotelson. B. Cephalon. C. Paratype, ♂, 7.3 mm (UMAR-PERA 549). Base of mouthparts. D. Antennule. E. Antenna and detail of flagellum. F. Maxilliped. G. Maxilla. H. Mandible and detail of the lacinia." figureDoi="http://doi.org/10.5281/zenodo.5121796" httpUri="https://zenodo.org/record/5121796/files/figure.png" pageId="24" pageNumber="85">Fig. 14H</figureCitation>
). Symmetrical; palp with three articles, article 1 sub-equal in length to the second with a long pectinate seta; article 2 with three pectinate setae and setulated scales in dorsal margin; article 3 small, 0.4 times as long as second, with four distal setae; incisor with four teeth; lacinia mobilis with 11 denticles and molars with two rounded tubercles.
</paragraph>
<paragraph id="044B36C3BA57FFB1FF669361FB19488D" blockId="24.[189,1399,1828,1961]" pageId="24" pageNumber="85">
PEREOPOD 1 (
<figureCitation id="9CCF2A46BA57FFB1FE869360FE44481A" box="[349,462,1828,1855]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[205,1372,287,1839]" captionTargetId="figure-16@26.[189,1398,265,1853]" captionTargetPageId="26" captionText="Fig. 15. Skuphonura oaxaquensis sp. nov., holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Pereopod 1. B. Pereopod 2 and detail of the carpus (b) and sensory spine. C. Pereopod 3. D. Pereopod 4. E. Pereopod 5. F. Pereopod 6. G. Pereopod 7 and detail of the sensory spines.." figureDoi="http://doi.org/10.5281/zenodo.5121798" httpUri="https://zenodo.org/record/5121798/files/figure.png" pageId="24" pageNumber="85">Fig. 15A</figureCitation>
). Subchelate; basis with dorsodistal angle lobate; carpus subtriangular, distal angle projected transversally; palm concave with a strong proximal tooth; proximal margin with six setae, distal margin with eight setae; internal margin of dactylus slightly concave with a cuticular ridge; unguis slightly more than half the length of dactylus with a slight curvature distally.
</paragraph>
<caption id="508B664BBA56FFB0FF66931EFBD848FB" ID-DOI="http://doi.org/10.5281/zenodo.5121796" ID-Zenodo-Dep="5121796" httpUri="https://zenodo.org/record/5121796/files/figure.png" pageId="25" pageNumber="86" startId="25.[189,232,1882,1908]" targetBox="[211,1358,286,1821]" targetPageId="25">
<paragraph id="044B36C3BA56FFB0FF66931EFBD848FB" blockId="25.[189,1399,1882,2015]" pageId="25" pageNumber="86">
<emphasis id="3680EAD1BA56FFB0FF66931EFE974851" bold="true" box="[189,285,1882,1908]" pageId="25" pageNumber="86">Fig. 14.</emphasis>
<taxonomicName id="C3F44D40BA56FFB0FEFC931FFDC54851" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[295,591,1883,1909]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA56FFB0FEFC931FFDC54851" box="[295,591,1883,1909]" italics="true" pageId="25" pageNumber="86">Skuphonura oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA56FFB0FD82931FFD3D4850" box="[601,695,1883,1909]" pageId="25" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="3680EAD1BA56FFB0FD1B931EFCCE4850" bold="true" box="[704,836,1882,1909]" pageId="25" pageNumber="86">A–B, D–H</emphasis>
. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). 
<emphasis id="3680EAD1BA56FFB0FD1A933AFD5C48BD" bold="true" box="[705,726,1918,1944]" pageId="25" pageNumber="86">A</emphasis>
. Habitus and detail of the pleotelson. 
<emphasis id="3680EAD1BA56FFB0FB08933AFB6D48BD" bold="true" box="[1235,1255,1918,1944]" pageId="25" pageNumber="86">B</emphasis>
. Cephalon. 
<emphasis id="3680EAD1BA56FFB0FF6693E5FF58489E" bold="true" box="[189,210,1953,1979]" pageId="25" pageNumber="86">C</emphasis>
. Paratype, ♂, 7.3 mm (UMAR-PERA 549). Base of mouthparts. 
<emphasis id="3680EAD1BA56FFB0FC1593E5FC69489E" bold="true" box="[974,995,1953,1979]" pageId="25" pageNumber="86">D</emphasis>
. Antennule. 
<emphasis id="3680EAD1BA56FFB0FBAC93E5FB01489E" bold="true" box="[1143,1163,1953,1979]" pageId="25" pageNumber="86">E</emphasis>
. Antenna and detail of flagellum. 
<emphasis id="3680EAD1BA56FFB0FE819380FEE648FB" bold="true" box="[346,364,1988,2014]" pageId="25" pageNumber="86">F</emphasis>
. Maxilliped. 
<emphasis id="3680EAD1BA56FFB0FDD19380FDAB48FB" bold="true" box="[522,545,1988,2014]" pageId="25" pageNumber="86">G</emphasis>
. Maxilla. 
<emphasis id="3680EAD1BA56FFB0FD419380FD3B48FB" bold="true" box="[666,689,1988,2014]" pageId="25" pageNumber="86">H</emphasis>
. Mandible and detail of the lacinia.
</paragraph>
</caption>
<caption id="508B664BBA55FFB3FF669320FE9C48CD" ID-DOI="http://doi.org/10.5281/zenodo.5121798" ID-Zenodo-Dep="5121798" httpUri="https://zenodo.org/record/5121798/files/figure.png" pageId="26" pageNumber="87" startId="26.[189,232,1892,1918]" targetBox="[205,1372,287,1839]" targetPageId="26">
<paragraph id="044B36C3BA55FFB3FF669320FE9C48CD" blockId="26.[188,1399,1892,2024]" pageId="26" pageNumber="87">
<emphasis id="3680EAD1BA55FFB3FF669320FE97485B" bold="true" box="[189,285,1892,1918]" pageId="26" pageNumber="87">Fig. 15.</emphasis>
<taxonomicName id="C3F44D40BA55FFB3FEFC9320FDDA485B" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[295,592,1892,1918]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="26" pageNumber="87" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA55FFB3FEFC9320FDDA485B" box="[295,592,1892,1918]" italics="true" pageId="26" pageNumber="87">Skuphonura oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA55FFB3FD819321FD30485A" box="[602,698,1893,1919]" pageId="26" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). 
<emphasis id="3680EAD1BA55FFB3FDD293C3FD944884" bold="true" box="[521,542,1927,1953]" pageId="26" pageNumber="87">A</emphasis>
. Pereopod 1. 
<emphasis id="3680EAD1BA55FFB3FD6793C3FD5A4884" bold="true" box="[700,720,1927,1953]" pageId="26" pageNumber="87">B</emphasis>
. Pereopod 2 and detail of the carpus (b) and sensory spine. 
<emphasis id="3680EAD1BA55FFB3FF6793EEFF5B48E1" bold="true" box="[188,209,1962,1988]" pageId="26" pageNumber="87">C</emphasis>
. Pereopod 3. 
<emphasis id="3680EAD1BA55FFB3FEA393EFFE0648E0" bold="true" box="[376,396,1963,1989]" pageId="26" pageNumber="87">D</emphasis>
. Pereopod 4. 
<emphasis id="3680EAD1BA55FFB3FDE993EFFDCF48E0" bold="true" box="[562,581,1963,1989]" pageId="26" pageNumber="87">E</emphasis>
. Pereopod 5. 
<emphasis id="3680EAD1BA55FFB3FD3193EFFD7148E0" bold="true" box="[746,763,1963,1989]" pageId="26" pageNumber="87">F</emphasis>
. Pereopod 6. 
<emphasis id="3680EAD1BA55FFB3FC7A93EEFC3D48E1" bold="true" box="[929,951,1962,1988]" pageId="26" pageNumber="87">G</emphasis>
. Pereopod 7 and detail of the sensory spines..
</paragraph>
</caption>
<paragraph id="044B36C3BA54FFB2FF66954DFB7B4E4F" blockId="27.[188,1398,265,362]" pageId="27" pageNumber="88">
PEREOPODS 2–3 (
<figureCitation id="9CCF2A46BA54FFB2FEA7954DFD834E01" box="[380,521,265,292]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[205,1372,287,1839]" captionTargetId="figure-16@26.[189,1398,265,1853]" captionTargetPageId="26" captionText="Fig. 15. Skuphonura oaxaquensis sp. nov., holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Pereopod 1. B. Pereopod 2 and detail of the carpus (b) and sensory spine. C. Pereopod 3. D. Pereopod 4. E. Pereopod 5. F. Pereopod 6. G. Pereopod 7 and detail of the sensory spines.." figureDoi="http://doi.org/10.5281/zenodo.5121798" httpUri="https://zenodo.org/record/5121798/files/figure.png" pageId="27" pageNumber="88">Fig. 15B–C</figureCitation>
). Similar in shape and size. Merus internal margin with cuticular ridges and numerous setae; carpus triangular, distal margin with a denticulate seta; palm with numerous setae and setulated scales, ventral angle with a denticulate sensory spine and a group of smooth setae.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF6695D5FBA24D30" blockId="27.[189,1399,401,533]" pageId="27" pageNumber="88">
PEREOPODS 4–6 (
<figureCitation id="9CCF2A46BA54FFB2FE5F95D5FD9E4E89" box="[388,532,401,428]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[205,1372,287,1839]" captionTargetId="figure-16@26.[189,1398,265,1853]" captionTargetPageId="26" captionText="Fig. 15. Skuphonura oaxaquensis sp. nov., holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Pereopod 1. B. Pereopod 2 and detail of the carpus (b) and sensory spine. C. Pereopod 3. D. Pereopod 4. E. Pereopod 5. F. Pereopod 6. G. Pereopod 7 and detail of the sensory spines.." figureDoi="http://doi.org/10.5281/zenodo.5121798" httpUri="https://zenodo.org/record/5121798/files/figure.png" pageId="27" pageNumber="88">Fig. 15D–F</figureCitation>
). Similar in shape and size. Merus margin with cuticular ridges; carpus subtriangular, truncated distally with a simple sensory spine and simple setae; propodus cylindrical, 2–2.1 times as long as wide in P4 and P5, 
<quantity id="C30C9B26BA54FFB2FD669593FC8B4ED4" box="[701,769,471,497]" metricMagnitude="-2" metricUnit="m" metricValue="5.842" pageId="27" pageNumber="88" unit="in" value="2.3">2.3 in</quantity>
P6, internal margin with setulated scales and simple setae, ventrodistal angle with a denticulate sensory spine and simple setae.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF669678FD7D4DB9" blockId="27.[189,1399,572,668]" pageId="27" pageNumber="88">
PEREOPOD 7 (
<figureCitation id="9CCF2A46BA54FFB2FEB99678FE5F4D72" box="[354,469,572,599]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[205,1372,287,1839]" captionTargetId="figure-16@26.[189,1398,265,1853]" captionTargetPageId="26" captionText="Fig. 15. Skuphonura oaxaquensis sp. nov., holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Pereopod 1. B. Pereopod 2 and detail of the carpus (b) and sensory spine. C. Pereopod 3. D. Pereopod 4. E. Pereopod 5. F. Pereopod 6. G. Pereopod 7 and detail of the sensory spines.." figureDoi="http://doi.org/10.5281/zenodo.5121798" httpUri="https://zenodo.org/record/5121798/files/figure.png" pageId="27" pageNumber="88">Fig. 15G</figureCitation>
). Slenderer than the preceding ones, propodus 3.3 times as long as wide, ventrodistal margin with setulated scales and one denticulate sensory spine and two pectinate spines; lateral margins of dactylus with setulated scales.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF669680FC694DFB" blockId="27.[189,995,708,734]" box="[189,995,708,734]" pageId="27" pageNumber="88">PEREOPODS 2–7. With smooth and plumose sensory setae at the basis.</paragraph>
<paragraph id="044B36C3BA54FFB2FF669742FE494C43" blockId="27.[189,1398,773,870]" pageId="27" pageNumber="88">
PLEOPOD 1 (
<figureCitation id="9CCF2A46BA54FFB2FE9D9741FE384C05" box="[326,434,773,800]" captionStart="Fig" captionStartId="28.[189,232,1915,1941]" captionTargetBox="[236,1351,290,1848]" captionTargetId="figure-16@28.[189,1398,275,1863]" captionTargetPageId="28" captionText="Fig. 16. Skuphonura oaxaquensis sp. nov.A–D. Holotype, ♀, 7.9 mm, Oaxaca, PuertoÁngel, Estacahuite Beach (UMAR-PERA 548). A. Pleopod 1. B. Pleopod 2. C. Uropodal endopod. D. Uropodal exopod. E–G. Paratype, ♂, 7.3 mm (UMAR-PERA 549). E. Antennule. F. Pereopod 1. G. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.5121800" httpUri="https://zenodo.org/record/5121800/files/figure.png" pageId="27" pageNumber="88">Fig. 16A</figureCitation>
). Protopod with three retinaculae; exopod operculiform, 2.2 times as long as wide, surrounded distally by 40 plumose setae; endopod 0.7 times the length of exopod, surrounded distally by five plumose setae.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF6697C9FAB84CEE" blockId="27.[189,1398,909,971]" pageId="27" pageNumber="88">
PLEOPOD 2 (
<figureCitation id="9CCF2A46BA54FFB2FE9C97C9FE3B4C8D" box="[327,433,909,936]" captionStart="Fig" captionStartId="28.[189,232,1915,1941]" captionTargetBox="[236,1351,290,1848]" captionTargetId="figure-16@28.[189,1398,275,1863]" captionTargetPageId="28" captionText="Fig. 16. Skuphonura oaxaquensis sp. nov.A–D. Holotype, ♀, 7.9 mm, Oaxaca, PuertoÁngel, Estacahuite Beach (UMAR-PERA 548). A. Pleopod 1. B. Pleopod 2. C. Uropodal endopod. D. Uropodal exopod. E–G. Paratype, ♂, 7.3 mm (UMAR-PERA 549). E. Antennule. F. Pereopod 1. G. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.5121800" httpUri="https://zenodo.org/record/5121800/files/figure.png" pageId="27" pageNumber="88">Fig. 16B</figureCitation>
). Oval and elongated; exopod 2.8 times as long as wide, surrounded distally by 12 plumose setae; endopod slightly shorter than exopod, surrounded distally by eight plumose setae.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF6697B6FB8D4B0A" blockId="27.[189,1398,1009,1071]" pageId="27" pageNumber="88">
UROPODAL PROTOPOD (
<figureCitation id="9CCF2A46BA54FFB2FE6697B5FDA34B29" box="[445,553,1009,1036]" captionStart="Fig" captionStartId="28.[189,232,1915,1941]" captionTargetBox="[236,1351,290,1848]" captionTargetId="figure-16@28.[189,1398,275,1863]" captionTargetPageId="28" captionText="Fig. 16. Skuphonura oaxaquensis sp. nov.A–D. Holotype, ♀, 7.9 mm, Oaxaca, PuertoÁngel, Estacahuite Beach (UMAR-PERA 548). A. Pleopod 1. B. Pleopod 2. C. Uropodal endopod. D. Uropodal exopod. E–G. Paratype, ♂, 7.3 mm (UMAR-PERA 549). E. Antennule. F. Pereopod 1. G. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.5121800" httpUri="https://zenodo.org/record/5121800/files/figure.png" pageId="27" pageNumber="88">Fig. 16C</figureCitation>
). Rectangular, 1.7 times as long as wide, 1.2 times as long as endopod, external margin with plumose setae, ventrodistal angle with a long seta.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF669013FDC14BB1" blockId="27.[189,1398,1110,1172]" pageId="27" pageNumber="88">
UROPODAL ENDOPOD (
<figureCitation id="9CCF2A46BA54FFB2FE6D9012FDAE4B54" box="[438,548,1110,1137]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[205,1372,287,1839]" captionTargetId="figure-16@26.[189,1398,265,1853]" captionTargetPageId="26" captionText="Fig. 15. Skuphonura oaxaquensis sp. nov., holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Pereopod 1. B. Pereopod 2 and detail of the carpus (b) and sensory spine. C. Pereopod 3. D. Pereopod 4. E. Pereopod 5. F. Pereopod 6. G. Pereopod 7 and detail of the sensory spines.." figureDoi="http://doi.org/10.5281/zenodo.5121798" httpUri="https://zenodo.org/record/5121798/files/figure.png" pageId="27" pageNumber="88">Fig. 15C</figureCitation>
). Conical, 1.9 times as long as basal width, apically rounded, external margin with numerous long setae.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF6690FFFDCA4BDC" blockId="27.[189,1398,1211,1273]" pageId="27" pageNumber="88">
UROPODAL EXOPOD (
<figureCitation id="9CCF2A46BA54FFB2FE7090FFFD964BF3" box="[427,540,1211,1238]" captionStart="Fig" captionStartId="28.[189,232,1915,1941]" captionTargetBox="[236,1351,290,1848]" captionTargetId="figure-16@28.[189,1398,275,1863]" captionTargetPageId="28" captionText="Fig. 16. Skuphonura oaxaquensis sp. nov.A–D. Holotype, ♀, 7.9 mm, Oaxaca, PuertoÁngel, Estacahuite Beach (UMAR-PERA 548). A. Pleopod 1. B. Pleopod 2. C. Uropodal endopod. D. Uropodal exopod. E–G. Paratype, ♂, 7.3 mm (UMAR-PERA 549). E. Antennule. F. Pereopod 1. G. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.5121800" httpUri="https://zenodo.org/record/5121800/files/figure.png" pageId="27" pageNumber="88">Fig. 16D</figureCitation>
). Suboval with a distal notch, margin with 17 long smooth setae and approximately 55 plumose setae.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF319164FBDE4A1F" blockId="27.[189,1108,1311,1373]" box="[234,1108,1311,1338]" pageId="27" pageNumber="88">
<emphasis id="3680EAD1BA54FFB2FF319164FEA04A1F" bold="true" box="[234,298,1312,1338]" pageId="27" pageNumber="88">Male</emphasis>
(dimorphic characters; based on the 
<typeStatus id="DB4F8861BA54FFB2FD009164FCCA4A1F" box="[731,832,1312,1338]" pageId="27" pageNumber="88" type="paratype">paratype</typeStatus>
<specimenCount id="12F2FD4ABA54FFB2FC939164FCEA4A1F" box="[840,864,1312,1338]" pageId="27" pageNumber="88" type="male">♂</specimenCount>
, UMAR-PERA 549)
</paragraph>
<paragraph id="044B36C3BA54FFB2FF669107FBCA4A78" blockId="27.[189,1108,1311,1373]" box="[189,1088,1347,1373]" pageId="27" pageNumber="88">
BODY. Mid-region of the cephalon (
<figureCitation id="9CCF2A46BA54FFB2FD859107FD4D4A78" box="[606,711,1347,1373]" captionStart="Fig" captionStartId="25.[189,232,1882,1908]" captionTargetBox="[211,1358,286,1821]" captionTargetId="figure-11@25.[193,1394,265,1841]" captionTargetPageId="25" captionText="Fig. 14. Skuphonura oaxaquensis sp. nov. A–B, D–H. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). A. Habitus and detail of the pleotelson. B. Cephalon. C. Paratype, ♂, 7.3 mm (UMAR-PERA 549). Base of mouthparts. D. Antennule. E. Antenna and detail of flagellum. F. Maxilliped. G. Maxilla. H. Mandible and detail of the lacinia." figureDoi="http://doi.org/10.5281/zenodo.5121796" httpUri="https://zenodo.org/record/5121796/files/figure.png" pageId="27" pageNumber="88">Fig. 14C</figureCitation>
) with a poorly developed spine.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF6691C0FECD4AC1" blockId="27.[188,1399,1412,1508]" pageId="27" pageNumber="88">
ANTENNULE (
<figureCitation id="9CCF2A46BA54FFB2FE8991C0FE314ABA" box="[338,443,1412,1439]" captionStart="Fig" captionStartId="28.[189,232,1915,1941]" captionTargetBox="[236,1351,290,1848]" captionTargetId="figure-16@28.[189,1398,275,1863]" captionTargetPageId="28" captionText="Fig. 16. Skuphonura oaxaquensis sp. nov.A–D. Holotype, ♀, 7.9 mm, Oaxaca, PuertoÁngel, Estacahuite Beach (UMAR-PERA 548). A. Pleopod 1. B. Pleopod 2. C. Uropodal endopod. D. Uropodal exopod. E–G. Paratype, ♂, 7.3 mm (UMAR-PERA 549). E. Antennule. F. Pereopod 1. G. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.5121800" httpUri="https://zenodo.org/record/5121800/files/figure.png" pageId="27" pageNumber="88">Fig. 16E</figureCitation>
). Peduncle with three articles; article 1 and 2 with plumose sensory setae; article 2 with six long setae in the external distal angle; flagellum with nine articles, distal article with three aesthetascs.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF669248FC344949" blockId="27.[189,1399,1548,1645]" pageId="27" pageNumber="88">
PEREOPOD 1 (
<figureCitation id="9CCF2A46BA54FFB2FE8D9248FE354902" box="[342,447,1548,1575]" captionStart="Fig" captionStartId="28.[189,232,1915,1941]" captionTargetBox="[236,1351,290,1848]" captionTargetId="figure-16@28.[189,1398,275,1863]" captionTargetPageId="28" captionText="Fig. 16. Skuphonura oaxaquensis sp. nov.A–D. Holotype, ♀, 7.9 mm, Oaxaca, PuertoÁngel, Estacahuite Beach (UMAR-PERA 548). A. Pleopod 1. B. Pleopod 2. C. Uropodal endopod. D. Uropodal exopod. E–G. Paratype, ♂, 7.3 mm (UMAR-PERA 549). E. Antennule. F. Pereopod 1. G. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.5121800" httpUri="https://zenodo.org/record/5121800/files/figure.png" pageId="27" pageNumber="88">Fig. 16F</figureCitation>
). Subchelate; carpus triangular, distally produced in a narrow triangular process; propodus oval, palm straight with a triangular proximal process, accompanied by eight setae; dactylus 1.4 times as long as unguis, distoventral angle with a small spine.
</paragraph>
<paragraph id="044B36C3BA54FFB2FF6692D0FAAF49D0" blockId="27.[188,1399,1684,1781]" pageId="27" pageNumber="88">
PLEOPOD 2 (
<figureCitation id="9CCF2A46BA54FFB2FE8E92D0FE42498A" box="[341,456,1684,1711]" captionStart="Fig" captionStartId="28.[189,232,1915,1941]" captionTargetBox="[236,1351,290,1848]" captionTargetId="figure-16@28.[189,1398,275,1863]" captionTargetPageId="28" captionText="Fig. 16. Skuphonura oaxaquensis sp. nov.A–D. Holotype, ♀, 7.9 mm, Oaxaca, PuertoÁngel, Estacahuite Beach (UMAR-PERA 548). A. Pleopod 1. B. Pleopod 2. C. Uropodal endopod. D. Uropodal exopod. E–G. Paratype, ♂, 7.3 mm (UMAR-PERA 549). E. Antennule. F. Pereopod 1. G. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.5121800" httpUri="https://zenodo.org/record/5121800/files/figure.png" pageId="27" pageNumber="88">Fig. 16G</figureCitation>
). Endopod and exopod sub-equal in length; exopod surrounded distally by 11 plumose setae; endopod surrounded distally by seven plumose setae; male appendix thin, longer than the branches, emerging from the middle proximal part of the endopod, ending in a pointed apex.
</paragraph>
</subSubSection>
<subSubSection id="4CEE6548BA54FFB2FF66935FFB754840" pageId="27" pageNumber="88" type="distribution">
<paragraph id="044B36C3BA54FFB2FF66935FFE644810" blockId="27.[189,494,1819,1846]" box="[189,494,1819,1846]" pageId="27" pageNumber="88">
<heading id="5F0381AFBA54FFB2FF66935FFE644810" bold="true" box="[189,494,1819,1846]" fontSize="11" level="3" pageId="27" pageNumber="88" reason="3">
<emphasis id="3680EAD1BA54FFB2FF66935FFE644810" bold="true" box="[189,494,1819,1846]" pageId="27" pageNumber="88">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="044B36C3BA54FFB2FF66930FFB754840" blockId="27.[189,1279,1866,1893]" box="[189,1279,1866,1893]" pageId="27" pageNumber="88">
Known only from the 
<typeStatus id="DB4F8861BA54FFB2FE19930FFE7F4840" box="[450,501,1867,1893]" pageId="27" pageNumber="88">type</typeStatus>
locality, Estacahuite Beach, 
<collectingRegion id="C630F821BA54FFB2FC9C930EFC294841" box="[839,931,1866,1892]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="88">Oaxaca</collectingRegion>
. In coral rock; subtidal (
<quantity id="C30C9B26BA54FFB2FB1B930EFB784840" box="[1216,1266,1866,1893]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="88" unit="m" value="4.0">4 m</quantity>
).
</paragraph>
</subSubSection>
<caption id="508B664BBA53FFB5FF66933FFA8A48F9" ID-DOI="http://doi.org/10.5281/zenodo.5121800" ID-Zenodo-Dep="5121800" httpUri="https://zenodo.org/record/5121800/files/figure.png" pageId="28" pageNumber="89" startId="28.[189,232,1915,1941]" targetBox="[236,1351,290,1848]" targetPageId="28">
<paragraph id="044B36C3BA53FFB5FF66933FFA8A48F9" blockId="28.[189,1399,1915,2012]" pageId="28" pageNumber="89">
<emphasis id="3680EAD1BA53FFB5FF66933FFE9248B0" bold="true" box="[189,280,1915,1941]" pageId="28" pageNumber="89">Fig. 16.</emphasis>
<taxonomicName id="C3F44D40BA53FFB5FEC6933FFDCA48B0" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[285,576,1915,1941]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="28" pageNumber="89" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA53FFB5FEC6933FFDCA48B0" box="[285,576,1915,1941]" italics="true" pageId="28" pageNumber="89">Skuphonura oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA53FFB5FD9E9338FD1748B3" box="[581,669,1916,1942]" pageId="28" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="3680EAD1BA53FFB5FD44933FFD5248B0" bold="true" box="[671,728,1915,1941]" pageId="28" pageNumber="89">A–D</emphasis>
. Holotype, ♀, 7.9 mm, Oaxaca, Puerto Ángel, Estacahuite Beach (UMAR-PERA 548). 
<emphasis id="3680EAD1BA53FFB5FDCF93DAFDA2489D" bold="true" box="[532,552,1950,1976]" pageId="28" pageNumber="89">A</emphasis>
. Pleopod 1. 
<emphasis id="3680EAD1BA53FFB5FD1A93DAFD59489D" bold="true" box="[705,723,1950,1976]" pageId="28" pageNumber="89">B</emphasis>
. Pleopod 2. 
<emphasis id="3680EAD1BA53FFB5FCB793DAFC0A489D" bold="true" box="[876,896,1950,1976]" pageId="28" pageNumber="89">C</emphasis>
. Uropodal endopod. 
<emphasis id="3680EAD1BA53FFB5FBA693DAFB1B489D" bold="true" box="[1149,1169,1950,1976]" pageId="28" pageNumber="89">D</emphasis>
. Uropodal exopod. 
<emphasis id="3680EAD1BA53FFB5FF659385FF7C48FE" bold="true" box="[190,246,1985,2011]" pageId="28" pageNumber="89">E–G</emphasis>
. Paratype, ♂, 7.3 mm (UMAR-PERA 549). 
<emphasis id="3680EAD1BA53FFB5FCDB9386FC9E48F9" bold="true" box="[768,788,1986,2012]" pageId="28" pageNumber="89">E</emphasis>
. Antennule. 
<emphasis id="3680EAD1BA53FFB5FC739386FC3048F9" bold="true" box="[936,954,1986,2012]" pageId="28" pageNumber="89">F</emphasis>
. Pereopod 1. 
<emphasis id="3680EAD1BA53FFB5FB859385FBFF48FE" bold="true" box="[1118,1141,1985,2011]" pageId="28" pageNumber="89">G</emphasis>
. Pleopod 2.
</paragraph>
</caption>
<caption id="508B664BBA52FFB4FF66954DFB594E04" ID-Table-UUID="508B664BBA52FFB4FF66954DFB594E04" box="[189,1235,265,289]" httpUri="http://table.plazi.org/id/508B664BBA52FFB4FF66954DFB594E04" pageId="29" pageNumber="90" startId="29.[189,252,265,289]" targetBox="[200,1366,348,1310]" targetIsTable="true" targetPageId="29">
<paragraph id="044B36C3BA52FFB4FF66954DFB594E04" blockId="29.[189,1235,265,289]" box="[189,1235,265,289]" pageId="29" pageNumber="90">
<emphasis id="3680EAD1BA52FFB4FF66954DFE9C4E04" bold="true" box="[189,278,265,289]" pageId="29" pageNumber="90">Table 2.</emphasis>
Comparison between 
<taxonomicName id="C3F44D40BA52FFB4FDDD954DFC9B4E04" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[518,785,265,289]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA52FFB4FDDD954DFC9B4E04" box="[518,785,265,289]" italics="true" pageId="29" pageNumber="90">Skuphonura oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA52FFB4FCC3954DFCE04E05" box="[792,874,265,288]" pageId="29" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
and the closest species of genera.
</paragraph>
</caption>
<paragraph id="044B36C3BA52FFB4FF13952EFAA24A24" pageId="29" pageNumber="90">
<table id="76F4C463BA520056FF139518FADC4A3B" box="[200,1366,348,1310]" gridcols="5" gridrows="14" pageId="29" pageNumber="90">
<tr id="BAC43481BA520056FF139518FADC4EAB" box="[200,1366,348,398]" gridrow="0" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF139518FE114EAB" box="[200,411,348,398]" gridcol="0" gridrow="0" pageId="29" pageNumber="90">
<emphasis id="3680EAD1BA52FFB4FF13952EFE004EA5" bold="true" box="[200,394,362,384]" pageId="29" pageNumber="90">Character / species</emphasis>
</th>
<th id="F9155DFDBA520056FE689518FD0A4EAB" box="[435,640,348,398]" gridcol="1" gridrow="0" pageId="29" pageNumber="90">
<taxonomicName id="C3F44D40BA52FFB4FE689518FDB54EAB" authority="Kensley, 1980" authorityName="Kensley" authorityYear="1980" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="3680EAD1BA52FFB4FE689518FDB54EAB" bold="true" pageId="29" pageNumber="90">
<emphasis id="3680EAD1BA52FFB4FE689518FDC44E57" bold="true" box="[435,590,348,370]" italics="true" pageId="29" pageNumber="90">S. ecuadorensis</emphasis>
Kensley, 1980
</emphasis>
</taxonomicName>
</th>
<th id="F9155DFDBA520056FD449518FCEC4EAB" box="[671,870,348,398]" gridcol="2" gridrow="0" pageId="29" pageNumber="90">
<taxonomicName id="C3F44D40BA52FFB4FD449518FCA14EAB" authority="Kensley, 1980" authorityName="Kensley" authorityYear="1980" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="itapuca">
<emphasis id="3680EAD1BA52FFB4FD449518FCA14EAB" bold="true" pageId="29" pageNumber="90">
<emphasis id="3680EAD1BA52FFB4FD449518FC8B4E54" bold="true" box="[671,769,348,369]" italics="true" pageId="29" pageNumber="90">S. itapuca</emphasis>
Kensley, 1980
</emphasis>
</taxonomicName>
</th>
<th id="F9155DFDBA520056FC509518FBF84EAB" box="[907,1138,348,398]" gridcol="3" gridrow="0" pageId="29" pageNumber="90">
<taxonomicName id="C3F44D40BA52FFB4FC509518FB814EAB" authority="Brusca &amp; Muller, 1991" authorityName="Brusca &amp; Muller" authorityYear="1991" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="3680EAD1BA52FFB4FC509518FB814EAB" bold="true" pageId="29" pageNumber="90">
<emphasis id="3680EAD1BA52FFB4FC509518FC7E4E57" bold="true" box="[907,1012,348,370]" italics="true" pageId="29" pageNumber="90">S. kensleyi</emphasis>
Brusca &amp; Müller, 1991
</emphasis>
</taxonomicName>
</th>
<th id="F9155DFDBA520056FB529518FADC4EAB" box="[1161,1366,348,398]" gridcol="4" gridrow="0" pageId="29" pageNumber="90">
<emphasis id="3680EAD1BA52FFB4FB529518FADC4EAB" bold="true" box="[1161,1366,348,398]" pageId="29" pageNumber="90">
<taxonomicName id="C3F44D40BA52FFB4FB529518FA8E4EAB" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[1161,1285,348,398]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA52FFB4FB529518FA8E4EAB" bold="true" box="[1161,1285,348,398]" italics="true" pageId="29" pageNumber="90">Skuphonura oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA52FFB4FAD1953DFADC4EAB" box="[1290,1366,377,398]" pageId="29" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="BAC43481BA520056FF1395E4FADC4EF7" box="[200,1366,416,466]" gridrow="1" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF1395E4FE114EF7" box="[200,411,416,466]" gridcol="0" gridrow="1" pageId="29" pageNumber="90">Locality</th>
<td id="F9155DFDBA520056FE6895E4FD0A4EF7" box="[435,640,416,466]" gridcol="1" gridrow="1" pageId="29" pageNumber="90">La Libertad, Ecuador</td>
<td id="F9155DFDBA520056FD4495E4FCEC4EF7" box="[671,870,416,466]" gridcol="2" gridrow="1" pageId="29" pageNumber="90">Nichteroy, Rio de Janeiro, Brazil</td>
<td id="F9155DFDBA520056FC5095E4FBF84EF7" box="[907,1138,416,466]" gridcol="3" gridrow="1" pageId="29" pageNumber="90">Santa Marta, Punta de Betin, Colombia</td>
<td id="F9155DFDBA520056FB5295E4FADC4EF7" box="[1161,1366,416,466]" gridcol="4" gridrow="1" pageId="29" pageNumber="90">Estacahuite Beach, Oaxaca</td>
</tr>
<tr id="BAC43481BA520056FF1395A7FADC4EDF" box="[200,1366,483,506]" gridrow="2" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF1395A7FE114EDF" box="[200,411,483,506]" gridcol="0" gridrow="2" pageId="29" pageNumber="90">Body length</th>
<td id="F9155DFDBA520056FE6895A7FD0A4EDF" box="[435,640,483,506]" gridcol="1" gridrow="2" pageId="29" pageNumber="90">7.2 mm</td>
<td id="F9155DFDBA520056FD4495A7FCEC4EDF" box="[671,870,483,506]" gridcol="2" gridrow="2" pageId="29" pageNumber="90">9.3 mm</td>
<td id="F9155DFDBA520056FC5095A7FBF84EDF" box="[907,1138,483,506]" gridcol="3" gridrow="2" pageId="29" pageNumber="90">5.7 mm</td>
<td id="F9155DFDBA520056FB5295A7FADC4EDF" box="[1161,1366,483,506]" gridcol="4" gridrow="2" pageId="29" pageNumber="90">7.9 mm</td>
</tr>
<tr id="BAC43481BA520056FF13966CFADC4D1B" box="[200,1366,552,574]" gridrow="3" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF13966CFE114D1B" box="[200,411,552,574]" gridcol="0" gridrow="3" pageId="29" pageNumber="90">Body</th>
<td id="F9155DFDBA520056FE68966CFD0A4D1B" box="[435,640,552,574]" gridcol="1" gridrow="3" pageId="29" pageNumber="90">Not pigmented</td>
<td id="F9155DFDBA520056FD44966CFCEC4D1B" box="[671,870,552,574]" gridcol="2" gridrow="3" pageId="29" pageNumber="90">Not pigmented</td>
<td id="F9155DFDBA520056FC50966CFBF84D1B" box="[907,1138,552,574]" gridcol="3" gridrow="3" pageId="29" pageNumber="90">Reticulated pigment</td>
<td id="F9155DFDBA520056FB52966CFADC4D1B" box="[1161,1366,552,574]" gridcol="4" gridrow="3" pageId="29" pageNumber="90">Partially pigmented</td>
</tr>
<tr id="BAC43481BA520056FF139628FADC4DBB" box="[200,1366,620,670]" gridrow="4" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF139628FE114DBB" box="[200,411,620,670]" gridcol="0" gridrow="4" pageId="29" pageNumber="90">Anterior length of cephalon</th>
<td id="F9155DFDBA520056FE689628FD0A4DBB" box="[435,640,620,670]" gridcol="1" gridrow="4" pageId="29" pageNumber="90">Wider than the posterior</td>
<td id="F9155DFDBA520056FD449628FCEC4DBB" box="[671,870,620,670]" gridcol="2" gridrow="4" pageId="29" pageNumber="90">Subequal as the posterior</td>
<td id="F9155DFDBA520056FC509628FBF84DBB" box="[907,1138,620,670]" gridcol="3" gridrow="4" pageId="29" pageNumber="90">As wide as the posterior</td>
<td id="F9155DFDBA520056FB529628FADC4DBB" box="[1161,1366,620,670]" gridcol="4" gridrow="4" pageId="29" pageNumber="90">As wide as the posterior</td>
</tr>
<tr id="BAC43481BA520056FF1396F4FADC4DC7" box="[200,1366,688,738]" gridrow="5" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF1396F4FE114DC7" box="[200,411,688,738]" gridcol="0" gridrow="5" pageId="29" pageNumber="90">Medioventral processes of cephalon</th>
<td id="F9155DFDBA520056FE6896F4FD0A4DC7" box="[435,640,688,738]" gridcol="1" gridrow="5" pageId="29" pageNumber="90">1 conical spine, 1 blunt tubercle</td>
<td id="F9155DFDBA520056FD4496F4FCEC4DC7" box="[671,870,688,738]" gridcol="2" gridrow="5" pageId="29" pageNumber="90">2 conical spine</td>
<td id="F9155DFDBA520056FC5096F4FBF84DC7" box="[907,1138,688,738]" gridcol="3" gridrow="5" pageId="29" pageNumber="90">–</td>
<td id="F9155DFDBA520056FB5296F4FADC4DC7" box="[1161,1366,688,738]" gridcol="4" gridrow="5" pageId="29" pageNumber="90">2 conical spine</td>
</tr>
<tr id="BAC43481BA520056FF1396B0FADC4C03" box="[200,1366,756,806]" gridrow="6" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF1396B0FE114C03" box="[200,411,756,806]" gridcol="0" gridrow="6" pageId="29" pageNumber="90">Articles of flagellum A1</th>
<td id="F9155DFDBA520056FE6896B0FD0A4C03" box="[435,640,756,806]" gridcol="1" gridrow="6" pageId="29" pageNumber="90">1</td>
<td id="F9155DFDBA520056FD4496B0FCEC4C03" box="[671,870,756,806]" gridcol="2" gridrow="6" pageId="29" pageNumber="90">1</td>
<td id="F9155DFDBA520056FC5096B0FBF84C03" box="[907,1138,756,806]" gridcol="3" gridrow="6" pageId="29" pageNumber="90">3</td>
<td id="F9155DFDBA520056FB5296B0FADC4C03" box="[1161,1366,756,806]" gridcol="4" gridrow="6" pageId="29" pageNumber="90">3</td>
</tr>
<tr id="BAC43481BA520056FF13977CFADC4C6B" box="[200,1366,824,846]" gridrow="7" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF13977CFE114C6B" box="[200,411,824,846]" gridcol="0" gridrow="7" pageId="29" pageNumber="90">Aesthetascs</th>
<td id="F9155DFDBA520056FE68977CFD0A4C6B" box="[435,640,824,846]" gridcol="1" gridrow="7" pageId="29" pageNumber="90">2</td>
<td id="F9155DFDBA520056FD44977CFCEC4C6B" box="[671,870,824,846]" gridcol="2" gridrow="7" pageId="29" pageNumber="90">2</td>
<td id="F9155DFDBA520056FC50977CFBF84C6B" box="[907,1138,824,846]" gridcol="3" gridrow="7" pageId="29" pageNumber="90">1</td>
<td id="F9155DFDBA520056FB52977CFADC4C6B" box="[1161,1366,824,846]" gridcol="4" gridrow="7" pageId="29" pageNumber="90">3</td>
</tr>
<tr id="BAC43481BA520056FF139738FADC4C8B" box="[200,1366,892,942]" gridrow="8" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF139738FE114C8B" box="[200,411,892,942]" gridcol="0" gridrow="8" pageId="29" pageNumber="90">Article of flagellum A2</th>
<td id="F9155DFDBA520056FE689738FD0A4C8B" box="[435,640,892,942]" gridcol="1" gridrow="8" pageId="29" pageNumber="90">1</td>
<td id="F9155DFDBA520056FD449738FCEC4C8B" box="[671,870,892,942]" gridcol="2" gridrow="8" pageId="29" pageNumber="90">1</td>
<td id="F9155DFDBA520056FC509738FBF84C8B" box="[907,1138,892,942]" gridcol="3" gridrow="8" pageId="29" pageNumber="90">1</td>
<td id="F9155DFDBA520056FB529738FADC4C8B" box="[1161,1366,892,942]" gridcol="4" gridrow="8" pageId="29" pageNumber="90">4</td>
</tr>
<tr id="BAC43481BA520056FF139784FADC4CD7" box="[200,1366,960,1010]" gridrow="9" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF139784FE114CD7" box="[200,411,960,1010]" gridcol="0" gridrow="9" pageId="29" pageNumber="90">
<emphasis id="3680EAD1BA52FFB4FF139784FED44CF0" box="[200,350,960,981]" italics="true" pageId="29" pageNumber="90">Lacinia mobilis</emphasis>
denticles
</th>
<td id="F9155DFDBA520056FE689784FD0A4CD7" box="[435,640,960,1010]" gridcol="1" gridrow="9" pageId="29" pageNumber="90">11</td>
<td id="F9155DFDBA520056FD449784FCEC4CD7" box="[671,870,960,1010]" gridcol="2" gridrow="9" pageId="29" pageNumber="90">9</td>
<td id="F9155DFDBA520056FC509784FBF84CD7" box="[907,1138,960,1010]" gridcol="3" gridrow="9" pageId="29" pageNumber="90">6</td>
<td id="F9155DFDBA520056FB529784FADC4CD7" box="[1161,1366,960,1010]" gridcol="4" gridrow="9" pageId="29" pageNumber="90">10</td>
</tr>
<tr id="BAC43481BA520056FF139040FADC4B76" box="[200,1366,1028,1107]" gridrow="10" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF139040FE114B76" box="[200,411,1028,1107]" gridcol="0" gridrow="10" pageId="29" pageNumber="90">Setae of the last article of mandibular palp</th>
<td id="F9155DFDBA520056FE689040FD0A4B76" box="[435,640,1028,1107]" gridcol="1" gridrow="10" pageId="29" pageNumber="90">3</td>
<td id="F9155DFDBA520056FD449040FCEC4B76" box="[671,870,1028,1107]" gridcol="2" gridrow="10" pageId="29" pageNumber="90">3</td>
<td id="F9155DFDBA520056FC509040FBF84B76" box="[907,1138,1028,1107]" gridcol="3" gridrow="10" pageId="29" pageNumber="90">3</td>
<td id="F9155DFDBA520056FB529040FADC4B76" box="[1161,1366,1028,1107]" gridcol="4" gridrow="10" pageId="29" pageNumber="90">4</td>
</tr>
<tr id="BAC43481BA520056FF139027FADC4BB3" box="[200,1366,1123,1174]" gridrow="11" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF139027FE114BB3" box="[200,411,1123,1174]" gridcol="0" gridrow="11" pageId="29" pageNumber="90">Setae of the 2 article of mandibular palp</th>
<td id="F9155DFDBA520056FE689027FD0A4BB3" box="[435,640,1123,1174]" gridcol="1" gridrow="11" pageId="29" pageNumber="90">4</td>
<td id="F9155DFDBA520056FD449027FCEC4BB3" box="[671,870,1123,1174]" gridcol="2" gridrow="11" pageId="29" pageNumber="90">3</td>
<td id="F9155DFDBA520056FC509027FBF84BB3" box="[907,1138,1123,1174]" gridcol="3" gridrow="11" pageId="29" pageNumber="90">2</td>
<td id="F9155DFDBA520056FB529027FADC4BB3" box="[1161,1366,1123,1174]" gridcol="4" gridrow="11" pageId="29" pageNumber="90">3</td>
</tr>
<tr id="BAC43481BA520056FF1390E3FADC4B98" box="[200,1366,1191,1213]" gridrow="12" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF1390E3FE114B98" box="[200,411,1191,1213]" gridcol="0" gridrow="12" pageId="29" pageNumber="90">Carpus of P1</th>
<td id="F9155DFDBA520056FE6890E3FD0A4B98" box="[435,640,1191,1213]" gridcol="1" gridrow="12" pageId="29" pageNumber="90">Well produced</td>
<td id="F9155DFDBA520056FD4490E3FCEC4B98" box="[671,870,1191,1213]" gridcol="2" gridrow="12" pageId="29" pageNumber="90">Slightly produced</td>
<td id="F9155DFDBA520056FC5090E3FBF84B98" box="[907,1138,1191,1213]" gridcol="3" gridrow="12" pageId="29" pageNumber="90">Well produced</td>
<td id="F9155DFDBA520056FB5290E3FADC4B98" box="[1161,1366,1191,1213]" gridcol="4" gridrow="12" pageId="29" pageNumber="90">Well produced</td>
</tr>
<tr id="BAC43481BA520056FF1390AFFADC4A3B" box="[200,1366,1259,1310]" gridrow="13" pageId="29" pageNumber="90">
<th id="F9155DFDBA520056FF1390AFFE114A3B" box="[200,411,1259,1310]" gridcol="0" gridrow="13" pageId="29" pageNumber="90">Palm spines of P1</th>
<td id="F9155DFDBA520056FE6890AFFD0A4A3B" box="[435,640,1259,1310]" gridcol="1" gridrow="13" pageId="29" pageNumber="90">2 distal</td>
<td id="F9155DFDBA520056FD4490AFFCEC4A3B" box="[671,870,1259,1310]" gridcol="2" gridrow="13" pageId="29" pageNumber="90">1 large proximal and 1 small distal</td>
<td id="F9155DFDBA520056FC5090AFFBF84A3B" box="[907,1138,1259,1310]" gridcol="3" gridrow="13" pageId="29" pageNumber="90">1 small medial</td>
<td id="F9155DFDBA520056FB5290AFFADC4A3B" box="[1161,1366,1259,1310]" gridcol="4" gridrow="13" pageId="29" pageNumber="90">1 large proximal</td>
</tr>
</table>
</paragraph>
<subSubSection id="4CEE6548BA52FFB7FF669123FDE64D84" lastPageId="30" lastPageNumber="91" pageId="29" pageNumber="90" type="discussion">
<paragraph id="044B36C3BA52FFB4FF669123FEA54AA4" blockId="29.[189,303,1383,1409]" box="[189,303,1383,1409]" pageId="29" pageNumber="90">
<heading id="5F0381AFBA52FFB4FF669123FEA54AA4" bold="true" box="[189,303,1383,1409]" fontSize="11" level="3" pageId="29" pageNumber="90" reason="3">
<emphasis id="3680EAD1BA52FFB4FF669123FEA54AA4" bold="true" box="[189,303,1383,1409]" pageId="29" pageNumber="90">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="044B36C3BA52FFB4FF6691D2FEFA4945" blockId="29.[189,1399,1430,1632]" pageId="29" pageNumber="90">
<taxonomicName id="C3F44D40BA52FFB4FF6691D2FE694A95" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[189,483,1430,1456]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA52FFB4FF6691D2FE694A95" box="[189,483,1430,1456]" italics="true" pageId="29" pageNumber="90">Skuphonura oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA52FFB4FE3091D2FDC24A95" box="[491,584,1430,1456]" pageId="29" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
, and three other species of genus: 
<taxonomicName id="C3F44D40BA52FFB4FC0791D2FAF84A95" authority="Brusca &amp; Muller, 1991" authorityName="Brusca &amp; Muller" authorityYear="1991" box="[988,1394,1430,1456]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="3680EAD1BA52FFB4FC0791D2FBD24A95" box="[988,1112,1430,1456]" italics="true" pageId="29" pageNumber="90">S. kensleyi</emphasis>
Brusca &amp; Müller, 1991
</taxonomicName>
, 
<taxonomicName id="C3F44D40BA52FFB4FF6691FDFE6B4AF6" authority="Kensley, 1980" authorityName="Kensley" authorityYear="1980" box="[189,481,1465,1491]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="itapuca">
<emphasis id="3680EAD1BA52FFB4FF6691FDFEB84AF6" box="[189,306,1465,1491]" italics="true" pageId="29" pageNumber="90">S. itapuca</emphasis>
Kensley, 1980
</taxonomicName>
and 
<taxonomicName id="C3F44D40BA52FFB4FDC191FDFC6B4AF6" authority="Menzies &amp; Kruczynski, 1983" authorityName="Menzies &amp; Kruczynski" authorityYear="1983" box="[538,993,1465,1491]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="lindae">
<emphasis id="3680EAD1BA52FFB4FDC191FDFD0A4AF6" box="[538,640,1465,1491]" italics="true" pageId="29" pageNumber="90">S. lindae</emphasis>
Menzies &amp; Kruczynski, 1983
</taxonomicName>
(
<tableCitation id="49760378BA52FFB4FC2B91FDFBC14AF6" box="[1008,1099,1465,1491]" captionStart="Table 2" captionStartId="29.[189,252,265,289]" captionTargetBox="[200,1366,348,1310]" captionTargetPageId="29" captionText="Table 2. Comparison between Skuphonura oaxaquensis sp. nov. and the closest species of genera." httpUri="http://table.plazi.org/id/508B664BBA52FFB4FF66954DFB594E04" pageId="29" pageNumber="90" tableUuid="508B664BBA52FFB4FF66954DFB594E04">Table 2</tableCitation>
), share the following two characters: anterior margin of the cephalon as wide as the posterior margin and the anterolateral lobes are sub-equal to the rostral process, while in 
<taxonomicName id="C3F44D40BA52FFB4FD049244FBC4493C" authority="Kensley, 1980" authorityName="Kensley" authorityYear="1980" box="[735,1102,1535,1562]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="3680EAD1BA52FFB4FD049244FC13493C" box="[735,921,1535,1562]" italics="true" pageId="29" pageNumber="90">S. ecuadorensis</emphasis>
Kensley, 1980
</taxonomicName>
and 
<taxonomicName id="C3F44D40BA52FFB4FB509244FF704918" authority="Barnard, 1925" authorityName="Barnard" authorityYear="1925" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="3680EAD1BA52FFB4FB509244FA8C493C" box="[1163,1286,1535,1562]" italics="true" pageId="29" pageNumber="90">S. laticeps</emphasis>
Barnard, 1925
</taxonomicName>
, the anterior margin of the cephalon is wider than the posterior and the lobes are longer than the rostral process.
</paragraph>
<paragraph id="044B36C3BA52FFB4FF6692C3FEC048FB" blockId="29.[188,1399,1671,2014]" pageId="29" pageNumber="90">
<taxonomicName id="C3F44D40BA52FFB4FF6692C3FE384984" authorityName="Kensley" authorityYear="1980" box="[189,434,1671,1697]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="itapuca">
<emphasis id="3680EAD1BA52FFB4FF6692C3FE384984" box="[189,434,1671,1697]" italics="true" pageId="29" pageNumber="90">Skuphonura itapuca</emphasis>
</taxonomicName>
, described from Nichteroy, 
<collectingRegion id="C630F821BA52FFB4FCC192C3FC5C4984" box="[794,982,1671,1697]" country="Brazil" name="Rio de Janeiro" pageId="29" pageNumber="90">Rio de Janeiro</collectingRegion>
, 
<collectingCountry id="7CE37653BA52FFB4FC3292C3FBBE4984" box="[1001,1076,1671,1697]" name="Brazil" pageId="29" pageNumber="90">Brazil</collectingCountry>
, is the closest species to 
<taxonomicName id="C3F44D40BA52FFB4FF6692EFFEFA49E1" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[189,368,1707,1733]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA52FFB4FF6692EFFEFA49E1" box="[189,368,1707,1733]" italics="true" pageId="29" pageNumber="90">S. oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA52FFB4FEA792EFFE5649E0" box="[380,476,1707,1733]" pageId="29" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
by the following characteristics: (1) absence of a mid-ventral spine in the pereonite 1, (2) mid-ventral region of cephalon with two processes, (3) second article of mandibular palp with three long setae. However, in 
<taxonomicName id="C3F44D40BA52FFB4FD5E92B5FCB9482E" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[645,819,1777,1803]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA52FFB4FD5E92B5FCB9482E" box="[645,819,1777,1803]" italics="true" pageId="29" pageNumber="90">S. oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA52FFB4FCE292B5FC18482E" box="[825,914,1777,1803]" pageId="29" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
the body is partially pigmented, cephalon slightly wider medially, flagellum of antennule with three articles, distal article with three aesthetascs, flagellum of antenna with four articles, and carpus of pereopod 1 is externally projected and palm with one proximal tooth; while, 
<taxonomicName id="C3F44D40BA52FFB4FD98931FFD314851" authorityName="Kensley" authorityYear="1980" box="[579,699,1883,1909]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="29" pageNumber="90" phylum="Arthropoda" rank="species" species="itapuca">
<emphasis id="3680EAD1BA52FFB4FD98931FFD314851" box="[579,699,1883,1909]" italics="true" pageId="29" pageNumber="90">S. itapuca</emphasis>
</taxonomicName>
lacks pigment, anterior margin of the cephalon is slightly wider than the posterior, flagellum of antennule with two article and two distal aesthetascs, flagellum of antenna with one article, carpus of pereopod 1 projected apically and palm with one proximal and one distal spine.
</paragraph>
<paragraph id="044B36C3BA51FFB7FF66954DFB3F4ED3" blockId="30.[186,1399,265,503]" pageId="30" pageNumber="91">
<taxonomicName id="C3F44D40BA51FFB7FF66954DFE6A4E06" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[189,480,265,291]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="30" pageNumber="91" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA51FFB7FF66954DFE6A4E06" box="[189,480,265,291]" italics="true" pageId="30" pageNumber="91">Skuphonura oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA51FFB7FE3E954EFDB74E01" box="[485,573,266,292]" pageId="30" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="C3F44D40BA51FFB7FDB5954DFD6D4E06" authorityName="Brusca &amp; Muller" authorityYear="1991" box="[622,743,265,291]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="30" pageNumber="91" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="3680EAD1BA51FFB7FDB5954DFD6D4E06" box="[622,743,265,291]" italics="true" pageId="30" pageNumber="91">S. kensleyi</emphasis>
</taxonomicName>
, described from Santa Marta, Punta de Betín, Colombian Caribbean, are similar by the following characteristics: (1) pigmentation pattern, (2) cephalon slightly wider medially, (3) antennular flagellum, (4) carpus of pereopod 1. However, in 
<taxonomicName id="C3F44D40BA51FFB7FB8E9514FB454E4C" authorityName="Brusca &amp; Muller" authorityYear="1991" box="[1109,1231,335,362]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="30" pageNumber="91" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="3680EAD1BA51FFB7FB8E9514FB454E4C" box="[1109,1231,335,362]" italics="true" pageId="30" pageNumber="91">S. kensleyi</emphasis>
</taxonomicName>
the pereonites and pleon have reticulated pigmentation, antennular flagellum with one aesthetasc, antennal flagellum with one article, the second article of the mandibular palp with two distal setae and three proximal setae, lacinia mobilis with six denticles, and palm of the pereopod 1 has a medial tooth; while, in 
<taxonomicName id="C3F44D40BA51FFB7FF669599FEE14ED3" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[189,363,477,503]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="30" pageNumber="91" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA51FFB7FF669599FEE14ED3" box="[189,363,477,503]" italics="true" pageId="30" pageNumber="91">S. oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA51FFB7FEA89599FE474ED2" box="[371,461,477,503]" pageId="30" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
the reticulated pigmentation is partial in pereonite 4 and pleon.
</paragraph>
<paragraph id="044B36C3BA51FFB7FF66965AFDE64D84" blockId="30.[188,1399,542,674]" pageId="30" pageNumber="91">
<taxonomicName id="C3F44D40BA51FFB7FF66965AFE674D12" authority="Jarquín-Martínez &amp; García-Madrigal, 2021" authorityName="Jarquín-Martínez &amp; García-Madrigal" authorityYear="2021" box="[189,493,542,568]" class="Malacostraca" family="Anthuridae" genus="Skuphonura" kingdom="Animalia" order="Isopoda" pageId="30" pageNumber="91" phylum="Arthropoda" rank="species" species="oaxaquensis" status="sp. nov.">
<emphasis id="3680EAD1BA51FFB7FF66965AFE674D12" box="[189,493,542,568]" italics="true" pageId="30" pageNumber="91">Skuphonura oaxaquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DB357AABA51FFB7FE24965AFDEE4D1D" box="[511,612,542,568]" pageId="30" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
differs from the preceding species by the following unique characteristics: (1) body pigmentation, (2) last article of mandibular palp with four setae, (3) lacinia mobilis with 11 denticles, (4) inner margin of the second article of maxillipedal palp sinus, and (5) palm of pereopod 1 with a proximal tooth.
</paragraph>
</subSubSection>
</treatment>
</document>