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<document id="3A9803039AB751890407D73C6688B1C9" ID-DOI="10.11646/zootaxa.5437.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10984997" ID-ZooBank="79196FA8-AC75-4768-AC43-9C9C1B515FB9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1713344825674" checkinUser="plazi" docAuthor="Lopes, Rogério B. &amp; Noll, Fernando B." docDate="2024" docId="5C338787FE704B0E82FD0EA8FAA7C0ED" docLanguage="en" docName="zootaxa.5437.3.3.pdf" docOrigin="Zootaxa 5437 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5437.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zethus (Zethoides) schadei Bohart &amp; Stange 1965" docType="treatment" docVersion="3" lastPageNumber="378" masterDocId="A00AFFFFFE7F4B01826A0F56FFBEC354" masterDocTitle="Taxonomic notes on the Zethus olmecus de Saussure, 1875 species group (Hymenoptera, Vespidae, Eumeninae), with the description of five new species" masterLastPageNumber="383" masterPageNumber="363" pageNumber="378" updateTime="1713815074103" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="B682C8AC1B3B172617C712C43121019C">Taxonomic notes on the Zethus olmecus de Saussure, 1875 species group (Hymenoptera, Vespidae, Eumeninae), with the description of five new species</mods:title>
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<mods:namePart id="C1FA06401BFD8A41F46882B7EEB1A38A">Lopes, Rogério B.</mods:namePart>
<mods:nameIdentifier id="363D1633F612F1C43804645C26EBFE60" type="ORCID">0000-0002-1217-1152</mods:nameIdentifier>
<mods:affiliation id="4B585E7FF6A0C0E531EC66B693FF691E">Universidade Estadual Paulista “ Júlio de Mesquita Filho ”, Instituto de Biociências, Letras e Ciências Exatas, Departamento de Ciências Biológicas</mods:affiliation>
<mods:nameIdentifier id="451DD51065DE66F790D9D9E94B562212" type="email">rbotlopes@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5CF4BA1B90BD0BC886078FFFFC828487">Noll, Fernando B.</mods:namePart>
<mods:affiliation id="8F1C613CAE87A4183ECFDE2C9FDD7709">Universidade Estadual Paulista “ Júlio de Mesquita Filho ”, Instituto de Biociências, Letras e Ciências Exatas, Departamento de Ciências Biológicas</mods:affiliation>
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<mods:title id="A483745282B4F37BCAFD2DDF02178E09">Zootaxa</mods:title>
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<mods:part id="9B8DB113397B1755E634DAA47C86C721">
<mods:date id="372F7904896B5E68BAFB9402274B3508">2024</mods:date>
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<mods:number id="9BDD700F530D70C289F46375E394F17A">2024-04-15</mods:number>
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<treatment id="5C338787FE704B0E82FD0EA8FAA7C0ED" LSID="urn:lsid:plazi:treatment:5C338787FE704B0E82FD0EA8FAA7C0ED" httpUri="http://treatment.plazi.org/id/5C338787FE704B0E82FD0EA8FAA7C0ED" lastPageNumber="378" pageId="15" pageNumber="378">
<subSubSection id="9C80651AFE704B0E82FD0EA8FCBCC14C" box="[151,770,510,537]" pageId="15" pageNumber="378" type="nomenclature">
<paragraph id="D4253691FE704B0E82FD0EA8FCBCC14C" blockId="15.[151,770,510,537]" box="[151,770,510,537]" pageId="15" pageNumber="378">
<heading id="8F6D81FDFE704B0E82FD0EA8FCBCC14C" bold="true" box="[151,770,510,537]" fontSize="11" level="1" pageId="15" pageNumber="378" reason="1">
<taxonomicName id="139A4D12FE704B0E82FD0EA8FCBCC14C" ID-CoL="5D3TH" authority="Bohart &amp; Stange, 1965" authorityName="Bohart &amp; Stange" authorityYear="1965" box="[151,770,510,537]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="378" phylum="Arthropoda" rank="species" species="schadei" subGenus="Zethoides">
<emphasis id="E6EEEA83FE704B0E82FD0EA8FCBCC14C" bold="true" box="[151,770,510,537]" pageId="15" pageNumber="378">
<emphasis id="E6EEEA83FE704B0E82FD0EA8FF54C14D" bold="true" box="[151,234,510,537]" italics="true" pageId="15" pageNumber="378">Zethus</emphasis>
(
<emphasis id="E6EEEA83FE704B0E82910EA8FECEC14D" bold="true" box="[251,368,510,537]" italics="true" pageId="15" pageNumber="378">Zethoides</emphasis>
)
<emphasis id="E6EEEA83FE704B0E83E80EA8FE63C14D" bold="true" box="[386,477,510,537]" italics="true" pageId="15" pageNumber="378">schadei</emphasis>
<bibRefCitation id="B00B4B60FE704B0E838E0EA9FCBCC14C" author="Bohart, R. M. &amp; Stange, L. A." box="[484,770,510,537]" pageId="15" pageNumber="378" pagination="1 - 208" refId="ref14458" refString="Bohart, R. M. &amp; Stange, L. A. (1965) A Revision of the genus Zethus Fabricius in the Western Hemisphere (Hymenoptera: Eumenidae). University of California Publications in Entomology, 40, 1 - 208." type="journal article" year="1965">Bohart &amp; Stange, 1965</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9C80651AFE704B0E82FD0D15FC0BC1CD" pageId="15" pageNumber="378" type="reference_group">
<paragraph id="D4253691FE704B0E82FD0D15FC0BC1CD" blockId="15.[151,1436,579,665]" pageId="15" pageNumber="378">
<treatmentCitationGroup id="F48A11BFFE704B0E82FD0D15FC0BC1CD" pageId="15" pageNumber="378">
<treatmentCitation id="553B1080FE704B0E82FD0D15FDF3C10F" author="Bohart, R. M. &amp; Stange, L. A." box="[151,589,579,603]" page="14" pageId="15" pageNumber="378" year="1965">
<taxonomicName id="139A4D12FE704B0E82FD0D15FDF3C10F" ID-CoL="5D3TH" authority="Bohart &amp; Stange, 1965: 14" authorityName="Bohart &amp; Stange" authorityPageNumber="14" authorityYear="1965" box="[151,589,579,603]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="378" phylum="Arthropoda" rank="species" species="schadei">
<emphasis id="E6EEEA83FE704B0E82FD0D15FE91C10F" box="[151,303,579,603]" italics="true" pageId="15" pageNumber="378">Zethus schadei</emphasis>
<bibRefCitation id="B00B4B60FE704B0E83520D15FDF3C10F" author="Bohart, R. M. &amp; Stange, L. A." box="[312,589,579,603]" pageId="15" pageNumber="378" pagination="1 - 208" refId="ref14458" refString="Bohart, R. M. &amp; Stange, L. A. (1965) A Revision of the genus Zethus Fabricius in the Western Hemisphere (Hymenoptera: Eumenidae). University of California Publications in Entomology, 40, 1 - 208." type="journal article" year="1965">Bohart &amp; Stange, 1965: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(nest, figs 4B, C), 17 (nest), 159 (key), 168 (fig. 156–158), 169 (male, female; in subgenus
<taxonomicName id="139A4D12FE704B0E83470D34FE2EC12E" box="[301,400,610,634]" pageId="15" pageNumber="378" rank="subGenus" subGenus="Zethoides">
<emphasis id="E6EEEA83FE704B0E83470D34FE2EC12E" box="[301,400,610,634]" italics="true" pageId="15" pageNumber="378">Zethoides</emphasis>
</taxonomicName>
;
<collectingCountry id="AC8D7601FE704B0E83F40D34FE41C12E" box="[414,511,610,634]" name="Paraguay" pageId="15" pageNumber="378">Paraguay</collectingCountry>
,
<collectingCountry id="AC8D7601FE704B0E80600D34FD83C12E" box="[522,573,610,634]" name="Peru" pageId="15" pageNumber="378">Peru</collectingCountry>
).
<treatmentCitation id="553B1080FE704B0E80250D34FC30C12E" author="Rasmussen, C. &amp; Asenjo, A." box="[591,910,610,634]" page="43" pageId="15" pageNumber="378" year="2009">
<bibRefCitation id="B00B4B60FE704B0E80250D34FC30C12E" author="Rasmussen, C. &amp; Asenjo, A." box="[591,910,610,634]" pageId="15" pageNumber="378" pagination="1 - 78" refId="ref15654" refString="Rasmussen, C. &amp; Asenjo, A. (2009) A checklist to the wasps of Peru (Hymenoptera, Aculeata). ZooKeys, 15, 1 - 78. https: // doi. org / 10.3897 / zookeys. 15.196" type="journal article" year="2009">Rasmussen &amp; Asenjo 2009: 43</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="AC8D7601FE704B0E81F70D34FC6FC12E" box="[925,977,610,634]" name="Peru" pageId="15" pageNumber="378">Peru</collectingCountry>
).
<treatmentCitation id="553B1080FE704B0E81890D34FAA2C12E" author="Pulgarin-Diaz, J. A. &amp; Cano Gallego, L. E. &amp; Herrera-Florez, A. F. &amp; Quiroz Gamboa, J. A." box="[995,1308,610,634]" page="1" pageId="15" pageNumber="378" year="2018">
<bibRefCitation id="B00B4B60FE704B0E81890D34FAA2C12E" author="Pulgarin-Diaz, J. A. &amp; Cano Gallego, L. E. &amp; Herrera-Florez, A. F. &amp; Quiroz Gamboa, J. A." box="[995,1308,610,634]" pageId="15" pageNumber="378" pagination="1 - 3" refId="ref15572" refString="Pulgarin-Diaz, J. A., Cano Gallego, L. E., Herrera-Florez, A. F. &amp; Quiroz Gamboa, J. A. (2018) First report of Zethus schadei (Hymenoptera: Vespidae) as natural enemy of Hypsipyla grandella Zeller (Lepidoptera: Pyralidae) from Colombia. Acta Zoologica Mexicana, Nueva Serie, 34, 1 - 3. https: // doi. org / 10.21829 / azm. 2018.3412119" type="journal article" year="2018">
Pulgarín-Díaz
<emphasis id="E6EEEA83FE704B0E86120D35FB13C12E" box="[1144,1197,610,634]" italics="true" pageId="15" pageNumber="378">et al.</emphasis>
2018: 1–3
</bibRefCitation>
</treatmentCitation>
(predator of pyralid caterpillar
<taxonomicName id="139A4D12FE704B0E83E80DD6FDECC1CC" ID-CoL="3P485" authorityName="Zeller" authorityYear="1848" box="[386,594,640,664]" class="Insecta" family="Pyralidae" genus="Hypsipyla" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="378" phylum="Arthropoda" rank="species" species="grandella">
<emphasis id="E6EEEA83FE704B0E83E80DD6FDECC1CC" box="[386,594,640,664]" italics="true" pageId="15" pageNumber="378">Hypsipyla grandella</emphasis>
</taxonomicName>
; misidentification, not
<taxonomicName id="139A4D12FE704B0E81540DD7FC18C1CC" ID-CoL="5D3TH" authorityName="Bohart &amp; Stange" authorityYear="1965" box="[830,934,640,664]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="378" phylum="Arthropoda" rank="species" species="schadei">
<emphasis id="E6EEEA83FE704B0E81540DD7FC18C1CC" box="[830,934,640,664]" italics="true" pageId="15" pageNumber="378">Z. schadei</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9C80651AFE704B0E82FD0D90FD31C07D" pageId="15" pageNumber="378" type="materials_examined">
<paragraph id="D4253691FE704B0E82FD0D90FD31C07D" blockId="15.[151,1437,710,953]" pageId="15" pageNumber="378">
<emphasis id="E6EEEA83FE704B0E82FD0D90FEC0C1B4" bold="true" box="[151,382,710,736]" pageId="15" pageNumber="378">Examined material</emphasis>
.
<materialsCitation id="64F23CCCFE704B0E83E40D91FC11C1B5" box="[398,943,711,739]" country="Peru" location="PERU" pageId="15" pageNumber="378" specimenCount="7" specimenCount-female="7">
<specimenCount id="C29CFD18FE704B0E83E40D91FE0CC1B5" box="[398,434,711,737]" count="7" pageId="15" pageNumber="378" type="female">7♀</specimenCount>
, 1♁ (see Supplementary Material 1).
<collectingCountry id="AC8D7601FE704B0E81090D91FC11C1B5" box="[867,943,711,737]" name="Peru" pageId="15" pageNumber="378">PERU</collectingCountry>
</materialsCitation>
:
<materialsCitation id="64F23CCCFE704B0E81D60D91FF54C051" collectionCode="BME" country="Peru" location="Valle Chanchamayo" pageId="15" pageNumber="378" specimenCount="1" specimenCount-female="1" stateProvince="Junin" typeStatus="paratype">
<specimenCount id="C29CFD18FE704B0E81D60D91FC60C1B5" box="[956,990,711,737]" count="1" pageId="15" pageNumber="378" type="female">1♀</specimenCount>
(
<typeStatus id="0B218833FE704B0E81850D91FBEBC1B5" box="[1007,1109,711,737]" pageId="15" pageNumber="378" type="paratype">paratype</typeStatus>
),
<collectingRegion id="165EF873FE704B0E86000D91FB17C1B5" box="[1130,1193,711,737]" country="Peru" name="Junin" pageId="15" pageNumber="378">Junín</collectingRegion>
,
<location id="D145604AFE704B0E86DD0D91FA22C1B5" LSID="urn:lsid:plazi:treatment:5C338787FE704B0E82FD0EA8FAA7C0ED:D145604AFE704B0E86DD0D91FA22C1B5" box="[1207,1436,711,737]" country="Peru" name="Valle Chanchamayo" pageId="15" pageNumber="378" stateProvince="Junin">Valle Chanchamayo</location>
[
<collectionCode id="B28BAE54FE704B0E82F40DBDFF5DC051" box="[158,227,747,773]" pageId="15" pageNumber="378">BME</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="64F23CCCFE704B0E829C0DBDFB53C051" box="[246,1261,747,775]" collectionCode="DZUP" country="Brazil" county="Amazonas" location="Caceres" municipality="Rio Puaquequara" pageId="15" pageNumber="378" specimenCount="4" specimenCount-female="4" stateProvince="Mato Grosso" typeStatus="paratype">
<collectingCountry id="AC8D7601FE704B0E829C0DBDFEDEC051" box="[246,352,747,773]" name="Brazil" pageId="15" pageNumber="378">BRAZIL</collectingCountry>
:
<specimenCount id="C29CFD18FE704B0E83010DBDFE2EC051" box="[363,400,747,773]" count="4" pageId="15" pageNumber="378" type="female">4♀</specimenCount>
,
<collectingCounty id="3D444E1DFE704B0E83F70DBDFDA8C051" box="[413,534,747,773]" pageId="15" pageNumber="378">Amazonas</collectingCounty>
,
<collectingMunicipality id="3441ACEBFE704B0E80490DBDFD5BC051" box="[547,741,747,773]" pageId="15" pageNumber="378">Rio Puaquequara</collectingMunicipality>
[
<collectionCode id="B28BAE54FE704B0E80990DBDFCFBC051" box="[755,837,747,773]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="378" type="University or college">DZUP</collectionCode>
]; 1♁,
<collectingRegion id="165EF873FE704B0E81E10DBDFB9EC051" box="[907,1056,747,773]" country="Brazil" name="Mato Grosso" pageId="15" pageNumber="378">Mato Grosso</collectingRegion>
,
<location id="D145604AFE704B0E86470DBDFB38C051" LSID="urn:lsid:plazi:treatment:5C338787FE704B0E82FD0EA8FAA7C0ED:D145604AFE704B0E86470DBDFB38C051" box="[1069,1158,747,773]" country="Brazil" county="Amazonas" municipality="Rio Puaquequara" name="Caceres" pageId="15" pageNumber="378" stateProvince="Mato Grosso">Cáceres</location>
[
<collectionCode id="B28BAE54FE704B0E86FE0DBDFB58C051" box="[1172,1254,747,773]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="378" type="University or college">DZUP</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="64F23CCCFE704B0E86930DBDFD35C07D" collectionCode="DZUP" country="Paraguay" location="Estancia San Luis" pageId="15" pageNumber="378" specimenCount="2" specimenCount-female="2" stateProvince="Concepcion" typeStatus="paratype">
<collectingCountry id="AC8D7601FE704B0E86930DBDFA27C051" box="[1273,1433,747,773]" name="Paraguay" pageId="15" pageNumber="378">PARAGUAY</collectingCountry>
:
<specimenCount id="C29CFD18FE704B0E82FD0C59FF02C07D" box="[151,188,783,809]" count="2" pageId="15" pageNumber="378" type="female">2♀</specimenCount>
,
<collectingRegion id="165EF873FE704B0E82A20C59FEEFC07D" box="[200,337,783,809]" country="Paraguay" name="Concepcion" pageId="15" pageNumber="378">Concepción</collectingRegion>
,
<location id="D145604AFE704B0E83360C59FD9BC07D" LSID="urn:lsid:plazi:treatment:5C338787FE704B0E82FD0EA8FAA7C0ED:D145604AFE704B0E83360C59FD9BC07D" box="[348,549,783,809]" country="Paraguay" name="Estancia San Luis" pageId="15" pageNumber="378" stateProvince="Concepcion">Estancia San Luis</location>
[
<collectionCode id="B28BAE54FE704B0E80580C59FD3AC07D" box="[562,644,783,809]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="15" pageNumber="378" type="University or college">DZUP</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9C80651AFE704B0E82AD0C64FAA7C0ED" pageId="15" pageNumber="378" type="distribution">
<paragraph id="D4253691FE704B0E82AD0C64FADEC019" blockId="15.[151,1437,710,953]" box="[199,1376,818,845]" pageId="15" pageNumber="378">
<emphasis id="E6EEEA83FE704B0E82AD0C64FEE7C018" bold="true" box="[199,345,818,844]" pageId="15" pageNumber="378">Distribution</emphasis>
.
<collectingCountry id="AC8D7601FE704B0E830D0C65FE23C019" box="[359,413,819,845]" name="Peru" pageId="15" pageNumber="378">Peru</collectingCountry>
, *
<collectingCountry id="AC8D7601FE704B0E83DE0C65FE45C019" box="[436,507,819,845]" name="Brazil" pageId="15" pageNumber="378">Brazil</collectingCountry>
(Amazonas,
<collectingRegion id="165EF873FE704B0E80FA0C65FC98C019" box="[656,806,819,845]" country="Brazil" name="Mato Grosso" pageId="15" pageNumber="378">Mato Grosso</collectingRegion>
),
<collectingCountry id="AC8D7601FE704B0E81520C65FC1EC019" box="[824,928,819,845]" name="Paraguay" pageId="15" pageNumber="378">Paraguay</collectingCountry>
(
<bibRefCitation id="B00B4B60FE704B0E81C50C65FB11C019" author="Bohart, R. M. &amp; Stange, L. A." box="[943,1199,818,845]" pageId="15" pageNumber="378" pagination="1 - 208" refId="ref14458" refString="Bohart, R. M. &amp; Stange, L. A. (1965) A Revision of the genus Zethus Fabricius in the Western Hemisphere (Hymenoptera: Eumenidae). University of California Publications in Entomology, 40, 1 - 208." type="journal article" year="1965">Bohart &amp; Stange 1965</bibRefCitation>
; and new data).
</paragraph>
<paragraph id="D4253691FE704B0E82AD0C00FAA7C0ED" blockId="15.[151,1437,710,953]" pageId="15" pageNumber="378">
<emphasis id="E6EEEA83FE704B0E82AD0C00FED8C024" bold="true" box="[199,358,854,880]" pageId="15" pageNumber="378">Observations</emphasis>
. Díaz
<emphasis id="E6EEEA83FE704B0E83D20C01FE4AC025" box="[440,500,855,881]" italics="true" pageId="15" pageNumber="378">et al.</emphasis>
(2018) reports
<taxonomicName id="139A4D12FE704B0E80C70C01FC9BC025" authorityName="Bohart &amp; Stange" authorityYear="1965" box="[685,805,855,881]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="378" phylum="Arthropoda" rank="species" species="schadei">
<emphasis id="E6EEEA83FE704B0E80C70C01FC9BC025" box="[685,805,855,881]" italics="true" pageId="15" pageNumber="378">Z. schadei</emphasis>
</taxonomicName>
as a natural enemy of a pyralid caterpillar. However, the image presented in the article certainly does not represent this wasp species. Identity cannot be certified, but apparently it belongs to the
<taxonomicName id="139A4D12FE704B0E83A50CC9FDF3C0ED" authorityName="Spinola" authorityYear="1841" box="[463,589,927,953]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="378" phylum="Arthropoda" rank="species" species="biglumis">
<emphasis id="E6EEEA83FE704B0E83A50CC9FDF3C0ED" box="[463,589,927,953]" italics="true" pageId="15" pageNumber="378">Z. biglumis</emphasis>
</taxonomicName>
species group and perhaps it is
<taxonomicName id="139A4D12FE704B0E81D80CC9FB6DC0ED" authority="Spinola, 1841" authorityName="Spinola" authorityYear="1841" box="[946,1235,927,953]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="378" phylum="Arthropoda" rank="species" species="biglumis">
<emphasis id="E6EEEA83FE704B0E81D80CC9FB8EC0ED" box="[946,1072,927,953]" italics="true" pageId="15" pageNumber="378">Z. biglumis</emphasis>
Spinola, 1841
</taxonomicName>
itself.
</paragraph>
</subSubSection>
</treatment>
</document>
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