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<document id="85A22D422334EC2F203B8855586B18FF" ID-DOI="10.11646/zootaxa.3682.1.2" ID-GBIF-Dataset="eaf497f5-d458-4e40-850d-40cb21f163e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="284116" ID-ZooBank="BCEA9C83-9664-4A40-9BC2-A7D56BB134B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460641719393" checkinUser="plazi" docAuthor="Marshall, S. A." docDate="2013" docId="03F78361FFA9FFB9FF44137DFB1DFF59" docLanguage="en" docName="zt03682p084.pdf" docOrigin="Zootaxa 3682 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Grallipeza marleyi Marshall, 2013, new species" docType="treatment" docVersion="6" lastPageNumber="64" masterDocId="FFCEFB19FFBBFFADFFD31719FFF5FFAE" masterDocTitle="Grallipeza Rondani (Diptera: Micropezidae: Taeniapterinae) of the Caribbean and North America" masterLastPageNumber="84" masterPageNumber="45" pageNumber="63" updateTime="1698335989015" updateUser="plazi">
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<mods:title id="13882C9F5B409C4B9779A3F67C89C168">Grallipeza Rondani (Diptera: Micropezidae: Taeniapterinae) of the Caribbean and North America</mods:title>
</mods:titleInfo>
<mods:name id="34CB99639E6FA9C4C7EC4FB8320B410D" type="personal">
<mods:role id="075D8A5DCB0E6F0D913CACE735C6D0A4">
<mods:roleTerm id="1C7C3BEE88B20FD0B33AE501259796C2">Author</mods:roleTerm>
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<mods:namePart id="F2C6EC516728A98F2DDB9166287A8F6A">Marshall, S. A.</mods:namePart>
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<mods:title id="51F1E0FD62D24B20D5D7624CE83014AA">Zootaxa</mods:title>
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<mods:part id="706F2B581DFA9B584E736E7962A06EC7">
<mods:date id="18D84D1E67082BE5996F5B7CD692FB85">2013</mods:date>
<mods:detail id="44876E12330577298E6AAD38F8F58716" type="volume">
<mods:number id="2DFA25E6DF7D088864F328167F9D1862">3682</mods:number>
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<mods:classification id="85077E377AE7E090004EA799B1583762">journal article</mods:classification>
<mods:identifier id="3A1BA2F79B3812F1002E67265F47EAA5" type="DOI">10.11646/zootaxa.3682.1.2</mods:identifier>
<mods:identifier id="23DEB1AEAB1F796F646D2B5326AC341F" type="GBIF-Dataset">eaf497f5-d458-4e40-850d-40cb21f163e2</mods:identifier>
<mods:identifier id="7C97A603247ED58AA42C12288143FEE6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F78361FFA9FFB9FF44137DFB1DFF59" ID-DOI="http://doi.org/10.5281/zenodo.6154150" ID-GBIF-Taxon="127664683" ID-Zenodo-Dep="6154150" LSID="urn:lsid:plazi:treatment:03F78361FFA9FFB9FF44137DFB1DFF59" httpUri="http://treatment.plazi.org/id/03F78361FFA9FFB9FF44137DFB1DFF59" lastPageId="20" lastPageNumber="64" pageId="18" pageNumber="63">
<subSubSection id="C34461FCFFA9FFBFFF44137DFEEEFB30" pageId="18" pageNumber="63" type="nomenclature">
<paragraph id="8BE13277FFA9FFBFFF44137DFDFBFBD3" blockId="18.[151,526,1123,1182]" box="[151,526,1123,1150]" pageId="18" pageNumber="63">
<heading id="D0A9851BFFA9FFBFFF44137DFDFBFBD3" bold="true" box="[151,526,1123,1150]" fontSize="11" level="1" pageId="18" pageNumber="63" reason="1">
<emphasis id="B92AEE65FFA9FFBFFF44137DFDFBFBD3" bold="true" box="[151,526,1123,1150]" pageId="18" pageNumber="63">
<taxonomicName id="4C5E49F4FFA9FFBFFF44137DFE82FBD0" ID-CoL="9HYPR" box="[151,375,1124,1150]" class="Insecta" family="Micropezidae" genus="Grallipeza" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="63" phylum="Arthropoda" rank="species" species="marleyi" status="sp. nov.">
<emphasis id="B92AEE65FFA9FFBFFF44137DFE82FBD0" bold="true" box="[151,375,1124,1150]" italics="true" pageId="18" pageNumber="63">Grallipeza marleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A219531EFFA9FFBFFEAD137AFDFBFBD3" box="[382,526,1123,1149]" pageId="18" pageNumber="63" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE13277FFA9FFBFFF44139FFEEEFB30" blockId="18.[151,526,1123,1182]" box="[151,283,1158,1182]" pageId="18" pageNumber="63">
<figureCitation id="13652EF2FFA9FFBFFF44139FFEEEFB30" box="[151,283,1158,1182]" captionStart="FIGURES 28 – 31" captionStartId="19.[151,264,1601,1624]" captionTargetBox="[151,1434,193,1579]" captionTargetId="figure@19.[151,1436,193,1580]" captionTargetPageId="19" captionText="FIGURES 28 – 31. Grallipeza marleyi. 28, female, right lateral; 29, spermathecae and ducts, partially concealed in matrix; 30, female head dorsal; 31, female head lateral." httpUri="https://zenodo.org/record/284123/files/figure.png" pageId="18" pageNumber="63">Figs. 28–31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34461FCFFA9FFBFFF4413D7FA78F83C" pageId="18" pageNumber="63" type="description">
<paragraph id="8BE13277FFA9FFBFFF4413D7FD55FA10" blockId="18.[151,1436,1230,2011]" pageId="18" pageNumber="63">
<emphasis id="B92AEE65FFA9FFBFFF4413D7FED4FB49" bold="true" box="[151,289,1230,1255]" pageId="18" pageNumber="63">Description</emphasis>
(females only). Size: Approximately
<quantity id="4CA69F92FFA9FFBFFD0313D6FCE3FB48" box="[720,790,1231,1254]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="63" unit="mm" value="7.0">7 mm</quantity>
. Colour: Head orange except shining brown clypeus and apically slightly darkened palpus (
<figureCitation id="13652EF2FFA9FFBFFDC413EBFD9EFAA4" box="[535,619,1266,1290]" captionStart="FIGURES 28 – 31" captionStartId="19.[151,264,1601,1624]" captionTargetBox="[151,1434,193,1579]" captionTargetId="figure@19.[151,1436,193,1580]" captionTargetPageId="19" captionText="FIGURES 28 – 31. Grallipeza marleyi. 28, female, right lateral; 29, spermathecae and ducts, partially concealed in matrix; 30, female head dorsal; 31, female head lateral." httpUri="https://zenodo.org/record/284123/files/figure.png" pageId="18" pageNumber="63">Fig. 28</figureCitation>
). Thorax with notum dark brown and pleuron yellow-orange. Fore femur orange, darkened on distal half, tarsomere one and tarsomere two off-white with white bristles, apex of tarsomere 2 and tarsomeres 3–5 brown with brown bristles. Mid and hind femora pale brown with an indistinct dark distal ring. Katepisternal bristles golden. Abdominal tergites two and three pale, tergites 1, 4, 5 pruinose brown, tergite 7 pruinose at base, otherwise shining brown. Upper third of female abdominal pleuron black on segments 3–5; oviscape shining brown with scattered setulae.
</paragraph>
<paragraph id="8BE13277FFA9FFBFFF1412D3FC05F984" blockId="18.[151,1436,1230,2011]" pageId="18" pageNumber="63">
Head: Arista conspicuously long-haired at least over basal 2/3, hairs as long as scape width (
<figureCitation id="13652EF2FFA9FFBFFB3312D3FAC0FA4C" box="[1248,1333,1482,1506]" captionStart="FIGURES 28 – 31" captionStartId="19.[151,264,1601,1624]" captionTargetBox="[151,1434,193,1579]" captionTargetId="figure@19.[151,1436,193,1580]" captionTargetPageId="19" captionText="FIGURES 28 – 31. Grallipeza marleyi. 28, female, right lateral; 29, spermathecae and ducts, partially concealed in matrix; 30, female head dorsal; 31, female head lateral." httpUri="https://zenodo.org/record/284123/files/figure.png" pageId="18" pageNumber="63">Fig. 30</figureCitation>
). Pedicel with long ventroapical bristles, one almost as long as first flagellomere. Frontal vitta broadened and slightly raised in front of ocelli; narrow, parallel-sided and slightly depressed behind ocelli.
</paragraph>
<paragraph id="8BE13277FFA9FFBFFF14112FFF36F970" blockId="18.[151,1436,1230,2011]" pageId="18" pageNumber="63">
Thorax: Cervical sclerite concave ventrally, concave surface densely covered with distinct pores. Fore femur with only small ventral setulae. One or two distinct dorsocentral bristles; suprahumeral bristles very large, exclinate, forming a distinct row of 4–5 black bristles at anterior end of an inconspicuous row of minute, pale dorsocentral setulae (
<figureCitation id="13652EF2FFA9FFBFFE5411BBFE29F914" box="[391,476,1698,1722]" captionStart="FIGURES 28 – 31" captionStartId="19.[151,264,1601,1624]" captionTargetBox="[151,1434,193,1579]" captionTargetId="figure@19.[151,1436,193,1580]" captionTargetPageId="19" captionText="FIGURES 28 – 31. Grallipeza marleyi. 28, female, right lateral; 29, spermathecae and ducts, partially concealed in matrix; 30, female head dorsal; 31, female head lateral." httpUri="https://zenodo.org/record/284123/files/figure.png" pageId="18" pageNumber="63">Fig. 31</figureCitation>
). Wing: Anal cell microsetulose, wing membrane lightly and evenly infuscated (
<figureCitation id="13652EF2FFA9FFBFFABD11BBFF42F970" captionStart="FIGURES 28 – 31" captionStartId="19.[151,264,1601,1624]" captionTargetBox="[151,1434,193,1579]" captionTargetId="figure@19.[151,1436,193,1580]" captionTargetPageId="19" captionText="FIGURES 28 – 31. Grallipeza marleyi. 28, female, right lateral; 29, spermathecae and ducts, partially concealed in matrix; 30, female head dorsal; 31, female head lateral." httpUri="https://zenodo.org/record/284123/files/figure.png" pageId="18" pageNumber="63">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="8BE13277FFA9FFBFFF1411F3FA78F83C" blockId="18.[151,1436,1230,2011]" pageId="18" pageNumber="63">
Female abdomen: Oviscape elongate, 3 times as long as tergite 6 and almost as long as preabdomen (
<figureCitation id="13652EF2FFA9FFBFFAE811F3FA65F8AC" box="[1339,1424,1770,1794]" captionStart="FIGURES 28 – 31" captionStartId="19.[151,264,1601,1624]" captionTargetBox="[151,1434,193,1579]" captionTargetId="figure@19.[151,1436,193,1580]" captionTargetPageId="19" captionText="FIGURES 28 – 31. Grallipeza marleyi. 28, female, right lateral; 29, spermathecae and ducts, partially concealed in matrix; 30, female head dorsal; 31, female head lateral." httpUri="https://zenodo.org/record/284123/files/figure.png" pageId="18" pageNumber="63">Fig. 28</figureCitation>
). Paired spermathecae elongate, three times as long as wide, expanded distally, on short ducts arising from a swollen oval distal part of the common duct, swollen part occupying distal third of common duct and three times as wide as spermatheca. Single spermatheca bilobed, duct uniform with a constriction at base of spermatheca (one of the two female
<typeStatus id="54E58CD5FFA9FFBFFF391062FED0F83D" box="[234,293,1915,1939]" pageId="18" pageNumber="63">types</typeStatus>
was dissected but incompletely cleared, spermathecae partially embedded in a hard matrix,
<figureCitation id="13652EF2FFA9FFBFFAFE1063FA75F83C" box="[1325,1408,1914,1938]" captionStart="FIGURES 28 – 31" captionStartId="19.[151,264,1601,1624]" captionTargetBox="[151,1434,193,1579]" captionTargetId="figure@19.[151,1436,193,1580]" captionTargetPageId="19" captionText="FIGURES 28 – 31. Grallipeza marleyi. 28, female, right lateral; 29, spermathecae and ducts, partially concealed in matrix; 30, female head dorsal; 31, female head lateral." httpUri="https://zenodo.org/record/284123/files/figure.png" pageId="18" pageNumber="63">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34461FCFFA9FFBFFF141086FAD1F874" pageId="18" pageNumber="63" type="materials_examined">
<paragraph id="8BE13277FFA9FFBFFF141086FAD1F874" blockId="18.[151,1436,1230,2011]" pageId="18" pageNumber="63">
<emphasis id="B92AEE65FFA9FFBFFF141086FE8CF819" bold="true" box="[199,377,1950,1975]" pageId="18" pageNumber="63">
<typeStatus id="54E58CD5FFA9FFBFFF141086FEF5F819" box="[199,256,1951,1975]" pageId="18" pageNumber="63">Type</typeStatus>
material:
</emphasis>
<typeStatus id="54E58CD5FFA9FFBFFE541087FE1AF818" box="[391,495,1950,1974]" pageId="18" pageNumber="63" type="holotype">Holotype</typeStatus>
female and one
<typeStatus id="54E58CD5FFA9FFBFFD681086FCEEF819" box="[699,795,1951,1975]" pageId="18" pageNumber="63" type="paratype">paratype</typeStatus>
female:
<collectingCountry id="F34972E7FFA9FFBFFC541087FC13F818" box="[903,998,1950,1974]" name="Jamaica" pageId="18" pageNumber="63">Jamaica</collectingCountry>
: Windsor,
<quantity id="4CA69F92FFA9FFBFFBB51087FB32F818" box="[1126,1223,1950,1974]" metricMagnitude="4" metricUnit="m" metricValue="1.55" pageId="18" pageNumber="63" unit="km" value="15.5">15.5 km</quantity>
S Falmouth, path through Cockpit country to Troy,
<date id="FFE014B7FFA9FFBFFDDC10DBFD8CF874" box="[527,633,1986,2010]" pageId="18" pageNumber="63" value="1983-04-02">2.iv.1983</date>
, J. Spott, wet limestone forest, debu00256921 (2 Ƥ,
<collectionCode id="ED4FAAB2FFA9FFBFFB1410DBFAECF874" box="[1223,1305,1986,2010]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="18" pageNumber="63">DEBU</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF2162FFFFA8FFBEFF441158FDA7F9D8" httpUri="https://zenodo.org/record/284123/files/figure.png" pageId="19" pageNumber="64" targetBox="[151,1434,193,1579]" targetPageId="19">
<paragraph id="8BE13277FFA8FFBEFF441158FDA7F9D8" blockId="19.[151,1436,1601,1654]" pageId="19" pageNumber="64">
<emphasis id="B92AEE65FFA8FFBEFF441158FDD6F9F9" bold="true" box="[151,547,1601,1624]" pageId="19" pageNumber="64">
FIGURES 28–31.
<emphasis id="B92AEE65FFA8FFBEFE881158FDD6F9F9" bold="true" box="[347,547,1601,1623]" italics="true" pageId="19" pageNumber="64">
<taxonomicName id="4C5E49F4FFA8FFBEFE881158FDEAF9F9" box="[347,543,1601,1623]" class="Insecta" family="Micropezidae" genus="Grallipeza" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="64" phylum="Arthropoda" rank="species" species="marleyi">Grallipeza marleyi</taxonomicName>
.
</emphasis>
</emphasis>
28, female, right lateral; 29, spermathecae and ducts, partially concealed in matrix; 30, female head dorsal; 31, female head lateral.
</paragraph>
</caption>
<subSubSection id="C34461FCFFA8FFB9FF141184FB1DFF59" lastPageId="20" lastPageNumber="65" pageId="19" pageNumber="64" type="discussion">
<paragraph id="8BE13277FFA8FFB9FF141184FB1DFF59" blockId="19.[151,1436,1693,2005]" lastBlockId="20.[151,1436,151,248]" lastPageId="20" lastPageNumber="65" pageId="19" pageNumber="64">
<emphasis id="B92AEE65FFA8FFBEFF141184FEBDF918" bold="true" box="[199,328,1693,1718]" pageId="19" pageNumber="64">Comments</emphasis>
:
<taxonomicName id="4C5E49F4FFA8FFBEFE8E1184FDCDF91B" box="[349,568,1693,1717]" class="Insecta" family="Micropezidae" genus="Grallipeza" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="64" phylum="Arthropoda" rank="species" species="marleyi">
<emphasis id="B92AEE65FFA8FFBEFE8E1184FDCDF91B" box="[349,568,1693,1717]" italics="true" pageId="19" pageNumber="64">Grallipeza marleyi</emphasis>
</taxonomicName>
is distinctively coloured and easily distinguished from other
<taxonomicName id="4C5E49F4FFA8FFBEFAF71184FA69F91B" box="[1316,1436,1693,1717]" class="Insecta" family="Micropezidae" genus="Grallipeza" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEE65FFA8FFBEFAF71184FA69F91B" box="[1316,1436,1693,1717]" italics="true" pageId="19" pageNumber="64">Grallipeza</emphasis>
</taxonomicName>
species, although the apparent difference in dorsocentral number between the two otherwise identical female
<typeStatus id="54E58CD5FFA8FFBEFABF11DBFA69F974" box="[1388,1436,1730,1754]" pageId="19" pageNumber="64">type</typeStatus>
specimens is unusual. The desclerotized tergites 2–3 suggest a close relationship to another Jamaican species,
<taxonomicName id="4C5E49F4FFA8FFBEFA5711FFFF3FF88F" class="Insecta" family="Micropezidae" genus="Grallipeza" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="64" phylum="Arthropoda" rank="species" species="cliffi">
<emphasis id="B92AEE65FFA8FFBEFA5711FFFF3FF88F" italics="true" pageId="19" pageNumber="64">G. cliffi</emphasis>
</taxonomicName>
, and to the Hispaniolan species
<emphasis id="B92AEE65FFA8FFBEFD901013FC6DF88F" box="[579,920,1801,1825]" italics="true" pageId="19" pageNumber="64">
<taxonomicName id="4C5E49F4FFA8FFBEFD901013FD39F88F" box="[579,716,1802,1825]" class="Insecta" family="Micropezidae" genus="Grallipeza" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="64" phylum="Arthropoda" rank="species" species="nigrivitta">G. nigrivitta</taxonomicName>
and
<taxonomicName id="4C5E49F4FFA8FFBEFCD91013FC61F88F" box="[778,916,1801,1825]" class="Insecta" family="Micropezidae" genus="Grallipeza" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="64" phylum="Arthropoda" rank="species" species="albiterga">G. albiterga</taxonomicName>
.
</emphasis>
The strongly swollen common spermathecal duct is characteristic of several members of the
<taxonomicName id="4C5E49F4FFA8FFBEFD6E1037FCC4F8E8" box="[701,817,1838,1862]" class="Insecta" family="Micropezidae" genus="Grallipeza" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="64" phylum="Arthropoda" rank="species" species="placida">
<emphasis id="B92AEE65FFA8FFBEFD6E1037FCC4F8E8" box="[701,817,1838,1862]" italics="true" pageId="19" pageNumber="64">G. placida</emphasis>
</taxonomicName>
group, but the relatively small, elongate, trumpet-like paired spermathecae appear to be unique. The oviscape was removed from both the female
<typeStatus id="54E58CD5FFA8FFBEFB431048FB07F8C7" box="[1168,1266,1873,1897]" pageId="19" pageNumber="64" type="holotype">holotype</typeStatus>
and the female
<typeStatus id="54E58CD5FFA8FFBEFF44106FFF0FF820" box="[151,250,1910,1934]" pageId="19" pageNumber="64" type="paratype">paratype</typeStatus>
, and in both cases the usual clearing process (boiling in potassium hydroxide) left the spermathecae and ducts embedded in opaque tissue that could not be teased away from the spermathecae. This is an unusual problem and it is remarkable that it occurred in two specimens cleared in different stock solution several weeks apart.
<figureCitation id="13652EF2FFA8FFB9FA8010A4FF46FF01" captionStart="FIGURES 28 – 31" captionStartId="19.[151,264,1601,1624]" captionTargetBox="[151,1434,193,1579]" captionTargetId="figure@19.[151,1436,193,1580]" captionTargetPageId="19" captionText="FIGURES 28 – 31. Grallipeza marleyi. 28, female, right lateral; 29, spermathecae and ducts, partially concealed in matrix; 30, female head dorsal; 31, female head lateral." httpUri="https://zenodo.org/record/284123/files/figure.png" lastPageId="20" lastPageNumber="65" pageId="19" pageNumber="64">Figure 29</figureCitation>
shows the stained
<typeStatus id="54E58CD5FFAFFFB9FE461781FE03FF1E" box="[405,502,152,176]" pageId="20" pageNumber="65" type="paratype">paratype</typeStatus>
spermathecae in which the distally swollen common duct is clearly visible; the
<typeStatus id="54E58CD5FFAFFFB9FF4417A5FF0CFF7A" box="[151,249,188,212]" pageId="20" pageNumber="65" type="holotype">holotype</typeStatus>
spermathecae are barely distinguishable from the surrounding tissue but appear to match the
<typeStatus id="54E58CD5FFAFFFB9FAC317A4FA81FF7B" box="[1296,1396,189,213]" pageId="20" pageNumber="65" type="paratype">paratype</typeStatus>
.
<emphasis id="B92AEE65FFAFFFB9FF1417F9FEBDFF56" bold="true" box="[199,328,224,248]" pageId="20" pageNumber="65">Etymology</emphasis>
:
<taxonomicName id="4C5E49F4FFAFFFB9FE8417F9FDDEFF56" box="[343,555,224,248]" class="Insecta" family="Micropezidae" genus="Grallipeza" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="65" phylum="Arthropoda" rank="species" species="marleyi">
<emphasis id="B92AEE65FFAFFFB9FE8417F9FDDEFF56" box="[343,555,224,248]" italics="true" pageId="20" pageNumber="65">Grallipeza marleyi</emphasis>
</taxonomicName>
is a patronym referring to the Jamaican musician Bob Marley.
</paragraph>
</subSubSection>
</treatment>
</document>
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