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<document id="47D7F3F02E81FA98D1F9D6DE39061BD8" ID-CLB-Dataset="54010" ID-DOI="10.5852/ejt.2023.873.2133" ID-GBIF-Dataset="6b60ec14-d9a3-4400-b636-bb3ee16094b7" ID-ISSN="2118-9773" ID-Zenodo-Dep="8038022" ID-ZooBank="099F6A49-6516-4F39-9E20-A077A5D60535" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1686745931301" checkinUser="felipe" docAuthor="Pham, Nhi Thi, Pham, Phu Van, Matsumoto, Rikio, Shimizu, So &amp; Broad, Gavin R." docDate="2023" docId="03E3AC44FFB8FFC397ABFEE004E58802" docLanguage="en" docName="ejt-873_1-151.pdf" docOrigin="European Journal of Taxonomy 873 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.873.2133" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Enicospilus hiepi Pham &amp; Pham &amp; Matsumoto &amp; Shimizu &amp; Broad 2023, sp. nov." docType="treatment" docUuid="85F24F01-1A78-46EB-BE6C-9A0DEA4C3767" docUuidSource="ZooBank" docVersion="8" lastPageNumber="74" masterDocId="FFDAD43CFFFFFF8A952BFFE8073A8A55" masterDocTitle="A review of the genus Enicospilus Stephens (Ichneumonidae: Ophioninae) from Vietnam, with descriptions of ten new species" masterLastPageNumber="151" masterPageNumber="1" pageNumber="72" updateTime="1699524284659" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5FD9D44703048DD139FA47AB301C4C65">A review of the genus Enicospilus Stephens (Ichneumonidae: Ophioninae) from Vietnam, with descriptions of ten new species</mods:title>
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<mods:namePart id="8C9072E3C7CCF1907CEB04D316AB52FD">Pham, Nhi Thi</mods:namePart>
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<mods:affiliation id="BC81E28CA500C9B832F816F1B14C84A3">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. Osaka Museum of Natural History, Nagai Park 1 - 23, Higashisumiyoshi-ku, Osaka, Japan. Institute for Agro-Environmental Sciences, NARO, Kannondai 3 - 1 - 3, Tsukuba 305 - 8604, Japan. Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Kobe, Hyogo, Japan. The Natural History Museum, Cromwell Road, London SW 7 BD, UK.</mods:affiliation>
<mods:nameIdentifier id="AB71890BF8444D7C2D87B38AFEC9B248" type="email">ptnhi2@yahoo.com</mods:nameIdentifier>
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<mods:name id="44AC217B8F416CC1791F8F3C5D284145" type="personal">
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<mods:namePart id="119587CB762CC4FEE633D8DAC4DFD29F">Pham, Phu Van</mods:namePart>
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<mods:affiliation id="E96F83BE5474F247B82DE79A1140494E">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. Osaka Museum of Natural History, Nagai Park 1 - 23, Higashisumiyoshi-ku, Osaka, Japan. Institute for Agro-Environmental Sciences, NARO, Kannondai 3 - 1 - 3, Tsukuba 305 - 8604, Japan. Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Kobe, Hyogo, Japan. The Natural History Museum, Cromwell Road, London SW 7 BD, UK.</mods:affiliation>
<mods:nameIdentifier id="1E90CE0BB3B017388FD6248796BF84A5" type="email">phamphu93.k56@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E3A188A1262B66EB2928215377BD2B40">Matsumoto, Rikio</mods:namePart>
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<mods:nameIdentifier id="D34547B05F760D95ADD6FE4B6A042E41" type="email">rikio@mus-nh.city.osaka.jp</mods:nameIdentifier>
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<mods:namePart id="3D8B3AF08F1F7A212E81ED8FA745A587">Shimizu, So</mods:namePart>
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<mods:nameIdentifier id="1C98E5898F6C46A94704FFD230B1D378" type="email">parasitoidwasp.sou@gmail.com</mods:nameIdentifier>
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<mods:namePart id="95E8BC17895F5A63E2AF1126CB824137">Broad, Gavin R.</mods:namePart>
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<mods:nameIdentifier id="AE9634384EFBD8FCF7B6EFFDB870ED3A" type="email">g.broad@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="95DA32761C2CE756AE2DD4136A7BA55B">European Journal of Taxonomy</mods:title>
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<mods:date id="5999C1C91E927D782E44A9F85C0A7648">2023</mods:date>
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<treatment id="03E3AC44FFB8FFC397ABFEE004E58802" ID-DOI="http://doi.org/10.5281/zenodo.8039562" ID-GBIF-Taxon="209689792" ID-Zenodo-Dep="8039562" LSID="urn:lsid:zoobank.org:act:85F24F01-1A78-46EB-BE6C-9A0DEA4C3767" httpUri="http://treatment.plazi.org/id/03E3AC44FFB8FFC397ABFEE004E58802" lastPageId="73" lastPageNumber="74" pageId="71" pageNumber="72">
<subSubSection id="C3504ED9FFB8FFCD97ABFEE0048E8B76" box="[640,948,264,291]" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="8BF51D52FFB8FFCD97ABFEE0048E8B76" blockId="71.[363,1224,264,359]" box="[640,948,264,291]" pageId="71" pageNumber="72">
<heading id="D0BDAA3EFFB8FFCD97ABFEE0048E8B76" box="[640,948,264,291]" centered="true" fontSize="11" level="2" pageId="71" pageNumber="72" reason="2">
<taxonomicName id="4C4A66D1FFB8FFCD97ABFEE0046B8B76" authority="Pham &amp; Pham &amp; Matsumoto &amp; Shimizu &amp; Broad, 2023" authorityName="Pham &amp; Pham &amp; Matsumoto &amp; Shimizu &amp; Broad" authorityYear="2023" box="[640,849,264,291]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="hiepi" status="sp. nov.">
<emphasis id="B93EC140FFB8FFCD97ABFEE0046B8B76" bold="true" box="[640,849,264,291]" italics="true" pageId="71" pageNumber="72">Enicospilus hiepi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D7C3BFFB8FFCD9672FEE1048E8B76" box="[857,948,265,291]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<uri id="FFDB1150FFB8FFCD9440FEC303F28B13" box="[363,1224,299,326]" pageId="71" pageNumber="72">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BF51D52FFB8FFCD97DBFEA504798B32" blockId="71.[363,1224,264,359]" box="[752,835,333,359]" pageId="71" pageNumber="72">
<figureCitation id="137101D7FFB8FFCD97DBFEA504798B32" box="[752,835,333,359]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3504ED9FFB8FFCD9596FE65040C88B8" pageId="71" pageNumber="72" type="diagnosis">
<paragraph id="8BF51D52FFB8FFCD9596FE65060C8BF2" blockId="71.[189,310,397,423]" box="[189,310,397,423]" pageId="71" pageNumber="72">
<heading id="D0BDAA3EFFB8FFCD9596FE65060C8BF2" bold="true" box="[189,310,397,423]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
<emphasis id="B93EC140FFB8FFCD9596FE65060C8BF2" bold="true" box="[189,310,397,423]" pageId="71" pageNumber="72">Diagnosis</emphasis>
</heading>
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<paragraph id="8BF51D52FFB8FFCD9596FE5306A08808" blockId="71.[189,1398,443,605]" pageId="71" pageNumber="72">Interocellar area reddish brown; clypeus convex, ventral margin impressed, acute; mandible twisted ca 30°, outer surface with a diagonal setose groove; scutellum with lateral longitudinal carina present on anterior 0.6; fore wing fenestra with proximal, central and distal sclerites, central sclerite large, shaped resembling footprint, pigmented, positioned in upper half of distal part of fenestra; mesoscutum with three black stripes.</paragraph>
<paragraph id="8BF51D52FFB8FFCD9596FD6C06FC88CB" blockId="71.[189,454,644,670]" box="[189,454,644,670]" pageId="71" pageNumber="72">
<heading id="D0BDAA3EFFB8FFCD9596FD6C06FC88CB" bold="true" box="[189,454,644,670]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
<emphasis id="B93EC140FFB8FFCD9596FD6C06FC88CB" bold="true" box="[189,454,644,670]" pageId="71" pageNumber="72">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF51D52FFB8FFCD9596FD59040C88B8" blockId="71.[189,1398,689,749]" pageId="71" pageNumber="72">The new species can be recognised from all other species by the combination of the shape of the alar sclerites and the short lateral carinae of the scutellum.</paragraph>
</subSubSection>
<subSubSection id="C3504ED9FFB8FFCD9596FCFC0266890E" box="[189,1372,788,860]" pageId="71" pageNumber="72" type="etymology">
<paragraph id="8BF51D52FFB8FFCD9596FCFC067E897B" blockId="71.[189,324,788,814]" box="[189,324,788,814]" pageId="71" pageNumber="72">
<heading id="D0BDAA3EFFB8FFCD9596FCFC067E897B" bold="true" box="[189,324,788,814]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
<emphasis id="B93EC140FFB8FFCD9596FCFC067E897B" bold="true" box="[189,324,788,814]" pageId="71" pageNumber="72">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF51D52FFB8FFCD9596FCA90266890E" blockId="71.[189,1372,833,860]" box="[189,1372,833,860]" pageId="71" pageNumber="72">
Named after MSc. Nguyen Duc Hiep, an entomologist from IEBR, who collected the
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specimen.
</paragraph>
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<subSubSection id="C3504ED9FFB8FFCD9596FC6906868E5B" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="8BF51D52FFB8FFCD9596FC69069089CE" blockId="71.[189,426,897,923]" box="[189,426,897,923]" pageId="71" pageNumber="72">
<heading id="D0BDAA3EFFB8FFCD9596FC69069089CE" bold="true" box="[189,426,897,923]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
<emphasis id="B93EC140FFB8FFCD9596FC69069089CE" bold="true" box="[189,426,897,923]" pageId="71" pageNumber="72">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B22170FFFB8FFCD95C1FC4706868E5B" ID-GBIF-Occurrence="4130444889" collectingDate="2021-05-08" collectingMethod="light trap" collectionCode="IEBR" collectorName="Nguyen D. H." country="Vietnam" elevation="791" location="Phia Oac" pageId="71" pageNumber="72" specimenCount="1" specimenCount-female="1" stateProvince="Cao Bang Province" typeStatus="holotype">
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<emphasis id="B93EC140FFB8FFCD95C1FC470666899C" bold="true" box="[234,348,943,969]" pageId="71" pageNumber="72">
<typeStatus id="54F1A3F0FFB8FFCD95C1FC470666899C" box="[234,348,943,969]" pageId="71" pageNumber="72" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF51D52FFB8FFCD9596FC3906868E5B" blockId="71.[189,1398,943,1038]" pageId="71" pageNumber="72">
<collectingCountry id="F35D5DC2FFB8FFCD9596FC39067E89BE" box="[189,324,977,1003]" name="Vietnam" pageId="71" pageNumber="72">VIETNAM</collectingCountry>
•
<specimenCount id="9D4CD6DBFFB8FFCD9476FC3A064E89B9" box="[349,372,978,1004]" pageId="71" pageNumber="72" type="female">♀</specimenCount>
;
<collectingRegion id="498ED3B0FFB8FFCD94A8FC39055089BE" box="[387,618,977,1004]" country="Vietnam" name="Cao Bang" pageId="71" pageNumber="72">Cao Bang Province</collectingRegion>
,
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-
<location id="8E954B89FFB8FFCD97D1FC3904AD89B9" LSID="urn:lsid:plazi:treatment:03E3AC44FFB8FFC397ABFEE004E58802:8E954B89FFB8FFCD97D1FC3904AD89B9" box="[762,919,977,1004]" country="Vietnam" name="Phia Den NP" pageId="71" pageNumber="72" stateProvince="Cao Bang Province">Phia Den NP</location>
;
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<quantity id="4CB2B0B7FFB8FFCD9688FC3904D489B9" box="[931,1006,977,1004]" metricMagnitude="2" metricUnit="m" metricValue="7.909999999999999" pageId="71" pageNumber="72" unit="m" value="791.0">791 m</quantity>
a.s.l.
</elevation>
;
<date id="FFF43B92FFB8FFCD9110FC3903F789BE" box="[1083,1229,977,1004]" pageId="71" pageNumber="72" value="2021-05-08">
<collectingDate id="EFB0C27AFFB8FFCD9110FC3903F789BE" box="[1083,1229,977,1004]" pageId="71" pageNumber="72" value="2021-05-08">8 May 2021</collectingDate>
</date>
;
<collectorName id="26BF7884FFB8FFCD91F2FC3A024C89B9" box="[1241,1398,978,1004]" pageId="71" pageNumber="72">Nguyen D.H.</collectorName>
leg.;
<collectingMethod id="520B6545FFB8FFCD95D3FC1B065C8E5B" box="[248,358,1011,1038]" pageId="71" pageNumber="72">light trap</collectingMethod>
;
<collectionCode id="ED5B8597FFB8FFCD9459FC1C06838E5B" box="[370,441,1012,1038]" pageId="71" pageNumber="72">IEBR</collectionCode>
.
</paragraph>
</materialsCitation>
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<emphasis id="B93EC140FFB8FFCD9596FBDB06748E18" bold="true" box="[189,334,1075,1101]" pageId="71" pageNumber="72">Description</emphasis>
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</paragraph>
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<emphasis id="B93EC140FFB8FFCD95C1FB89067F8E2E" bold="true" box="[234,325,1121,1147]" pageId="71" pageNumber="72">Female</emphasis>
(
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)
</paragraph>
<paragraph id="8BF51D52FFB8FFCD9596FB6B04F18ECB" blockId="71.[189,971,1121,1182]" box="[189,971,1155,1182]" pageId="71" pageNumber="72">
MEASUREMENTS. Body length
<quantity id="4CB2B0B7FFB8FFCD9730FB6B05BD8ECB" box="[539,647,1155,1182]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="71" pageNumber="72" unit="mm" value="25.4">25.4 mm</quantity>
, fore wing length 18.0 mm.
</paragraph>
<paragraph id="8BF51D52FFB8FFCD9596FB2C034C8F99" blockId="71.[189,1399,1219,1485]" pageId="71" pageNumber="72">
HEAD with FI = 0.7, GOI = 2.0 (
<figureCitation id="137101D7FFB8FFCD9704FB2B05AF8E88" box="[559,661,1219,1246]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38d</figureCitation>
). Lower face 0.8× as wide as high, densely punctate (
<figureCitation id="137101D7FFB8FFCD9028FB2B02538E88" box="[1283,1385,1219,1245]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38b</figureCitation>
). Clypeus 1.4× as wide as high, convex in profile, more densely punctate than face, ventral margin impressed, acute (
<figureCitation id="137101D7FFB8FFCD94B9FAE006C38F77" box="[402,505,1288,1314]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38b</figureCitation>
). Malar space 0.2× as long as basal mandibular width (
<figureCitation id="137101D7FFB8FFCD91A2FAE003CB8F77" box="[1161,1265,1288,1314]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38b</figureCitation>
). Mandible twisted by ca 30°, moderately long, evenly narrowed, outer surface with diagonal setose groove (
<figureCitation id="137101D7FFB8FFCD902CFAC202508F11" box="[1287,1386,1322,1348]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38b</figureCitation>
). Upper tooth of mandible depressed, about 2 × as long as lower tooth (
<figureCitation id="137101D7FFB8FFCD96C7FAA403698F33" box="[1004,1107,1356,1382]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38b</figureCitation>
). Frons short, vertex and gena with fine setae (
<figureCitation id="137101D7FFB8FFCD9490FA86057A8FDD" box="[443,576,1390,1416]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38c–d</figureCitation>
). Posterior ocellus close to eye (
<figureCitation id="137101D7FFB8FFCD96E9FA8603138FDD" box="[962,1065,1390,1416]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38c</figureCitation>
). Occipital carina complete, ventral end meeting hypostomal carina at about 0.2 × basal mandible width from base of mandible. Antenna with 70 flagellomeres; F1 1.9× as long as F2; F20 2.4 × as long as wide.
</paragraph>
<paragraph id="8BF51D52FFB8FFCD9596FA1D03D98CA8" blockId="71.[187,1399,1524,1790]" pageId="71" pageNumber="72">
MESOSOMA. Moderately shiny (
<figureCitation id="137101D7FFB8FFCD970DFA1C05B08C5A" box="[550,650,1524,1551]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38e</figureCitation>
). Pronotum finely punctate (
<figureCitation id="137101D7FFB8FFCD96F9FA1C030F8C5A" box="[978,1077,1524,1551]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38e</figureCitation>
). Mesoscutum 1.4× as long as maximum width, evenly rounded in profile. Notaulus absent. Scutellum strongly convex, punctate, without any rugae, lateral longitudinal carinae present on anterior 0.6. Epicnemium densely punctate (
<figureCitation id="137101D7FFB8FFCD95EDF9B306138C20" box="[198,297,1627,1653]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38e</figureCitation>
). Epicnemial carina moderately strong, dorsal end bent to reach anterior margin of mesopleuron (
<figureCitation id="137101D7FFB8FFCD95EDF99506128CC2" box="[198,296,1661,1687]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38e</figureCitation>
). Mesopleuron punctate and puncto-striate (
<figureCitation id="137101D7FFB8FFCD960AF99504BE8CC2" box="[801,900,1661,1687]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38e</figureCitation>
). Submetapleural carina slightly broadened anteriorly (
<figureCitation id="137101D7FFB8FFCD9469F97706968CEC" box="[322,428,1695,1721]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38e</figureCitation>
). Metapleuron finely punctate (
<figureCitation id="137101D7FFB8FFCD9601F97704A88CEC" box="[810,914,1695,1721]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38e</figureCitation>
). Propodeum evenly rounded in profile; anterior transverse carina complete; anterior area longitudinally striate; spiracular area smooth; posterior area rugose; propodeal spiracle elliptical and not joining pleural carina by ridge (
<figureCitation id="137101D7FFB8FFCD915BF90B03EF8CA8" box="[1136,1237,1763,1789]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38e</figureCitation>
).
</paragraph>
<paragraph id="8BF51D52FFB8FFC29596F8CC044F8B13" blockId="71.[189,1399,1828,2025]" lastBlockId="72.[189,1398,265,327]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
WINGS (
<figureCitation id="137101D7FFB8FFCD9437F8CC06418D6B" box="[284,379,1828,1854]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38f</figureCitation>
). Fore wing with AI = 0.59, CI = 0.50, DI = 0.35, ICI = 0.51, SDI = 1.31, SI = 0.13,
<collectingRegion id="498ED3B0FFB8FFCD9018F8CC025A8D6B" box="[1331,1376,1828,1854]" country="United Kingdom" name="Surrey" pageId="71" pageNumber="72">SRI</collectingRegion>
= 0.23; vein 1m-cu&amp;M strongly sinuate, angled medially; vein 2r&amp;RS slightly sinuate, abruptly narrow distally; vein RS curved; fenestra and sclerites of discosubmarginal cell as in
<figureCitation id="137101D7FFB8FFCD916EF880039F8DD7" box="[1093,1189,1896,1922]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="71" pageNumber="72">Fig. 38f</figureCitation>
; proximal sclerite triangular, confluent with distal sclerite, strongly pigmented; central sclerite large, somewhat resembling footprint in shape, pigmented, positioned in upper half of distal part of fenestra; distal sclerite pigmented; proximal corner of marginal cell sparsely setose; posterodistal corner of second discal cell ca 80° and that of subbasal cell ca 70°; vein 1cu-a antefurcal to M&amp;RS by 0.3× length of 1cu-a. Hind wing with NI = 1.7; vein RS straight; vein RA with 7 uniform hamuli.
</paragraph>
<paragraph id="8BF51D52FFB7FFC29596FE8604E38BFE" blockId="72.[189,1398,366,427]" pageId="72" pageNumber="73">LEGS. Hind leg with coxa in profile 2.0× as long as high; basitarsus 2.1× as long as second tarsomere; fourth tarsomere 3.3× as long as wide; tarsal claw simply pectinate.</paragraph>
<paragraph id="8BF51D52FFB7FFC29596FE3A05268845" blockId="72.[189,1398,466,528]" pageId="72" pageNumber="73">METASOMA. With DMI = 1.2, PI = 3.1, THI = 2.8; thyridium oval; ovipositor sheath not longer than posterior height of metasoma.</paragraph>
<caption id="DF354DDAFFB7FFC29596F897024D8DE8" ID-DOI="http://doi.org/10.5281/zenodo.8038166" ID-Zenodo-Dep="8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="72" pageNumber="73" startId="72.[189,232,1919,1945]" targetBox="[189,1399,591,1881]" targetPageId="72">
<paragraph id="8BF51D52FFB7FFC29596F897024D8DE8" blockId="72.[189,1399,1919,1981]" pageId="72" pageNumber="73">
<emphasis id="B93EC140FFB7FFC29596F89706228DCC" bold="true" box="[189,280,1919,1945]" pageId="72" pageNumber="73">Fig. 38.</emphasis>
<taxonomicName id="4C4A66D1FFB7FFC29436F89706DD8DCC" authority="Pham &amp; Pham &amp; Matsumoto &amp; Shimizu &amp; Broad, 2023" authorityName="Pham &amp; Pham &amp; Matsumoto &amp; Shimizu &amp; Broad" authorityYear="2023" box="[285,487,1919,1945]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="hiepi" status="sp. nov.">
<emphasis id="B93EC140FFB7FFC29436F89706DD8DCC" box="[285,487,1919,1945]" italics="true" pageId="72" pageNumber="73">Enicospilus hiepi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D7C3BFFB7FFC294C7F86805728DCF" box="[492,584,1920,1946]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (IEBR).
<emphasis id="B93EC140FFB7FFC2966EF897046E8DCC" bold="true" box="[837,852,1919,1945]" pageId="72" pageNumber="73">a</emphasis>
. Habitus.
<emphasis id="B93EC140FFB7FFC296E2F89704E38DCC" bold="true" box="[969,985,1919,1945]" pageId="72" pageNumber="73">b</emphasis>
. Head, frontal view.
<emphasis id="B93EC140FFB7FFC291E1F89703ED8DCC" bold="true" box="[1226,1239,1919,1945]" pageId="72" pageNumber="73">c</emphasis>
. Head, dorsal view.
<emphasis id="B93EC140FFB7FFC29429F84B06288DE8" bold="true" box="[258,274,1955,1981]" pageId="72" pageNumber="73">d</emphasis>
. Head, lateral view.
<emphasis id="B93EC140FFB7FFC2972BF84B05378DE8" bold="true" box="[512,525,1955,1981]" pageId="72" pageNumber="73">e</emphasis>
. Mesosoma, lateral view.
<emphasis id="B93EC140FFB7FFC29616F84A047D8DE9" bold="true" box="[829,839,1954,1980]" pageId="72" pageNumber="73">f</emphasis>
. Central part of fore wing. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BF51D52FFB6FFC39596FEE106C58B27" blockId="73.[189,1398,265,370]" pageId="73" pageNumber="74">
COLOUR (
<figureCitation id="137101D7FFB6FFC39406FEE106B48B76" box="[301,398,265,291]" captionStart="Fig" captionStartId="72.[189,232,1919,1945]" captionTargetBox="[189,1399,591,1881]" captionTargetId="figure-102@72.[189,1399,591,1881]" captionTargetPageId="72" captionText="Fig. 38. Enicospilus hiepi sp. nov., holotype, ♀ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038166" httpUri="https://zenodo.org/record/8038166/files/figure.png" pageId="73" pageNumber="74">Fig. 38a</figureCitation>
). Reddish brown except for apex of mandible, three longitudinal stripes of mesoscutum, and ventral parts of T5 onwards black. Wings hyaline; sclerites pigmented and amber; veins dark brown, pterostigma reddish brown.
</paragraph>
<paragraph id="8BF51D52FFB6FFC395C1FE7506108BE2" blockId="73.[189,313,413,479]" box="[234,298,413,439]" pageId="73" pageNumber="74">
<emphasis id="B93EC140FFB6FFC395C1FE7506108BE2" bold="true" box="[234,298,413,439]" pageId="73" pageNumber="74">Male</emphasis>
</paragraph>
<paragraph id="8BF51D52FFB6FFC39596FE2D06038B8A" blockId="73.[189,313,413,479]" box="[189,313,453,479]" pageId="73" pageNumber="74">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3504ED9FFB6FFC39596FDE204E58802" box="[189,991,522,599]" pageId="73" pageNumber="74" type="distribution">
<paragraph id="8BF51D52FFB6FFC39596FDE2066C8871" blockId="73.[189,342,522,548]" box="[189,342,522,548]" pageId="73" pageNumber="74">
<heading id="D0BDAA3EFFB6FFC39596FDE2066C8871" bold="true" box="[189,342,522,548]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="B93EC140FFB6FFC39596FDE2066C8871" bold="true" box="[189,342,522,548]" pageId="73" pageNumber="74">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF51D52FFB6FFC39596FDD504E58802" blockId="73.[189,991,573,599]" box="[189,991,573,599]" pageId="73" pageNumber="74">
Currently known only from
<collectingRegion id="498ED3B0FFB6FFC39723FDD505D58802" box="[520,751,573,599]" country="Vietnam" name="Cao Bang" pageId="73" pageNumber="74">Cao Bang Province</collectingRegion>
, Northeast
<collectingCountry id="F35D5DC2FFB6FFC39658FDD504E08802" box="[883,986,573,599]" name="Vietnam" pageId="73" pageNumber="74">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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