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<document id="3D643C23483A2FEC91FB3F40482FF88F" ID-DOI="10.1649/0010-065X-72.3.433" ID-ISSN="1938-4394" ID-Zenodo-Dep="5383463" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630296642708" checkinUser="diego" docAuthor="Gillett, Conrad P. D. T. &amp; Barr, Iain" docDate="2018" docId="03E32325FF803362879D3CECFD8EFB19" docLanguage="en" docName="0010-065X-72.3.433.pdf" docOrigin="The Coleopterists Bulletin 72 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-72.3.433" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Copris fidius" docType="treatment" docVersion="4" lastPageNumber="436" masterDocId="FFDA5B5DFF84336687633D48FFE8FFB5" masterDocTitle="New Country Records of Scarab Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae) in Swaziland and Zambia" masterLastPageNumber="438" masterPageNumber="433" pageNumber="436" updateTime="1699219109578" updateUser="plazi">
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<mods:title id="FED79ECEE9DA0E62A97AC6AE55D04530">New Country Records of Scarab Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae) in Swaziland and Zambia</mods:title>
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<mods:namePart id="28FC6829C9229F20A0F56DB1E0FDE9F4">Gillett, Conrad P. D. T.</mods:namePart>
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<mods:namePart id="14CDC6DCBF465B0A84F72802AF108878">Barr, Iain</mods:namePart>
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<taxonomicName id="4C4AE9B0FF803362879D3CECFDF0FE0F" ID-CoL="7ZMH8" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[254,536,420,442]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="436" phylum="Arthropoda" rank="species" species="fidius">
<emphasis id="B93E4E21FF803362879D3CECFDF0FE0F" bold="true" box="[254,536,420,442]" pageId="4" pageNumber="436">
<emphasis id="B93E4E21FF803362879D3CECFE96FE0C" bold="true" box="[254,382,420,441]" italics="true" pageId="4" pageNumber="436">Copris fidius</emphasis>
(Olivier, 1789)
</emphasis>
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<subSubSection id="C350C1B8FF803362863D3CF7FD8EFB19" pageId="4" pageNumber="436" type="description">
<paragraph id="8BF59233FF803362863D3CF7FE5FFE61" blockId="4.[254,536,420,468]" box="[350,439,447,468]" pageId="4" pageNumber="436">
(
<figureCitation id="13718EB6FF80336286063CF7FE58FE61" box="[357,432,447,468]" captionStart="Figs" captionStartId="3.[197,236,1616,1635]" captionTargetBox="[220,1145,196,1598]" captionTargetId="figure-1@3.[218,1147,194,1600]" captionTargetPageId="3" captionText="Figs. 13. 1) Maps depicting the approximate locations of the main sampling sites (numbered and named) in A) Zambia and B) Swaziland; 2) Habitats at sampling localities A) Savanna-forest at Mbuluzi Game Reserve, Lubombo, Swaziland, March 2016, B) Montane grassland at Malolotja Nature Reserve, Hhohho, Swaziland, March 2016, C) Miombo woodland, 30 km SSW of Kitwe, Copperbelt Province, Zambia, December 2011; 3) Newly recorded species of scarabaeine dung beetles from Swaziland A) Copris fidius, B) Catharsius tricornutus, C) Proagoderus lanista. All photographs by Conrad P. D. T. Gillett." figureDoi="http://doi.org/10.5281/zenodo.5383465" httpUri="https://zenodo.org/record/5383465/files/figure.png" pageId="4" pageNumber="436">Fig. 3A</figureCitation>
)
</paragraph>
<paragraph id="8BF59233FF80336287CC3CA2FDEBFDFA" blockId="4.[151,639,490,1016]" pageId="4" pageNumber="436">
Previously recorded from
<collectingCountry id="F35DD2A3FF80336286D73CA2FE83FDAC" name="Democratic Republic of the Congo" pageId="4" pageNumber="436">The Democratic Republic of the Congo</collectingCountry>
(
<collectingCountry id="F35DD2A3FF803362861D3F4CFE5BFDAC" box="[382,435,516,537]" name="Democratic Republic of the Congo" pageId="4" pageNumber="436">DRC</collectingCountry>
),
<collectingCountry id="F35DD2A3FF80336286A83F4CFDA3FDAC" box="[459,587,516,537]" name="Mozambique" pageId="4" pageNumber="436">Mozambique</collectingCountry>
, and
<collectingCountry id="F35DD2A3FF80336287F43F57FEFAFD81" box="[151,274,543,564]" name="South Africa" pageId="4" pageNumber="436">South Africa</collectingCountry>
(
<bibRefCitation id="EFDBEFC2FF803362867C3F57FE4EFD81" author="Ferreira, M. C." box="[287,422,543,564]" pageId="4" pageNumber="436" pagination="5 - 1088" refId="ref3778" refString="Ferreira, M. C. 1972. Os escarab'ideos de Africa (sul do Saara). I. Revista de Entomologia de Mocambique 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972</bibRefCitation>
;
<bibRefCitation id="EFDBEFC2FF80336286D33F57FD91FD81" author="Nguyen-Phung, T." box="[432,633,543,564]" pageId="4" pageNumber="436" pagination="5 - 15" refId="ref3997" refString="Nguyen-Phung, T. 1988. Revision des especes Afrotropicales du genre Copris Muller, 1764. IV. Le groupe fidius. Revue Francaise d' Entomologie 10 (1): 5 - 15." type="journal article" year="1988">Nguyen-Phung 1988</bibRefCitation>
;
<bibRefCitation id="EFDBEFC2FF80336287F43F72FEFCFDFA" author="Davis, A. L. V." box="[151,276,570,591]" pageId="4" pageNumber="436" refId="ref3432" refString="Davis, A. L. V. 2013 a. Copris fidius. The IUCN Red List of Threatened Species 2013: e. T 138095 A 536036. dx. doi. org / 10.2305 / IUCN. UK. 2013 - 2. RLTS. T 138095 A 536036. en (accessed 8 November 2017)." type="book" year="2013">Davis 2013a</bibRefCitation>
;
<bibRefCitation id="EFDBEFC2FF803362867C3F72FE10FDFA" author="Deschodt, C. M." box="[287,504,569,591]" pageId="4" pageNumber="436" pagination="431 - 438" refId="ref3714" refString="Deschodt, C. M., A. L. Davis, and C. H. Scholtz. 2015 b. A new synonymy in the fidius group of Copris Muller 1764 (Coleoptera: Scarabaeidae: Scarabaeinae) and a new species from the highland grasslands of South Africa. Zootaxa, 3949 (3): 431 - 438." type="journal article" year="2015">
Deschodt
<emphasis id="B93E4E21FF80336286E33F72FE5AFDFA" box="[384,434,569,591]" italics="true" pageId="4" pageNumber="436">et al.</emphasis>
2015b
</bibRefCitation>
).
</paragraph>
<paragraph id="8BF59233FF80336287CC3F1CFDBDFC4D" blockId="4.[151,639,490,1016]" pageId="4" pageNumber="436">
<emphasis id="B93E4E21FF80336287CC3F1CFE96FDDC" bold="true" box="[175,382,596,617]" pageId="4" pageNumber="436">New country record</emphasis>
for
<collectingCountry id="F35DD2A3FF80336286C93F1CFDE6FDDC" box="[426,526,596,617]" name="Swaziland" pageId="4" pageNumber="436">Swaziland</collectingCountry>
: Ten specimens, Mbuluzi Game Reserve,
<date id="FFF4B4F3FF80336286A03F27FD93FD31" box="[451,635,622,644]" pageId="4" pageNumber="436" value="2016-03-20" valueMax="2016-03-30" valueMin="2016-03-20">2030 March 2016</date>
. Nocturnal. We found this species in overnight pitfall traps baited with fresh wildebeest dung, in company with
<taxonomicName id="4C4AE9B0FF80336287A83FF6FE28FD66" authority="Klug. It" authorityName="Klug. It" box="[203,448,702,724]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="436" phylum="Arthropoda" rank="species" species="elphenor">
<emphasis id="B93E4E21FF80336287A83FF6FE81FD61" box="[203,361,702,724]" italics="true" pageId="4" pageNumber="436">Copris elphenor</emphasis>
Klug. It
</taxonomicName>
was captured only during
<date id="FFF4B4F3FF80336287B93F91FEA6FD5B" box="[218,334,729,750]" pageId="4" pageNumber="436" value="2016-03">March 2016</date>
, which was a comparatively wet period compared to
<date id="FFF4B4F3FF80336286373FBBFE54FCBD" box="[340,444,755,776]" pageId="4" pageNumber="436" value="2015-04">April 2015</date>
, when no specimens were found despite many pitfall traps being operated. The species is recorded as being “centred on forest patches, both along the eastern coastline of
<collectingCountry id="F35DD2A3FF80336285243E0BFF3AFCC6" name="South Africa" pageId="4" pageNumber="436">South Africa</collectingCountry>
as far as the extreme south of
<collectingCountry id="F35DD2A3FF803362869E3E16FD93FCC6" box="[509,635,862,883]" name="Mozambique" pageId="4" pageNumber="436">Mozambique</collectingCountry>
, and at higher altitude along the edge of the eastern escarpment in the north of its South African range” (
<bibRefCitation id="EFDBEFC2FF80336287FD3EE5FE68FC77" author="Deschodt, C. M." box="[158,384,941,963]" pageId="4" pageNumber="436" pagination="431 - 438" refId="ref3714" refString="Deschodt, C. M., A. L. Davis, and C. H. Scholtz. 2015 b. A new synonymy in the fidius group of Copris Muller 1764 (Coleoptera: Scarabaeidae: Scarabaeinae) and a new species from the highland grasslands of South Africa. Zootaxa, 3949 (3): 431 - 438." type="journal article" year="2015">
Deschodt
<emphasis id="B93E4E21FF80336286613EE6FEDFFC76" box="[258,311,941,963]" italics="true" pageId="4" pageNumber="436">et al.</emphasis>
2015b
</bibRefCitation>
). It was evaluated as belonging to the category of least concern in the IUCN Red List of Threatened Species (
<bibRefCitation id="EFDBEFC2FF80336286AC3EABFDA3FC4D" author="Davis, A. L. V." box="[463,587,995,1016]" pageId="4" pageNumber="436" refId="ref3432" refString="Davis, A. L. V. 2013 a. Copris fidius. The IUCN Red List of Threatened Species 2013: e. T 138095 A 536036. dx. doi. org / 10.2305 / IUCN. UK. 2013 - 2. RLTS. T 138095 A 536036. en (accessed 8 November 2017)." type="book" year="2013">Davis 2013a</bibRefCitation>
).
</paragraph>
<paragraph id="8BF59233FF80336287AB395AFDA5FB92" blockId="4.[151,638,1042,1196]" box="[200,589,1042,1064]" pageId="4" pageNumber="436">
Genus
<taxonomicName id="4C4AE9B0FF803362866E395AFDA5FB92" authority="Boucomont, 1920" authorityName="Boucomont" authorityYear="1920" box="[269,589,1042,1064]" class="Insecta" family="Scarabaeidae" genus="Copridaspidus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B93E4E21FF803362866E395AFE71FB9D" box="[269,409,1042,1064]" italics="true" pageId="4" pageNumber="436">Copridaspidus</emphasis>
Boucomont, 1920
</taxonomicName>
</paragraph>
<paragraph id="8BF59233FF80336287CC3965FD8EFB19" blockId="4.[151,638,1042,1196]" pageId="4" pageNumber="436">
This monotypic genus contains only the following species, which is rarely collected and known only from disjunct regions of moist savanna from the north and south of the
<collectingCountry id="F35DD2A3FF80336286483935FE82FB27" box="[299,362,1149,1170]" name="Democratic Republic of the Congo" pageId="4" pageNumber="436">Congo</collectingCountry>
Basin (
<bibRefCitation id="EFDBEFC2FF80336286CE3935FDA4FB27" author="Davis, A. L. V." box="[429,588,1148,1170]" pageId="4" pageNumber="436" refId="ref3615" refString="Davis, A. L. V., A. V. Frolov, and C. H. Scholtz. 2008. The African Dung Beetle Genera. Protea Book House, Pretoria, South Africa." type="book" year="2008">
Davis
<emphasis id="B93E4E21FF803362868A3935FDFEFB27" box="[489,534,1148,1170]" italics="true" pageId="4" pageNumber="436">et al.</emphasis>
2008
</bibRefCitation>
).The nesting behavior of the genus remains unknown.
</paragraph>
</subSubSection>
</treatment>
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