149 lines
17 KiB
XML
149 lines
17 KiB
XML
<document id="18777A88A600C6B76338FA53198BFA46" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio & Müller, Patrick" docDate="2022" docId="03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus leniae Fanti & Müller, 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="358" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="356" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="FF190B339D107E460364C184766D493C">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="B9B7005C3FD28AB776A30FB5A32E8BC7">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="5060308350BF4A87CAD6C9E1C87E8E95">Müller, Patrick</mods:namePart>
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<mods:typeOfResource id="B30F442901673F5D31CB490A9D72B9D0">text</mods:typeOfResource>
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<mods:title id="5B54EDC86B937A2B60CCC7DDFF38EA61">Baltic Journal of Coleopterology</mods:title>
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<mods:part id="57A7B9C474008CF0B4A9353B57C4CE6E">
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<mods:date id="1F913748548DC64CE39F3AC3FCF2999D">2022</mods:date>
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<mods:detail id="D9C7911A3413E68BC6ECC2D700DDEA11" type="volume">
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<mods:number id="EB01D70EB8AA6CFFF8CA2B38C59E38B4">22</mods:number>
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<mods:number id="1808CED4CAE37ECBDAC45EE4E218037A">2</mods:number>
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<mods:identifier id="479B3D1762D88CEB057DD1E0AB26D5D5" type="DOI">10.5281/zenodo.13203160</mods:identifier>
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<mods:identifier id="DD058A6E33F0AFEFEA0996232FF5CCAB" type="ISSN">1407-8619</mods:identifier>
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<treatment id="03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1" ID-DOI="http://doi.org/10.5281/zenodo.13199821" ID-Zenodo-Dep="13199821" LSID="urn:lsid:plazi:treatment:03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1" httpUri="http://treatment.plazi.org/id/03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1" lastPageId="27" lastPageNumber="358" pageId="25" pageNumber="356">
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<subSubSection id="C36937B6FFBCFFF3FFEBBE8A5EEFA2BF" box="[113,425,951,976]" pageId="25" pageNumber="356" type="nomenclature">
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<paragraph id="8BCC643DFFBCFFF3FFEBBE8A5EEFA2BF" blockId="25.[113,425,951,1007]" box="[113,425,951,976]" pageId="25" pageNumber="356">
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<heading id="D084D351FFBCFFF3FFEBBE8A5EEFA2BF" bold="true" box="[113,425,951,976]" fontSize="10" level="2" pageId="25" pageNumber="356" reason="4">
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<emphasis id="B907B82FFFBCFFF3FFEBBE8A5EEFA2BF" bold="true" box="[113,425,951,976]" pageId="25" pageNumber="356">
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<taxonomicName id="4C731FBEFFBCFFF3FFEBBE8A5E08A2A0" authorityName="Fanti & Müller" authorityYear="2022" box="[113,334,952,976]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="356" phylum="Arthropoda" rank="species" species="leniae" status="sp. nov.">
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<emphasis id="B907B82FFFBCFFF3FFEBBE8A5E08A2A0" bold="true" box="[113,334,952,976]" italics="true" pageId="25" pageNumber="356">Sanaungulus leniae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFBCFFF3FECEBE855EEFA2BF" box="[340,425,951,975]" pageId="25" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFBCFFF3FFEBBEEA5F88A29F" box="[113,206,984,1007]" pageId="25" pageNumber="356" type="description">
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<paragraph id="8BCC643DFFBCFFF3FFEBBEEA5F88A29F" blockId="25.[113,425,951,1007]" box="[113,206,984,1007]" pageId="25" pageNumber="356">
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(
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<figureCitation id="134878B8FFBCFFF3FFE3BEEA5F80A29F" box="[121,198,984,1007]" captionStart="Fig" captionStartId="26.[151,190,1638,1661]" captionTargetBox="[262,1066,151,1605]" captionTargetId="figure-49@26.[262,1066,636,1118]" captionTargetPageId="26" captionText="Fig. 14. Sanaungulus leniae sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of right metathoracic leg and right antenna." figureDoi="http://doi.org/10.5281/zenodo.13203205" httpUri="https://zenodo.org/record/13203205/files/figure.png" pageId="25" pageNumber="356">Fig. 14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFBCFFF3FFEBB93D5ED6A428" pageId="25" pageNumber="356" type="materials_examined">
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<paragraph id="8BCC643DFFBCFFF3FFEBB93D5EFBA5D9" blockId="25.[113,605,1039,1193]" pageId="25" pageNumber="356">
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<materialsCitation id="3B1B6E60FFBCFFF3FFEBB93D5EFBA5D9" collectionCode="BUB" county="Sex" location="Catalog" municipality="Burmese" pageId="25" pageNumber="356" specimenCode="BUB4315" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="B907B82FFFBCFFF3FFEBB93D5FA6A557" bold="true" box="[113,224,1039,1063]" pageId="25" pageNumber="356">
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<typeStatus id="54C8DA9FFFBCFFF3FFEBB93D5F9DA557" box="[113,219,1039,1063]" pageId="25" pageNumber="356" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62AD1CB1FFBCFFF3FF7DB93D5E49A556" box="[231,271,1039,1062]" pageId="25" pageNumber="356">Sex</collectingCounty>
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undefined, adult specimen in a
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<collectingMunicipality id="6BA8FE47FFBCFFF3FFEBB9025F8BA537" box="[113,205,1072,1095]" pageId="25" pageNumber="356">Burmese</collectingMunicipality>
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(
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<collectingRegion id="49B7AADFFFBCFFF3FF43B9025E61A537" box="[217,295,1072,1095]" country="Myanmar" name="Kachin" pageId="25" pageNumber="356">Kachin</collectingRegion>
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) amber piece:
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<location id="8EAC32E6FFBCFFF3FE23B9025D4FA537" LSID="urn:lsid:plazi:treatment:03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1:8EAC32E6FFBCFFF3FE23B9025D4FA537" box="[441,521,1072,1095]" county="Sex" municipality="Burmese" name="Catalog" pageId="25" pageNumber="356" stateProvince="Kachin">Catalog</location>
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number SNHM
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<quantity id="4C8BC9D8FFBCFFF3FF50B9625E67A517" box="[202,289,1104,1127]" metricMagnitude="2" metricUnit="m" metricValue="1.5260319999999998" pageId="25" pageNumber="356" unit="in" value="6008.0">6008 in</quantity>
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the
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<location id="8EAC32E6FFBCFFF3FEC0B9625E8FA5F8" LSID="urn:lsid:plazi:treatment:03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1:8EAC32E6FFBCFFF3FEC0B9625E8FA5F8" county="Sex" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="25" pageNumber="356" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
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(ex
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<location id="8EAC32E6FFBCFFF3FD9AB9435D1AA5F8" LSID="urn:lsid:plazi:treatment:03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1:8EAC32E6FFBCFFF3FD9AB9435D1AA5F8" box="[512,604,1137,1160]" county="Sex" municipality="Burmese" name="Muller's" pageId="25" pageNumber="356" stateProvince="Kachin">Müller’s</location>
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collection number:
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<specimenCode id="DBD5CC46FFBCFFF3FEDAB9A05EF7A5D9" box="[320,433,1170,1193]" collectionCode="BUB" pageId="25" pageNumber="356">BUB4315</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCC643DFFBCFFF3FFEBB9FA5E82A470" blockId="25.[113,605,1224,1280]" pageId="25" pageNumber="356">
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<emphasis id="B907B82FFFBCFFF3FFEBB9FA5E4CA590" bold="true" box="[113,266,1224,1248]" pageId="25" pageNumber="356">Type locality.</emphasis>
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<collectingCountry id="F36424ADFFBCFFF3FE89B9FB5E39A590" box="[275,383,1225,1248]" name="Myanmar" pageId="25" pageNumber="356">Myanmar</collectingCountry>
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:
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<collectingRegion id="49B7AADFFFBCFFF3FE10B9FB5D52A590" box="[394,532,1225,1248]" country="Myanmar" name="Kachin" pageId="25" pageNumber="356">Kachin state</collectingRegion>
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, Myitkyina District, Hukawng Valley.
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</paragraph>
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<paragraph id="8BCC643DFFBCFFF3FFEBB8125ED6A428" blockId="25.[113,605,1312,1368]" pageId="25" pageNumber="356">
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<emphasis id="B907B82FFFBCFFF3FFEBB8125E43A448" bold="true" box="[113,261,1312,1336]" pageId="25" pageNumber="356">Type horizon.</emphasis>
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Lowermost Cenomanian (98.79 ± 0.62 Ma), midCretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFBCFFF3FFEBB84A5FF2A7E4" pageId="25" pageNumber="356" type="diagnosis">
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<paragraph id="8BCC643DFFBCFFF3FFEBB84A5FF2A7E4" blockId="25.[113,605,1400,1684]" pageId="25" pageNumber="356">
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<emphasis id="B907B82FFFBCFFF3FFEBB84A5E2CA4E0" bold="true" box="[113,362,1400,1424]" pageId="25" pageNumber="356">Differential diagnosis.</emphasis>
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The nondehiscent and wide elytra make this new species easily distinguishable from all other
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<taxonomicName id="4C731FBEFFBCFFF3FE0CB8885D66A4A1" authorityName="Fanti, Damgaard & Ellenberger" authorityYear="2018" box="[406,544,1466,1489]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="356" phylum="Arthropoda" rank="genus">
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<emphasis id="B907B82FFFBCFFF3FE0CB8885D66A4A1" box="[406,544,1466,1489]" italics="true" pageId="25" pageNumber="356">Sanaungulus</emphasis>
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</taxonomicName>
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. This character could easily make one think we this is a new genus; unfortunately, the specimen has a crumpled abdomen and we cannot evaluate the length of the elytra. Only the discovery of one or more similar specimens will clarify the taxon.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFBCFFF1FD19BDA95EA8A007" lastPageId="27" lastPageNumber="358" pageId="25" pageNumber="356" type="description">
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<paragraph id="8BCC643DFFBCFFF3FD19BDA95C67A183" blockId="25.[643,1134,155,243]" pageId="25" pageNumber="356">
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<emphasis id="B907B82FFFBCFFF3FD19BDA95C4BA1C3" bold="true" box="[643,781,155,179]" pageId="25" pageNumber="356">Description.</emphasis>
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Adult, winged, rather robust. Sex undefined. Body length
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<quantity id="4C8BC9D8FFBCFFF3FC1CBD895C9AA1A2" box="[902,988,187,210]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="356" unit="mm" value="2.1">2.1 mm</quantity>
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. Entirely reddishbrownish.
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</paragraph>
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<paragraph id="8BCC643DFFBCFFF1FD19BC205EA8A007" blockId="25.[643,1137,274,1685]" lastBlockId="27.[113,605,155,375]" lastPageId="27" lastPageNumber="358" pageId="25" pageNumber="356">Head transverse, wide, slightly convex in the apex, constricted behind the eyes, covered by pronotum, smooth with short pubescence. Eyes prominent, large, convex, roundishsubelliptic, inserted in the lateralupper part of the head, interocular dorsal distance about 3.9 times greater than the eye diameter. Mandibles falciform, slender. Maxillary palps 4 segmented with the last palpomere securiform. Labial palps 3 segmented. Antennae 11 segmented, filiform, short, not reaching the half of elytra, antennal insertion in frons, far from eyes; scape elongated, robust, curved and distorted; antennomere II short, robust, enlarged and globular from the middle to the apex; antennomere III thinner and longer than second; antennomere IV slightly shorter than antennomere III; antennomeres V– VIII filiform and subequal in length; antennomeres IX–XI globularelongated, sturdier than previous ones, with antennomere IX clavate, antennomere X shorter than previous one and antennomere XI rounded at apex; all the antennomeres with very short pubescence. Pronotum transverse, as long as head (eyes are slightly extruded), anterior margin very slightly curved and not bordered, posterior margin almost straight and not bordered, sides strongly curved, corners rounded, pronotum disc smooth and flat andpubescent.Scutellumsmall, triangularshaped, with pointed apex. Elytra elongate, wider than pronotum and head, subparallelsided with apex enlarged, strongly rounded apexes, surface smooth without punctaction, and sligthly pubescent equipped with short and not erected setae. Metathoracic wings transparent, almost completely covered by elytra, as long as elytra or very slightly longer. Sternum very elongated and subquadrate with posterior margin almost straight, sternites transverse, last tergite elongated, narrower than previous ones and rounded at apex. Legs thin, rather long, pubescent; coxae massive, elongated, curved; trochanters elongated, curved, roundish at apex; femora very sturdier than tibiae, nearly straight; tibiae cylindrical, thin, without spurs near the apex, metatibiae longer than metafemora. Tarsal formula 5 5 5; first tarsomere very elongated and thin; second tarsomere about 1.3 times shorter than first, third tarsomere shorter than second; fourth slightly bilobed; fifth tarsomere elongated, thin, flat and curved; claws simple, without denticle at the base.</paragraph>
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</subSubSection>
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<caption id="DF0C34B5FFBFFFF0FF0DBB545C83A7ED" ID-DOI="http://doi.org/10.5281/zenodo.13203205" ID-Zenodo-Dep="13203205" httpUri="https://zenodo.org/record/13203205/files/figure.png" pageId="26" pageNumber="357" startId="26.[151,190,1638,1661]" targetBox="[262,1066,151,1605]" targetPageId="26" targetType="figure">
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<paragraph id="8BCC643DFFBFFFF0FF0DBB545C83A7ED" blockId="26.[151,1172,1638,1693]" pageId="26" pageNumber="357">
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Fig. 14.
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<taxonomicName id="4C731FBEFFBFFFF0FF6ABB545E84A70D" authorityName="Fanti & Müller" authorityYear="2022" box="[240,450,1638,1661]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="357" phylum="Arthropoda" rank="species" species="leniae" status="sp. nov.">
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<emphasis id="B907B82FFFBFFFF0FF6ABB545E84A70D" box="[240,450,1638,1661]" italics="true" pageId="26" pageNumber="357">Sanaungulus leniae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFBFFFF0FE52BB545D5DA70D" box="[456,539,1638,1661]" pageId="26" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
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in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of right metathoracic leg and right antenna.
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</paragraph>
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</caption>
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<subSubSection id="C36937B6FFBEFFF1FFEBBCA55E0AA358" pageId="27" pageNumber="358" type="etymology">
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<paragraph id="8BCC643DFFBEFFF1FFEBBCA55D5EA0BF" blockId="27.[113,605,407,463]" pageId="27" pageNumber="358">
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<emphasis id="B907B82FFFBEFFF1FFEBBCA55FB4A0DF" bold="true" box="[113,242,407,431]" pageId="27" pageNumber="358">Etymology.</emphasis>
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Named in honor of Leni Müller, daughter of the second author (Patrick).
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</paragraph>
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<paragraph id="8BCC643DFFBEFFF1FFEBBCDD5E0AA358" blockId="27.[113,605,495,552]" pageId="27" pageNumber="358">
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<emphasis id="B907B82FFFBEFFF1FFEBBCDD5E52A377" bold="true" box="[113,276,495,519]" pageId="27" pageNumber="358">Syninclusions.</emphasis>
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Two Psocoptera, one
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<taxonomicName id="4C731FBEFFBEFFF1FD9CBCC25D1FA377" box="[518,601,496,519]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
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, air bubbles, detritus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFBEFFF1FFEBBF7A5D1BA3D1" pageId="27" pageNumber="358" type="discussion">
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<paragraph id="8BCC643DFFBEFFF1FFEBBF7A5D1BA3D1" blockId="27.[113,605,584,673]" pageId="27" pageNumber="358">
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<emphasis id="B907B82FFFBEFFF1FFEBBF7A5F98A310" bold="true" box="[113,222,584,608]" pageId="27" pageNumber="358">Remarks.</emphasis>
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Piece of clear, golden amber in Cabochon cut 27x14x
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<quantity id="4C8BC9D8FFBEFFF1FEBCBF5B5E21A3F0" box="[294,359,617,640]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="358" unit="mm" value="5.0">5 mm</quantity>
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. The inclusion is complete, but its abdomen is folded and crumpled.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |