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<document id="027A081ABF2DA51B6DEDA43EDB907A84" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Müller, Patrick" docDate="2022" docId="03DAD52BFF86FFCFFD19B9C95CE8A0AF" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus peteriruedeli Fanti &amp; Müller, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="368" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="366" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="25AAFEAF59C78E79830ABB6F978C4C70">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="F8EBABAC31079DA82925165D6DB6D9A4">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="3326B7238848BD093D470601406C4EE2">Müller, Patrick</mods:namePart>
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<mods:title id="E4B97E687ADE5F3167DC53AFBE704E2A">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="E8F8BA691A3351D0FCBA489B4C814A38">2022</mods:date>
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<mods:number id="334EB881393B126983A882231023FFD5">22</mods:number>
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<treatment id="03DAD52BFF86FFCFFD19B9C95CE8A0AF" LSID="urn:lsid:plazi:treatment:03DAD52BFF86FFCFFD19B9C95CE8A0AF" httpUri="http://treatment.plazi.org/id/03DAD52BFF86FFCFFD19B9C95CE8A0AF" lastPageId="37" lastPageNumber="368" pageId="35" pageNumber="366">
<subSubSection id="C36937B6FF86FFC9FD19B9C95B44A463" box="[643,1026,1275,1299]" pageId="35" pageNumber="366" type="nomenclature">
<paragraph id="8BCC643DFF86FFC9FD19B9C95B44A463" blockId="35.[643,1026,1275,1330]" box="[643,1026,1275,1299]" pageId="35" pageNumber="366">
<heading id="D084D351FF86FFC9FD19B9C95B44A463" bold="true" box="[643,1026,1275,1299]" fontSize="10" level="2" pageId="35" pageNumber="366" reason="4">
<emphasis id="B907B82FFF86FFC9FD19B9C95B44A463" bold="true" box="[643,1026,1275,1299]" pageId="35" pageNumber="366">
<taxonomicName id="4C731FBEFF86FFC9FD19B9C95CE1A463" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[643,935,1275,1299]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="366" phylum="Arthropoda" rank="species" species="peteriruedeli" status="sp. nov.">
<emphasis id="B907B82FFF86FFC9FD19B9C95CE1A463" bold="true" box="[643,935,1275,1299]" italics="true" pageId="35" pageNumber="366">Sanaungulus peteriruedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF86FFC9FC37B9C95B44A463" box="[941,1026,1275,1299]" pageId="35" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF86FFC9FD19B8295DA6A442" box="[643,736,1307,1330]" pageId="35" pageNumber="366" type="description">
<paragraph id="8BCC643DFF86FFC9FD19B8295DA6A442" blockId="35.[643,1026,1275,1330]" box="[643,736,1307,1330]" pageId="35" pageNumber="366">
(
<figureCitation id="134878B8FF86FFC9FD11B8295D9EA442" box="[651,728,1307,1330]" captionStart="Fig" captionStartId="36.[151,190,1407,1430]" captionTargetBox="[151,1172,151,1376]" captionTargetId="figure-39@36.[151,1172,151,1376]" captionTargetPageId="36" captionText="Fig. 20. Sanaungulus peteriruedeli sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13203221" httpUri="https://zenodo.org/record/13203221/files/figure.png" pageId="35" pageNumber="366">Fig. 20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF86FFCEFD19B8605B00A1F0" lastPageId="36" lastPageNumber="367" pageId="35" pageNumber="366" type="materials_examined">
<paragraph id="8BCC643DFF86FFC9FD19B8605C88A49A" blockId="35.[643,1134,1362,1514]" pageId="35" pageNumber="366">
<materialsCitation id="3B1B6E60FF86FFC9FD19B8605C88A49A" collectionCode="BUB" county="Sex" location="Catalog" municipality="Burmese" pageId="35" pageNumber="366" specimenCode="BUB4460" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFF86FFC9FD19B8605DB7A41A" bold="true" box="[643,753,1362,1386]" pageId="35" pageNumber="366">
<typeStatus id="54C8DA9FFF86FFC9FD19B8605DABA41A" box="[643,749,1362,1386]" pageId="35" pageNumber="366" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FF86FFC9FD62B8605C66A419" box="[760,800,1362,1385]" pageId="35" pageNumber="366">Sex</collectingCounty>
undefined, adult specimen in a
<collectingMunicipality id="6BA8FE47FF86FFC9FD19B8405D99A4F9" box="[643,735,1394,1417]" pageId="35" pageNumber="366">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFF86FFC9FD70B8405C7EA4F9" box="[746,824,1394,1417]" country="Myanmar" name="Kachin" pageId="35" pageNumber="366">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FF86FFC9FC50B8405B5CA4F9" LSID="urn:lsid:plazi:treatment:03DAD52BFF86FFCFFD19B9C95CE8A0AF:8EAC32E6FF86FFC9FC50B8405B5CA4F9" box="[970,1050,1394,1417]" county="Sex" municipality="Burmese" name="Catalog" pageId="35" pageNumber="366" stateProvince="Kachin">Catalog</location>
number SNHM­
<quantity id="4C8BC9D8FF86FFC9FD41B8A15C75A4DA" box="[731,819,1427,1450]" metricMagnitude="2" metricUnit="m" metricValue="1.527048" pageId="35" pageNumber="366" unit="in" value="6012.0">6012 in</quantity>
the
<location id="8EAC32E6FF86FFC9FCF1B8A15C9CA4BA" LSID="urn:lsid:plazi:treatment:03DAD52BFF86FFCFFD19B9C95CE8A0AF:8EAC32E6FF86FFC9FCF1B8A15C9CA4BA" county="Sex" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="35" pageNumber="366" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FF86FFC9FB88B8815B28A4BA" LSID="urn:lsid:plazi:treatment:03DAD52BFF86FFCFFD19B9C95CE8A0AF:8EAC32E6FF86FFC9FB88B8815B28A4BA" box="[1042,1134,1459,1482]" county="Sex" municipality="Burmese" name="Muller's" pageId="35" pageNumber="366" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FF86FFC9FCCBB8E15C84A49A" box="[849,962,1491,1514]" collectionCode="BUB" pageId="35" pageNumber="366">BUB4460</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFF86FFC9FD19BB385C93A731" blockId="35.[643,1134,1546,1601]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FD19BB385C5AA752" bold="true" box="[643,796,1546,1570]" pageId="35" pageNumber="366">Type locality.</emphasis>
<collectingCountry id="F36424ADFF86FFC9FCBEBB385CD6A751" box="[804,912,1546,1569]" name="Myanmar" pageId="35" pageNumber="366">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFF86FFC9FC01BB385B63A751" box="[923,1061,1546,1569]" country="Myanmar" name="Kachin" pageId="35" pageNumber="366">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFF86FFC9FD19BB535CE4A7E9" blockId="35.[643,1134,1633,1689]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FD19BB535C50A709" bold="true" box="[643,790,1633,1657]" pageId="35" pageNumber="366">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
<paragraph id="8BCC643DFF81FFCEFF7CBD5F5B00A1F0" blockId="36.[230,1094,109,128]" box="[230,1094,109,128]" pageId="36" pageNumber="367">
<emphasis id="B907B82FFF81FFCEFF7CBD5F5B00A1F0" box="[230,1094,109,128]" italics="true" pageId="36" pageNumber="367">
Fossil
<taxonomicName id="4C731FBEFF81FFCEFEBBBD5F5ECBA1F0" authorityName="Imhoff" authorityYear="1856" box="[289,397,109,128]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="367" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
from the Cretaceous Burmese (
<collectingRegion id="49B7AADFFF81FFCEFD3ABD5F5D99A1F0" box="[672,735,109,128]" country="Myanmar" name="Kachin" pageId="36" pageNumber="367">Kachin</collectingRegion>
) amber of the Patrick Müller collection..
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF0C34B5FF81FFCEFF0DB84D5ED6A4C6" ID-DOI="http://doi.org/10.5281/zenodo.13203221" ID-Zenodo-Dep="13203221" httpUri="https://zenodo.org/record/13203221/files/figure.png" pageId="36" pageNumber="367" startId="36.[151,190,1407,1430]" targetBox="[151,1172,151,1376]" targetPageId="36" targetType="figure">
<paragraph id="8BCC643DFF81FFCEFF0DB84D5ED6A4C6" blockId="36.[151,1172,1407,1462]" pageId="36" pageNumber="367">
Fig. 20.
<taxonomicName id="4C731FBEFF81FFCEFF6CB84D5D52A4E6" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[246,532,1407,1430]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="367" phylum="Arthropoda" rank="species" species="peteriruedeli" status="sp. nov.">
<emphasis id="B907B82FFF81FFCEFF6CB84D5D52A4E6" box="[246,532,1407,1430]" italics="true" pageId="36" pageNumber="367">Sanaungulus peteriruedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF81FFCEFD84B84D5D35A4E6" box="[542,627,1407,1430]" pageId="36" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FF81FFCEFF0DB8DE5EB5A715" pageId="36" pageNumber="367" type="diagnosis">
<paragraph id="8BCC643DFF81FFCEFF0DB8DE5EB5A715" blockId="36.[151,643,1516,1637]" pageId="36" pageNumber="367">
<emphasis id="B907B82FFF81FFCEFF0DB8DE5ED6A774" bold="true" box="[151,400,1516,1540]" pageId="36" pageNumber="367">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFF81FFCEFE0FB8DE5F8DA754" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="367" phylum="Arthropoda" rank="species" species="peteriruedeli" status="sp. nov.">
<emphasis id="B907B82FFF81FFCEFE0FB8DE5F8DA754" italics="true" pageId="36" pageNumber="367">Sanaungulus peteriruedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF81FFCEFF49BB3F5E61A754" box="[211,295,1549,1572]" pageId="36" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by its antennae with very large and robust antennomeres, in particular antennomeres IIIVII.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF81FFCFFD32B8D95F89A7E7" lastPageId="37" lastPageNumber="368" pageId="36" pageNumber="367" type="description">
<paragraph id="8BCC643DFF81FFCEFD32B8D95BD3A752" blockId="36.[680,1173,1515,1570]" pageId="36" pageNumber="367">
<emphasis id="B907B82FFF81FFCEFD32B8D95C75A773" bold="true" box="[680,819,1515,1539]" pageId="36" pageNumber="367">Description.</emphasis>
Adult, winged, rather robust. Sex undefined. Body length
<quantity id="4C8BC9D8FF81FFCEFC06BB395CAAA752" box="[924,1004,1547,1570]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="36" pageNumber="367" unit="mm" value="4.1">4.1 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFF81FFCFFD32BB705F89A7E7" blockId="36.[680,1172,1602,1689]" lastBlockId="37.[113,606,155,1687]" lastPageId="37" lastPageNumber="368" pageId="36" pageNumber="367">Head transverse, constricted (triangular­shaped) behind the eyes, surface irregular. Eyes prominent, very large, roundish, inserted in the upper and lateral part of the head. Mandibles elongated, robust, falciform. Maxillary palps 4­ segmented with the last palpomere strongly elongated and securiform. Labial palps 3­ segmented, with the last palpomere securiform. Antennae 11­ segmented, slightly surpassing half of elytra and not reaching half of abdomen, antennal insertion far from the eyes; scape club­shaped, elongated, robust; antennomere II very short, small, swollen apically; antennomeres IIIIV extremely wide and robust, flat, stouter than scape; antennomeres VVI robust, less wide and slightly longer than previous ones; antennomeres VIIX robust, narrower than previous ones; antennomere XI robust, with rounded apex; all the antennomeres pubescent. Pronotum very elongated, slightly narrower than head, anterior margin undulate and not bordered, posterior margin slightly curved in the middle and slightly bordered, sides straight except for a small tooth near the apex and not bordered, corners obtuse, pronotum disc irregular with large drafts. Scutellum triangular­shaped, wide basally with rounded apex. Elytra short which reveals some abdominal segments, at humeri strongly wider than pronotum, posteriorly parallel­sided and slightly dehiscent apically, rounded apexes, surface smooth (only few and very small punctuation) with pubescence. Metathoracic wings transparent, approximatively as long as elytra. Sternum very elongated and slender, sternites transverse and pubescent, last sternite small, penultimate abdominal segment with a long lobe at the apical corners. Legs thin, long, slightly pubescent; coxae massive; trochanters elongated, robust, with thin and rounded apex; femora curved, cylindrical, slightly sturdier than tibiae; tibiae, thin, subcylindrical, long, with a small spur near the apex, pro­ and mesotibiae slightly longer than pro­ and mesofemora, metatibiae longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 1.1­ 1.2 times longer than second; third tarsomere short, triangular­shaped; fourth strongly bilobed; fifth tarsomere extremely elongated, thin, cylindrical and curved; claws simple, long, pointed, without denticle at the base.</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF80FFCFFD19BDA85B51A038" pageId="37" pageNumber="368" type="etymology">
<paragraph id="8BCC643DFF80FFCFFD19BDA85C68A182" blockId="37.[643,1134,154,242]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19BDA85C45A1C2" bold="true" box="[643,771,154,178]" pageId="37" pageNumber="368">Etymology.</emphasis>
Named in honor of Peter Rüdel, friend of the second author (Patrick), and a great amber collector.
</paragraph>
<paragraph id="8BCC643DFF80FFCFFD19BC235B51A038" blockId="37.[643,1134,273,328]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19BC235C60A059" bold="true" box="[643,806,273,297]" pageId="37" pageNumber="368">Syninclusions.</emphasis>
Detritus, air bubbles (small), a specimen of
<taxonomicName id="4C731FBEFF80FFCFFC90BC035C39A038" authorityName="Handlirsch" authorityYear="1906" box="[778,895,305,328]" class="Insecta" family="Berothidae" kingdom="Animalia" order="Neuroptera" pageId="37" pageNumber="368" phylum="Arthropoda" rank="family">Berothidae</taxonomicName>
(
<taxonomicName id="4C731FBEFF80FFCFFC17BC035B4AA038" box="[909,1036,305,328]" class="Insecta" kingdom="Animalia" order="Neuroptera" pageId="37" pageNumber="368" phylum="Arthropoda" rank="order">Neuroptera</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF80FFCFFD19BC5A5CE8A0AF" pageId="37" pageNumber="368" type="discussion">
<paragraph id="8BCC643DFF80FFCFFD19BC5A5CE8A0AF" blockId="37.[643,1134,360,479]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19BC5A5DB7A0F0" bold="true" box="[643,753,360,384]" pageId="37" pageNumber="368">Remarks.</emphasis>
Piece of brownish Burmese amber that measures 14x9x
<quantity id="4C8BC9D8FF80FFCFFCCBBCBA5CC8A0EF" box="[849,910,392,415]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="368" unit="mm" value="4.0">4 mm</quantity>
. The inclusion is complete and only opacified in the last sternites due to the presence of emulsion.
</paragraph>
</subSubSection>
</treatment>
</document>