361 lines
42 KiB
XML
361 lines
42 KiB
XML
<document id="1D9FA64AC1B22AFD40B6174497F9DD48" ID-DOI="10.11646/zootaxa.4012.1.9" ID-GBIF-Dataset="4c0d8376-71b9-42d1-8dbc-08e0ff16338a" ID-ISSN="1175-5326" ID-Zenodo-Dep="244632" ID-ZooBank="55822610-6FE6-4218-A412-A080F3218F51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461239579310" checkinUser="plazi" docAuthor="Keith, Denis, Sabatinelli, Guido & Uliana, Marco" docDate="2015" docId="03DA87884C3BFFE9FF0FB463FE0A6C0B" docLanguage="en" docName="zt04012p180.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pharaonus (Pharaonus) farsensis Keith, Sabatinelli" docType="treatment" docVersion="8" lastPageNumber="176" masterDocId="FFE3FFF04C3FFFE0FF98B26BFF826853" masterDocTitle="Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa" masterLastPageNumber="180" masterPageNumber="167" pageNumber="171" updateTime="1698605019073" updateUser="plazi">
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<mods:mods id="BAED55FE2E827659C60ABE1DB2109FF4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="12089CD8C3712822491E80A4837DE7F3">
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<mods:title id="09D4CFE658E8EB8A25E85FDC4CB0DBAB">Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa</mods:title>
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<mods:name id="81B954EDCABACA656B86E530DA4D0D8E" type="personal">
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<mods:role id="9F2BB706C350BF8D51177EB2EA4650C8">
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<mods:roleTerm id="A5BE4902238F8F00789B0CA3176AF3BC">Author</mods:roleTerm>
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<mods:namePart id="8A77CA06A14673EE5115B8A886BF81AC">Keith, Denis</mods:namePart>
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<mods:namePart id="A45D5D8AF62A2FC66415C9A230F4060D">Sabatinelli, Guido</mods:namePart>
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</mods:name>
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<mods:name id="62F7395600D3F75BB6724AB3BC0D9C9E" type="personal">
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<mods:roleTerm id="9C2A0EDDD43ED4588BA76066FE76D499">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="82B309B9E0D88A978475745A22DC1A72">Uliana, Marco</mods:namePart>
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<mods:typeOfResource id="D555C06B5D89FB0C61EB2AFE5A6C3CA9">text</mods:typeOfResource>
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<mods:title id="06B9AB1FC8E0403870B857982BA9BB52">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="76097EE737D3FFAB932C0B786CD104F7">
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<mods:date id="A7653E5AB3DDB22DF5A7A8087D0CC066">2015</mods:date>
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<mods:detail id="EFCF134B5B30B91DA27673DE039D6C4D" type="volume">
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<mods:number id="D6A811293617104953E6DD1C39CA688A">4012</mods:number>
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<mods:detail id="02F8D9B04AED256068E2AC031B0F8584" type="issue">
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<mods:number id="1BA5B377C956B79077C1C520F124AD34">1</mods:number>
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<mods:classification id="07BDEA60D95E9E25A803D373FD1BC55C">journal article</mods:classification>
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<mods:identifier id="E54C18228EEA28A921C22531BF14F02E" type="DOI">10.11646/zootaxa.4012.1.9</mods:identifier>
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<mods:identifier id="23461513F58F01D10838020881A86C37" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DA87884C3BFFE9FF0FB463FE0A6C0B" ID-DOI="http://doi.org/10.5281/zenodo.5694706" ID-GBIF-Taxon="119632616" ID-Zenodo-Dep="5694706" LSID="urn:lsid:plazi:treatment:03DA87884C3BFFE9FF0FB463FE0A6C0B" httpUri="http://treatment.plazi.org/id/03DA87884C3BFFE9FF0FB463FE0A6C0B" lastPageId="9" lastPageNumber="176" pageId="4" pageNumber="171">
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<subSubSection id="C36965154C3BFFE4FF0FB463FEC86E10" pageId="4" pageNumber="171" type="nomenclature">
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<paragraph id="8BCC369E4C3BFFE4FF0FB463FBCA6E71" blockId="4.[151,1096,1544,1603]" box="[151,1096,1544,1570]" pageId="4" pageNumber="171">
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<heading id="D08481F24C3BFFE4FF0FB463FBCA6E71" bold="true" box="[151,1096,1544,1570]" fontSize="11" level="1" pageId="4" pageNumber="171" reason="1">
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<emphasis id="B907EA8C4C3BFFE4FF0FB463FBCA6E71" bold="true" box="[151,1096,1544,1570]" pageId="4" pageNumber="171">
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<taxonomicName id="4C734D1D4C3BFFE4FF0FB463FC916E71" authority="Keith, Sabatinelli" authorityName="Keith, Sabatinelli" box="[151,787,1544,1570]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="farsensis" subGenus="Pharaonus">
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<emphasis id="B907EA8C4C3BFFE4FF0FB463FE9F6E71" bold="true" box="[151,285,1544,1570]" italics="true" pageId="4" pageNumber="171">Pharaonus</emphasis>
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(
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<emphasis id="B907EA8C4C3BFFE4FEB5B463FE306E71" bold="true" box="[301,434,1544,1570]" italics="true" pageId="4" pageNumber="171">Pharaonus</emphasis>
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)
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<emphasis id="B907EA8C4C3BFFE4FE5CB463FDB26E71" bold="true" box="[452,560,1544,1570]" italics="true" pageId="4" pageNumber="171">farsensis</emphasis>
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Keith, Sabatinelli
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</taxonomicName>
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, and Uliana, new species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCC369E4C3BFFE4FF0FB441FEC86E10" blockId="4.[151,1096,1544,1603]" box="[151,330,1578,1603]" pageId="4" pageNumber="171">
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(
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<figureCitation id="13482A1B4C3BFFE4FF07B441FF6C6E10" box="[159,238,1578,1603]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1953,1975]" captionTargetBox="[212,1376,196,1915]" captionTargetId="figure@3.[201,1386,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Habitus of Pharaonus (Pharaonus) species. 3: P. adelphus, male paratype (Kerman, Iran). 4: P. adelphus, female paratype (Kerman, Bakram-djord [= Bahramjerd], Iran). 5: P. caucasicus, male (Gorovan sands, Armenia). 6: P. farsensis, male paratype (Darab, Iran)." httpUri="https://zenodo.org/record/244634/files/figure.png" pageId="4" pageNumber="171">Figs. 6</figureCitation>
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,
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<figureCitation id="13482A1B4C3BFFE4FF62B440FE9A6E10" box="[250,280,1579,1603]" captionStart="FIGURES 13 – 14" captionStartId="7.[151,264,1550,1572]" captionTargetBox="[169,1432,195,1512]" captionTargetId="figure@7.[155,1432,195,1528]" captionTargetPageId="7" captionText="FIGURES 13 – 14. Different elytral setation in evidence. 13: P. adelphus. 14: P. farsensis." httpUri="https://zenodo.org/record/244637/files/figure.png" pageId="4" pageNumber="171">14</figureCitation>
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,
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<figureCitation id="13482A1B4C3BFFE4FEBDB440FEC36E10" box="[293,321,1579,1603]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="4" pageNumber="171">18</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36965154C3BFFE4FF0FB419FDE16F48" pageId="4" pageNumber="171" type="diagnosis">
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<paragraph id="8BCC369E4C3BFFE4FF0FB419FDE16F48" blockId="4.[151,1437,1650,2035]" pageId="4" pageNumber="171">
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<emphasis id="B907EA8C4C3BFFE4FF0FB419FE936ED8" bold="true" box="[151,273,1650,1675]" pageId="4" pageNumber="171">Diagnosis.</emphasis>
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a species of
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<taxonomicName id="4C734D1D4C3BFFE4FE31B419FD386ED8" box="[425,698,1650,1675]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="genus">
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<emphasis id="B907EA8C4C3BFFE4FE31B419FDA66ED8" box="[425,548,1650,1675]" italics="true" pageId="4" pageNumber="171">Pharaonus</emphasis>
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(
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<emphasis id="B907EA8C4C3BFFE4FDAEB419FD336ED8" box="[566,689,1650,1675]" italics="true" pageId="4" pageNumber="171">Pharaonus</emphasis>
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)
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</taxonomicName>
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with dark body integuments, except for yellowish-brown elytra. Clypeus strongly transverse (2.1–2.4 times as wide as long), with right angles. Head and pronotum with long, erect pilosity, elytra with scattered long pilosity, more dense on odd intervals. Pronotum impressed along the midline, punctation dense and coarse. Pygidium with a large impunctate area. Antennal club of the male 1.13–1.16 as long as the clypeus width. Protibiae bidentate.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36965154C3BFFE4FF5FB54DFCE56F30" pageId="4" pageNumber="171" type="materials_examined">
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<paragraph id="8BCC369E4C3BFFE4FF5FB54DFCE56F30" blockId="4.[151,1437,1650,2035]" pageId="4" pageNumber="171">
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<emphasis id="B907EA8C4C3BFFE4FF5FB54DFE356F6C" bold="true" box="[199,439,1830,1855]" pageId="4" pageNumber="171">Examined material.</emphasis>
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<typeStatus id="54C8883C4C3BFFE4FE5BB54DFDB26F6C" box="[451,560,1830,1855]" pageId="4" pageNumber="171" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F364760E4C3BFFE4FDA6B54CFDEC6F6D" box="[574,622,1831,1854]" name="Iran" pageId="4" pageNumber="171">Iran</collectingCountry>
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, Darab,
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<date id="FFCD105E4C3BFFE4FD4CB54DFCB16F6C" box="[724,819,1830,1855]" pageId="4" pageNumber="171" value="1960-06">VI.1960</date>
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, s/blé [= on wheat], [leg.] Rémaudière, ♂ (
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<collectionCode id="ED62AE5B4C3BFFE4FAADB54CFA136F6C" LSID="urn:lsid:biocol.org:col:34988" box="[1333,1425,1831,1855]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="171">MNHN</collectionCode>
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).
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<typeStatus id="54C8883C4C3BFFE4FF0FB520FE8B6F30" box="[151,265,1867,1891]" pageId="4" pageNumber="171" type="paratype">Paratypes</typeStatus>
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: same data of the
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<typeStatus id="54C8883C4C3BFFE4FE48B521FDB76F30" box="[464,565,1866,1891]" pageId="4" pageNumber="171" type="holotype">holotype</typeStatus>
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,
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<specimenCount id="9D75FD174C3BFFE4FDD8B520FDE86F31" box="[576,618,1867,1891]" pageId="4" pageNumber="171" type="generic">3 ♂</specimenCount>
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(1
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<collectionCode id="ED62AE5B4C3BFFE4FD17B520FD626F30" box="[655,736,1867,1891]" pageId="4" pageNumber="171">DKCC</collectionCode>
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, 2
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<collectionCode id="ED62AE5B4C3BFFE4FC98B520FCD96F30" LSID="urn:lsid:biocol.org:col:34988" box="[768,859,1867,1891]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="171">MNHN</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36965154C3BFFE9FF5FB505FDC46913" lastPageId="9" lastPageNumber="176" pageId="4" pageNumber="171" type="description">
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<paragraph id="8BCC369E4C3BFFE4FF5FB505FF766FF9" blockId="4.[151,1437,1650,2035]" pageId="4" pageNumber="171">
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<emphasis id="B907EA8C4C3BFFE4FF5FB505FDD46FD4" bold="true" box="[199,598,1902,1927]" pageId="4" pageNumber="171">
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Description of the male
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<typeStatus id="54C8883C4C3BFFE4FE70B505FDD36FD4" box="[488,593,1902,1927]" pageId="4" pageNumber="171" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Length (from the anterior margin of the clypeus to the apex of the elytra)
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<quantity id="4C8B9B7B4C3BFFE4FF0FB5F8FF726FF9" box="[151,240,1939,1963]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="4" pageNumber="171" unit="mm" value="9.2">9.2 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BCC369E4C3BFFE4FF5FB5DDFA996FA0" blockId="4.[151,1437,1650,2035]" pageId="4" pageNumber="171">Head, pronotum and underside black; antenna light brown with apex of antennomere 1 and club darker; legs black with dark brown tarsi; elytra yellowish-brown, darker along the suture and the epipleural margin.</paragraph>
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<caption id="DF0C66164C3AFFE5FF0FB5C8FF6C6FA5" httpUri="https://zenodo.org/record/244635/files/figure.png" pageId="5" pageNumber="172" targetBox="[211,1377,193,1914]" targetPageId="5">
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<paragraph id="8BCC369E4C3AFFE5FF0FB5C8FF6C6FA5" blockId="5.[151,1436,1955,2038]" pageId="5" pageNumber="172">
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<emphasis id="B907EA8C4C3AFFE5FF0FB5C8FECA6FEA" bold="true" box="[151,328,1955,1977]" pageId="5" pageNumber="172">FIGURES 7–10.</emphasis>
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Habitus of
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<taxonomicName id="4C734D1D4C3AFFE5FE5AB5C8FD3B6FEA" box="[450,697,1955,1977]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="genus">
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<emphasis id="B907EA8C4C3AFFE5FE5AB5C8FDB36FEA" box="[450,561,1955,1977]" italics="true" pageId="5" pageNumber="172">Pharaonus</emphasis>
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||
(
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<emphasis id="B907EA8C4C3AFFE5FDD9B5C8FD326FEA" box="[577,688,1955,1977]" italics="true" pageId="5" pageNumber="172">Pharaonus</emphasis>
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)
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</taxonomicName>
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species. 7:
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<emphasis id="B907EA8C4C3AFFE5FCAAB5CFFC4E6FEA" box="[818,972,1955,1977]" italics="true" pageId="5" pageNumber="172">
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<taxonomicName id="4C734D1D4C3AFFE5FCAAB5CFFC4A6FEA" box="[818,968,1955,1977]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="fasciculatus">P. fasciculatus</taxonomicName>
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,
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</emphasis>
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male (Egypt). 8:
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<taxonomicName id="4C734D1D4C3AFFE5FB1AB5CFFB7C6FEA" box="[1154,1278,1955,1977]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="saharicus">
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<emphasis id="B907EA8C4C3AFFE5FB1AB5CFFB7C6FEA" box="[1154,1278,1955,1977]" italics="true" pageId="5" pageNumber="172">P. saharicus</emphasis>
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</taxonomicName>
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, male paratype (Tamrit, Tassili n’Ajjer, Algeria). 9:
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<taxonomicName id="4C734D1D4C3AFFE5FD88B5A9FD2B6F84" box="[528,681,1985,2007]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="varicoloreus">
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<emphasis id="B907EA8C4C3AFFE5FD88B5A9FD2B6F84" box="[528,681,1985,2007]" italics="true" pageId="5" pageNumber="172">P. varicoloreus</emphasis>
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</taxonomicName>
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, male (environs of Palymra, Syria). 10:
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<taxonomicName id="4C734D1D4C3AFFE5FBD5B5A9FB656F84" box="[1101,1255,1985,2007]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="varicoloreus">
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<emphasis id="B907EA8C4C3AFFE5FBD5B5A9FB656F84" box="[1101,1255,1985,2007]" italics="true" pageId="5" pageNumber="172">P. varicoloreus</emphasis>
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</taxonomicName>
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, female (Birecik, Turkey).
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</paragraph>
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</caption>
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<caption id="DF0C66164C39FFE6FF0FB602FC326CCD" httpUri="https://zenodo.org/record/244636/files/figure.png" pageId="6" pageNumber="173" targetBox="[159,1427,207,1105]" targetPageId="6">
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<paragraph id="8BCC369E4C39FFE6FF0FB602FC326CCD" blockId="6.[151,1436,1129,1182]" pageId="6" pageNumber="173">
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<emphasis id="B907EA8C4C39FFE6FF0FB602FED66C2C" bold="true" box="[151,340,1129,1151]" pageId="6" pageNumber="173">FIGURES 11–12.</emphasis>
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Habitus of
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<taxonomicName id="4C734D1D4C39FFE6FE4AB602FD5A6C2C" authority="Tamerlanius" authorityName="Tamerlanius" box="[466,728,1129,1151]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="173" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907EA8C4C39FFE6FE4AB602FDC36C2C" box="[466,577,1129,1151]" italics="true" pageId="6" pageNumber="173">Pharaonus</emphasis>
|
||
(
|
||
<emphasis id="B907EA8C4C39FFE6FDCAB602FD4D6C2C" box="[594,719,1129,1151]" italics="true" pageId="6" pageNumber="173">Tamerlanius</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species. 11:
|
||
<emphasis id="B907EA8C4C39FFE6FCF9B601FC366C2C" box="[865,948,1129,1151]" italics="true" pageId="6" pageNumber="173">P. l e de r i</emphasis>
|
||
, male (Koron 15 km Ekoc. Dzejran, Bukhara, Uzbekistan). 12:
|
||
<taxonomicName id="4C734D1D4C39FFE6FEDCB6E2FE436CCD" box="[324,449,1161,1182]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="173" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="B907EA8C4C39FFE6FEDCB6E2FE436CCD" box="[324,449,1161,1182]" italics="true" pageId="6" pageNumber="173">P. semenowi</emphasis>
|
||
</taxonomicName>
|
||
, male (Katta-kum, Turkmenkuduk, Uzbekistan).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC369E4C39FFE6FF5FB6AEFB756D89" blockId="6.[151,1437,1221,2038]" pageId="6" pageNumber="173">Head: clypeus very transverse (2.3 times as wide as long), almost rectangular, with margins distinctly raised. Anterior margin straight; anterior angles right, broadly rounded. Integument glabrous with large, shallow, irregular punctures; defining a coarse and reticulate sculpture. Frontoclypeal suture almost invisible but marked by a dense tuft of long, yellowish gray, erect pilosity. Frons with sculpture similar to that of clypeus, and with pilosity becoming shorter and sparser going from the frontoclypeal suture towards the vertex. Ocular canthus strong; eyes small, not strongly protruding. Antennae with 9 antennomeres; club with 3 antennomeres, 2.0 times longer than funicle (antennomeres 2–6) and 1.12 times as long as clypeus width. Scape noticeably enlarged at the apex; antennomere 2 wider than long; antennomeres 3, 4, 5 increasing in length; antennomere 6 discoidal.</paragraph>
|
||
<paragraph id="8BCC369E4C39FFE6FF5FB78DFC2F6E85" blockId="6.[151,1437,1221,2038]" pageId="6" pageNumber="173">Pronotum transverse, 1.45 times wider than long. Sides subparallel in the basal two thirds, strongly convergent in the anterior third; slightly concave in the basal portion, almost straight in the distal one. Anterior angles acute, posterior angles rounded and right. Base broadly truncate in front of scutellum. Anterior margin unconspicuous; lateral and basal margins distinct, basal margins broader laterally and visible in front of the scutellum. Punctation dense, forming irregular series of rough punctures, open dorsally. Surface with long, greyish-yellow, erect pilosity. Median, longitudinal furrow weakly defined over the entire length. Scutellum semi-elliptical, with sculpture and pilosity similar to those of the pronotum, punctures evenly distributed.</paragraph>
|
||
<paragraph id="8BCC369E4C39FFE6FF5FB48AFDD96FA5" blockId="6.[151,1437,1221,2038]" pageId="6" pageNumber="173">Elytra quite flat with 7 poorly defined striae of punctures, with sparse punctures over entire surface, punctures less dense than on the pronotum. First (sutural) interval raised over entire length. Humeral callus strong, more finely punctate than the surrounding integument. Apical callus also distinct, but weaker. Surface with sparse, raised setae; the longest setae being about half the length of the pronotal setae; setae denser and forming weak rows on interstriae 3, 5, 7. Epipleural margin broad and inflated under the humeral callus, then flattened and narrowing until the middle of the elytra; from the middle of the elytra to the apex again with relief in the form of a narrow border. Membranous margin of the epipleura gradually widening from the basal third to the apex, where it is conspicuous. Suture prolonged in a short apical spine.</paragraph>
|
||
<caption id="DF0C66164C38FFE7FF0FB465FBA26E77" box="[151,1056,1550,1572]" httpUri="https://zenodo.org/record/244637/files/figure.png" pageId="7" pageNumber="174" targetBox="[169,1432,195,1512]" targetPageId="7">
|
||
<paragraph id="8BCC369E4C38FFE7FF0FB465FBA26E77" blockId="7.[151,1056,1550,1572]" box="[151,1056,1550,1572]" pageId="7" pageNumber="174">
|
||
<emphasis id="B907EA8C4C38FFE7FF0FB465FED16E77" bold="true" box="[151,339,1550,1572]" pageId="7" pageNumber="174">FIGURES 13–14.</emphasis>
|
||
Different elytral setation in evidence. 13:
|
||
<taxonomicName id="4C734D1D4C38FFE7FD67B464FCF66E77" box="[767,884,1550,1572]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C38FFE7FD67B464FCF66E77" box="[767,884,1550,1572]" italics="true" pageId="7" pageNumber="174">P. adelphus</emphasis>
|
||
</taxonomicName>
|
||
. 14:
|
||
<taxonomicName id="4C734D1D4C38FFE7FC3FB464FB986E77" box="[935,1050,1550,1572]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="B907EA8C4C38FFE7FC3FB464FB986E77" box="[935,1050,1550,1572]" italics="true" pageId="7" pageNumber="174">P. farsensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC369E4C38FFE7FF5FB43AFEE56E86" blockId="7.[151,1437,1617,2038]" pageId="7" pageNumber="174">Propygidium punctate over entire surface, without apical fringe of pilosity. Pygidium convex, with a glabrous central band; central band smooth at the base and sparsely punctate on its apical half. Distinct punctate triangular area on either side of central band bearing a patch of almost adpressed, greyish-white, long pilosity. Apical margin with erect pilosity.</paragraph>
|
||
<paragraph id="8BCC369E4C38FFE7FF5FB48AFEFD6FD9" blockId="7.[151,1437,1617,2038]" pageId="7" pageNumber="174">Protibiae bidentate, basal tooth obtuse, apical tooth slightly curved. Apical inner spur short, inconspicuous. Protarsomeres 1–4 slightly and gradually widened. Protarsomere 5 approximately of the same length as protarsomeres 1–4 combined. Inner claw of protarsomeres broad, bifid; outer claw narrow. Mesotibiae and metatibiae clearly enlarged in the middle, with two rows of spiniform setae. Claws of mesotarsi and metatarsi simple and elongate.</paragraph>
|
||
<paragraph id="8BCC369E4C38FFE7FF5FB5FEFED26F81" blockId="7.[151,1437,1617,2038]" pageId="7" pageNumber="174">Labrum appearing weakly bilobate. Terminal maxillary palpomere elongate, fusiform, with a sulcus in the basal two thirds.</paragraph>
|
||
<paragraph id="8BCC369E4C38FFE7FF5FB5B6FE4C6FA5" blockId="7.[151,1437,1617,2038]" box="[199,462,2013,2038]" pageId="7" pageNumber="174">
|
||
Aedeagus as in
|
||
<figureCitation id="13482A1B4C38FFE7FEEFB5B6FE486FA5" box="[375,458,2013,2038]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="7" pageNumber="174">Fig. 18</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF0C66164C37FFE8FF0FB55CFC746FB5" httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="8" pageNumber="175" targetBox="[174,1402,241,1792]" targetPageId="8">
|
||
<paragraph id="8BCC369E4C37FFE8FF0FB55CFC746FB5" blockId="8.[151,1436,1847,2022]" pageId="8" pageNumber="175">
|
||
<emphasis id="B907EA8C4C37FFE8FF0FB55CFED16F1E" bold="true" box="[151,339,1847,1869]" pageId="8" pageNumber="175">FIGURES 15–23.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName id="4C734D1D4C37FFE8FE4FB55CFDC46F1E" box="[471,582,1847,1869]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907EA8C4C37FFE8FE4FB55CFDC46F1E" box="[471,582,1847,1869]" italics="true" pageId="8" pageNumber="175">Pharaonus</emphasis>
|
||
</taxonomicName>
|
||
. 15–19: aedeagus in dorsal, ventral, and lateral view of
|
||
<taxonomicName id="4C734D1D4C37FFE8FBE2B55CFAF36F1E" box="[1146,1393,1847,1869]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907EA8C4C37FFE8FBE2B55CFB6B6F1E" box="[1146,1257,1847,1869]" italics="true" pageId="8" pageNumber="175">Pharaonus</emphasis>
|
||
(
|
||
<emphasis id="B907EA8C4C37FFE8FB6FB55CFAE46F1E" box="[1271,1382,1847,1869]" italics="true" pageId="8" pageNumber="175">Pharaonus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
: 15:
|
||
<taxonomicName id="4C734D1D4C37FFE8FF0FB53CFE886F38" box="[151,266,1877,1900]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C37FFE8FF0FB53CFE886F38" box="[151,266,1877,1900]" italics="true" pageId="8" pageNumber="175">P. adelphus</emphasis>
|
||
</taxonomicName>
|
||
, paratype (a-c) and holotype (d) (Kerman, Iran); 16:
|
||
<taxonomicName id="4C734D1D4C37FFE8FCB9B53CFC2B6F3F" box="[801,937,1879,1900]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="caucasicus">
|
||
<emphasis id="B907EA8C4C37FFE8FCB9B53CFC2B6F3F" box="[801,937,1879,1900]" italics="true" pageId="8" pageNumber="175">P. caucasicus</emphasis>
|
||
</taxonomicName>
|
||
(Gorovan, Armenia); 17:
|
||
<taxonomicName id="4C734D1D4C37FFE8FB28B53CFA1A6F38" authority="(Egypt)" baseAuthorityName="Egypt" box="[1200,1432,1877,1900]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="fasciculatus">
|
||
<emphasis id="B907EA8C4C37FFE8FB28B53CFAC06F38" box="[1200,1346,1877,1900]" italics="true" pageId="8" pageNumber="175">P. fasciculatus</emphasis>
|
||
(Egypt)
|
||
</taxonomicName>
|
||
; 18:
|
||
<taxonomicName id="4C734D1D4C37FFE8FF5AB51EFEB86FD9" box="[194,314,1908,1930]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="B907EA8C4C37FFE8FF5AB51EFEB86FD9" box="[194,314,1908,1930]" italics="true" pageId="8" pageNumber="175">P. farsensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype (Darab, Iran); 19:
|
||
<emphasis id="B907EA8C4C37FFE8FDEEB51EFD7E6FD9" box="[630,764,1908,1930]" italics="true" pageId="8" pageNumber="175">
|
||
<taxonomicName id="4C734D1D4C37FFE8FDEEB51EFD7B6FD9" box="[630,761,1908,1930]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="saharicus">P. saharicus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
paratype (Tamrit, Tassili n’Ajjer, Algeria); 20:
|
||
<taxonomicName id="4C734D1D4C37FFE8FB67B51EFA1E6FD9" box="[1279,1436,1908,1930]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="varicoloreus">
|
||
<emphasis id="B907EA8C4C37FFE8FB67B51EFA1E6FD9" box="[1279,1436,1908,1930]" italics="true" pageId="8" pageNumber="175">P. varicoloreus</emphasis>
|
||
</taxonomicName>
|
||
(Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of
|
||
<taxonomicName id="4C734D1D4C37FFE8FCD1B5FFFC3E6FFA" box="[841,956,1939,1961]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C37FFE8FCD1B5FFFC3E6FFA" box="[841,956,1939,1961]" italics="true" pageId="8" pageNumber="175">P. adelphus</emphasis>
|
||
</taxonomicName>
|
||
(Kerman, Bakram-djord [= Bahramjerd], Iran). 22–23: aedeagus in dorsal, ventral, and lateral view of
|
||
<taxonomicName id="4C734D1D4C37FFE8FD57B5D9FC556F9B" authority="(Tamerlanius)" authorityName="Tamerlanius" baseAuthorityName="Tamerlanius" box="[719,983,1970,1992]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907EA8C4C37FFE8FD57B5D9FCBC6F9B" box="[719,830,1970,1992]" italics="true" pageId="8" pageNumber="175">Pharaonus</emphasis>
|
||
(
|
||
<emphasis id="B907EA8C4C37FFE8FCD6B5D9FC4E6F9B" box="[846,972,1970,1992]" italics="true" pageId="8" pageNumber="175">Tamerlanius</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
: 22:
|
||
<taxonomicName id="4C734D1D4C37FFE8FB94B5D8FBE36F9B" box="[1036,1121,1970,1992]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="lederi">
|
||
<emphasis id="B907EA8C4C37FFE8FB94B5D8FBE36F9B" box="[1036,1121,1970,1992]" italics="true" pageId="8" pageNumber="175">P. lederi</emphasis>
|
||
</taxonomicName>
|
||
(Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23:
|
||
<taxonomicName id="4C734D1D4C37FFE8FE3EB5BAFDA16FB5" box="[422,547,2001,2022]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="B907EA8C4C37FFE8FE3EB5BAFDA16FB5" box="[422,547,2001,2022]" italics="true" pageId="8" pageNumber="175">P. semenowi</emphasis>
|
||
</taxonomicName>
|
||
(Kul'dzhuktau, Kizilkum desert, Uzbekistan).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC369E4C36FFE9FF5FB2FCFE1F68E3" blockId="9.[151,1436,151,1112]" box="[199,413,151,176]" pageId="9" pageNumber="176">
|
||
<emphasis id="B907EA8C4C36FFE9FF5FB2FCFE9C68E3" bold="true" box="[199,286,151,176]" pageId="9" pageNumber="176">Female</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C36FFE9FF5FB2D7FDC46913" blockId="9.[151,1436,151,1112]" pageId="9" pageNumber="176">
|
||
<emphasis id="B907EA8C4C36FFE9FF5FB2D7FECF6886" bold="true" box="[199,333,188,213]" pageId="9" pageNumber="176">Variability.</emphasis>
|
||
Size ranging between
|
||
<quantity id="4C8B9B7B4C36FFE9FDCEB2D7FD606887" box="[598,738,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" metricValueMax="9.3" metricValueMin="9.1" pageId="9" pageNumber="176" unit="mm" value="9.2" valueMax="9.3" valueMin="9.1">9.1–9.3 mm</quantity>
|
||
; clypeus 2.1–2.4 times as wide as long, antennal club of the male 1.13–1.16 as long as the clypeus width. Scutellum with punctures evenly distributed or with punctures smaller and denser laterally. Protibiae basal tooth either acute or obtuse. No other relevant variation was observed on the small number of examined specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C36FFE9FF5FB327FB2A6937" box="[199,1192,332,357]" pageId="9" pageNumber="176" type="etymology">
|
||
<paragraph id="8BCC369E4C36FFE9FF5FB327FB2A6937" blockId="9.[151,1436,151,1112]" box="[199,1192,332,357]" pageId="9" pageNumber="176">
|
||
<emphasis id="B907EA8C4C36FFE9FF5FB327FECC6936" bold="true" box="[199,334,332,357]" pageId="9" pageNumber="176">Etymology.</emphasis>
|
||
Based on the Iranian province of
|
||
<collectingCountry id="F364760E4C36FFE9FD51B326FD7E6937" box="[713,764,333,356]" name="Iran" pageId="9" pageNumber="176">Fars</collectingCountry>
|
||
, where this species is known to occur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C36FFE9FF5FB304FE0A6C0B" pageId="9" pageNumber="176" type="discussion">
|
||
<paragraph id="8BCC369E4C36FFE9FF5FB304FC336A6E" blockId="9.[151,1436,151,1112]" pageId="9" pageNumber="176">
|
||
<emphasis id="B907EA8C4C36FFE9FF5FB304FECE69DB" bold="true" box="[199,332,367,392]" pageId="9" pageNumber="176">Discussion.</emphasis>
|
||
Based on their habitus, the two new species are close to
|
||
<taxonomicName id="4C734D1D4C36FFE9FBA7B304FABE69DB" box="[1087,1340,367,392]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="caucasicus">
|
||
<emphasis id="B907EA8C4C36FFE9FBA7B304FABE69DB" box="[1087,1340,367,392]" italics="true" pageId="9" pageNumber="176">Pharaonus caucasicus</emphasis>
|
||
</taxonomicName>
|
||
and to
|
||
<taxonomicName id="4C734D1D4C36FFE9FF0FB3FEFEBA69FE" box="[151,312,404,429]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="fasciculatus">
|
||
<emphasis id="B907EA8C4C36FFE9FF0FB3FEFEBA69FE" box="[151,312,404,429]" italics="true" pageId="9" pageNumber="176">P. fasciculatus</emphasis>
|
||
</taxonomicName>
|
||
. From the former, they can be easily distinguished by the punctation, which is much stronger and rougher over the entire dorsal surface, with coalescent punctures (finer and not coalescent in
|
||
<taxonomicName id="4C734D1D4C36FFE9FB4CB3D2FAE86983" box="[1236,1386,441,464]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="caucasicus">
|
||
<emphasis id="B907EA8C4C36FFE9FB4CB3D2FAE86983" box="[1236,1386,441,464]" italics="true" pageId="9" pageNumber="176">P. caucasicus</emphasis>
|
||
</taxonomicName>
|
||
). In addition, the two new species have the proximal tooth of the protibiae quite similar to the distal one (broader and less elongate in
|
||
<taxonomicName id="4C734D1D4C36FFE9FED3B06AFE606A4B" box="[331,482,513,536]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="caucasicus">
|
||
<emphasis id="B907EA8C4C36FFE9FED3B06AFE606A4B" box="[331,482,513,536]" italics="true" pageId="9" pageNumber="176">P. caucasicus</emphasis>
|
||
</taxonomicName>
|
||
), and the longitudinal glabrous band of the pygidium much more developed and less punctate than
|
||
<taxonomicName id="4C734D1D4C36FFE9FEACB04EFE486A6F" box="[308,458,549,572]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="caucasicus">
|
||
<emphasis id="B907EA8C4C36FFE9FEACB04EFE486A6F" box="[308,458,549,572]" italics="true" pageId="9" pageNumber="176">P. caucasicus</emphasis>
|
||
</taxonomicName>
|
||
, and with thinner patches of setae laterally.
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C36FFE9FF5FB02CFBB86AF4" blockId="9.[151,1436,151,1112]" pageId="9" pageNumber="176">
|
||
<taxonomicName id="4C734D1D4C36FFE9FF5FB02CFE2F6A33" box="[199,429,583,608]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C36FFE9FF5FB02CFE2F6A33" box="[199,429,583,608]" italics="true" pageId="9" pageNumber="176">Pharaonus adelphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C734D1D4C36FFE9FE7BB022FDE06A33" box="[483,610,584,608]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="B907EA8C4C36FFE9FE7BB022FDE06A33" box="[483,610,584,608]" italics="true" pageId="9" pageNumber="176">P. farsensis</emphasis>
|
||
</taxonomicName>
|
||
are very similar to
|
||
<taxonomicName id="4C734D1D4C36FFE9FCA3B022FC5E6A33" box="[827,988,583,608]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="fasciculatus">
|
||
<emphasis id="B907EA8C4C36FFE9FCA3B022FC5E6A33" box="[827,988,583,608]" italics="true" pageId="9" pageNumber="176">P. fasciculatus</emphasis>
|
||
</taxonomicName>
|
||
, and are distinct mainly by the presence of a smooth region in the medial area of the pygidium and by the presence of a longitudinal impression along the midline of the pronotum. In addition, the protarsi and the inner proclaw are stouter.
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C36FFE9FF5FB0DFFE0A6C0B" blockId="9.[151,1436,151,1112]" pageId="9" pageNumber="176">
|
||
<taxonomicName id="4C734D1D4C36FFE9FF5FB0DFFE306A9E" box="[199,434,692,717]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="aldephus">
|
||
<emphasis id="B907EA8C4C36FFE9FF5FB0DFFE306A9E" box="[199,434,692,717]" italics="true" pageId="9" pageNumber="176">Pharaonus aldephus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C734D1D4C36FFE9FE6AB0DEFDF36A9F" box="[498,625,692,716]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="B907EA8C4C36FFE9FE6AB0DEFDF36A9F" box="[498,625,692,716]" italics="true" pageId="9" pageNumber="176">P. farsensis</emphasis>
|
||
</taxonomicName>
|
||
are very similar each other and occur in close proximity (their
|
||
<typeStatus id="54C8883C4C36FFE9FAF4B0DEFA1E6A9E" box="[1388,1436,693,717]" pageId="9" pageNumber="176">type</typeStatus>
|
||
localities in southeastern
|
||
<collectingCountry id="F364760E4C36FFE9FE27B0B3FE6D6ABC" box="[447,495,728,751]" name="Iran" pageId="9" pageNumber="176">Iran</collectingCountry>
|
||
, are separated by a distance of about
|
||
<quantity id="4C8B9B7B4C36FFE9FC2AB0B3FB8D6AA3" box="[946,1039,727,752]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="176" unit="km" value="300.0">300 km</quantity>
|
||
), nevertheless the two species are clearly distinct by the different state of elytral pilosity (see
|
||
<figureCitation id="13482A1B4C36FFE9FCCFB097FC626B47" box="[855,992,764,789]" captionStart="FIGURES 13 – 14" captionStartId="7.[151,264,1550,1572]" captionTargetBox="[169,1432,195,1512]" captionTargetId="figure@7.[155,1432,195,1528]" captionTargetPageId="7" captionText="FIGURES 13 – 14. Different elytral setation in evidence. 13: P. adelphus. 14: P. farsensis." httpUri="https://zenodo.org/record/244637/files/figure.png" pageId="9" pageNumber="176">Figs. 13–14</figureCitation>
|
||
) and by the different lengths of male antennal club. The latter difference appears trivial, but is statistically significant: a Student’s
|
||
<emphasis id="B907EA8C4C36FFE9FB38B14AFB2A6B6B" box="[1184,1192,801,824]" italics="true" pageId="9" pageNumber="176">t</emphasis>
|
||
-test calculated on the ratio length of antennal club/width of clypeus of the available males (two-tailed test for groups with unequal variance). The results were
|
||
<emphasis id="B907EA8C4C36FFE9FE53B102FE5E6BD3" box="[459,476,873,896]" italics="true" pageId="9" pageNumber="176">P</emphasis>
|
||
= 0.0012, which strongly support the hypothesis that there is a difference between the two groups. A further slight difference was noted in the different degree of inflation of the sutural interval, which appear more raised in
|
||
<taxonomicName id="4C734D1D4C36FFE9FE0BB1DAFD906B9B" box="[403,530,944,968]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="B907EA8C4C36FFE9FE0BB1DAFD906B9B" box="[403,530,944,968]" italics="true" pageId="9" pageNumber="176">P. farsensis</emphasis>
|
||
</taxonomicName>
|
||
, in particular in the basal half. This character may be just intraspecific variation and needs to be tested using larger samples. The parameres of
|
||
<taxonomicName id="4C734D1D4C36FFE9FCC0B1BEFC5A6BBF" box="[856,984,980,1004]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="B907EA8C4C36FFE9FCC0B1BEFC5A6BBF" box="[856,984,980,1004]" italics="true" pageId="9" pageNumber="176">P. farsensis</emphasis>
|
||
</taxonomicName>
|
||
appear to be more slender than those of
|
||
<taxonomicName id="4C734D1D4C36FFE9FF0FB192FE956C43" box="[151,279,1015,1040]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C36FFE9FF0FB192FE956C43" box="[151,279,1015,1040]" italics="true" pageId="9" pageNumber="176">P. adelphus</emphasis>
|
||
</taxonomicName>
|
||
in lateral view. However the intraspecific variation observed in the small sample of
|
||
<taxonomicName id="4C734D1D4C36FFE9FB52B192FAC86C43" box="[1226,1354,1015,1040]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C36FFE9FB52B192FAC86C43" box="[1226,1354,1015,1040]" italics="true" pageId="9" pageNumber="176">P. adelphus</emphasis>
|
||
</taxonomicName>
|
||
appear considerable in comparison with the interspecific variation, suggesting caution in the evaluation of this character (see
|
||
<figureCitation id="13482A1B4C36FFE9FF53B654FEA46C0B" box="[203,294,1087,1112]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="9" pageNumber="176">Figs. 15</figureCitation>
|
||
c, d, 18).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |