247 lines
30 KiB
XML
247 lines
30 KiB
XML
<document id="BE76A56102B90F56B5FC315382248980" ID-CLB-Dataset="29014" ID-DOI="10.11646/zootaxa.4459.3.3" ID-GBIF-Dataset="7cf2f2b2-90cc-46f2-a511-9000a6c927ac" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458796" ID-ZooBank="E764FED3-CFC9-4120-8A72-917D46D1EDED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539289892809" checkinUser="plazi" docAuthor="Kovačić, Marcelo, Bogorodsky, Sergey V., Mal, Ahmad O. & Alpermann, Tilman J." docDate="2018" docId="03DA4E03FFDE7C03FF03F947B0A0FD4A" docLanguage="en" docName="zootaxa.4459.3.3.pdf" docOrigin="Zootaxa 4459 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Koumansetta Whitley 1940" docType="treatment" docVersion="8" lastPageNumber="456" masterDocId="FFE3367BFFDC7C07FF94FFAEB20FFFBA" masterDocTitle="Redescription of the genus Koumansetta (Teleostei: Gobiidae), with description of a new species from the Red Sea" masterLastPageNumber="481" masterPageNumber="453" pageNumber="455" updateTime="1698578740455" updateUser="ExternalLinkService">
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<mods:title id="60860839AAEE7439227CB2BB5796F0BF">Redescription of the genus Koumansetta (Teleostei: Gobiidae), with description of a new species from the Red Sea</mods:title>
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<mods:name id="5A314F8EF1F61A7D50772F5A159D4EF4" type="personal">
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<mods:namePart id="2E04F82B5BC873FD8199CF34B6A52E5B">Kovačić, Marcelo</mods:namePart>
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<mods:roleTerm id="DE24778D9B59AEA627A565979ED0AF40">Author</mods:roleTerm>
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<mods:namePart id="520122796109CA2C22B3F5AC23CEB160">Bogorodsky, Sergey V.</mods:namePart>
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<mods:roleTerm id="7EE9828F2C20ED98FD681FC99784830E">Author</mods:roleTerm>
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<mods:namePart id="F0407D9F832FA949C072898210A25813">Mal, Ahmad O.</mods:namePart>
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<mods:name id="880EB261B863CE5A5F7821431ACA05B6" type="personal">
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<mods:roleTerm id="8C2BECC2A4C611432599E759EF5E8E8B">Author</mods:roleTerm>
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<mods:namePart id="FD96B5DB5795FAB62CB9F9755D7003F6">Alpermann, Tilman J.</mods:namePart>
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<mods:typeOfResource id="AA43F460F42A1AEBD1A183ACD4838C3C">text</mods:typeOfResource>
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<mods:title id="070CB61694AB575E2074B4AB771D2BA6">Zootaxa</mods:title>
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<mods:part id="BC219DB8C0F88D8254A1F02A32BB43F8">
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<mods:date id="5014E37FA72A607FFB2C1131BF4D7316">2018</mods:date>
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<mods:number id="D61049BC0B30416BB861C8AE69FEBDA2">2018-08-16</mods:number>
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<mods:number id="8BED5C9A9908469A9D3CC7DA310F09F7">4459</mods:number>
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<treatment id="03DA4E03FFDE7C03FF03F947B0A0FD4A" ID-DOI="http://doi.org/10.5281/zenodo.5959372" ID-GBIF-Taxon="148692815" ID-Zenodo-Dep="5959372" LSID="urn:lsid:plazi:treatment:03DA4E03FFDE7C03FF03F947B0A0FD4A" httpUri="http://treatment.plazi.org/id/03DA4E03FFDE7C03FF03F947B0A0FD4A" lastPageId="4" lastPageNumber="456" pageId="2" pageNumber="455">
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<subSubSection id="C369AC9EFFDE7C05FF03F947B3ECF8B9" box="[151,483,1769,1795]" pageId="2" pageNumber="455" type="nomenclature">
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<paragraph id="8BCCFF15FFDE7C05FF03F947B3ECF8B9" blockId="2.[151,483,1769,1795]" box="[151,483,1769,1795]" pageId="2" pageNumber="455">
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<heading id="D0844879FFDE7C05FF03F947B3ECF8B9" bold="true" box="[151,483,1769,1795]" fontSize="11" level="1" pageId="2" pageNumber="455" reason="1">
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<taxonomicName id="4C738496FFDE7C05FF03F947B3ECF8B9" ID-CoL="592H" ID-ENA="1647184" authority="Whitley 1940" authorityName="Whitley" authorityYear="1940" box="[151,483,1769,1795]" class="Actinopterygii" family="Gobiidae" genus="Koumansetta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="455" phylum="Chordata" rank="genus">
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<emphasis id="B9072307FFDE7C05FF03F947B3ECF8B9" bold="true" box="[151,483,1769,1795]" pageId="2" pageNumber="455">
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<emphasis id="B9072307FFDE7C05FF03F947B339F8B9" bold="true" box="[151,310,1769,1795]" italics="true" pageId="2" pageNumber="455">Koumansetta</emphasis>
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<bibRefCitation id="EFE282E4FFDE7C05FEA9F947B3ECF8B9" box="[317,483,1769,1795]" pageId="2" pageNumber="455" refId="ref19135" refString="Whitley, G. P. (1940) Illustrations of some Australian fishes. Australian Zoologist, 9 (4), 397 - 428." type="journal article">Whitley 1940</bibRefCitation>
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</emphasis>
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</taxonomicName>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C369AC9EFFDE7C05FF03F89FB705F8DE" pageId="2" pageNumber="455" type="reference_group">
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<paragraph id="8BCCFF15FFDE7C05FF03F89FB705F8DE" blockId="2.[151,1401,1840,1892]" pageId="2" pageNumber="455">
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<treatmentCitationGroup id="AB63D83BFFDE7C05FF03F89FB705F8DE" pageId="2" pageNumber="455">
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<taxonomicName id="4C738496FFDE7C05FF03F89FB3ECF8FC" ID-CoL="592H" authority="Whitley 1940: 425" authorityName="Whitley" authorityPageNumber="425" authorityYear="1940" box="[151,483,1840,1862]" class="Actinopterygii" family="Gobiidae" genus="Koumansetta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="455" phylum="Chordata" rank="genus">
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<emphasis id="B9072307FFDE7C05FF03F89FB312F8FC" box="[151,285,1841,1862]" italics="true" pageId="2" pageNumber="455">Koumansetta</emphasis>
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<treatmentCitation id="0AD2D904FFDE7C05FEB7F89EB3ECF8FC" author="" box="[291,483,1840,1862]" page="425" pageId="2" pageNumber="455">
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<bibRefCitation id="EFE282E4FFDE7C05FEB7F89EB3BEF8FC" box="[291,433,1840,1862]" pageId="2" pageNumber="455" refId="ref19135" refString="Whitley, G. P. (1940) Illustrations of some Australian fishes. Australian Zoologist, 9 (4), 397 - 428." type="journal article">Whitley 1940</bibRefCitation>
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: 425
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</treatmentCitation>
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</taxonomicName>
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,
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<typeStatus id="54C841B7FFDE7C05FE79F89FB016F8FC" box="[493,537,1841,1862]" pageId="2" pageNumber="455">type</typeStatus>
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species
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<taxonomicName id="4C738496FFDE7C05FDFAF89FB1E7F8FC" ID-CoL="3RDR2" authority="Whitley 1940" authorityName="Whitley" authorityYear="1940" box="[622,1000,1840,1862]" class="Actinopterygii" family="Gobiidae" genus="Koumansetta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="455" phylum="Chordata" rank="species" species="rainfordi">
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<emphasis id="B9072307FFDE7C05FDFAF89FB15AF8FC" box="[622,853,1840,1862]" italics="true" pageId="2" pageNumber="455">Koumansetta rainfordi</emphasis>
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<bibRefCitation id="EFE282E4FFDE7C05FCCFF89EB1E7F8FC" box="[859,1000,1840,1862]" pageId="2" pageNumber="455" refId="ref19135" refString="Whitley, G. P. (1940) Illustrations of some Australian fishes. Australian Zoologist, 9 (4), 397 - 428." type="journal article">Whitley 1940</bibRefCitation>
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</taxonomicName>
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, by original designation and monotypy.
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<taxonomicName id="4C738496FFDE7C05FF03F8E0B3BBF8DE" authority="Smith 1957: 726" authorityName="Smith" authorityPageNumber="726" authorityYear="1957" box="[151,436,1870,1892]" class="Actinopterygii" family="Gobiidae" genus="Seychellea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="455" phylum="Chordata" rank="genus">
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<emphasis id="B9072307FFDE7C05FF03F8E0B30CF8DE" box="[151,259,1870,1892]" italics="true" pageId="2" pageNumber="455">Seychellea</emphasis>
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<treatmentCitation id="0AD2D904FFDE7C05FE9DF8E0B3BBF8DE" author="" box="[265,436,1870,1892]" page="726" pageId="2" pageNumber="455">
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<bibRefCitation id="EFE282E4FFDE7C05FE9DF8E0B38DF8DE" box="[265,386,1870,1892]" pageId="2" pageNumber="455" refId="ref18898" refString="Smith, J. L. B. (1957) The fishes of Aldabra. Part V. Annals and Magazine of Natural History, Series 12, 9 (106), Art. 93, 721 - 729." type="journal article">Smith 1957</bibRefCitation>
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: 726
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</treatmentCitation>
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||
</taxonomicName>
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,
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<typeStatus id="54C841B7FFDE7C05FE2BF8E1B3E4F8DE" box="[447,491,1871,1892]" pageId="2" pageNumber="455">type</typeStatus>
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species
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<taxonomicName id="4C738496FFDE7C05FDD4F8E0B177F8DE" ID-CoL="4X3NT" authority="Smith 1957" authorityName="Smith" authorityYear="1957" box="[576,888,1870,1892]" class="Actinopterygii" family="Gobiidae" genus="Seychellea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="455" phylum="Chordata" rank="species" species="hectori">
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<emphasis id="B9072307FFDE7C05FDD4F8E0B0F4F8DE" box="[576,763,1870,1892]" italics="true" pageId="2" pageNumber="455">Seychellea hectori</emphasis>
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<bibRefCitation id="EFE282E4FFDE7C05FC95F8E0B177F8DE" box="[769,888,1870,1892]" pageId="2" pageNumber="455" refId="ref18898" refString="Smith, J. L. B. (1957) The fishes of Aldabra. Part V. Annals and Magazine of Natural History, Series 12, 9 (106), Art. 93, 721 - 729." type="journal article">Smith 1957</bibRefCitation>
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</taxonomicName>
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, by original designation and monotypy.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C369AC9EFFDE7C04FF03F83FB138FD3F" lastPageId="3" lastPageNumber="456" pageId="2" pageNumber="455" type="diagnosis">
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<paragraph id="8BCCFF15FFDE7C04FF03F83FB0F0FEDF" blockId="2.[151,1436,1937,2037]" lastBlockId="3.[151,1436,151,645]" lastPageId="3" lastPageNumber="456" pageId="2" pageNumber="455">
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<emphasis id="B9072307FFDE7C05FF03F83FB305F810" bold="true" box="[151,266,1937,1962]" pageId="2" pageNumber="455">Diagnosis</emphasis>
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. (1) pelvic fins united to form a complete pointed pelvic disc; (2) cephalic suborbital row
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<emphasis id="B9072307FFDE7C05FA9AF83CB713F813" box="[1294,1308,1938,1961]" italics="true" pageId="2" pageNumber="455">a</emphasis>
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absent; (3) oculoscapular transverse rows
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<emphasis id="B9072307FFDE7C05FE62F817B018F86A" box="[502,535,1977,2000]" italics="true" pageId="2" pageNumber="455">trp</emphasis>
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and
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<emphasis id="B9072307FFDE7C05FDC6F817B07CF86A" box="[594,627,1977,2000]" italics="true" pageId="2" pageNumber="455">tra</emphasis>
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long, extending dorsally well above level of rows
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<date id="FFCDD9D5FFDE7C05FB54F817B72BF879" box="[1216,1316,1973,2001]" pageId="2" pageNumber="455">
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<emphasis id="B9072307FFDE7C05FB54F817B6DAF879" box="[1216,1237,1973,2000]" italics="true" pageId="2" pageNumber="455">x1</emphasis>
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and
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<emphasis id="B9072307FFDE7C05FA84F817B72BF879" box="[1296,1316,1973,2000]" italics="true" pageId="2" pageNumber="455">x2</emphasis>
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</date>
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; (4) snout pointed, prominent, long,
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<quantity id="4C8B52F0FFDE7C05FE5CF872B031F84F" box="[456,574,2012,2037]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.7779999999999998" pageId="2" pageNumber="455" unit="in" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 in</quantity>
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eye diameter, 2.8–3.0 in head length, with gently sloping dorsal profile, overhanging mouth; (5) mouth subterminal; (6) the upper limb of first gill arch with 1–2 slender, weak and soft gill rakers anteriorly, followed by 1–5 short, also soft, broad structures (
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<figureCitation id="1348E390FFDF7C04FC0FFF12B1D2FF6E" box="[923,989,188,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1493,1515]" captionTargetBox="[151,1436,700,1469]" captionTargetId="figure@3.[151,1436,700,1469]" captionTargetPageId="3" captionText="FIGURE 1. Gill rakers and structures of Koumansetta hoesei sp. nov.: A) gill rakers on lower limb, B) gill rakers on upper limb, C) gill structures on upper limb. Non-type material, SMF 35872, 32.4 + 8.3 mm, Al Lith, Saudi Arabia." httpUri="https://zenodo.org/record/1458798/files/figure.png" pageId="3" pageNumber="456">Fig. 1</figureCitation>
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); (7) first two dorsal-fin spines longest and elongate, remaining spines progressively shorter; (8) pelvic frenum absent; (9) color mostly brown to browngreen with narrow yellow or orange longitudinal stripes on body and head and two black spots posteriorly, one ocellated, partly or completely on the second dorsal fin, and another spot dorsoposteriorly on caudal peduncle; (10) scales mostly ctenoid, predorsal area scaled at midline.
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</paragraph>
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<paragraph id="8BCCFF15FFDF7C04FF53FEC1B138FD3F" blockId="3.[151,1436,151,645]" pageId="3" pageNumber="456">
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This diagnosis distinguishes members of the genus
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<taxonomicName id="4C738496FFDF7C04FC87FEDFB1A8FE32" authorityName="Whitley" authorityYear="1940" box="[787,935,369,392]" class="Actinopterygii" family="Gobiidae" genus="Koumansetta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">
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<emphasis id="B9072307FFDF7C04FC87FEDFB1A8FE32" box="[787,935,369,392]" italics="true" pageId="3" pageNumber="456">Koumansetta</emphasis>
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</taxonomicName>
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from 15 valid species presently assigned to
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<taxonomicName id="4C738496FFDF7C04FF03FE3AB326FE17" box="[151,297,404,429]" class="Actinopterygii" family="Gobiidae" genus="Amblygobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">
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<emphasis id="B9072307FFDF7C04FF03FE3AB326FE17" box="[151,297,404,429]" italics="true" pageId="3" pageNumber="456">Amblygobius</emphasis>
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</taxonomicName>
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, and from two other genera of the “
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<taxonomicName id="4C738496FFDF7C04FD4BFE3AB17DFE17" authorityName="Bleeker" authorityYear="1856" box="[735,882,404,429]" class="Actinopterygii" family="Gobiidae" genus="Valenciennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">
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||
<emphasis id="B9072307FFDF7C04FD4BFE3AB17DFE17" box="[735,882,404,429]" italics="true" pageId="3" pageNumber="456">Valenciennea</emphasis>
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</taxonomicName>
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-lineage” (
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<taxonomicName id="4C738496FFDF7C04FC64FE3AB67BFE17" authorityName="Hoese & Allen" authorityYear="1977" box="[1008,1140,404,429]" class="Actinopterygii" family="Gobiidae" genus="Signigobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">
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<emphasis id="B9072307FFDF7C04FC64FE3AB67BFE17" box="[1008,1140,404,429]" italics="true" pageId="3" pageNumber="456">Signigobius</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C738496FFDF7C04FB20FE3AB746FE17" authorityName="Bleeker" authorityYear="1856" box="[1204,1353,404,429]" class="Actinopterygii" family="Gobiidae" genus="Valenciennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">
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<emphasis id="B9072307FFDF7C04FB20FE3AB746FE17" box="[1204,1353,404,429]" italics="true" pageId="3" pageNumber="456">Valenciennea</emphasis>
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</taxonomicName>
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) sensu Agorretta
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<emphasis id="B9072307FFDF7C04FE9AFE17B34BFE6A" box="[270,324,439,464]" italics="true" pageId="3" pageNumber="456">et al</emphasis>
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. (2013). Characters (1) and (2) distinguishes the genus from
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<emphasis id="B9072307FFDF7C04FB8BFE19B6B4FE6A" box="[1055,1211,439,464]" italics="true" pageId="3" pageNumber="456">
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<taxonomicName id="4C738496FFDF7C04FB8BFE19B6B8FE6A" authorityName="Bleeker" authorityYear="1856" box="[1055,1207,439,464]" class="Actinopterygii" family="Gobiidae" genus="Valenciennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">Valenciennea</taxonomicName>
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,
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</emphasis>
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character (2) from
|
||
<taxonomicName id="4C738496FFDF7C04FF03FE72B314FE4F" authorityName="Hoese & Allen" authorityYear="1977" box="[151,283,476,501]" class="Actinopterygii" family="Gobiidae" genus="Signigobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9072307FFDF7C04FF03FE72B314FE4F" box="[151,283,476,501]" italics="true" pageId="3" pageNumber="456">Signigobius</emphasis>
|
||
</taxonomicName>
|
||
. The diagnosis was confirmed to be different compared to all comparative material examined of 13 valid species presently assigned to
|
||
<taxonomicName id="4C738496FFDF7C04FDB5FE51B0BCFDA2" box="[545,691,511,536]" class="Actinopterygii" family="Gobiidae" genus="Amblygobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9072307FFDF7C04FDB5FE51B0BCFDA2" box="[545,691,511,536]" italics="true" pageId="3" pageNumber="456">Amblygobius</emphasis>
|
||
</taxonomicName>
|
||
and compared to published data in the species descriptions of two additional recently described
|
||
<taxonomicName id="4C738496FFDF7C04FE60FD8AB089FD87" box="[500,646,548,573]" class="Actinopterygii" family="Gobiidae" genus="Amblygobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9072307FFDF7C04FE60FD8AB089FD87" box="[500,646,548,573]" italics="true" pageId="3" pageNumber="456">Amblygobius</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="EFE282E4FFDF7C04FD6DFD8AB618FD87" box="[761,1047,548,573]" pageId="3" pageNumber="456" refId="ref16283" refString="Allen, G. R. & Erdmann, M. V. (2016) Descriptions of two new gobies (Gobiidae: Amblygobius) from the tropical western Pacific Ocean. Journal of the Ocean Science Foundation, 24, 10 - 23. https: // doi. org / 10.5281 / zenodo. 167891" type="journal article">Allen & Erdmann 2016</bibRefCitation>
|
||
) in characters (3)–(9) (
|
||
<tableCitation id="C6F1CAAEFFDF7C04FAA5FD8AB782FD86" box="[1329,1421,548,573]" captionStart="TABLE 3" captionStartId="21.[147,223,616,638]" captionText="TABLE 3. Diagnostic characters of Koumansetta compared to Amblygobius species." pageId="3" pageNumber="456">Table 3</tableCitation>
|
||
). Character (10) distinguishes
|
||
<taxonomicName id="4C738496FFDF7C04FE78FDE7B08FFDDA" authorityName="Whitley" authorityYear="1940" box="[492,640,585,608]" class="Actinopterygii" family="Gobiidae" genus="Koumansetta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9072307FFDF7C04FE78FDE7B08FFDDA" box="[492,640,585,608]" italics="true" pageId="3" pageNumber="456">Koumansetta</emphasis>
|
||
</taxonomicName>
|
||
only from "
|
||
<taxonomicName id="4C738496FFDF7C04FC8EFDE9B625FDDA" baseAuthorityName="Herre" baseAuthorityYear="1945" box="[794,1066,583,608]" class="Actinopterygii" family="Gobiidae" genus="Amblygobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="species" species="nocturnus">
|
||
<emphasis id="B9072307FFDF7C04FC8EFDE9B625FDDA" box="[794,1066,583,608]" italics="true" pageId="3" pageNumber="456">Amblygobius nocturnus</emphasis>
|
||
</taxonomicName>
|
||
species group”
|
||
<emphasis id="B9072307FFDF7C04FB7BFDE7B722FDDA" box="[1263,1325,585,608]" italics="true" pageId="3" pageNumber="456">sensu</emphasis>
|
||
<bibRefCitation id="EFE282E4FFDF7C04FAAFFDE9B340FD3F" pageId="3" pageNumber="456" refId="ref16283" refString="Allen, G. R. & Erdmann, M. V. (2016) Descriptions of two new gobies (Gobiidae: Amblygobius) from the tropical western Pacific Ocean. Journal of the Ocean Science Foundation, 24, 10 - 23. https: // doi. org / 10.5281 / zenodo. 167891" type="journal article">Allen & Erdmann (2016)</bibRefCitation>
|
||
and
|
||
<taxonomicName id="4C738496FFDF7C04FE11FDC2B13DFD3F" authority="(Playfair 1867)" baseAuthorityName="Playfair" baseAuthorityYear="1867" box="[389,818,620,645]" class="Actinopterygii" family="Gobiidae" genus="Amblygobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="species" species="sewardii">
|
||
<emphasis id="B9072307FFDF7C04FE11FDC2B071FD3F" box="[389,638,620,645]" italics="true" pageId="3" pageNumber="456">Amblygobius sewardii</emphasis>
|
||
(Playfair 1867)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0CAF9DFFDF7C04FF03FA7BB6E4F9B0" httpUri="https://zenodo.org/record/1458798/files/figure.png" pageId="3" pageNumber="456" startId="3.[151,250,1493,1515]" targetBox="[151,1436,700,1469]" targetPageId="3">
|
||
<paragraph id="8BCCFF15FFDF7C04FF03FA7BB6E4F9B0" blockId="3.[151,1436,1493,1546]" pageId="3" pageNumber="456">
|
||
<emphasis id="B9072307FFDF7C04FF03FA7BB31BFA51" bold="true" box="[151,276,1493,1515]" pageId="3" pageNumber="456">FIGURE 1.</emphasis>
|
||
Gill rakers and structures of
|
||
<taxonomicName id="4C738496FFDF7C04FDD2FA78B11BFA51" authority="Kovačić & Bogorodsky & Mal & Alpermann, 2018" authorityName="Kovačić & Bogorodsky & Mal & Alpermann" authorityYear="2018" box="[582,788,1493,1515]" class="Actinopterygii" family="Gobiidae" genus="Koumansetta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="456" phylum="Chordata" rank="species" species="hoesei" status="sp. nov.">
|
||
<emphasis id="B9072307FFDF7C04FDD2FA78B11BFA51" box="[582,788,1493,1515]" italics="true" pageId="3" pageNumber="456">Koumansetta hoesei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2349E7CFFDF7C04FC88FA7BB161FA51" box="[796,878,1493,1515]" pageId="3" pageNumber="456" rank="species">
|
||
<emphasis id="B9072307FFDF7C04FC88FA7BB161FA51" bold="true" box="[796,878,1493,1515]" pageId="3" pageNumber="456">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: A) gill rakers on lower limb, B) gill rakers on upper limb, C) gill structures on upper limb. Non-type material, SMF 35872, 32.4 + 8.3 mm, Al Lith, Saudi Arabia.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C369AC9EFFDF7C03FF53F99FB0A0FD4A" lastPageId="4" lastPageNumber="457" pageId="3" pageNumber="456" type="description">
|
||
<paragraph id="8BCCFF15FFDF7C03FF53F99FB0A0FD4A" blockId="3.[151,1437,1585,2006]" lastBlockId="4.[151,1437,151,752]" lastPageId="4" lastPageNumber="457" pageId="3" pageNumber="456">
|
||
<emphasis id="B9072307FFDF7C04FF53F99FB35EF9F0" bold="true" box="[199,337,1585,1610]" pageId="3" pageNumber="456">Description</emphasis>
|
||
. Body moderately elongate, laterally compressed. Depth at anal-fin origin 4.8 (4.5–5.2) in SL. Head slightly laterally compressed. Snout pointed, prominent, long,
|
||
<quantity id="4C8B52F0FFDF7C04FC29F9F8B63CF9D4" box="[957,1075,1621,1646]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.7779999999999998" pageId="3" pageNumber="456" unit="in" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 in</quantity>
|
||
eye diameter, 2.8–3.0 in head length, with gently sloping dorsal profile, overhanging mouth. Eyes lateral, interorbital broad. Anterior nostril short, tubular, erect, without process from rim; posterior nostril slit-like, with a low, raised ridge, closer to anterior nostril than to anterior margin of eye. Mouth subterminal, moderately small, oblique. Posterior angle of jaws below anterior edge of eye or slightly in front or behind the vertical. Teeth in upper and lower jaws unicuspid and pointed, more or less curved inward. Upper jaw with caniniform teeth of similar size at the front, followed with inner row of smaller slender sharp teeth laterally; lower jaw with an outer row of caniniform teeth of similar size at the front ending laterally with single large curved canine and the inner row of smaller slender sharp teeth. Tongue reduced, adnate, truncate in front. Chin without fold or barbels. Cranial roof covered by dorsal axial musculature. Gill openings lateral, ending forward at pectoral-fin base. Branchiostegal membranes attached to entire lateral margin of isthmus. Gill rakers poorly developed, rakers on lower limb of anterior arch slender, weak and soft,
|
||
<quantity id="4C8B52F0FFDF7C04FAC6F813B793F86C" box="[1362,1436,1981,2006]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="3" pageNumber="456" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
|
||
number, the upper limb of first gill arch with 1–2 slender, weak and soft gill rakers anteriorly, followed by 1–5 short, also soft, broad structures (
|
||
<figureCitation id="1348E390FFD87C03FDB6FF12B067FF6E" box="[546,616,188,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1493,1515]" captionTargetBox="[151,1436,700,1469]" captionTargetId="figure@3.[151,1436,700,1469]" captionTargetPageId="3" captionText="FIGURE 1. Gill rakers and structures of Koumansetta hoesei sp. nov.: A) gill rakers on lower limb, B) gill rakers on upper limb, C) gill structures on upper limb. Non-type material, SMF 35872, 32.4 + 8.3 mm, Al Lith, Saudi Arabia." httpUri="https://zenodo.org/record/1458798/files/figure.png" pageId="4" pageNumber="457">Fig. 1</figureCitation>
|
||
). Two dorsal fins, first fin with VI spines, first two dorsal-fin spines elongate, remaining spines progressively shorter, second fin
|
||
<date id="FFCDD9D5FFD87C03FCC0FF71B1A1FF42" box="[852,942,223,248]" bridgedPair="/" pageId="4" pageNumber="457">I/15–17</date>
|
||
; anal fin
|
||
<date id="FFCDD9D5FFD87C03FB8BFF71B671FF42" box="[1055,1150,223,248]" bridgedPair="/" pageId="4" pageNumber="457">I/15–17.</date>
|
||
Pectoral-fin rays 15–18. Pectoral girdle without flaps on anterior edge. Uppermost rays of pectoral fin within membrane. Pelvic fin united to form a complete pelvic disc, pointed. Pelvic frenum absent. Caudal fin rounded, shorter than head length, with 13 branched and 16–17 segmented rays. Body covered with small, mainly ctenoid scales,
|
||
<quantity id="4C8B52F0FFD87C03FB06FEE2B6F6FEDF" box="[1170,1273,332,357]" metricMagnitude="0" metricUnit="m" metricValue="1.3969999999999998" metricValueMax="1.5493999999999999" metricValueMin="1.2446000000000002" pageId="4" pageNumber="457" unit="in" value="55.0" valueMax="61.0" valueMin="49.0">49–61 in</quantity>
|
||
lateral series; transverse scale series 14–19. Predorsal area, including nape, breast and belly with cycloid scales, the area between first dorsal-fin base and pectoral fin also with cycloid scales.
|
||
<collectingRegion id="49B731F7FFD87C03FCD8FE3BB161FE16" box="[844,878,405,428]" country="Maldives" name="Uthuru" pageId="4" pageNumber="457">No</collectingRegion>
|
||
scales on cheek and opercle (a few small cycloid scales occasionally present on upper opercle). Predorsal area scaled, scales not reaching the vertical through the posterior edge of eye, predorsal scales 16–26. Head with anterior oculoscapular canal with pores B’, C (single), D (single), E, F, G, and H’; posterior oculoscapular canal with K’ and L’; preopercular canal with M’, N, and O’. Suborbital rows of sensory papillae: row
|
||
<emphasis id="B9072307FFD87C03FDFFFD8BB076FD86" box="[619,633,549,572]" italics="true" pageId="4" pageNumber="457">a</emphasis>
|
||
absent, row
|
||
<emphasis id="B9072307FFD87C03FC9FFD8AB116FD87" box="[779,793,548,573]" italics="true" pageId="4" pageNumber="457">b</emphasis>
|
||
longitudinal, short, beginning anteriorly below posterior part of eye, not reaching above row
|
||
<emphasis id="B9072307FFD87C03FDD3FDE9B05AFDDA" box="[583,597,583,608]" italics="true" pageId="4" pageNumber="457">d</emphasis>
|
||
, row
|
||
<emphasis id="B9072307FFD87C03FD09FDE7B0A6FDDA" box="[669,681,585,608]" italics="true" pageId="4" pageNumber="457">c</emphasis>
|
||
as 10–16 transverse sub
|
||
<emphasis id="B9072307FFD87C03FC5FFDE9B1DBFDDA" bold="true" box="[971,980,583,608]" pageId="4" pageNumber="457">-</emphasis>
|
||
orbital rows of sensory papillae, row
|
||
<emphasis id="B9072307FFD87C03FA1AFDE9B793FDDA" box="[1422,1436,583,608]" italics="true" pageId="4" pageNumber="457">d</emphasis>
|
||
longitudinal. Oculoscapular transverse rows
|
||
<emphasis id="B9072307FFD87C03FD00FDC3B0BAFD3E" box="[660,693,621,644]" italics="true" pageId="4" pageNumber="457">trp</emphasis>
|
||
and
|
||
<emphasis id="B9072307FFD87C03FD64FDC3B11EFD3E" box="[752,785,621,644]" italics="true" pageId="4" pageNumber="457">tra</emphasis>
|
||
long, extending dorsally well above level of rows
|
||
<emphasis id="B9072307FFD87C03FAC9FDC3B766FD3E" box="[1373,1385,621,644]" italics="true" pageId="4" pageNumber="457">x</emphasis>
|
||
.
|
||
<collectingRegion id="49B731F7FFD87C03FAEEFDC3B793FD3E" box="[1402,1436,621,644]" country="Maldives" name="Uthuru" pageId="4" pageNumber="457">No</collectingRegion>
|
||
interorbital papillae. Head and body brown, reddish brown to greenish brown or brownish to greenish grey with narrow yellow or orange longitudinal stripes, black ocellus entirely on or partially on the second dorsal fin, and black spot dorsoposteriorly on caudal peduncle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |