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<document id="4A7BA02D268B7113F2A5CD2F32EEE7BF" ID-DOI="10.11646/zootaxa.4103.4.5" ID-GBIF-Dataset="c49bfd75-07b9-4df8-ad33-e52ebce75cfe" ID-ISSN="1175-5326" ID-Zenodo-Dep="266659" ID-ZooBank="040A6B20-353A-4A58-9771-FB1B50402323" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460617596129" checkinUser="plazi" docAuthor="Couri, Márcia &amp; Pont, Adrian" docDate="2016" docId="03D987DC7909E866FF0CAA9430F22C24" docLanguage="en" docName="zootaxa.4103.4.5.pdf.imf" docOrigin="Zootaxa 4103 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helina mediomaculata Couri &amp; Pont, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="379" masterDocId="FFE0FFA4790CE863FF9BAA03326C2B56" masterDocTitle="New African species of Helina Robineau-Desvoidy (Diptera, Muscidae)" masterLastPageNumber="382" masterPageNumber="374" pageNumber="379" updateTime="1698332138420" updateUser="plazi">
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<mods:titleInfo id="2CC1ADD8E2102A259432CCA8D98575A1">
<mods:title id="D30A754F09F9607E44828B6B7E840A47">New African species of Helina Robineau-Desvoidy (Diptera, Muscidae)</mods:title>
</mods:titleInfo>
<mods:name id="375478335D5B5CDD97530ED039C67EE9" type="personal">
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<mods:namePart id="4A30C9F88DDDF1EE63438F0CA2F2CB4E">Couri, Márcia</mods:namePart>
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<mods:namePart id="F071173BECF7DDBBB9973CE500DE96B3">Pont, Adrian</mods:namePart>
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<mods:typeOfResource id="5A0825867D4E748745D44EEF602FCC18">text</mods:typeOfResource>
<mods:relatedItem id="E5554DFC729FFC0825A967EA2AC3A934" type="host">
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<mods:title id="2A7C05B0FEB6334203B1E97901F0BB91">Zootaxa</mods:title>
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<mods:part id="0CC64246318F8099265E43D244D8063F">
<mods:date id="4E3C2815CF65B772CDAE67585307ADB3">2016</mods:date>
<mods:detail id="F26A6C5DB96A2FC79E5C46246C8CAC55" type="volume">
<mods:number id="CAA186B66197026FAB6E2BC84D865245">4103</mods:number>
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<mods:number id="4F4D0A8786554F1587AFAC1C571B5003">4</mods:number>
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<mods:identifier id="D49A7BD456A3C6000956C6DD3CB7DC06" type="DOI">10.11646/zootaxa.4103.4.5</mods:identifier>
<mods:identifier id="99C068A9A26DCEF4645E13CD6C754487" type="GBIF-Dataset">c49bfd75-07b9-4df8-ad33-e52ebce75cfe</mods:identifier>
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<treatment id="03D987DC7909E866FF0CAA9430F22C24" ID-DOI="http://doi.org/10.5281/zenodo.6087901" ID-GBIF-Taxon="119569088" ID-Zenodo-Dep="6087901" LSID="urn:lsid:plazi:treatment:03D987DC7909E866FF0CAA9430F22C24" httpUri="http://treatment.plazi.org/id/03D987DC7909E866FF0CAA9430F22C24" lastPageNumber="379" pageId="5" pageNumber="379">
<subSubSection id="C36A65417909E866FF0CAA9433252B85" pageId="5" pageNumber="379" type="nomenclature">
<paragraph id="8BCF36CA7909E866FF0CAA9430632BE4" blockId="5.[151,527,151,211]" box="[151,527,151,178]" pageId="5" pageNumber="379">
<heading id="D08781A67909E866FF0CAA9430632BE4" bold="true" box="[151,527,151,178]" fontSize="11" level="1" pageId="5" pageNumber="379" reason="1">
<emphasis id="B904EAD87909E866FF0CAA9430632BE4" bold="true" box="[151,527,151,178]" pageId="5" pageNumber="379">
<taxonomicName id="4C704D497909E866FF0CAA9433C62BE7" box="[151,426,151,177]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="mediomaculata" status="sp. nov.">
<emphasis id="B904EAD87909E866FF0CAA9433C62BE7" bold="true" box="[151,426,151,177]" italics="true" pageId="5" pageNumber="379">Helina mediomaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23757A37909E866FE29AA9B30632BE4" box="[434,527,152,178]" pageId="5" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36CA7909E866FF0CAAB933252B85" blockId="5.[151,527,151,211]" box="[151,329,186,211]" pageId="5" pageNumber="379">
(
<figureCitation id="134B2A4F7909E866FF04AAB932812B85" box="[159,237,186,211]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1937,1960]" captionTargetBox="[211,1373,210,1890]" captionTargetId="figure@2.[180,1392,193,1899]" captionTargetPageId="2" captionText="FIGURES 1 4. Dorsal view of holotypes: (1) Helina duocolorata sp. nov., holotype male; (2) Helina longicerca sp. nov., holotype male; (3) Helina mediomaculata sp. nov., holotype male; (4) Helina sexnotata sp. nov., holotype female." httpUri="https://zenodo.org/record/266660/files/figure.png" pageId="5" pageNumber="379">Figs. 3</figureCitation>
,
<figureCitation id="134B2A4F7909E866FF61AAB9332D2B85" box="[250,321,186,211]" captionStart="FIGURES 13 20" captionStartId="6.[151,264,1407,1430]" captionTargetBox="[154,1430,196,1374]" captionTargetId="figure@6.[151,1435,195,1386]" captionTargetPageId="6" captionText="FIGURES 13 20. (13 16) Helina mediomaculata sp. nov., male: (13) sternite 5, dorsal view; (14) male cercal plate and surstyli, dorsal view; (15) male cercal plate and surstyli, lateral view; (16) aedeagal complex, lateral view. (17 20) Helina sexnotata sp. nov., male: (17) sternite 5, dorsal view; (18) cercal plate and surstyli, dorsal view; (19) cercal plate and surstyli, lateral view; (20) male aedeagal complex, lateral view." httpUri="https://zenodo.org/record/266662/files/figure.png" pageId="5" pageNumber="379">1316</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36A65417909E866FF0CAB0131912A69" pageId="5" pageNumber="379" type="materials_examined">
<paragraph id="8BCF36CA7909E866FF0CAB0131912A69" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF0CAB0133A92A4D" bold="true" box="[151,453,258,283]" pageId="5" pageNumber="379">
<typeStatus id="54CB88687909E866FF0CAB0132BE2A4D" box="[151,210,258,283]" pageId="5" pageNumber="379">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="54CB88687909E866FE55AB01305A2A4D" box="[462,566,258,283]" pageId="5" pageNumber="379" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F367765A7909E866FD1CAB00316E2A4D" box="[647,770,259,283]" name="Angola" pageId="5" pageNumber="379">ANGOLA</collectingCountry>
: (A26), Salazar, I.I.A.A,
<date id="FFCE100A7909E866FBBDAB0136D02A4D" box="[1062,1212,258,283]" pageId="5" pageNumber="379" value="1972-03-09" valueMax="1972-03-15" valueMin="1972-03-09">915.iii.1972</date>
/ Southern African Exp., B. M. 1972-1.
<typeStatus id="54CB88687909E866FEE5AB2433822A69" box="[382,494,295,319]" pageId="5" pageNumber="379" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D76FD437909E866FE62AB2430082A69" box="[505,612,294,319]" pageId="5" pageNumber="379" type="female">2 females</specimenCount>
(1
<collectionCode id="ED61AE0F7909E866FD11AB2430B52A69" box="[650,729,295,319]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="5" pageNumber="379">MNRJ</collectionCode>
), same labels as
<typeStatus id="54CB88687909E866FC08AB2531952A69" box="[915,1017,294,319]" pageId="5" pageNumber="379" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A65417909E866FF5CAB4936EB2AFD" pageId="5" pageNumber="379" type="diagnosis">
<paragraph id="8BCF36CA7909E866FF5CAB4936EB2AFD" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CAB49332D2A35" bold="true" box="[199,321,330,355]" pageId="5" pageNumber="379">Diagnosis.</emphasis>
Both postsutural intraalar setae present, the anterior one short and the posterior one very long; lower katepisternal seta inserted closer to the posterior than to the anterior seta; fore tibia with a posterior median seta; mid tibia without anterodorsal seta; hind tibia without posterodorsal seta; wing clear.
</paragraph>
</subSubSection>
<subSubSection id="C36A65417909E866FF5CABB536332DE8" pageId="5" pageNumber="379" type="description">
<paragraph id="8BCF36CA7909E866FF5CABB5308E29B9" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CABB533352A99" bold="true" box="[199,345,438,463]" pageId="5" pageNumber="379">Description.</emphasis>
<emphasis id="B904EAD87909E866FEC4ABB530292A99" box="[351,581,438,463]" italics="true" pageId="5" pageNumber="379">General colouration</emphasis>
. Brown and yellow with grey pollinosity. Frons brown; ocellar triangle with grey pollinosity; fronto-orbital plate, face, parafacial and gena silvery-grey pollinose; antenna with scape yellow and postpedicel brown; arista light brown; palpus yellow; scutum presuturally with two brown vittae between the dorsocentrals; postsuturally with a median brown and light brown spot from the suture to a little beyond the second pair of dorsocentrals and two broad lateral vittae (
<figureCitation id="134B2A4F7909E866FD5DA84531652909" box="[710,777,582,607]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1937,1960]" captionTargetBox="[211,1373,210,1890]" captionTargetId="figure@2.[180,1392,193,1899]" captionTargetPageId="2" captionText="FIGURES 1 4. Dorsal view of holotypes: (1) Helina duocolorata sp. nov., holotype male; (2) Helina longicerca sp. nov., holotype male; (3) Helina mediomaculata sp. nov., holotype male; (4) Helina sexnotata sp. nov., holotype female." httpUri="https://zenodo.org/record/266660/files/figure.png" pageId="5" pageNumber="379">Fig. 3</figureCitation>
); postpronotum yellow with whitish pollinosity; scutellum yellow; pleura mostly yellow, with grey pollinosity on anepisternum and basal part of katepimeron; calypters white; haltere yellow; wing clear; legs yellow, tarsi brown; abdomen of male mostly yellow with two round brown spots on tergite 4, close to the distal margin; in female, the abdomen with grey pollinosity, and brown spots on tergites 3 and 4; tergite 5 with a brown median vitta.
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CA8F930912844" blockId="5.[151,1437,258,1906]" box="[199,765,762,787]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CA8F9336D2845" box="[199,257,762,787]" italics="true" pageId="5" pageNumber="379">Male</emphasis>
. Body length:
<quantity id="4C889B2F7909E866FE3CA8F9306C2844" box="[423,512,762,787]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="5" pageNumber="379" unit="mm" value="6.2">6.2 mm</quantity>
; wing length: 6.0 mm.
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CA91D37F128F5" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CA91D33682861" box="[199,260,798,823]" italics="true" pageId="5" pageNumber="379">Head</emphasis>
. Eye bare; frons at level of anterior ocellus about one-fifth of head-width; 4 pairs of frontal setae inserted on less than lower half of frons, the one closest to lunule longer and the last one ciliform; inner and outer vertical setae moderate and similar in size; ocellar setae long; antenna inserted a little above the mid-level of eye; postpedicel long, about 4 times the length of pedicel, reaching mouth-margin; arista long-plumose; palpus filiform.
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CA9AD30672F2D" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CA9AD33742891" box="[199,280,942,967]" italics="true" pageId="5" pageNumber="379">Thorax</emphasis>
. Dorsocentral setae 2+3, the anterior presutural seta shorter than the others; presutural acrostichals hairlike; 2 postpronotals; 1 presutural and 2 postsutural intraalars, the anterior one very short and the posterior one very long; 1 supraalar; 1 postalar; notopleuron with 2 setae, the posterior slightly shorter than anterior; anepisternum with a series of 6 long setae, and a strong setula in dorsal anterior corner; lower katepisternal seta inserted closer to posterior than to anterior seta. Scutellum with 1 long basal and 1 long apical pair of setae, similar in length; margins and ventral surface bare.
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CAE8437F12ECD" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CAE8432902FC8" box="[199,252,1159,1182]" italics="true" pageId="5" pageNumber="379">Legs</emphasis>
. Fore femur with rows of posteroventral, posterior and posterodorsal setae; fore tibia with a median posterior seta, 1 preapical dorsal and 1 ventral apical seta; mid femur with a row of anterodorsal setae on basal half and 3 well-spaced ventral setae on apical half, 1 anterodorsal on apical fifth and 2 preapical dorsals; mid tibia with 2 posterior setae inserted at the limits of the thirds and a shorter one at basal third, 1 preapical anterodorsal and 1 posterodorsal, apical setae on dorsal, posterodorsal, anterior and ventral surfaces, the latter longer; hind femur with a well-spaced row of anterodorsal setae, and 4 anteroventral setae inserted along the length of the femur, posterodorsal surface with a seta at apical fifth; hind tibia with 2 anterodorsals on middle third and 2 anteroventrals, one median and one submedian, 1 anterodorsal and 1 posterodorsal preapicals, and 1 ventral apical.
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CAFA433152EB3" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CAFA432932EE8" box="[199,255,1447,1470]" italics="true" pageId="5" pageNumber="379">Wing</emphasis>
. Veins bare; veins R4+5 and M parallel at wing-tip; costal spine distinct; lower calypter about twice the length of upper one.
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CAFF237332D78" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CAFF2335D2D5C" box="[199,305,1521,1546]" italics="true" pageId="5" pageNumber="379">Abdomen</emphasis>
. Syntergite 1+2 and tergite 3 with laterals and a marginal row of long setae; tergites 4 and 5 each with a marginal and discal row of long setae; sternite 1 bare; sternite 5 with one longer seta on each lobe (
<figureCitation id="134B2A4F7909E866FB65AC16373E2D78" box="[1278,1362,1557,1582]" captionStart="FIGURES 13 20" captionStartId="6.[151,264,1407,1430]" captionTargetBox="[154,1430,196,1374]" captionTargetId="figure@6.[151,1435,195,1386]" captionTargetPageId="6" captionText="FIGURES 13 20. (13 16) Helina mediomaculata sp. nov., male: (13) sternite 5, dorsal view; (14) male cercal plate and surstyli, dorsal view; (15) male cercal plate and surstyli, lateral view; (16) aedeagal complex, lateral view. (17 20) Helina sexnotata sp. nov., male: (17) sternite 5, dorsal view; (18) cercal plate and surstyli, dorsal view; (19) cercal plate and surstyli, lateral view; (20) male aedeagal complex, lateral view." httpUri="https://zenodo.org/record/266662/files/figure.png" pageId="5" pageNumber="379">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CAC3A37352D04" blockId="5.[151,1437,258,1906]" box="[199,1369,1593,1618]" pageId="5" pageNumber="379">
<taxonomicName id="4C704D497909E866FF5CAC3A332D2D04" box="[199,321,1593,1618]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="379" phylum="Tracheophyta" rank="genus">
<emphasis id="B904EAD87909E866FF5CAC3A332D2D04" box="[199,321,1593,1618]" italics="true" pageId="5" pageNumber="379">Terminalia</emphasis>
</taxonomicName>
. Cercal plate as long as wide and surstylus as in
<figureCitation id="134B2A4F7909E866FCFFAC3A31892D04" box="[868,997,1593,1618]" captionStart="FIGURES 13 20" captionStartId="6.[151,264,1407,1430]" captionTargetBox="[154,1430,196,1374]" captionTargetId="figure@6.[151,1435,195,1386]" captionTargetPageId="6" captionText="FIGURES 13 20. (13 16) Helina mediomaculata sp. nov., male: (13) sternite 5, dorsal view; (14) male cercal plate and surstyli, dorsal view; (15) male cercal plate and surstyli, lateral view; (16) aedeagal complex, lateral view. (17 20) Helina sexnotata sp. nov., male: (17) sternite 5, dorsal view; (18) cercal plate and surstyli, dorsal view; (19) cercal plate and surstyli, lateral view; (20) male aedeagal complex, lateral view." httpUri="https://zenodo.org/record/266662/files/figure.png" pageId="5" pageNumber="379">Figs 1415</figureCitation>
. Aedeagal complex as in
<figureCitation id="134B2A4F7909E866FA98AC3A37392D04" box="[1283,1365,1593,1618]" captionStart="FIGURES 13 20" captionStartId="6.[151,264,1407,1430]" captionTargetBox="[154,1430,196,1374]" captionTargetId="figure@6.[151,1435,195,1386]" captionTargetPageId="6" captionText="FIGURES 13 20. (13 16) Helina mediomaculata sp. nov., male: (13) sternite 5, dorsal view; (14) male cercal plate and surstyli, dorsal view; (15) male cercal plate and surstyli, lateral view; (16) aedeagal complex, lateral view. (17 20) Helina sexnotata sp. nov., male: (17) sternite 5, dorsal view; (18) cercal plate and surstyli, dorsal view; (19) cercal plate and surstyli, lateral view; (20) male aedeagal complex, lateral view." httpUri="https://zenodo.org/record/266662/files/figure.png" pageId="5" pageNumber="379">Fig. 16</figureCitation>
.
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CAC5E31142D23" blockId="5.[151,1437,258,1906]" box="[199,888,1629,1654]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CAC5E33772D20" box="[199,283,1629,1654]" italics="true" pageId="5" pageNumber="379">Female</emphasis>
. Body length: 5.86.0 mm; wing length:
<quantity id="4C889B2F7909E866FD71AC5E31182D23" box="[746,884,1629,1654]" metricMagnitude="-3" metricUnit="m" metricValue="5.699999999999999" metricValueMax="5.8" metricValueMin="5.6" pageId="5" pageNumber="379" unit="mm" value="5.699999999999999" valueMax="5.8" valueMin="5.6">5.65.8 mm</quantity>
.
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CAC8236332DE8" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">Similar to male, except for: head dichoptic; frons at level of anterior ocellus about one-third of head-width; 3 pairs of frontal setae, the one closest to lunule long, and two pairs of reclinate orbitals.</paragraph>
</subSubSection>
<subSubSection id="C36A65417909E866FF5CACCA30412DB4" box="[199,557,1737,1762]" pageId="5" pageNumber="379" type="distribution">
<paragraph id="8BCF36CA7909E866FF5CACCA30412DB4" blockId="5.[151,1437,258,1906]" box="[199,557,1737,1762]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CACCA330C2DB4" bold="true" box="[199,352,1737,1762]" pageId="5" pageNumber="379">Distribution.</emphasis>
<collectingCountry id="F367765A7909E866FEFCACCA33D52DB4" box="[359,441,1737,1762]" name="Angola" pageId="5" pageNumber="379">Angola</collectingCountry>
(Salazar).
</paragraph>
</subSubSection>
<subSubSection id="C36A65417909E866FF5CACEE30F22C24" pageId="5" pageNumber="379" type="etymology">
<paragraph id="8BCF36CA7909E866FF5CACEE31F92C7F" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CACEE33222C50" bold="true" box="[199,334,1773,1798]" pageId="5" pageNumber="379">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C704D497909E866FDDBACEE30812C50" box="[576,749,1773,1798]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="mediomaculata">
<emphasis id="B904EAD87909E866FDDBACEE30812C50" box="[576,749,1773,1798]" italics="true" pageId="5" pageNumber="379">mediomaculata</emphasis>
</taxonomicName>
is derived from the Latin
<emphasis id="B904EAD87909E866FB86ACEE36002C50" box="[1053,1132,1773,1798]" italics="true" pageId="5" pageNumber="379">medius</emphasis>
(“middle”) and
<emphasis id="B904EAD87909E866FAB2ACEE37F02C50" box="[1321,1436,1773,1798]" italics="true" pageId="5" pageNumber="379">maculatus</emphasis>
(“marked”), and refers to the dark brown central spot on the scutum.
</paragraph>
<paragraph id="8BCF36CA7909E866FF5CAD3630F22C24" blockId="5.[151,1437,258,1906]" pageId="5" pageNumber="379">
<emphasis id="B904EAD87909E866FF5CAD36337D2C18" bold="true" box="[199,273,1845,1870]" pageId="5" pageNumber="379">Notes.</emphasis>
The new species runs to couplet 10 and
<taxonomicName id="4C704D497909E866FD45AD3431E22C18" authority="Curran" authorityName="Curran" box="[734,910,1845,1870]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="379" phylum="Arthropoda" rank="species" species="lenta">
<emphasis id="B904EAD87909E866FD45AD3431552C18" box="[734,825,1845,1870]" italics="true" pageId="5" pageNumber="379">H. lenta</emphasis>
Curran
</taxonomicName>
in Emdens (1951) key, but the general colour of the body easily separates these two species.
</paragraph>
</subSubSection>
</treatment>
</document>