427 lines
68 KiB
XML
427 lines
68 KiB
XML
<document id="A0F517FCD6BED4B022BDA4AD6912C792" ID-CLB-Dataset="99517" ID-DOI="10.11646/zootaxa.5165.4.1" ID-GBIF-Dataset="cd84b880-d77f-401a-bf4c-a98021a5e166" ID-ISSN="1175-5326" ID-Zenodo-Dep="6853942" ID-ZooBank="DBE812FF-7B25-4522-97FE-45A86ECAEFD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658127043518" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi & Pathomwattananurak, Wuttipon" docDate="2022" docId="03D9879BE84A17767EFDF8C969AFA34F" docLanguage="en" docName="zootaxa.5165.4.1.pdf" docOrigin="Zootaxa 5165 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eophileurus sidereus Yang & Pathomwattananurak 2022, new species" docType="treatment" docVersion="7" lastPageNumber="476" masterDocId="FFE0FFE3E85D176C7E6AFFEE6B79A22A" masterDocTitle="Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species" masterLastPageNumber="485" masterPageNumber="451" pageNumber="474" updateTime="1699360166553" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A68347C04D3E08768169D457B3CCE203">
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<mods:title id="2BF420DFC06DC87DB633ED36ACF66DAD">Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species</mods:title>
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<mods:name id="F1F991EEFCD54B465BBE0FE0B415BF8A" type="personal">
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<mods:roleTerm id="F94DBD8FBCDF3822369B06E76E56A699">Author</mods:roleTerm>
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<mods:namePart id="3BD5DB41AD6E8950F23E121AC8F0E3E4">Yang, Qiao-Zhi</mods:namePart>
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<mods:affiliation id="48D575249872BD20452994723DFEBA03">3 B, Pingguoshequ, Baiziwan Road, Chaoyang Dist., Beijing, 100022, P. R. China.</mods:affiliation>
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<mods:roleTerm id="8300A1ABDCBD64D6E80410EA957B286E">Author</mods:roleTerm>
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<mods:namePart id="04EB15CF7408702DE8B24783067FF192">Pathomwattananurak, Wuttipon</mods:namePart>
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<mods:affiliation id="1787256A310A287B26B457F2D4AC3184">1051 / 38 Wiang, Mueang Dist., Chiang Rai Province, 57000, Thailand.</mods:affiliation>
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<mods:typeOfResource id="FA5F095EA3C6FF32639F90177513F865">text</mods:typeOfResource>
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<mods:title id="EB2F210AF9394626E7B97B20A796AAC0">Zootaxa</mods:title>
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<mods:part id="8449E71535EB8AB8B654AFF05DD2D5B2">
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<mods:date id="F49F3772CBD11D5B5F66B63DA44704AF">2022</mods:date>
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<mods:number id="0D23FB874A120213DFF40917A541CC3B">2022-07-18</mods:number>
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<treatment id="03D9879BE84A17767EFDF8C969AFA34F" ID-DOI="http://doi.org/10.5281/zenodo.6857354" ID-GBIF-Taxon="197819088" ID-Zenodo-Dep="6857354" LSID="urn:lsid:plazi:treatment:03D9879BE84A17767EFDF8C969AFA34F" httpUri="http://treatment.plazi.org/id/03D9879BE84A17767EFDF8C969AFA34F" lastPageId="26" lastPageNumber="476" pageId="23" pageNumber="474">
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<subSubSection id="C36A6506E84A177B7EFDF8C968C1A568" box="[151,952,1831,1858]" pageId="23" pageNumber="474" type="nomenclature">
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<paragraph id="8BCF368DE84A177B7EFDF8C968C1A568" blockId="23.[151,952,1831,1895]" box="[151,952,1831,1858]" pageId="23" pageNumber="474">
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<heading id="D08781E1E84A177B7EFDF8C968C1A568" bold="true" box="[151,952,1831,1858]" fontSize="11" level="1" pageId="23" pageNumber="474" reason="1">
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<emphasis id="B904EA9FE84A177B7EFDF8C968C1A568" bold="true" box="[151,952,1831,1858]" pageId="23" pageNumber="474">
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<taxonomicName id="4C704D0EE84A177B7EFDF8C96862A568" authority="Yang & Pathomwattananurak" authorityName="Yang & Pathomwattananurak" authorityYear="2022" box="[151,795,1831,1858]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="sidereus" status="sp. nov.">
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<emphasis id="B904EA9FE84A177B7EFDF8C96AEEA568" bold="true" box="[151,407,1831,1858]" italics="true" pageId="23" pageNumber="474">Eophileurus sidereus</emphasis>
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Yang & Pathomwattananurak
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23757E4E84A177B7D42F8C668C1A568" box="[808,952,1832,1858]" pageId="23" pageNumber="474" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C36A6506E84A177B7EFDF8A269E1A54C" box="[151,664,1868,1895]" pageId="23" pageNumber="474" type="description">
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<paragraph id="8BCF368DE84A177B7EFDF8A269E1A54C" blockId="23.[151,952,1831,1895]" box="[151,664,1868,1895]" pageId="23" pageNumber="474">
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(
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<figureCitation id="134B2A08E84A177B7EF5F8A26A3DA54C" box="[159,324,1868,1894]" captionStart="FIGURES 103–108" captionStartId="24.[151,264,1591,1615]" captionTargetBox="[159,1425,250,1560]" captionTargetId="figure-41@24.[151,1435,237,1568]" captionTargetPageId="24" captionText="FIGURES 103–108. Habitus of Eophileurus sidereus Yang & Pathomwattananurak, new species. 103–104, holotype; 105, male paratype; 106–108, female paratypes; 103, 105, 106, 108, dorsal view; 104, 107, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853976" httpUri="https://zenodo.org/record/6853976/files/figure.png" pageId="23" pageNumber="474">Figs. 103–108</figureCitation>
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,
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<figureCitation id="134B2A08E84A177B7F25F8A26AC8A54D" box="[335,433,1868,1895]" captionStart="FIGURES109–116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109–116.HabitusofEophileurusArrow,1908species(obliqueview).109–110,E.minorYang&Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111–112, E. prelli Yang & Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113–114, E. quadratifovealis Yang & Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115–116, E. sidereus Yang & Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="23" pageNumber="474">115–116</figureCitation>
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,
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<figureCitation id="134B2A08E84A177B7FD7F8A2695BA54C" box="[445,546,1868,1894]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral & Strnad, 1992 (Longli); 169–172, E. minor Yang & Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang & Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang & Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang & Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="23" pageNumber="474">181–184</figureCitation>
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,
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<figureCitation id="134B2A08E84A177B7C47F8A269E9A54C" box="[557,656,1868,1894]" captionStart="FIGURES 185–196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185–196. Parameres of Eophileurus Arrow, 1908. 185–192, E. minor Yang & Pathomwattananurak, new species, 185–187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189–190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193–196, E. sidereus Yang & Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="23" pageNumber="474">193–196</figureCitation>
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)
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</paragraph>
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<subSubSection id="C36A6506E84A17757EFDF87A681BA673" lastPageId="25" lastPageNumber="476" pageId="23" pageNumber="474" type="materials_examined">
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<emphasis id="B904EA9FE84A177B7EFDF87A6A3BA584" bold="true" box="[151,322,1940,1966]" pageId="23" pageNumber="474">Type material.</emphasis>
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(
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<specimenCount id="9D76FD04E84A177B7F25F87A6AF3A584" box="[335,394,1940,1966]" pageId="23" pageNumber="474" type="male">6♂♂</specimenCount>
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,
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<specimenCount id="9D76FD04E84A177B7FF9F87A6AA3A584" box="[403,474,1940,1966]" pageId="23" pageNumber="474" type="female">12♀♀</specimenCount>
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).
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<typeStatus id="54CB882FE84A177B7F87F87A6927A584" box="[493,606,1940,1966]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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(
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<specimenCount id="9D76FD04E84A177B7C1BF87A69F1A584" box="[625,648,1940,1966]" pageId="23" pageNumber="474" type="male">♂</specimenCount>
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,
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<collectionCode id="ED61AE48E84A177B7CFBF87B6981A585" box="[657,760,1941,1967]" pageId="23" pageNumber="474">THNHM</collectionCode>
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<emphasis id="B904EA9FE84A177B7C92F87A6878A584" bold="true" box="[760,769,1940,1966]" pageId="23" pageNumber="474">)</emphasis>
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, “
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<specimenCode id="DBD69EF6E84A177B7D72F87A684FA584" box="[792,822,1940,1966]" collectionCode="THNHM" pageId="23" pageNumber="474">F1</specimenCode>
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generation of female from
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<location id="8EAF6056E84A177B7EFDF8576A50A5F8" LSID="urn:lsid:plazi:treatment:03D9879BE84A17767EFDF8C969AFA34F:8EAF6056E84A177B7EFDF8576A50A5F8" box="[151,297,1976,2003]" country="Thailand" latitude="5.8854165" longLatPrecision="1" longitude="101.02156" name="Betong Dist." pageId="23" pageNumber="474" stateProvince="Yala">Betong Dist.</location>
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,
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<collectingRegion id="49B4F86FE84A177B7F5EF8566AD2A5F9" box="[308,427,1976,2003]" country="Thailand" name="Yala" pageId="23" pageNumber="474">Yala Prov.</collectingRegion>
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</date>
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,
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<collectorName id="2685535BE84A177B7CC7F8566807A5F8" box="[685,894,1976,2002]" pageId="23" pageNumber="474">Suradet Sae-Liang</collectorName>
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leg., adult emerged in
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<date id="FFCE104DE84A177B7AE9F8566FA3A5F8" box="[1155,1242,1976,2002]" pageId="23" pageNumber="474" value="2021-02">
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<collectingDate id="EF8AE9A5E84A177B7AE9F8566FA3A5F8" box="[1155,1242,1976,2002]" pageId="23" pageNumber="474" value="2021-02">II.2021</collectingDate>
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</date>
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”
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</materialsCitation>
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.
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<materialsCitation id="3B183CD0E84A177B7A9AF8566EEEA5DC" ID-GBIF-Occurrence="3863916384" collectingDate="2020-10-18" collectionCode="THNHM" collectorName="Suradet Sae-Liang" country="Thailand" latitude="5.8854165" location="Betong Dist." longLatPrecision="1" longitude="101.02156" pageId="23" pageNumber="474" specimenCount="1" specimenCount-female="1" stateProvince="Yala" typeStatus="paratype">
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<emphasis id="B904EA9FE84A177B7A9AF8566E08A5F8" bold="true" box="[1264,1393,1976,2002]" pageId="23" pageNumber="474">
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<typeStatus id="54CB882FE84A177B7A9AF8566E15A5F8" box="[1264,1388,1976,2002]" pageId="23" pageNumber="474" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D76FD04E84A177B7B13F8566EE5A5F8" box="[1401,1436,1976,2002]" pageId="23" pageNumber="474" type="female">1♀</specimenCount>
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(
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<collectionCode id="ED61AE48E84A177B7EF5F8326A75A5DC" box="[159,268,2012,2038]" pageId="23" pageNumber="474">THNHM</collectionCode>
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), “
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<geoCoordinate id="EE44504AE84A177B7F40F8326AC1A5DD" box="[298,440,2012,2039]" degrees="5" direction="north" minutes="53" orientation="latitude" pageId="23" pageNumber="474" precision="1" seconds="07.5" value="5.8854165">5°53'07.5"N</geoCoordinate>
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,
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,
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<location id="8EAF6056E84A177B7C1CF833687EA5DC" LSID="urn:lsid:plazi:treatment:03D9879BE84A17767EFDF8C969AFA34F:8EAF6056E84A177B7C1CF833687EA5DC" box="[630,775,2012,2039]" country="Thailand" latitude="5.8854165" longLatPrecision="1" longitude="101.02156" name="Betong Dist." pageId="23" pageNumber="474" stateProvince="Yala">Betong Dist.</location>
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,
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<collectingRegion id="49B4F86FE84A177B7D7BF83268FEA5DD" box="[785,903,2012,2039]" country="Thailand" name="Yala" pageId="23" pageNumber="474">Yala Prov.</collectingRegion>
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,
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<collectingCountry id="F367761DE84A177B7DF8F8326881A5DC" box="[914,1016,2012,2038]" name="Thailand" pageId="23" pageNumber="474">Thailand</collectingCountry>
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,
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<date id="FFCE104DE84A177B7A6EF8326F04A5DC" box="[1028,1149,2012,2038]" pageId="23" pageNumber="474" value="2020-10-18">
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<collectingDate id="EF8AE9A5E84A177B7A6EF8326F04A5DC" box="[1028,1149,2012,2038]" pageId="23" pageNumber="474" value="2020-10-18">18.X.2020</collectingDate>
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</date>
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,
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<collectorName id="2685535BE84A177B7AE2F8326E21A5DC" box="[1160,1368,2012,2038]" pageId="23" pageNumber="474">Suradet Sae-Liang</collectorName>
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leg.”
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</materialsCitation>
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;
|
||
<materialsCitation id="3B183CD0E84517747EFDFF796900A29B" ID-GBIF-Occurrence="3863916361" box="[151,633,151,177]" collectionCode="THNHM" pageId="24" pageNumber="475" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<specimenCount id="9D76FD04E84517747EFDFF796BC3A29B" box="[151,186,151,177]" pageId="24" pageNumber="475" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED61AE48E84517747EA0FF796A4EA29B" box="[202,311,151,177]" pageId="24" pageNumber="475">THNHM</collectionCode>
|
||
), same label as the
|
||
<typeStatus id="54CB882FE84517747C79FF796900A29B" box="[531,633,151,177]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B183CD0E84517747CEFFF796FDFA29B" ID-GBIF-Occurrence="3863916369" box="[645,1190,151,177]" collectionCode="CWT" pageId="24" pageNumber="475" specimenCount="10" specimenCount-female="7" specimenCount-male="3" typeStatus="holotype">
|
||
<specimenCount id="9D76FD04E84517747CEFFF7969C6A29B" box="[645,703,151,177]" pageId="24" pageNumber="475" type="male">3♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D76FD04E84517747CA6FF79687DA29B" box="[716,772,151,177]" pageId="24" pageNumber="475" type="female">7♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED61AE48E84517747D7EFF796821A29B" box="[788,856,151,177]" pageId="24" pageNumber="475">CWT</collectionCode>
|
||
), same labels as the
|
||
<typeStatus id="54CB882FE84517747A2AFF796FDFA29B" box="[1088,1190,151,177]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B183CD0E84517747ADBFF796AB7A2FF" ID-GBIF-Occurrence="3863916375" collectionCode="ZQZY" pageId="24" pageNumber="475" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="holotype">
|
||
<specimenCount id="9D76FD04E84517747ADBFF796F95A29B" box="[1201,1260,151,177]" pageId="24" pageNumber="475" type="male">2♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D76FD04E84517747A92FF796E49A29B" box="[1272,1328,151,177]" pageId="24" pageNumber="475" type="female">3♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED61AE48E84517747B2AFF796EE9A29B" box="[1344,1424,151,177]" pageId="24" pageNumber="475">ZQZY</collectionCode>
|
||
), same labels as the
|
||
<typeStatus id="54CB882FE84517747F02FF556AB7A2FF" box="[360,462,187,213]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF0F6605E84517747EFDF9D96FFEA459" ID-DOI="http://doi.org/10.5281/zenodo.6853976" ID-Zenodo-Dep="6853976" httpUri="https://zenodo.org/record/6853976/files/figure.png" pageId="24" pageNumber="475" startId="24.[151,264,1591,1615]" targetBox="[159,1425,250,1560]" targetPageId="24">
|
||
<paragraph id="8BCF368DE84517747EFDF9D96FFEA459" blockId="24.[151,1436,1591,1652]" pageId="24" pageNumber="475">
|
||
<emphasis id="B904EA9FE84517747EFDF9D96A09A465" bold="true" box="[151,368,1591,1615]" pageId="24" pageNumber="475">FIGURES 103–108.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C704D0EE84517747F85F9D66F7DA47A" authority="Yang & Pathomwattananurak" authorityName="Yang & Pathomwattananurak" authorityYear="2022" box="[495,1028,1592,1616]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="sidereus" status="sp. nov.">
|
||
<emphasis id="B904EA9FE84517747F85F9D669BEA47A" box="[495,711,1592,1616]" italics="true" pageId="24" pageNumber="475">Eophileurus sidereus</emphasis>
|
||
Yang & Pathomwattananurak
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B904EA9FE84517747A7AF9D66FECA465" bold="true" box="[1040,1173,1591,1616]" pageId="24" pageNumber="475">
|
||
<taxonomicNameLabel id="A23757E4E84517747A7AF9D66FE8A465" box="[1040,1169,1591,1616]" pageId="24" pageNumber="475" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</emphasis>
|
||
103–104, holotype; 105, male paratype; 106–108, female paratypes; 103, 105, 106, 108, dorsal view; 104, 107, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCF368DE84517757EADF94C6895A0DB" blockId="24.[151,1437,1698,2012]" lastBlockId="25.[151,1437,151,2013]" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475">
|
||
<emphasis id="B904EA9FE84517747EADF94C6A4DA496" bold="true" box="[199,308,1698,1724]" pageId="24" pageNumber="475">
|
||
<typeStatus id="54CB882FE84517747EADF94C6A4DA496" box="[199,308,1698,1724]" pageId="24" pageNumber="475" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(male).
|
||
<emphasis id="B904EA9FE84517747FFCF94C6A8FA496" bold="true" box="[406,502,1698,1724]" pageId="24" pageNumber="475">General</emphasis>
|
||
(
|
||
<figureCitation id="134B2A08E84517747C6DF94C69D4A496" box="[519,685,1698,1724]" captionStart="FIGURES 103–108" captionStartId="24.[151,264,1591,1615]" captionTargetBox="[159,1425,250,1560]" captionTargetId="figure-41@24.[151,1435,237,1568]" captionTargetPageId="24" captionText="FIGURES 103–108. Habitus of Eophileurus sidereus Yang & Pathomwattananurak, new species. 103–104, holotype; 105, male paratype; 106–108, female paratypes; 103, 105, 106, 108, dorsal view; 104, 107, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853976" httpUri="https://zenodo.org/record/6853976/files/figure.png" pageId="24" pageNumber="475">Figs. 103–104</figureCitation>
|
||
,
|
||
<figureCitation id="134B2A08E84517747CD0F94C699DA497" box="[698,740,1698,1725]" captionStart="FIGURES109–116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109–116.HabitusofEophileurusArrow,1908species(obliqueview).109–110,E.minorYang&Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111–112, E. prelli Yang & Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113–114, E. quadratifovealis Yang & Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115–116, E. sidereus Yang & Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="24" pageNumber="475">115</figureCitation>
|
||
): Body dark reddish-brown, rather oblong, flattened, surface shiny; ventral surface clads with slightly dense, long reddish-yellow setae.
|
||
<emphasis id="B904EA9FE84517747D83F9286F4BA4CA" bold="true" box="[1001,1074,1734,1760]" pageId="24" pageNumber="475">Head:</emphasis>
|
||
Clypeus with very sparse small punctures, with a long and backward horn at middle, as well as a ridge from horn to apex; clypeal apex slightly rounded. Frons depressed, disk almost impunctate, only with a few punctures at each side. Mandible sinuate at outer margin.
|
||
<emphasis id="B904EA9FE84517747F53F8DC6AC5A566" bold="true" box="[313,444,1842,1868]" pageId="24" pageNumber="475">Pronotum:</emphasis>
|
||
Surface with dense and large punctures, punctures becoming denser close to margins. Anterior region with a large, deep, round fovea; punctures in fovea semi-rounded, moderately sparse; margin with a weak protuberance at each side posteriorly. Side roundly curved; all margins beaded except before the fovea, more strongly at sides. Anterior angle acute, apex slightly rounded, posterior angle obtusely angulate.
|
||
<emphasis id="B904EA9FE84517747B71F8706EE5A592" bold="true" box="[1307,1436,1950,1976]" pageId="24" pageNumber="475">Scutellum:</emphasis>
|
||
Surface with only a few punctures close to base.
|
||
<emphasis id="B904EA9FE84517747CD1F82C6869A5F6" bold="true" box="[699,784,1986,2012]" pageId="24" pageNumber="475">Elytra:</emphasis>
|
||
Surface with dense micropunctures, each with scale-liked seta, and dense, sub-U-shape, large punctures. Interstice 1 with punctures in three irregular rows combined to two irregular rows at anterior third, and to one irregular row at middle, interstice 2 with one irregular row. Primary costa A and primary costa B rather distinct. Subapical umbones slightly prominent. Margin of elytron moderately dilated horizontally from base to posterior three-seventh.
|
||
<emphasis id="B904EA9FE84417757CB3FEEC682CA336" bold="true" box="[729,853,258,284]" pageId="25" pageNumber="476">Pygidium:</emphasis>
|
||
slightly convex, with dense punctures throughout, punctures denser near base and sparser near apex, each with scale-liked seta.
|
||
<emphasis id="B904EA9FE84417757D8EFEC96FF6A36B" bold="true" box="[996,1167,295,321]" pageId="25" pageNumber="476">Metasternum:</emphasis>
|
||
Surface mainly reddish, with black region near sides and posterior margin; center with a small, shallow depression; punctures large and dense close to all sides; median region with sparser, slightly smaller punctures; setae yellow, short, at anterior region and each side.
|
||
<emphasis id="B904EA9FE84417757F61FE7C6970A386" bold="true" box="[267,521,402,428]" pageId="25" pageNumber="476">Abdominal ventrites:</emphasis>
|
||
Surfaces with dense and small punctures throughout, punctures becoming slightly denser at each side; ventrite 6 with rugose and large punctures at anterior region, and a very shallow, horizontally oval depression at center.
|
||
<emphasis id="B904EA9FE84417757FDBFE356A8BA3DF" bold="true" box="[433,498,475,501]" pageId="25" pageNumber="476">Legs:</emphasis>
|
||
Protibia tridentate, protarsi strongly thickened, inner protibial claw strongly enlarged and elongated. Inner metatibial spur long, apex slightly curved outward, outer metatibial spur shorter, very slightly curved outward.
|
||
<emphasis id="B904EA9FE84417757F3DFDCD6AA1A017" bold="true" box="[343,472,547,573]" pageId="25" pageNumber="476">Parameres</emphasis>
|
||
(
|
||
<figureCitation id="134B2A08E84417757F83FDCD69F4A017" box="[489,653,547,573]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral & Strnad, 1992 (Longli); 169–172, E. minor Yang & Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang & Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang & Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang & Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="25" pageNumber="476">Figs. 181–182</figureCitation>
|
||
): In front view (
|
||
<figureCitation id="134B2A08E84417757D24FDCD68C8A017" box="[846,945,547,573]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral & Strnad, 1992 (Longli); 169–172, E. minor Yang & Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang & Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang & Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang & Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="25" pageNumber="476">Fig. 181</figureCitation>
|
||
), base very weakly overlapped; outer sides curved outward at anterior one-third, and curved inward at anterior two-third, with a pair of long, acute, outward processes near apex; inner sides curved outward near anterior one-third, with a pair of overlapped, triangular, inward processes at anterior two-third; apex rounded; in lateral view (
|
||
<figureCitation id="134B2A08E84417757D37FD6168C6A083" box="[861,959,655,681]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral & Strnad, 1992 (Longli); 169–172, E. minor Yang & Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang & Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang & Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang & Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="25" pageNumber="476">Fig. 182</figureCitation>
|
||
), base extending backward; apex strongly bent backward, with a process near apex slightly backwards, and an upward process at upper margin; in ventral view, ventral base with a pair of short, outward, flat, semicircular processes.
|
||
</paragraph>
|
||
<paragraph id="8BCF368DE84417757EADFD156A73A1AB" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
|
||
<emphasis id="B904EA9FE84417757EADFD156A46A13F" bold="true" box="[199,319,763,789]" pageId="25" pageNumber="476">
|
||
<typeStatus id="54CB882FE84417757EADFD156A46A13F" box="[199,319,763,789]" pageId="25" pageNumber="476" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(male). Extremely similar to
|
||
<typeStatus id="54CB882FE84417757CEFFD156993A13F" box="[645,746,763,789]" pageId="25" pageNumber="476" type="holotype">holotype</typeStatus>
|
||
, characters mainly stable, except clypeal horn slightly shorter; fovea more sparsely punctate; scutellum sometimes impunctate; micropunctures on elytra slightly sparser (
|
||
<figureCitation id="134B2A08E84417757B05FCF16BBCA177" captionStart="FIGURES 103–108" captionStartId="24.[151,264,1591,1615]" captionTargetBox="[159,1425,250,1560]" captionTargetId="figure-41@24.[151,1435,237,1568]" captionTargetPageId="24" captionText="FIGURES 103–108. Habitus of Eophileurus sidereus Yang & Pathomwattananurak, new species. 103–104, holotype; 105, male paratype; 106–108, female paratypes; 103, 105, 106, 108, dorsal view; 104, 107, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853976" httpUri="https://zenodo.org/record/6853976/files/figure.png" pageId="25" pageNumber="476">Fig. 105</figureCitation>
|
||
); processes at inner sides of parameres extended less upward in front view and less upward in lateral view (
|
||
<figureCitation id="134B2A08E84417757B0EFCAD6B87A1AB" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral & Strnad, 1992 (Longli); 169–172, E. minor Yang & Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang & Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang & Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang & Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="25" pageNumber="476">Figs. 183–184</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCF368DE84417757EADFC65681BA673" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
|
||
<emphasis id="B904EA9FE84417757EADFC656A46A18F" bold="true" box="[199,319,907,933]" pageId="25" pageNumber="476">
|
||
<typeStatus id="54CB882FE84417757EADFC656A46A18F" box="[199,319,907,933]" pageId="25" pageNumber="476" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(female). Similar to male, except clypeus and frons much more densely rugopunctate throughout; clypeal horn much shorter; fovea absent, with an extremely shallow and narrow furrow on anterior region of pronotum; pronotum densely rugopunctate anteriorly; scutellum sometimes impunctate; elytral margin slightly more dilated; pygidium slightly flattened at center, with a small protuberance close to apex; ventrite 6 without depression, with slightly larger punctures throughout, rugopunctures denser near anterior margin; protarsi not thickened, inner protarsal claw not enlarged and elongated (
|
||
<figureCitation id="134B2A08E84417757C13FBD16864A673" box="[633,797,1087,1113]" captionStart="FIGURES 103–108" captionStartId="24.[151,264,1591,1615]" captionTargetBox="[159,1425,250,1560]" captionTargetId="figure-41@24.[151,1435,237,1568]" captionTargetPageId="24" captionText="FIGURES 103–108. Habitus of Eophileurus sidereus Yang & Pathomwattananurak, new species. 103–104, holotype; 105, male paratype; 106–108, female paratypes; 103, 105, 106, 108, dorsal view; 104, 107, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853976" httpUri="https://zenodo.org/record/6853976/files/figure.png" pageId="25" pageNumber="476">Figs. 106–108</figureCitation>
|
||
,
|
||
<figureCitation id="134B2A08E84417757D43FBD1682FA673" box="[809,854,1087,1113]" captionStart="FIGURES109–116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109–116.HabitusofEophileurusArrow,1908species(obliqueview).109–110,E.minorYang&Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111–112, E. prelli Yang & Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113–114, E. quadratifovealis Yang & Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115–116, E. sidereus Yang & Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="25" pageNumber="476">116</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6506E84417757EADFB8D6811A68B" pageId="25" pageNumber="476" type="description">
|
||
<paragraph id="8BCF368DE84417757EADFB8D6811A68B" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
|
||
<emphasis id="B904EA9FE84417757EADFB8D6A07A657" bold="true" box="[199,382,1123,1149]" pageId="25" pageNumber="476">Measurements.</emphasis>
|
||
Body length: male
|
||
<quantity id="4C889B68E84417757C0EFB8C6873A657" box="[612,778,1122,1149]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" metricValueMax="2.61" metricValueMin="2.51" pageId="25" pageNumber="476" unit="mm" value="25.6" valueMax="26.1" valueMin="25.1">25.1–26.1 mm</quantity>
|
||
(
|
||
<typeStatus id="54CB882FE84417757D70FB8D6807A657" box="[794,894,1123,1149]" pageId="25" pageNumber="476" type="holotype">holotype</typeStatus>
|
||
<quantity id="4C889B68E84417757DEDFB8D688BA657" box="[903,1010,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="25" pageNumber="476" unit="mm" value="26.1">26.1 mm</quantity>
|
||
), female
|
||
<quantity id="4C889B68E84417757A37FB8C6E7CA657" box="[1117,1285,1122,1149]" metricMagnitude="-2" metricUnit="m" metricValue="2.405" metricValueMax="2.57" metricValueMin="2.2399999999999998" pageId="25" pageNumber="476" unit="mm" value="24.049999999999997" valueMax="25.7" valueMin="22.4">22.4–25.7 mm</quantity>
|
||
; body width: male
|
||
<quantity id="4C889B68E84417757EBFFB696A01A68B" box="[213,376,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" metricValueMax="1.38" metricValueMin="1.24" pageId="25" pageNumber="476" unit="mm" value="13.1" valueMax="13.8" valueMin="12.4">12.4–13.8 mm</quantity>
|
||
(
|
||
<typeStatus id="54CB882FE84417757FECFB696A93A68B" box="[390,490,1159,1185]" pageId="25" pageNumber="476" type="holotype">holotype</typeStatus>
|
||
<quantity id="4C889B68E84417757F9BFB696920A68B" box="[497,601,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="25" pageNumber="476" unit="mm" value="13.2">13.2 mm</quantity>
|
||
), female
|
||
<quantity id="4C889B68E84417757CD5FB68681AA68B" box="[703,867,1158,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" metricValueMax="1.25" metricValueMin="1.13" pageId="25" pageNumber="476" unit="mm" value="11.9" valueMax="12.5" valueMin="11.3">11.3–12.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6506E84417757EADFB44683DA47B" pageId="25" pageNumber="476" type="diagnosis">
|
||
<paragraph id="8BCF368DE84417757EADFB44683DA47B" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
|
||
<emphasis id="B904EA9FE84417757EADFB446A3BA6EE" bold="true" box="[199,322,1194,1220]" pageId="25" pageNumber="476">Diagnosis.</emphasis>
|
||
This species is related to
|
||
<taxonomicName id="4C704D0EE84417757C0DFB4568C7A6EF" authority="Endrodi, 1977" authorityName="Endrodi" authorityYear="1977" box="[615,958,1195,1221]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="nicobarensis">
|
||
<emphasis id="B904EA9FE84417757C0DFB45686CA6EF" box="[615,789,1195,1221]" italics="true" pageId="25" pageNumber="476">E. nicobarensis</emphasis>
|
||
<bibRefCitation id="EFE14B7CE84417757D77FB4568C7A6EF" author="Endrodi, S." box="[797,958,1195,1221]" pageId="25" pageNumber="476" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134B2A08E84417757DA4FB456F21A6EF" box="[974,1112,1195,1221]" captionStart="FIGURES 79–86" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[180,1404,199,1908]" captionTargetId="figure-18@16.[169,1419,181,1920]" captionTargetPageId="16" captionText="FIGURES 79–86. Type specimens of Eophileurus nicobarensis Endrődi, 1977 (images ©HNHM/Ottó Merkl). 69–83, holotype, male, 79, habitus, dorsal; 80, habitus, lateral; 81, parameres, apical; 82, parameres, lateral; 83, labels; 84–86, allotype, female, 84, habitus, dorsal; 85, habitus, lateral; 86, labels." figureDoi="http://doi.org/10.5281/zenodo.6853968" httpUri="https://zenodo.org/record/6853968/files/figure.png" pageId="25" pageNumber="476">Figs. 79–86</figureCitation>
|
||
), but differs in the following points: clypeal horn much larger; pronotum with denser and larger punctures throughout; fovea much larger, with denser and larger punctures; elytra with smaller and denser punctures and much denser micropunctures; elytral margin more strongly dilated (
|
||
<figureCitation id="134B2A08E84417757F81FAF969F6A71B" box="[491,655,1303,1329]" captionStart="FIGURES 103–108" captionStartId="24.[151,264,1591,1615]" captionTargetBox="[159,1425,250,1560]" captionTargetId="figure-41@24.[151,1435,237,1568]" captionTargetPageId="24" captionText="FIGURES 103–108. Habitus of Eophileurus sidereus Yang & Pathomwattananurak, new species. 103–104, holotype; 105, male paratype; 106–108, female paratypes; 103, 105, 106, 108, dorsal view; 104, 107, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853976" httpUri="https://zenodo.org/record/6853976/files/figure.png" pageId="25" pageNumber="476">Figs. 103–108</figureCitation>
|
||
,
|
||
<figureCitation id="134B2A08E84417757CF0FAF86987A71B" box="[666,766,1302,1329]" captionStart="FIGURES109–116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109–116.HabitusofEophileurusArrow,1908species(obliqueview).109–110,E.minorYang&Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111–112, E. prelli Yang & Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113–114, E. quadratifovealis Yang & Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115–116, E. sidereus Yang & Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="25" pageNumber="476">115–116</figureCitation>
|
||
); parameres wider, base much less significantly overlapped, apex slightly narrower; processes near apex smaller, apex of each process less backwardly extended (
|
||
<figureCitation id="134B2A08E84417757B49FAD56BBAA753" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral & Strnad, 1992 (Longli); 169–172, E. minor Yang & Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang & Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang & Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang & Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="25" pageNumber="476">Figs. 181– 184</figureCitation>
|
||
,
|
||
<figureCitation id="134B2A08E84417757EBAFAB16A4FA753" box="[208,310,1375,1401]" captionStart="FIGURES 185–196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185–196. Parameres of Eophileurus Arrow, 1908. 185–192, E. minor Yang & Pathomwattananurak, new species, 185–187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189–190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193–196, E. sidereus Yang & Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="25" pageNumber="476">193–196</figureCitation>
|
||
), inner overlapped processes absent in
|
||
<taxonomicName id="4C704D0EE84417757C98FAB168D8A753" authorityName="Endrodi" authorityYear="1977" box="[754,929,1375,1401]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="nicobarensis">
|
||
<emphasis id="B904EA9FE84417757C98FAB168D8A753" box="[754,929,1375,1401]" italics="true" pageId="25" pageNumber="476">E. nicobarensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134B2A08E84417757DDBFAB16F45A753" box="[945,1084,1375,1401]" captionStart="FIGURES 79–86" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[180,1404,199,1908]" captionTargetId="figure-18@16.[169,1419,181,1920]" captionTargetPageId="16" captionText="FIGURES 79–86. Type specimens of Eophileurus nicobarensis Endrődi, 1977 (images ©HNHM/Ottó Merkl). 69–83, holotype, male, 79, habitus, dorsal; 80, habitus, lateral; 81, parameres, apical; 82, parameres, lateral; 83, labels; 84–86, allotype, female, 84, habitus, dorsal; 85, habitus, lateral; 86, labels." figureDoi="http://doi.org/10.5281/zenodo.6853968" httpUri="https://zenodo.org/record/6853968/files/figure.png" pageId="25" pageNumber="476">Figs. 81–82</figureCitation>
|
||
).
|
||
<taxonomicName id="4C704D0EE84417757A3AFAB16FB4A753" authority="Yang & Pathomwattananurak, 2022" authorityName="Yang & Pathomwattananurak" authorityYear="2022" box="[1104,1229,1375,1401]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="sidereus" status="sp. nov.">
|
||
<emphasis id="B904EA9FE84417757A3AFAB16FB4A753" box="[1104,1229,1375,1401]" italics="true" pageId="25" pageNumber="476">E. sidereus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B904EA9FE84417757AB6FAB16E1FA752" bold="true" box="[1244,1382,1374,1401]" pageId="25" pageNumber="476">
|
||
<taxonomicNameLabel id="A23757E4E84417757AB6FAB16E1FA752" box="[1244,1382,1374,1401]" pageId="25" pageNumber="476" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
also resembles
|
||
<taxonomicName id="4C704D0EE84417757F65FA6D6971A7B7" authority="Prell, 1913" authorityName=", Prell Determined" authorityYear="1913" box="[271,520,1411,1437]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B904EA9FE84417757F65FA6D6AFCA7B7" box="[271,389,1411,1437]" italics="true" pageId="25" pageNumber="476">E. gracilis</emphasis>
|
||
<bibRefCitation id="EFE14B7CE84417757FE7FA6D6971A7B7" author="Prell, H." box="[397,520,1411,1437]" pageId="25" pageNumber="476" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
for having similar external characters (
|
||
<figureCitation id="134B2A08E84417757DACFA6D6F29A7B7" box="[966,1104,1410,1437]" captionStart="FIGURES 54–59" captionStartId="12.[151,264,1933,1957]" captionTargetBox="[158,1426,643,1893]" captionTargetId="figure-215@12.[151,1435,633,1909]" captionTargetPageId="12" captionText="FIGURES 54–59. Lectotype of Eophileurus gracilis Prell, 1913. 54, habitus, dorsal; 55, habitus, lateral; 56, habitus, ventral; 57, parameres, apical; 58, parameres, lateral; 59, labels." figureDoi="http://doi.org/10.5281/zenodo.6853960" httpUri="https://zenodo.org/record/6853960/files/figure.png" pageId="25" pageNumber="476">Figs. 54–56</figureCitation>
|
||
,
|
||
<figureCitation id="134B2A08E84417757A31FA6D6FDAA7B7" box="[1115,1187,1411,1437]" captionStart="FIGURES 60–64" captionStartId="13.[151,264,1304,1328]" captionTargetBox="[159,1434,424,1278]" captionTargetId="figure-125@13.[151,1435,417,1280]" captionTargetPageId="13" captionText="FIGURES 60–64. Lectotype of Eophileurus andamanicus Arrow, 1914 (images by Keita Matsumoto © Natural History Museum, London). 60, habitus, dorsal; 61, habitus, lateral; 62, parameres, apical; 63, parameres, lateral; 64, labels." figureDoi="http://doi.org/10.5281/zenodo.6853962" httpUri="https://zenodo.org/record/6853962/files/figure.png" pageId="25" pageNumber="476">60–61</figureCitation>
|
||
) and similarly shaped overlapped processes at the inner sides of the parameres. Although having these similarities, the new species can be easily distinguished from
|
||
<taxonomicName id="4C704D0EE84417757FD1FA256948A7CF" authorityName=", Prell Determined" authorityYear="1913" box="[443,561,1483,1509]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B904EA9FE84417757FD1FA256948A7CF" box="[443,561,1483,1509]" italics="true" pageId="25" pageNumber="476">E. gracilis</emphasis>
|
||
</taxonomicName>
|
||
by the rest of the parameres: the base of the parameres extending backwards, overlapped processes at the inner sides near base absent, strongly backwardly curved process at the outer sides absent, but with an additional pair of outward processes at the outer sides near the apex (
|
||
<figureCitation id="134B2A08E84417757A0CF9FD6E73A407" box="[1126,1290,1555,1581]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral & Strnad, 1992 (Longli); 169–172, E. minor Yang & Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang & Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang & Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang & Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="25" pageNumber="476">Figs. 181–184</figureCitation>
|
||
,
|
||
<figureCitation id="134B2A08E84417757B7EF9FD6E0FA407" box="[1300,1398,1555,1581]" captionStart="FIGURES 185–196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185–196. Parameres of Eophileurus Arrow, 1908. 185–192, E. minor Yang & Pathomwattananurak, new species, 185–187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189–190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193–196, E. sidereus Yang & Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="25" pageNumber="476">193–196</figureCitation>
|
||
for
|
||
<taxonomicName id="4C704D0EE84417757EFDF9D96A6BA47B" authority="Yang & Pathomwattananurak, 2022" authorityName="Yang & Pathomwattananurak" authorityYear="2022" box="[151,274,1591,1617]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="sidereus" status="sp. nov.">
|
||
<emphasis id="B904EA9FE84417757EFDF9D96A6BA47B" box="[151,274,1591,1617]" italics="true" pageId="25" pageNumber="476">E. sidereus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B904EA9FE84417757F4BF9D96AD3A47A" bold="true" box="[289,426,1590,1617]" pageId="25" pageNumber="476">
|
||
<taxonomicNameLabel id="A23757E4E84417757F4BF9D96AD3A47A" box="[289,426,1590,1617]" pageId="25" pageNumber="476" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="134B2A08E84417757FD2F9D96946A47B" box="[440,575,1590,1617]" captionStart="FIGURES 54–59" captionStartId="12.[151,264,1933,1957]" captionTargetBox="[158,1426,643,1893]" captionTargetId="figure-215@12.[151,1435,633,1909]" captionTargetPageId="12" captionText="FIGURES 54–59. Lectotype of Eophileurus gracilis Prell, 1913. 54, habitus, dorsal; 55, habitus, lateral; 56, habitus, ventral; 57, parameres, apical; 58, parameres, lateral; 59, labels." figureDoi="http://doi.org/10.5281/zenodo.6853960" httpUri="https://zenodo.org/record/6853960/files/figure.png" pageId="25" pageNumber="476">Figs. 57–58</figureCitation>
|
||
,
|
||
<figureCitation id="134B2A08E84417757C20F9D969E8A47B" box="[586,657,1591,1617]" captionStart="FIGURES 60–64" captionStartId="13.[151,264,1304,1328]" captionTargetBox="[159,1434,424,1278]" captionTargetId="figure-125@13.[151,1435,417,1280]" captionTargetPageId="13" captionText="FIGURES 60–64. Lectotype of Eophileurus andamanicus Arrow, 1914 (images by Keita Matsumoto © Natural History Museum, London). 60, habitus, dorsal; 61, habitus, lateral; 62, parameres, apical; 63, parameres, lateral; 64, labels." figureDoi="http://doi.org/10.5281/zenodo.6853962" httpUri="https://zenodo.org/record/6853962/files/figure.png" pageId="25" pageNumber="476">62–63</figureCitation>
|
||
for
|
||
<taxonomicName id="4C704D0EE84417757CD5F9D9684DA47B" authorityName=", Prell Determined" authorityYear="1913" box="[703,820,1591,1617]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B904EA9FE84417757CD5F9D9684DA47B" box="[703,820,1591,1617]" italics="true" pageId="25" pageNumber="476">E. gracilis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6506E84417757EADF9B56AFFA497" pageId="25" pageNumber="476" type="etymology">
|
||
<paragraph id="8BCF368DE84417757EADF9B56AFFA497" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
|
||
<emphasis id="B904EA9FE84417757EADF9B56A37A45F" bold="true" box="[199,334,1627,1653]" pageId="25" pageNumber="476">Etymology.</emphasis>
|
||
This new species is named for its shiny elytra with dense punctures and micropunctures, with the reflection of light onto the punctures and micropunctures giving the impression of a starry sky. The name “sidereus” means “full of stars”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6506E84417757EADF92868BFA567" pageId="25" pageNumber="476" type="distribution">
|
||
<paragraph id="8BCF368DE84417757EADF9286AAAA4CB" blockId="25.[151,1437,151,2013]" box="[199,467,1734,1761]" pageId="25" pageNumber="476">
|
||
<emphasis id="B904EA9FE84417757EADF9286A18A4CA" bold="true" box="[199,353,1734,1760]" pageId="25" pageNumber="476">Distribution.</emphasis>
|
||
<collectingCountry id="F367761DE84417757F0DF9296AB7A4CB" box="[359,462,1735,1761]" name="Thailand" pageId="25" pageNumber="476">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCF368DE84417757EADF90468BFA567" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="476">
|
||
<emphasis id="B904EA9FE84417757EADF9046A33A52E" bold="true" box="[199,330,1770,1796]" pageId="25" pageNumber="476">Variations.</emphasis>
|
||
This species also has parameres with distinct individual variations. With
|
||
<specimenCount id="9D76FD04E84417757AD3F9056E53A52F" box="[1209,1322,1771,1797]" pageId="25" pageNumber="476" type="male">four male</specimenCount>
|
||
offspring from the same wild female all having slightly differently shaped parameres, as shown in
|
||
<figureCitation id="134B2A08E84417757AE9F8E16E5EA503" box="[1155,1319,1807,1833]" captionStart="FIGURES 185–196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185–196. Parameres of Eophileurus Arrow, 1908. 185–192, E. minor Yang & Pathomwattananurak, new species, 185–187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189–190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193–196, E. sidereus Yang & Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="25" pageNumber="476">Figs. 193–196</figureCitation>
|
||
. The most significant variation here is the inner processes pointing different angles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6506E84417767EADF8B969AFA34F" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476" type="discussion">
|
||
<paragraph id="8BCF368DE84417767EADF8B969AFA34F" blockId="25.[151,1437,151,2013]" lastBlockId="26.[151,1437,151,357]" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476">
|
||
<emphasis id="B904EA9FE84417757EADF8B96A42A55B" bold="true" box="[199,315,1879,1905]" pageId="25" pageNumber="476">Remarks.</emphasis>
|
||
This species resembles both
|
||
<taxonomicName id="4C704D0EE84417757CECF8B968A4A55B" authority="Endrodi, 1977" authorityName="Endrodi" authorityYear="1977" box="[646,989,1879,1905]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="nicobarensis">
|
||
<emphasis id="B904EA9FE84417757CECF8B9684DA55B" box="[646,820,1879,1905]" italics="true" pageId="25" pageNumber="476">E. nicobarensis</emphasis>
|
||
<bibRefCitation id="EFE14B7CE84417757D56F8B968A4A55B" author="Endrodi, S." box="[828,989,1879,1905]" pageId="25" pageNumber="476" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C704D0EE84417757A7FF8B96E68A55B" authority="Prell, 1913" authorityName=", Prell Determined" authorityYear="1913" box="[1045,1297,1879,1905]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B904EA9FE84417757A7FF8B96FF2A55B" box="[1045,1163,1879,1905]" italics="true" pageId="25" pageNumber="476">E. gracilis</emphasis>
|
||
<bibRefCitation id="EFE14B7CE84417757AF8F8B96E68A55B" author="Prell, H." box="[1170,1297,1879,1905]" pageId="25" pageNumber="476" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
, but is more closely related to the former for an essential reason: the base of the parameres of
|
||
<taxonomicName id="4C704D0EE84417757A51F8956FC1A5BF" authority="Yang & Pathomwattananurak, 2022" authorityName="Yang & Pathomwattananurak" authorityYear="2022" box="[1083,1208,1915,1941]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="sidereus" status="sp. nov.">
|
||
<emphasis id="B904EA9FE84417757A51F8956FC1A5BF" box="[1083,1208,1915,1941]" italics="true" pageId="25" pageNumber="476">E. sidereus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B904EA9FE84417757AA2F8956E2AA5BE" bold="true" box="[1224,1363,1914,1941]" pageId="25" pageNumber="476">
|
||
<taxonomicNameLabel id="A23757E4E84417757AA2F8956E2AA5BE" box="[1224,1363,1914,1941]" pageId="25" pageNumber="476" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="134B2A08E84417757B0EF8956B85A593" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral & Strnad, 1992 (Longli); 169–172, E. minor Yang & Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang & Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang & Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang & Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="25" pageNumber="476">Figs. 181–184</figureCitation>
|
||
,
|
||
<figureCitation id="134B2A08E84417757F61F8716A17A593" box="[267,366,1951,1977]" captionStart="FIGURES 185–196" captionStartId="31.[151,264,1633,1657]" captionTargetBox="[184,1415,199,1580]" captionTargetId="figure-18@31.[151,1435,181,1609]" captionTargetPageId="31" captionText="FIGURES 185–196. Parameres of Eophileurus Arrow, 1908. 185–192, E. minor Yang & Pathomwattananurak, new species, 185–187 from male offspring of the same wild female from Saraburi, 188 from Dong Tien; 189–190 from Bua Yai; 191 from Mueang Nakhon Ratchasima; 192 from Phanat Nikhom; 193–196, E. sidereus Yang & Pathomwattananurak, new species, male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853992" httpUri="https://zenodo.org/record/6853992/files/figure.png" pageId="25" pageNumber="476">193–196</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C704D0EE84417757FDFF871691FA593" authorityName="Endrodi" authorityYear="1977" box="[437,614,1951,1977]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="nicobarensis">
|
||
<emphasis id="B904EA9FE84417757FDFF871691FA593" box="[437,614,1951,1977]" italics="true" pageId="25" pageNumber="476">E. nicobarensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134B2A08E84417757C10F871687DA593" box="[634,772,1951,1977]" captionStart="FIGURES 79–86" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[180,1404,199,1908]" captionTargetId="figure-18@16.[169,1419,181,1920]" captionTargetPageId="16" captionText="FIGURES 79–86. Type specimens of Eophileurus nicobarensis Endrődi, 1977 (images ©HNHM/Ottó Merkl). 69–83, holotype, male, 79, habitus, dorsal; 80, habitus, lateral; 81, parameres, apical; 82, parameres, lateral; 83, labels; 84–86, allotype, female, 84, habitus, dorsal; 85, habitus, lateral; 86, labels." figureDoi="http://doi.org/10.5281/zenodo.6853968" httpUri="https://zenodo.org/record/6853968/files/figure.png" pageId="25" pageNumber="476">Figs. 81–82</figureCitation>
|
||
) both backwardly extended, but not in
|
||
<taxonomicName id="4C704D0EE84417757AB2F8716E28A593" authorityName=", Prell Determined" authorityYear="1913" box="[1240,1361,1951,1977]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B904EA9FE84417757AB2F8716E28A593" box="[1240,1361,1951,1977]" italics="true" pageId="25" pageNumber="476">E. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134B2A08E84417757B0EF8716BA6A5F7" captionStart="FIGURES 54–59" captionStartId="12.[151,264,1933,1957]" captionTargetBox="[158,1426,643,1893]" captionTargetId="figure-215@12.[151,1435,633,1909]" captionTargetPageId="12" captionText="FIGURES 54–59. Lectotype of Eophileurus gracilis Prell, 1913. 54, habitus, dorsal; 55, habitus, lateral; 56, habitus, ventral; 57, parameres, apical; 58, parameres, lateral; 59, labels." figureDoi="http://doi.org/10.5281/zenodo.6853960" httpUri="https://zenodo.org/record/6853960/files/figure.png" pageId="25" pageNumber="476">Figs. 57–58</figureCitation>
|
||
,
|
||
<figureCitation id="134B2A08E84417757E87F82D6A41A5F7" box="[237,312,1987,2013]" captionStart="FIGURES 60–64" captionStartId="13.[151,264,1304,1328]" captionTargetBox="[159,1434,424,1278]" captionTargetId="figure-125@13.[151,1435,417,1280]" captionTargetPageId="13" captionText="FIGURES 60–64. Lectotype of Eophileurus andamanicus Arrow, 1914 (images by Keita Matsumoto © Natural History Museum, London). 60, habitus, dorsal; 61, habitus, lateral; 62, parameres, apical; 63, parameres, lateral; 64, labels." figureDoi="http://doi.org/10.5281/zenodo.6853962" httpUri="https://zenodo.org/record/6853962/files/figure.png" pageId="25" pageNumber="476">62–63</figureCitation>
|
||
). The rest of the parameres of the new species also shows much more similarity to
|
||
<taxonomicName id="4C704D0EE84417757A84F82D6EE5A5F7" authorityName="Endrodi" authorityYear="1977" box="[1262,1436,1987,2013]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="nicobarensis">
|
||
<emphasis id="B904EA9FE84417757A84F82D6EE5A5F7" box="[1262,1436,1987,2013]" italics="true" pageId="25" pageNumber="476">E. nicobarensis</emphasis>
|
||
</taxonomicName>
|
||
than to
|
||
<taxonomicName id="4C704D0EE84717767E87FF796A1AA29B" authorityName=", Prell Determined" authorityYear="1913" box="[237,355,151,177]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B904EA9FE84717767E87FF796A1AA29B" box="[237,355,151,177]" italics="true" pageId="26" pageNumber="477">E. gracilis</emphasis>
|
||
</taxonomicName>
|
||
as stated in the diagnosis. The morphology of the parameres also suggest relation to the insular species such as
|
||
<taxonomicName id="4C704D0EE84717767F2FFF556942A2FF" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[325,571,187,213]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="javanus">
|
||
<emphasis id="B904EA9FE84717767F2FFF556AC3A2FE" box="[325,442,187,212]" italics="true" pageId="26" pageNumber="477">E. javanus</emphasis>
|
||
<bibRefCitation id="EFE14B7CE84717767FABFF556942A2FF" author="Prell, H." box="[449,571,187,213]" pageId="26" pageNumber="477" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
due to them having extended backward base of the parameres, and the similar curvature of the middle and apex region of the parameres in lateral view, but since the overlapping of the base of the parameres rarely appears on any insular species, they are therefore less closely related than to the aforementioned two species. Overall,
|
||
<taxonomicName id="4C704D0EE84717767FE3FEC9697AA36B" authority="Yang & Pathomwattananurak, 2022" authorityName="Yang & Pathomwattananurak" authorityYear="2022" box="[393,515,295,321]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="sidereus" status="sp. nov.">
|
||
<emphasis id="B904EA9FE84717767FE3FEC9697AA36B" box="[393,515,295,321]" italics="true" pageId="26" pageNumber="477">E. sidereus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B904EA9FE84717767C78FEC969E2A36A" bold="true" box="[530,667,294,321]" pageId="26" pageNumber="477">
|
||
<taxonomicNameLabel id="A23757E4E84717767C78FEC969E2A36A" box="[530,667,294,321]" pageId="26" pageNumber="477" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
might be an intermediating species between Indochinese and insular species since its parameres show similarity to both.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |