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<document id="AAA781EC5BDB081EC6FEFDE61E7016E5" ID-CLB-Dataset="99517" ID-DOI="10.11646/zootaxa.5165.4.1" ID-GBIF-Dataset="cd84b880-d77f-401a-bf4c-a98021a5e166" ID-ISSN="1175-5326" ID-Zenodo-Dep="6853942" ID-ZooBank="DBE812FF-7B25-4522-97FE-45A86ECAEFD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658127043518" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi &amp; Pathomwattananurak, Wuttipon" docDate="2022" docId="03D9879BE848177B7EFDFA246E02A4CB" docLanguage="en" docName="zootaxa.5165.4.1.pdf" docOrigin="Zootaxa 5165 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eophileurus quadratifovealis Yang &amp; Pathomwattananurak 2022, new species" docType="treatment" docVersion="7" lastPageNumber="474" masterDocId="FFE0FFE3E85D176C7E6AFFEE6B79A22A" masterDocTitle="Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species" masterLastPageNumber="485" masterPageNumber="451" pageNumber="472" updateTime="1699360166553" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A981EE8446C5030D04A39140AD081CEF">Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species</mods:title>
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<mods:name id="47579861B703D8F32EF282DD797ADDC1" type="personal">
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<mods:roleTerm id="8D7D9D21BBAAD9B877648ECC10E87D5D">Author</mods:roleTerm>
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<mods:namePart id="9BEE6195F0DF197F15B00B63AE28C0F5">Yang, Qiao-Zhi</mods:namePart>
<mods:affiliation id="00A21A3AC1C5CC5F6B00CD2C927F4919">3 B, Pingguoshequ, Baiziwan Road, Chaoyang Dist., Beijing, 100022, P. R. China.</mods:affiliation>
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</mods:role>
<mods:namePart id="FEC5F7F8CEF58D54685801019DF7C101">Pathomwattananurak, Wuttipon</mods:namePart>
<mods:affiliation id="6B487860CFE149924CE6EB9E92F18AC1">1051 / 38 Wiang, Mueang Dist., Chiang Rai Province, 57000, Thailand.</mods:affiliation>
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<mods:title id="699AB94F977C825057749A202C430C26">Zootaxa</mods:title>
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<mods:part id="8503BCF7543A1CC624C1F88AC353803D">
<mods:date id="00D9515AF3702889686759AE47D20773">2022</mods:date>
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<mods:number id="B830060A7CEB98104B5D0C33A5328C74">2022-07-18</mods:number>
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<mods:classification id="49E66578F960134362105494B4E95836">journal article</mods:classification>
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<mods:identifier id="98804AFC870A414DA109193E80074116" type="DOI">10.11646/zootaxa.5165.4.1</mods:identifier>
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<treatment id="03D9879BE848177B7EFDFA246E02A4CB" ID-DOI="http://doi.org/10.5281/zenodo.6853940" ID-GBIF-Taxon="197819089" ID-Zenodo-Dep="6853940" LSID="urn:lsid:plazi:treatment:03D9879BE848177B7EFDFA246E02A4CB" httpUri="http://treatment.plazi.org/id/03D9879BE848177B7EFDFA246E02A4CB" lastPageId="23" lastPageNumber="474" pageId="21" pageNumber="472">
<subSubSection id="C36A6506E84817797EFDFA246F61A7CE" box="[151,1048,1482,1509]" pageId="21" pageNumber="472" type="nomenclature">
<paragraph id="8BCF368DE84817797EFDFA246F61A7CE" blockId="21.[151,1048,1482,1545]" box="[151,1048,1482,1509]" pageId="21" pageNumber="472">
<heading id="D08781E1E84817797EFDFA246F61A7CE" bold="true" box="[151,1048,1482,1509]" fontSize="11" level="1" pageId="21" pageNumber="472" reason="1">
<emphasis id="B904EA9FE84817797EFDFA246F61A7CE" bold="true" box="[151,1048,1482,1509]" pageId="21" pageNumber="472">
<taxonomicName id="4C704D0EE84817797EFDFA246802A7CF" authority="Yang &amp; Pathomwattananurak" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[151,891,1482,1509]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="quadratifovealis" status="sp. nov.">
<emphasis id="B904EA9FE84817797EFDFA246A81A7CF" bold="true" box="[151,504,1482,1509]" italics="true" pageId="21" pageNumber="472">Eophileurus quadratifovealis</emphasis>
Yang &amp; Pathomwattananurak
</taxonomicName>
,
<taxonomicNameLabel id="A23757E4E84817797DE2FA256F61A7CE" box="[904,1048,1482,1509]" pageId="21" pageNumber="472" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84817797EFDFA01693DA423" box="[151,580,1518,1545]" pageId="21" pageNumber="472" type="description">
<paragraph id="8BCF368DE84817797EFDFA01693DA423" blockId="21.[151,1048,1482,1545]" box="[151,580,1518,1545]" pageId="21" pageNumber="472">
(
<figureCitation id="134B2A08E84817797EF5FA016B85A423" box="[159,252,1519,1545]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1853]" captionTargetBox="[151,1435,1065,1805]" captionTargetId="figure-436@2.[151,1435,1065,1805]" captionTargetPageId="2" captionText="FIGURE 1. Elytra of Eophileurus Arrow, 1908 species. a, E. quadrigeminatus Arrow, 1914; b, E. quadratifovealis Yang &amp; Pathomwattananurak, new species; c, E. felschei Prell, 1913. Primary costae marked in yellow and labeled in capital letters, interstices labeled in numbers without additional coloring." figureDoi="http://doi.org/10.5281/zenodo.6853944" httpUri="https://zenodo.org/record/6853944/files/figure.png" pageId="21" pageNumber="472">Figs. 1b</figureCitation>
,
<figureCitation id="134B2A08E84817797F63FA016A26A423" box="[265,351,1519,1545]" captionStart="FIGURES 99–102" captionStartId="22.[151,264,1943,1967]" captionTargetBox="[176,1407,194,1900]" captionTargetId="figure-18@22.[160,1428,181,1920]" captionTargetPageId="22" captionText="FIGURES 99–102. Habitus of Eophileurus quadratifovealis Yang &amp; Pathomwattananurak, new species. 99–100, holotype; 101–102, paratype; 99, 101 dorsal view; 100, 102, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853974" httpUri="https://zenodo.org/record/6853974/files/figure.png" pageId="21" pageNumber="472">99–102</figureCitation>
,
<figureCitation id="134B2A08E84817797F01FA016AB4A423" box="[363,461,1519,1545]" captionStart="FIGURES109–116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109–116.HabitusofEophileurusArrow,1908species(obliqueview).109–110,E.minorYang&amp;Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111–112, E. prelli Yang &amp; Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113–114, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115–116, E. sidereus Yang &amp; Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="21" pageNumber="472">113–114</figureCitation>
,
<figureCitation id="134B2A08E84817797FB3FA006945A423" box="[473,572,1518,1545]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169–172, E. minor Yang &amp; Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="21" pageNumber="472">175–180</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E848177B7EFDF9D96ADBA673" lastPageId="23" lastPageNumber="474" pageId="21" pageNumber="472" type="materials_examined">
<paragraph id="8BCF368DE84817797EFDF9D96FD9A503" blockId="21.[151,1437,1590,2013]" pageId="21" pageNumber="472">
<emphasis id="B904EA9FE84817797EFDF9D96A31A47A" bold="true" box="[151,328,1590,1617]" pageId="21" pageNumber="472">Type material.</emphasis>
(
<specimenCount id="9D76FD04E84817797F36F9D96AEEA47B" box="[348,407,1591,1617]" pageId="21" pageNumber="472" type="male">3♂♂</specimenCount>
).
<materialsCitation id="3B183CD0E84817797FC5F9D96804A45F" ID-GBIF-Occurrence="3863916378" collectingDate="2019-03-31" collectionCode="THNHM" collectorName="T. Unnahachote &amp; T. Hongsuwong" country="Thailand" location="Pluang Subdist." pageId="21" pageNumber="472" specimenCount="1" specimenCount-male="1" stateProvince="Khao Khitchakut Dist." typeStatus="holotype">
<emphasis id="B904EA9FE84817797FC5F9D9695CA47B" bold="true" box="[431,549,1591,1617]" pageId="21" pageNumber="472">
<typeStatus id="54CB882FE84817797FC5F9D96959A47B" box="[431,544,1591,1617]" pageId="21" pageNumber="472" type="holotype">Holotype</typeStatus>
:
</emphasis>
(
<specimenCount id="9D76FD04E84817797C53F9D96929A47B" box="[569,592,1591,1617]" pageId="21" pageNumber="472" type="male">♂</specimenCount>
,
<collectionCode id="ED61AE48E84817797C35F9D969B3A47B" box="[607,714,1591,1617]" pageId="21" pageNumber="472">THNHM</collectionCode>
), “
<location id="8EAF6056E84817797C80F9D968DAA47B" LSID="urn:lsid:plazi:treatment:03D9879BE848177B7EFDFA246E02A4CB:8EAF6056E84817797C80F9D968DAA47B" box="[746,931,1591,1617]" country="Thailand" name="Pluang Subdist." pageId="21" pageNumber="472" stateProvince="Khao Khitchakut Dist.">Pluang Subdist.</location>
,
<collectingRegion id="49B4F86FE84817797DDBF9D96FC1A47B" box="[945,1208,1591,1617]" pageId="21" pageNumber="472">Khao Khitchakut Dist.</collectingRegion>
,
<collectingRegion id="49B4F86FE84817797AACF9D96EE0A47B" box="[1222,1433,1591,1617]" country="Thailand" name="Chanthaburi" pageId="21" pageNumber="472">Chanthaburi Prov.</collectingRegion>
,
<collectingCountry id="F367761DE84817797EFDF9B56B84A45F" box="[151,253,1627,1653]" name="Thailand" pageId="21" pageNumber="472">Thailand</collectingCountry>
,
<date id="FFCE104DE84817797F64F9B56AF6A45F" box="[270,399,1627,1653]" pageId="21" pageNumber="472" value="2019-03-31">
<collectingDate id="EF8AE9A5E84817797F64F9B56AF6A45F" box="[270,399,1627,1653]" pageId="21" pageNumber="472" value="2019-03-31">31.III.2019</collectingDate>
</date>
,
<collectorName id="2685535BE84817797FF5F9B5692FA45F" box="[415,598,1627,1653]" pageId="21" pageNumber="472">T. Unnahachote</collectorName>
&amp;
<collectorName id="2685535BE84817797CE9F9B56842A45F" box="[643,827,1627,1653]" pageId="21" pageNumber="472">T. Hongsuwong</collectorName>
leg.”
</materialsCitation>
<emphasis id="B904EA9FE84817797DE0F9B56F79A45F" bold="true" box="[906,1024,1627,1653]" pageId="21" pageNumber="472">
<typeStatus id="54CB882FE84817797DE0F9B56882A45F" box="[906,1019,1627,1653]" pageId="21" pageNumber="472" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D76FD04E84817797A66F9B56F56A45F" box="[1036,1071,1627,1653]" pageId="21" pageNumber="472" type="male">1♂</specimenCount>
(
<collectionCode id="ED61AE48E84817797A2EF9B56FF1A45F" box="[1092,1160,1627,1653]" pageId="21" pageNumber="472">CWT</collectionCode>
), “Mu Si Subdist., Pak Chong Dist.,
<collectingRegion id="49B4F86FE84817797F5DF9916911A4B3" box="[311,616,1663,1689]" country="Thailand" name="Nakhon Ratchasima" pageId="21" pageNumber="472">Nakhon Ratchasima Prov.</collectingRegion>
,
<collectingCountry id="F367761DE84817797C1DF99169A4A4B3" box="[631,733,1663,1689]" name="Thailand" pageId="21" pageNumber="472">Thailand</collectingCountry>
,
<date id="FFCE104DE84817797C84F99168FBA4B3" box="[750,898,1663,1689]" pageId="21" pageNumber="472" value="2019-08-12">12.VIII.2019</date>
, W. Pathomwattananurak leg.”;
<specimenCount id="9D76FD04E84817797A95F9916E5BA4B3" box="[1279,1314,1663,1689]" pageId="21" pageNumber="472" type="male">1♂</specimenCount>
(
<collectionCode id="ED61AE48E84817797B5CF9916EE9A4B3" box="[1334,1424,1663,1689]" pageId="21" pageNumber="472">BMNH</collectionCode>
),
<typeStatus id="54CB882FE84817797EFDF94D6A56A497" box="[151,303,1699,1725]" pageId="21" pageNumber="472" type="paralectotype">Paralectotype</typeStatus>
of
<taxonomicName id="4C704D0EE84817797F30F94D698DA497" authority="Arrow, 1914" authorityName="Arrow" authorityYear="1914" box="[346,756,1699,1725]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B904EA9FE84817797F30F94D6920A496" box="[346,601,1699,1725]" italics="true" pageId="21" pageNumber="472">Eophileurus siamensis</emphasis>
<bibRefCitation id="EFE14B7CE84817797C0BF94D698DA497" author="Arrow, G. J." box="[609,756,1699,1725]" pageId="21" pageNumber="472" pagination="257 - 276" refId="ref16776" refString="Arrow, G. J. (1914) Some further notes on Lamellicorn beetles of the sub-family Dynastinae. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (14), 257 - 276. https: // doi. org / 10.1080 / 00222931408693571" type="journal article" year="1914">Arrow, 1914</bibRefCitation>
</taxonomicName>
, labeled: “SYN-TYPE // Chatabon S. S. Flower. 99-243. //
<collectionCode id="ED61AE48E84817797EFDF9296A7DA4CB" box="[151,260,1735,1761]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="21" pageNumber="472" type="Museum">NHMUK</collectionCode>
014380701”, two additional red labels will be provided labeling: “
<typeStatus id="54CB882FE84817797D93F9296FEDA4CB" box="[1017,1172,1735,1761]" pageId="21" pageNumber="472" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C704D0EE84817797AF7F9296A55A52F" authority="Arrow, 1914" authorityName="Arrow" authorityYear="1914" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B904EA9FE84817797AF7F9296EE5A4CA" box="[1181,1436,1735,1761]" italics="true" pageId="21" pageNumber="472">Eophileurus siamensis</emphasis>
<bibRefCitation id="EFE14B7CE84817797EFDF9056A55A52F" author="Arrow, G. J." box="[151,300,1771,1797]" pageId="21" pageNumber="472" pagination="257 - 276" refId="ref16776" refString="Arrow, G. J. (1914) Some further notes on Lamellicorn beetles of the sub-family Dynastinae. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (14), 257 - 276. https: // doi. org / 10.1080 / 00222931408693571" type="journal article" year="1914">Arrow, 1914</bibRefCitation>
</taxonomicName>
des. Yang &amp; Pathomwattananurak 2022” and “
<typeStatus id="54CB882FE84817797D04F90568AAA52F" box="[878,979,1771,1797]" pageId="21" pageNumber="472" type="paratype">Paratype</typeStatus>
<taxonomicName id="4C704D0EE84817797D8BF9056AB6A503" authority="Yang &amp; Pathomwattananurak, 2022" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="quadratifovealis">
<emphasis id="B904EA9FE84817797D8BF9056E49A52F" box="[993,1328,1771,1797]" italics="true" pageId="21" pageNumber="472">Eophileurus quadratifovealis</emphasis>
Yang &amp; Pathomwattananurak, 2022
</taxonomicName>
”, examined through photographs provided by Keita Matsumoto.
</paragraph>
<paragraph id="8BCF368DE848177B7EADF8DD682DA1AB" blockId="21.[151,1437,1590,2013]" lastBlockId="23.[151,1437,151,1761]" lastPageId="23" lastPageNumber="474" pageId="21" pageNumber="472">
<emphasis id="B904EA9FE84817797EADF8DD6A4DA567" bold="true" box="[199,308,1843,1869]" pageId="21" pageNumber="472">
<typeStatus id="54CB882FE84817797EADF8DD6A4DA567" box="[199,308,1843,1869]" pageId="21" pageNumber="472" type="holotype">Holotype</typeStatus>
</emphasis>
(male).
<emphasis id="B904EA9FE84817797FF9F8DC6A8AA566" bold="true" box="[403,499,1842,1868]" pageId="21" pageNumber="472">General</emphasis>
(
<figureCitation id="134B2A08E84817797C69F8DD69E0A567" box="[515,665,1843,1869]" captionStart="FIGURES 99–102" captionStartId="22.[151,264,1943,1967]" captionTargetBox="[176,1407,194,1900]" captionTargetId="figure-18@22.[160,1428,181,1920]" captionTargetPageId="22" captionText="FIGURES 99–102. Habitus of Eophileurus quadratifovealis Yang &amp; Pathomwattananurak, new species. 99–100, holotype; 101–102, paratype; 99, 101 dorsal view; 100, 102, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853974" httpUri="https://zenodo.org/record/6853974/files/figure.png" pageId="21" pageNumber="472">Figs. 99–100</figureCitation>
,
<figureCitation id="134B2A08E84817797CCFF8DD69B6A567" box="[677,719,1843,1869]" captionStart="FIGURES109–116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109–116.HabitusofEophileurusArrow,1908species(obliqueview).109–110,E.minorYang&amp;Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111–112, E. prelli Yang &amp; Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113–114, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115–116, E. sidereus Yang &amp; Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="21" pageNumber="472">113</figureCitation>
): Body black with slight reddish color, rather oblong, flattened, surface shiny; ventral surface clads with extremely sparse, short reddish-yellow setae.
<emphasis id="B904EA9FE84817797AECF8B96FB6A55B" bold="true" box="[1158,1231,1879,1905]" pageId="21" pageNumber="472">Head:</emphasis>
Clypeus sparsely punctate, punctures minute, with a short, backward horn at center and a ridge from horn to apex; clypeal apex slightly rounded. Frons depressed, disk almost impunctate, with punctures at each side anteriorly only. Mandible slightly sinuate at outer margin.
<emphasis id="B904EA9FE84817797C63F82D69F2A5F7" bold="true" box="[521,651,1987,2013]" pageId="21" pageNumber="472">Pronotum:</emphasis>
Surface with dense and large punctures, becoming denser and larger close to all margins. Anterior region with a large, slightly deep, square-shape fovea; punctures in fovea spare and large, transverse, coalescent at anterior part, slightly denser and smaller at posterior region; sides of fovea almost parallel, with a very weak protuberance at each side at posterior part. Sides parallel at middle; all margins beaded except before fovea, more strongly at sides. Anterior angle acute, apex slightly rounded, posterior angle obtusely angulate.
<emphasis id="B904EA9FE84A177B7F6EFEC86AFCA36A" bold="true" box="[260,389,294,320]" pageId="23" pageNumber="474">Scutellum:</emphasis>
Surface with dense punctures except region close to apex.
<emphasis id="B904EA9FE84A177B7A73FEC96F16A36B" bold="true" box="[1049,1135,295,321]" pageId="23" pageNumber="474">Elytra:</emphasis>
Surface with dense micropunctures, and dense, U-shape, large punctures. Interstice 1 with punctures in four irregular rows combined at middle to two irregular rows, interstice 2 with chaotic punctures, rows unrecognizable. Primary costa A highly distinct, primary costa B not distinct. Subapical umbones moderately prominent. Margins of elytra moderately horizontally dilated from base to middle.
<emphasis id="B904EA9FE84A177B7FBFFE586928A3FA" bold="true" box="[469,593,438,464]" pageId="23" pageNumber="474">Pygidium:</emphasis>
convex, with fine punctures throughout, punctures denser and smaller close to all margins, and slightly sparser near apex.
<emphasis id="B904EA9FE84A177B7CCDFE35682BA3DF" bold="true" box="[679,850,475,501]" pageId="23" pageNumber="474">Metasternum:</emphasis>
Surface mainly black, with reddish color at central region; punctures dense and large near all sides; central portion with sparse and small punctures only; setae yellow, very sparse and short, only at posterior region close to each side.
<emphasis id="B904EA9FE84A177B7D1FFDCC6F0AA016" bold="true" box="[885,1139,546,572]" pageId="23" pageNumber="474">Abdominal ventrites:</emphasis>
Surfaces with micropunctures throughout and larger punctures at each side; last ventrite with rugose, large punctures at anterior region, micropunctures denser and larger than on other ventrites; ventrites 5 and 6 with very shallow, round depressions at center.
<emphasis id="B904EA9FE84A177B7E83FD616A53A083" bold="true" box="[233,298,655,681]" pageId="23" pageNumber="474">Legs:</emphasis>
Protibia tridentate, protarsi strongly thickened, inner protibial claw strongly enlarged and elongated. Inner metatibial spur long, slightly curved outward, outer metatibial spur shorter, more strongly curved outward.
<emphasis id="B904EA9FE84A177B7EFDFD396A61A0DB" bold="true" box="[151,280,727,753]" pageId="23" pageNumber="474">Parameres</emphasis>
(
<figureCitation id="134B2A08E84A177B7F43FD396AB7A0DB" box="[297,462,726,753]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169–172, E. minor Yang &amp; Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="23" pageNumber="474">Figs. 175–176</figureCitation>
): In frontal view (
<figureCitation id="134B2A08E84A177B7CCEFD396871A0DB" box="[676,776,726,753]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169–172, E. minor Yang &amp; Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="23" pageNumber="474">Fig. 175</figureCitation>
), base overlapped, extending upward; outer sides sinuate, prominent outward at posterior two-fifth, anterior one-seventh and near apex; inner sides also sinuate, prominent inward posterior to opening; in lateral view (
<figureCitation id="134B2A08E84A177B7CF6FCF16879A113" box="[668,768,799,825]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169–172, E. minor Yang &amp; Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="23" pageNumber="474">Fig. 176</figureCitation>
), base projected upward and slightly backward; apex bent strongly backward, with a tiny, backward, triangular process near apex of each paramere; in ventral view, ventral base with a pair of short, outward, flat, semicircular processes.
</paragraph>
<caption id="DF0F6605E84B177A7EFDF8796866A5FE" ID-DOI="http://doi.org/10.5281/zenodo.6853974" ID-Zenodo-Dep="6853974" httpUri="https://zenodo.org/record/6853974/files/figure.png" pageId="22" pageNumber="473" startId="22.[151,264,1943,1967]" targetBox="[176,1407,194,1900]" targetPageId="22">
<paragraph id="8BCF368DE84B177A7EFDF8796866A5FE" blockId="22.[151,1436,1943,2004]" pageId="22" pageNumber="473">
<emphasis id="B904EA9FE84B177A7EFDF8796A1AA585" bold="true" box="[151,355,1943,1967]" pageId="22" pageNumber="473">FIGURES 99–102.</emphasis>
Habitus of
<taxonomicName id="4C704D0EE84B177A7F88F8766F3EA59A" authority="Yang &amp; Pathomwattananurak" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[482,1095,1944,1968]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="quadratifovealis" status="sp. nov.">
<emphasis id="B904EA9FE84B177A7F88F8766875A59A" box="[482,780,1944,1968]" italics="true" pageId="22" pageNumber="473">Eophileurus quadratifovealis</emphasis>
Yang &amp; Pathomwattananurak
</taxonomicName>
,
<emphasis id="B904EA9FE84B177A7A39F8766FA1A585" bold="true" box="[1107,1240,1943,1968]" pageId="22" pageNumber="473">
<taxonomicNameLabel id="A23757E4E84B177A7A39F8766FADA585" box="[1107,1236,1943,1968]" pageId="22" pageNumber="473" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
99–100, holotype; 101–102, paratype; 99, 101 dorsal view; 100, 102, ventral view.
</paragraph>
</caption>
<paragraph id="8BCF368DE84A177B7EADFC656E65A61F" blockId="23.[151,1437,151,1761]" pageId="23" pageNumber="474">
<emphasis id="B904EA9FE84A177B7EADFC656A46A18F" bold="true" box="[199,319,907,933]" pageId="23" pageNumber="474">
<typeStatus id="54CB882FE84A177B7EADFC656A46A18F" box="[199,319,907,933]" pageId="23" pageNumber="474" type="paratype">Paratypes</typeStatus>
</emphasis>
(male). Underdeveloped, but characters mainly stable, except clypeal horn much shorter, clypeus and frons slightly denser punctate; Fovea much smaller and shallower, rather rounded, with denser punctures; micropunctures on elytra denser (
<figureCitation id="134B2A08E84A177B7C61FC3D69D7A1C7" box="[523,686,979,1005]" captionStart="FIGURES 99–102" captionStartId="22.[151,264,1943,1967]" captionTargetBox="[176,1407,194,1900]" captionTargetId="figure-18@22.[160,1428,181,1920]" captionTargetPageId="22" captionText="FIGURES 99–102. Habitus of Eophileurus quadratifovealis Yang &amp; Pathomwattananurak, new species. 99–100, holotype; 101–102, paratype; 99, 101 dorsal view; 100, 102, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853974" httpUri="https://zenodo.org/record/6853974/files/figure.png" pageId="23" pageNumber="474">Figs. 101–102</figureCitation>
,
<figureCitation id="134B2A08E84A177B7CD2FC3D699CA1C7" box="[696,741,979,1005]" captionStart="FIGURES109–116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109–116.HabitusofEophileurusArrow,1908species(obliqueview).109–110,E.minorYang&amp;Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111–112, E. prelli Yang &amp; Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113–114, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115–116, E. sidereus Yang &amp; Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="23" pageNumber="474">114</figureCitation>
); Base of parameres wider and less upwardly extended in front view, region from middle to apex of parameres narrower, ventral basal processes visible in front view (
<figureCitation id="134B2A08E84A177B7B46FC196EF6A63B" box="[1324,1423,1015,1041]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169–172, E. minor Yang &amp; Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="23" pageNumber="474">Fig. 177</figureCitation>
), base of parameres more prominent and processes close to apex almost absent in lateral view (
<figureCitation id="134B2A08E84A177B7AC4FBF56E69A61F" box="[1198,1296,1051,1077]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169–172, E. minor Yang &amp; Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="23" pageNumber="474">Fig. 178</figureCitation>
).
</paragraph>
<paragraph id="8BCF368DE84A177B7EADFBD16ADBA673" blockId="23.[151,1437,151,1761]" box="[199,418,1087,1113]" pageId="23" pageNumber="474">
<emphasis id="B904EA9FE84A177B7EADFBD16A5CA673" bold="true" box="[199,293,1087,1113]" pageId="23" pageNumber="474">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84A177B7EADFB8D6988A68B" pageId="23" pageNumber="474" type="description">
<paragraph id="8BCF368DE84A177B7EADFB8D6988A68B" blockId="23.[151,1437,151,1761]" pageId="23" pageNumber="474">
<emphasis id="B904EA9FE84A177B7EADFB8D6A07A657" bold="true" box="[199,382,1123,1149]" pageId="23" pageNumber="474">Measurements.</emphasis>
Body length: male 21.8–25.0 mm (
<typeStatus id="54CB882FE84A177B7D73FB8D6804A657" box="[793,893,1123,1149]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
25.0 mm), female unknown; body width: male
<quantity id="4C889B68E84A177B7EFDFB686A43A68B" box="[151,314,1158,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" metricValueMax="1.21" metricValueMin="1.05" pageId="23" pageNumber="474" unit="mm" value="11.3" valueMax="12.1" valueMin="10.5">10.5–12.1 mm</quantity>
(
<typeStatus id="54CB882FE84A177B7F22FB696AD5A68B" box="[328,428,1159,1185]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
<quantity id="4C889B68E84A177B7FD9FB696962A68B" box="[435,539,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="23" pageNumber="474" unit="mm" value="12.1">12.1 mm</quantity>
), female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84A177B7EADFB446AC3A71B" pageId="23" pageNumber="474" type="diagnosis">
<paragraph id="8BCF368DE84A177B7EADFB446AC3A71B" blockId="23.[151,1437,151,1761]" pageId="23" pageNumber="474">
<emphasis id="B904EA9FE84A177B7EADFB446A3BA6EE" bold="true" box="[199,322,1194,1220]" pageId="23" pageNumber="474">Diagnosis.</emphasis>
This new species is most similar to
<taxonomicName id="4C704D0EE84A177B7C98FB456881A6EF" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[754,1016,1195,1221]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE84A177B7C98FB456814A6EF" box="[754,877,1195,1221]" italics="true" pageId="23" pageNumber="474">E. confinis</emphasis>
<bibRefCitation id="EFE14B7CE84A177B7D1DFB456881A6EF" author="Prell, H." box="[887,1016,1195,1221]" pageId="23" pageNumber="474" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
, but differs by: middle of pronotum parallel (
<figureCitation id="134B2A08E84A177B7E94FB216AE0A6C3" box="[254,409,1231,1257]" captionStart="FIGURES 99–102" captionStartId="22.[151,264,1943,1967]" captionTargetBox="[176,1407,194,1900]" captionTargetId="figure-18@22.[160,1428,181,1920]" captionTargetPageId="22" captionText="FIGURES 99–102. Habitus of Eophileurus quadratifovealis Yang &amp; Pathomwattananurak, new species. 99–100, holotype; 101–102, paratype; 99, 101 dorsal view; 100, 102, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853974" httpUri="https://zenodo.org/record/6853974/files/figure.png" pageId="23" pageNumber="474">Figs. 99, 101</figureCitation>
) whereas usually not parallel in
<taxonomicName id="4C704D0EE84A177B7D71FB2168EEA6C3" authorityName="Prell" authorityYear="1913" box="[795,919,1231,1257]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE84A177B7D71FB2168EEA6C3" box="[795,919,1231,1257]" italics="true" pageId="23" pageNumber="474">E. confinis</emphasis>
</taxonomicName>
; fovea square-shape for normally developed individual; interval 1 wider; parameres with different shape, especially without a pair of processes at the middle of the sides (
<figureCitation id="134B2A08E84A177B7F62FAF96AD7A71B" box="[264,430,1302,1329]" captionStart="FIGURES 165–184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165–184. Parameres of Eophileurus Arrow, 1908 species. 165–166, E. confinis Prell, 1913 (Bao Loc); 167–168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169–172, E. minor Yang &amp; Pathomwattananurak, new species, 169–170, holotype, 171–172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173–174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174–180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175–176, holotype, 177–178, paratype (Mu Si), 179–180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181–184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181–182, holotype, 183–184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="23" pageNumber="474">Figs. 175–180</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84A177B7EADFAD56913A7B7" pageId="23" pageNumber="474" type="etymology">
<paragraph id="8BCF368DE84A177B7EADFAD56913A7B7" blockId="23.[151,1437,151,1761]" pageId="23" pageNumber="474">
<emphasis id="B904EA9FE84A177B7EADFAD56A37A77F" bold="true" box="[199,334,1339,1365]" pageId="23" pageNumber="474">Etymology.</emphasis>
This new species is named for the unique square-shaped fovea of the
<typeStatus id="54CB882FE84A177B7A3AFAD56FCCA77F" box="[1104,1205,1339,1365]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
, which is not present in other Indochina
<taxonomicName id="4C704D0EE84A177B7F1AFAB16A80A753" authorityName=", Endrodi" authorityYear="1977" box="[368,505,1375,1401]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="B904EA9FE84A177B7F1AFAB16A80A753" box="[368,505,1375,1401]" italics="true" pageId="23" pageNumber="474">Eophileurus</emphasis>
</taxonomicName>
. The species epithet is a combination of the Latin adjective “
<emphasis id="B904EA9FE84A177B7AD3FAB16E67A753" box="[1209,1310,1375,1401]" italics="true" pageId="23" pageNumber="474">quadrate</emphasis>
” and noun “
<emphasis id="B904EA9FE84A177B7ECEFA6D6B98A7B7" box="[164,225,1411,1437]" italics="true" pageId="23" pageNumber="474">fovea</emphasis>
”, to be used as noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84A177B7EADFA486AAAA7EB" box="[199,467,1446,1473]" pageId="23" pageNumber="474" type="distribution">
<paragraph id="8BCF368DE84A177B7EADFA486AAAA7EB" blockId="23.[151,1437,151,1761]" box="[199,467,1446,1473]" pageId="23" pageNumber="474">
<emphasis id="B904EA9FE84A177B7EADFA486A18A7EA" bold="true" box="[199,353,1446,1472]" pageId="23" pageNumber="474">Distribution.</emphasis>
<collectingCountry id="F367761DE84A177B7F0DFA496AB7A7EB" box="[359,462,1447,1473]" name="Thailand" pageId="23" pageNumber="474">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84A177B7EADFA256E02A4CB" pageId="23" pageNumber="474" type="discussion">
<paragraph id="8BCF368DE84A177B7EADFA256E02A4CB" blockId="23.[151,1437,151,1761]" pageId="23" pageNumber="474">
<emphasis id="B904EA9FE84A177B7EADFA256A42A7CF" bold="true" box="[199,315,1483,1509]" pageId="23" pageNumber="474">Remarks.</emphasis>
This species is most closely related to
<taxonomicName id="4C704D0EE84A177B7D7BFA256F63A7CF" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[785,1050,1483,1509]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE84A177B7D7BFA2568F4A7CF" box="[785,909,1483,1509]" italics="true" pageId="23" pageNumber="474">E. confinis</emphasis>
<bibRefCitation id="EFE14B7CE84A177B7DF2FA256F63A7CF" author="Prell, H." box="[920,1050,1483,1509]" pageId="23" pageNumber="474" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
, whose distribution surrounds its distribution in a large scale. Currently, whether they are sympatric is unknown. With only
<specimenCount id="9D76FD04E84A177B7A1DFA016E53A423" box="[1143,1322,1519,1545]" pageId="23" pageNumber="474" type="generic">three specimens</specimenCount>
examined from adjacent but separated mountain ranges, their parameres show considerable variations. This is likely to be in similar situation as
<taxonomicName id="4C704D0EE84A177B7F18F9D96A92A47B" authorityName="Prell" authorityYear="1913" box="[370,491,1591,1617]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE84A177B7F18F9D96A92A47B" box="[370,491,1591,1617]" italics="true" pageId="23" pageNumber="474">E. confinis</emphasis>
</taxonomicName>
where individuals tend to have distinct variation on the parameres both individually and geographically. Since the overall structures of the new species’ parameres can be traced, the three individuals are treated as the same species. Additionally, due to the high similarity between the males of
<taxonomicName id="4C704D0EE84A177B7AD7F9916EEFA4B3" authority="Yang &amp; Pathomwattananurak, 2022" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[1213,1430,1663,1689]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="quadratifovealis" status="sp. nov.">
<emphasis id="B904EA9FE84A177B7AD7F9916EEFA4B3" box="[1213,1430,1663,1689]" italics="true" pageId="23" pageNumber="474">E. quadratifovealis</emphasis>
</taxonomicName>
,
<emphasis id="B904EA9FE84A177B7EFDF94D6A66A496" bold="true" box="[151,287,1698,1725]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="A23757E4E84A177B7EFDF94D6A66A496" box="[151,287,1698,1725]" pageId="23" pageNumber="474" rank="species">new species</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C704D0EE84A177B7F3EF94D6AB2A497" authorityName="Prell" authorityYear="1913" box="[340,459,1699,1725]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE84A177B7F3EF94D6AB2A497" box="[340,459,1699,1725]" italics="true" pageId="23" pageNumber="474">E. confinis</emphasis>
</taxonomicName>
, the author speculate that the females of this species might also be highly similar to each other and have elytral margin more strongly dilated than the males, as commonly seen in females of
<taxonomicName id="4C704D0EE84A177B7A97F9296E0CA4CB" authorityName="Prell" authorityYear="1913" box="[1277,1397,1735,1761]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE84A177B7A97F9296E0CA4CB" box="[1277,1397,1735,1761]" italics="true" pageId="23" pageNumber="474">E. confinis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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