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<document id="66B51C0303AAC728DFBEE11336CF54D6" ID-DOI="10.11646/zootaxa.5227.5.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7523293" ID-ZooBank="E0446DE0-C6A5-4A2B-88B8-EC2336990CB3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673389231651" checkinUser="plazi" docAuthor="Lin, Shuanghu, Zou, Hongfen, Huang, Min &amp; Zhang, Yalin" docDate="2023" docId="03D987850E3A5B7FFF57FBC26FD6F8B6" docLanguage="en" docName="zootaxa.5227.5.4.pdf" docOrigin="Zootaxa 5227 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dentaneura biangulata Lin &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="569" masterDocId="FFE0FFFD0E3B5B7EFFC0FF9A6D3EFFF4" masterDocTitle="Five new species and two new records of Erythroneurini from China (Hemiptera, Cicadellidae, Typhlocybinae)" masterLastPageNumber="582" masterPageNumber="568" pageNumber="569" updateTime="1699446224892" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2BE7C82F202087E5F95ED2C23267282F">Five new species and two new records of Erythroneurini from China (Hemiptera, Cicadellidae, Typhlocybinae)</mods:title>
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<mods:namePart id="B8BE378C8F495DCEB8CF4CD179E7E29C">Lin, Shuanghu</mods:namePart>
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<mods:affiliation id="7762F413545E5560D3D8B8461C1FCE71">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; 509082999 @ qq. com; https: // orcid. org / 0000 - 0003 - 0625 - 5381</mods:affiliation>
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<mods:namePart id="EF5EE8B2EC764F0C069B80881CD90DEC">Zou, Hongfen</mods:namePart>
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<mods:affiliation id="E226B48E3053E31471D0FFDD8B88C332">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; 2097853970 @ qq. com; https: // orcid. org / 0000 - 0002 - 3292 - 0341</mods:affiliation>
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<mods:namePart id="9593035B364AB895C4370F71A45E0660">Huang, Min</mods:namePart>
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<mods:affiliation id="B652C48CECB79DBC31ECC88A9AF4173C">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; huangmin @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0001 - 7621 - 4863</mods:affiliation>
<mods:nameIdentifier id="E31084A9181786F9ED72E2974AAEAB5D" type="email">huangmin@nwsuaf.edu.cn</mods:nameIdentifier>
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<mods:namePart id="FA2BDF158EE9AD86FA60FA661C095CD7">Zhang, Yalin</mods:namePart>
<mods:affiliation id="9EF4CD3A7E9C65E78C512289652AAB65">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, Northwest A &amp; F University, Yangling, Shaanxi 712100, China</mods:affiliation>
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<mods:date id="DDE4ED4DCDB0B7D5AAD5DFDE5126A4C2">2023</mods:date>
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<treatment id="03D987850E3A5B7FFF57FBC26FD6F8B6" ID-DOI="http://doi.org/10.5281/zenodo.7523220" ID-Zenodo-Dep="7523220" LSID="urn:lsid:plazi:treatment:03D987850E3A5B7FFF57FBC26FD6F8B6" httpUri="http://treatment.plazi.org/id/03D987850E3A5B7FFF57FBC26FD6F8B6" lastPageNumber="569" pageId="1" pageNumber="569">
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<paragraph id="8BCF36930E3A5B7FFF57FBC26FFDFB86" blockId="1.[151,707,1111,1174]" box="[151,707,1111,1138]" pageId="1" pageNumber="569">
<heading id="D08781FF0E3A5B7FFF57FBC26FFDFB86" bold="true" box="[151,707,1111,1138]" fontSize="11" level="1" pageId="1" pageNumber="569" reason="1">
<emphasis id="B904EA810E3A5B7FFF57FBC26FFDFB86" bold="true" box="[151,707,1111,1138]" pageId="1" pageNumber="569">
<taxonomicName id="4C704D100E3A5B7FFF57FBC26F60FB86" ID-CoL="a651b3ab-193e-460b-b4fa-148c380bb8f4" authority="Lin &amp; Zhang" authorityName="Lin &amp; Zhang" authorityYear="2023" box="[151,606,1111,1138]" class="Insecta" family="Cicadellidae" genus="Dentaneura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="biangulata" status="sp. nov.">
<emphasis id="B904EA810E3A5B7FFF57FBC26C8CFB86" bold="true" box="[151,434,1111,1138]" italics="true" pageId="1" pageNumber="569">Dentaneura biangulata</emphasis>
Lin &amp; Zhang
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<taxonomicNameLabel id="A23757FA0E3A5B7FFDA5FBC26FFDFB86" box="[613,707,1112,1138]" pageId="1" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C36A65180E3A5B7FFF57FBE66E25FA0A" pageId="1" pageNumber="569" type="description">
<paragraph id="8BCF36930E3A5B7FFF57FBE66C0DFB62" blockId="1.[151,707,1111,1174]" box="[151,307,1148,1174]" pageId="1" pageNumber="569">
(
<figureCitation id="134B2A160E3A5B7FFF5FFBE66C2FFB62" box="[159,273,1148,1174]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[199,1390,190,1913]" captionTargetId="figure-19@2.[186,1401,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. ad. Dentaneura biangulata Lin &amp; Zhang sp. nov. eh. Motaga rokfaoides Lin &amp; Zhang sp. nov. il. Seriana paraequata Lin &amp; Zhang sp. nov. mp. Shibinga graciliprocessa Lin &amp; Zhang sp. nov. qt. Shibinga grossiprocessa Lin &amp; Zhang sp. nov. a, c, e, g, i, k, m, o, q, s. dorsal view; b, f, j, n, r. lateral view; d, h, l, p, t. face." figureDoi="http://doi.org/10.5281/zenodo.7523295" httpUri="https://zenodo.org/record/7523295/files/figure.png" pageId="1" pageNumber="569">Figs 1ad</figureCitation>
,
<figureCitation id="134B2A160E3A5B7FFEDCFBE66C14FB62" box="[284,298,1148,1174]" captionStart="FIGURE 3" captionStartId="4.[151,250,1703,1727]" captionTargetBox="[156,1426,184,1667]" captionTargetId="figure-19@4.[151,1436,181,1679]" captionTargetPageId="4" captionText="FIGURE 3. Dentaneura biangulata Lin &amp; Zhang sp. nov. a. pygofer and anal tube, lateral view; b. abdominal apodemes; c. pygofer dorsal appendage, lateral view; d. connective; e. style, ventral view; f. style, lateral view; g. aedeagus, lateral view; h. aedeagus, ventral view; i. subapical protrusion of subgenital plate, posterior view; j. anal appendage, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7523299" httpUri="https://zenodo.org/record/7523299/files/figure.png" pageId="1" pageNumber="569">3</figureCitation>
)
</paragraph>
<paragraph id="8BCF36930E3A5B7FFF57FB5E6870FB2A" blockId="1.[151,1437,1220,1859]" box="[151,1358,1220,1247]" pageId="1" pageNumber="569">
<emphasis id="B904EA810E3A5B7FFF57FB5E6C17FB2A" bold="true" box="[151,297,1220,1246]" pageId="1" pageNumber="569">Description.</emphasis>
Crown with pair of dark spots. The other characters as in generic description (
<figureCitation id="134B2A160E3A5B7FFB58FB5E687EFB2A" box="[1176,1344,1220,1247]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[199,1390,190,1913]" captionTargetId="figure-19@2.[186,1401,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. ad. Dentaneura biangulata Lin &amp; Zhang sp. nov. eh. Motaga rokfaoides Lin &amp; Zhang sp. nov. il. Seriana paraequata Lin &amp; Zhang sp. nov. mp. Shibinga graciliprocessa Lin &amp; Zhang sp. nov. qt. Shibinga grossiprocessa Lin &amp; Zhang sp. nov. a, c, e, g, i, k, m, o, q, s. dorsal view; b, f, j, n, r. lateral view; d, h, l, p, t. face." figureDoi="http://doi.org/10.5281/zenodo.7523295" httpUri="https://zenodo.org/record/7523295/files/figure.png" pageId="1" pageNumber="569">Figs 1a, b, c, d</figureCitation>
).
</paragraph>
<paragraph id="8BCF36930E3A5B7FFF07FB726DC1FAD2" blockId="1.[151,1437,1220,1859]" pageId="1" pageNumber="569">
Abdominal apodemes not extended to sternite IV (
<figureCitation id="134B2A160E3A5B7FFD3FFB726E6CFAF6" box="[767,850,1256,1282]" captionStart="FIGURE 3" captionStartId="4.[151,250,1703,1727]" captionTargetBox="[156,1426,184,1667]" captionTargetId="figure-19@4.[151,1436,181,1679]" captionTargetPageId="4" captionText="FIGURE 3. Dentaneura biangulata Lin &amp; Zhang sp. nov. a. pygofer and anal tube, lateral view; b. abdominal apodemes; c. pygofer dorsal appendage, lateral view; d. connective; e. style, ventral view; f. style, lateral view; g. aedeagus, lateral view; h. aedeagus, ventral view; i. subapical protrusion of subgenital plate, posterior view; j. anal appendage, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7523299" httpUri="https://zenodo.org/record/7523299/files/figure.png" pageId="1" pageNumber="569">Fig. 3b</figureCitation>
). Anal tube short, with poorly developed appendage (
<figureCitation id="134B2A160E3A5B7FFF5FFA966DCFFAD2" box="[159,241,1292,1318]" captionStart="FIGURE 3" captionStartId="4.[151,250,1703,1727]" captionTargetBox="[156,1426,184,1667]" captionTargetId="figure-19@4.[151,1436,181,1679]" captionTargetPageId="4" captionText="FIGURE 3. Dentaneura biangulata Lin &amp; Zhang sp. nov. a. pygofer and anal tube, lateral view; b. abdominal apodemes; c. pygofer dorsal appendage, lateral view; d. connective; e. style, ventral view; f. style, lateral view; g. aedeagus, lateral view; h. aedeagus, ventral view; i. subapical protrusion of subgenital plate, posterior view; j. anal appendage, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7523299" httpUri="https://zenodo.org/record/7523299/files/figure.png" pageId="1" pageNumber="569">Fig. 3a</figureCitation>
).
</paragraph>
<paragraph id="8BCF36930E3A5B7FFF07FAAA6F75FA2E" blockId="1.[151,1437,1220,1859]" pageId="1" pageNumber="569">
Pygofer with few short microsetae near posterior margin and group of macrosetae at cephalo-ventral angle (
<figureCitation id="134B2A160E3A5B7FFAAFFAAA6D8BFA9A" captionStart="FIGURE 3" captionStartId="4.[151,250,1703,1727]" captionTargetBox="[156,1426,184,1667]" captionTargetId="figure-19@4.[151,1436,181,1679]" captionTargetPageId="4" captionText="FIGURE 3. Dentaneura biangulata Lin &amp; Zhang sp. nov. a. pygofer and anal tube, lateral view; b. abdominal apodemes; c. pygofer dorsal appendage, lateral view; d. connective; e. style, ventral view; f. style, lateral view; g. aedeagus, lateral view; h. aedeagus, ventral view; i. subapical protrusion of subgenital plate, posterior view; j. anal appendage, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7523299" httpUri="https://zenodo.org/record/7523299/files/figure.png" pageId="1" pageNumber="569">Fig. 3a</figureCitation>
). Pygofer dorsal appendage not exceeding posterior margin of pygofer lobe (
<figureCitation id="134B2A160E3A5B7FFBE2FACE69ABFA9A" box="[1058,1173,1364,1390]" captionStart="FIGURE 3" captionStartId="4.[151,250,1703,1727]" captionTargetBox="[156,1426,184,1667]" captionTargetId="figure-19@4.[151,1436,181,1679]" captionTargetPageId="4" captionText="FIGURE 3. Dentaneura biangulata Lin &amp; Zhang sp. nov. a. pygofer and anal tube, lateral view; b. abdominal apodemes; c. pygofer dorsal appendage, lateral view; d. connective; e. style, ventral view; f. style, lateral view; g. aedeagus, lateral view; h. aedeagus, ventral view; i. subapical protrusion of subgenital plate, posterior view; j. anal appendage, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7523299" httpUri="https://zenodo.org/record/7523299/files/figure.png" pageId="1" pageNumber="569">Figs 3a, c</figureCitation>
). Subgenital plate with about 5 macrosetae scattered on lateral surface near outer margin, 2 angled processes apically, 34 peg-like on each angled process (
<figureCitation id="134B2A160E3A5B7FFE8FFA066C9CFA42" box="[335,418,1436,1462]" captionStart="FIGURE 3" captionStartId="4.[151,250,1703,1727]" captionTargetBox="[156,1426,184,1667]" captionTargetId="figure-19@4.[151,1436,181,1679]" captionTargetPageId="4" captionText="FIGURE 3. Dentaneura biangulata Lin &amp; Zhang sp. nov. a. pygofer and anal tube, lateral view; b. abdominal apodemes; c. pygofer dorsal appendage, lateral view; d. connective; e. style, ventral view; f. style, lateral view; g. aedeagus, lateral view; h. aedeagus, ventral view; i. subapical protrusion of subgenital plate, posterior view; j. anal appendage, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7523299" httpUri="https://zenodo.org/record/7523299/files/figure.png" pageId="1" pageNumber="569">Fig. 3a</figureCitation>
). Connective V-shaped, manubrium short (
<figureCitation id="134B2A160E3A5B7FFC49FA066EE0FA42" box="[905,990,1436,1462]" captionStart="FIGURE 3" captionStartId="4.[151,250,1703,1727]" captionTargetBox="[156,1426,184,1667]" captionTargetId="figure-19@4.[151,1436,181,1679]" captionTargetPageId="4" captionText="FIGURE 3. Dentaneura biangulata Lin &amp; Zhang sp. nov. a. pygofer and anal tube, lateral view; b. abdominal apodemes; c. pygofer dorsal appendage, lateral view; d. connective; e. style, ventral view; f. style, lateral view; g. aedeagus, lateral view; h. aedeagus, ventral view; i. subapical protrusion of subgenital plate, posterior view; j. anal appendage, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7523299" httpUri="https://zenodo.org/record/7523299/files/figure.png" pageId="1" pageNumber="569">Fig. 3d</figureCitation>
). Aedeagal shaft compressed distinctly, gourd-like in ventral view (
<figureCitation id="134B2A160E3A5B7FFE0AFA5A6F03FA2E" box="[458,573,1472,1498]" captionStart="FIGURE 3" captionStartId="4.[151,250,1703,1727]" captionTargetBox="[156,1426,184,1667]" captionTargetId="figure-19@4.[151,1436,181,1679]" captionTargetPageId="4" captionText="FIGURE 3. Dentaneura biangulata Lin &amp; Zhang sp. nov. a. pygofer and anal tube, lateral view; b. abdominal apodemes; c. pygofer dorsal appendage, lateral view; d. connective; e. style, ventral view; f. style, lateral view; g. aedeagus, lateral view; h. aedeagus, ventral view; i. subapical protrusion of subgenital plate, posterior view; j. anal appendage, posterior view." figureDoi="http://doi.org/10.5281/zenodo.7523299" httpUri="https://zenodo.org/record/7523299/files/figure.png" pageId="1" pageNumber="569">
Figs
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, h
</figureCitation>
).
</paragraph>
<paragraph id="8BCF36930E3A5B7FFF07FA7E6E25FA0A" blockId="1.[151,1437,1220,1859]" box="[199,795,1508,1535]" pageId="1" pageNumber="569">
<emphasis id="B904EA810E3A5B7FFF07FA7E6C4DFA0A" bold="true" box="[199,371,1508,1534]" pageId="1" pageNumber="569">Measurement.</emphasis>
Male
<quantity id="4C889B760E3A5B7FFE7AFA7E6F67FA0B" box="[442,601,1508,1535]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.75" metricValueMin="2.65" pageId="1" pageNumber="569" unit="mm" value="2.7" valueMax="2.75" valueMin="2.65">2.652.75mm</quantity>
, female
<quantity id="4C889B760E3A5B7FFD77FA7E6E29FA0A" box="[695,791,1508,1534]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="569" unit="mm" value="2.7">2.70mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A65180E3A5B7FFF07F9926CABF9B2" pageId="1" pageNumber="569" type="materials_examined">
<paragraph id="8BCF36930E3A5B7FFF07F9926CABF9B2" blockId="1.[151,1437,1220,1859]" pageId="1" pageNumber="569">
<emphasis id="B904EA810E3A5B7FFF07F9926C8CF9D6" bold="true" box="[199,434,1544,1570]" pageId="1" pageNumber="569">Material examined.</emphasis>
<materialsCitation id="3B183CCE0E3A5B7FFE7AF99269F8F9D6" box="[442,1222,1544,1572]" collectingDate="2013-07-27" country="China" elevation="1950" location="Qiannan" pageId="1" pageNumber="569" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<typeStatus id="54CB88310E3A5B7FFE7AF9926F19F9D6" box="[442,551,1544,1570]" pageId="1" pageNumber="569" type="holotype">Holotype</typeStatus>
: ♁,
<collectingCountry id="F36776030E3A5B7FFD96F9926F8CF9D6" box="[598,690,1544,1570]" name="China" pageId="1" pageNumber="569">CHINA</collectingCountry>
,
<collectingRegion id="49B4F8710E3A5B7FFD7EF9926E5AF9D7" box="[702,868,1544,1571]" country="China" name="Guizhou" pageId="1" pageNumber="569">Guizhou Prov.</collectingRegion>
,
<location id="8EAF60480E3A5B7FFCAFF9926EEFF9D6" LSID="urn:lsid:plazi:treatment:03D987850E3A5B7FFF57FBC26FD6F8B6:8EAF60480E3A5B7FFCAFF9926EEFF9D6" box="[879,977,1544,1570]" country="China" name="Qiannan" pageId="1" pageNumber="569" stateProvince="Guizhou">Qiannan</location>
,
<quantity id="4C889B760E3A5B7FFC1DF9926910F9D7" box="[989,1070,1544,1571]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="569" unit="m" value="1950.0">
<elevation id="005DD1A00E3A5B7FFC1DF9926910F9D7" box="[989,1070,1544,1571]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="569" unit="m" value="1950.0">1950m</elevation>
</quantity>
,
<date id="FFCE10530E3A5B7FFBFAF99269F8F9D6" box="[1082,1222,1544,1570]" pageId="1" pageNumber="569" value="2013-07-27">
<collectingDate id="EF8AE9BB0E3A5B7FFBFAF99269F8F9D6" box="[1082,1222,1544,1570]" pageId="1" pageNumber="569" value="2013-07-27">2013.VII.27</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B183CCE0E3A5B7FFB12F9936CACF9B2" collectingDate="2013-07-27" country="China" elevation="1950" location="Qiannan" pageId="1" pageNumber="569" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="54CB88310E3A5B7FFB12F993687DF9D7" box="[1234,1347,1545,1571]" pageId="1" pageNumber="569" type="paratype">Paratypes</typeStatus>
: 4♁
<specimenCount id="9D76FD1A0E3A5B7FFAB7F99368A8F9D0" box="[1399,1430,1545,1572]" pageId="1" pageNumber="569" type="female">1♀</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A65180E3A5B7FFF07F9CA6E75F90E" pageId="1" pageNumber="569" type="discussion">
<paragraph id="8BCF36930E3A5B7FFF07F9CA6E75F90E" blockId="1.[151,1437,1220,1859]" pageId="1" pageNumber="569">
<emphasis id="B904EA810E3A5B7FFF07F9CA6C05F99E" bold="true" box="[199,315,1616,1642]" pageId="1" pageNumber="569">Remarks.</emphasis>
This new species is similar to
<taxonomicName id="4C704D100E3A5B7FFD4DF9CB6976F99E" authority="Song, Li &amp; Dai" authorityName="Song, Li &amp; Dai" authorityYear="2016" box="[653,1096,1616,1642]" class="Insecta" family="Cicadellidae" genus="Dentaneura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="henanensis">
<emphasis id="B904EA810E3A5B7FFD4DF9CB6EABF99E" box="[653,917,1616,1642]" italics="true" pageId="1" pageNumber="569">Dentaneura henanensis</emphasis>
Song, Li &amp; Dai
</taxonomicName>
, but has the pygofer dorsal appendage not exceeding the posterior margin, the subgenital plate with 2 angled processes apically, and the aedeagal shaft obviously expanded medially in ventral view.
<taxonomicName id="4C704D100E3A5B7FFD1CF90369A5F946" authority="Song, Li &amp; Dai" authorityName="Song, Li &amp; Dai" authorityYear="2016" box="[732,1179,1688,1714]" class="Insecta" family="Cicadellidae" genus="Dentaneura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="henanensis">
<emphasis id="B904EA810E3A5B7FFD1CF9036ED8F946" box="[732,998,1688,1714]" italics="true" pageId="1" pageNumber="569">Dentaneura henanensis</emphasis>
Song, Li &amp; Dai
</taxonomicName>
has the pygofer dorsal appendage distinctly exceeding the posterior margin, the subgenital plate with only 1 angled process apically, and the aedeagal shaft slightly expanded medially in ventral view.
</paragraph>
</subSubSection>
<subSubSection id="C36A65180E3A5B7FFF07F89E6FD6F8B6" pageId="1" pageNumber="569" type="etymology">
<paragraph id="8BCF36930E3A5B7FFF07F89E6FD6F8B6" blockId="1.[151,1437,1220,1859]" pageId="1" pageNumber="569">
<emphasis id="B904EA810E3A5B7FFF07F89E6C70F8EA" bold="true" box="[199,334,1796,1822]" pageId="1" pageNumber="569">Etymology.</emphasis>
The specific epithet is derived from Latin prefix “bi-” and the Latin word “angulatus”, referring to the subgenital plate with 2 angled processes apically.
</paragraph>
</subSubSection>
</treatment>
</document>