treatments-xml/data/03/D4/B9/03D4B94FFFC72039FC32FF15FBF0C697.xml
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<document id="2C1A64C29BA33AF8B5F92F93F01D2098" ID-CLB-Dataset="24150" ID-DOI="10.5252/zoosystema2020v42a3" ID-GBIF-Dataset="7bdbb02c-ce5d-4df5-846a-b45c54fedc5a" ID-Zenodo-Dep="3652791" ID-ZooBank="urn:lsid:zoobank.org:pub:27C94F0F-BB9B-40A3-B615-4DB19C94F042" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581020160253" checkinUser="plazi" docAuthor="Kantor, Yuri, Kosyan, Alisa &amp; Sorokin, Pavel" docDate="2020" docId="03D4B94FFFC72039FC32FF15FBF0C697" docLanguage="en" docName="zoosystema2020v42a3.pdf" docOrigin="Zoosystema 42 (3)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Phaenomenella nicoi Kantor &amp; Kosyan &amp; Sorokin 2020, n. sp." docType="treatment" docUuid="BC0FCA38-4FC5-40F8-9D8C-159F76CA46BD" docUuidSource="ZooBank" docVersion="8" lastPageNumber="48" masterDocId="FFEDC137FFCB2028FFEFFFCDFFA9C206" masterDocTitle="On the taxonomic position of Phaenomenella Fraussen &amp; Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species" masterLastPageNumber="55" masterPageNumber="33" pageNumber="43" updateTime="1698780820290" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EFCB12E66B8BF873A5E456F25C4BF58A">On the taxonomic position of Phaenomenella Fraussen &amp; Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species</mods:title>
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<mods:namePart id="1480713C2C3D2BD21D65F12900561E53">Kantor, Yuri</mods:namePart>
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<mods:namePart id="DC915C2F26D91A64CC041BE78A3EDBBB">Kosyan, Alisa</mods:namePart>
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<mods:namePart id="E556C8DFBE7F3F4155F7672F7564AD38">Sorokin, Pavel</mods:namePart>
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<mods:title id="E5607B56791B8D3A394948F831AF7883">Zoosystema</mods:title>
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<mods:date id="C03EC0B1CCACFB08D5220AE21FD786F0">2020</mods:date>
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<taxonomicName id="4C7D73DAFFC72024FC32FF15FB14C2F5" ID-CoL="7SNZR" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[989,1213,216,243]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="43" phylum="Mollusca" rank="species" species="nicoi" status="sp. nov.">
<emphasis id="B909D44BFFC72024FC32FF15FB14C2F5" bold="true" box="[989,1213,216,243]" italics="true" pageId="12" pageNumber="43">Phaenomenella nicoi</emphasis>
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(
<figureCitation id="134614DCFFC72024FBF5FF34FBF3C312" box="[1050,1114,249,276]" captionStart="FIG" captionStartId="13.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1835]" captionTargetId="figure@13.[132,1455,215,1836]" captionTargetPageId="13" captionText="FIG. 5. — Radulae of Phaenomenella spp.A, B, Phaenomenella samadiae n. sp.; holotype,MNHN-IM-2013-61617 (shell on Fig. 4A-C) (A); MNHN-IM-2013-61674 (shell on Fig. 4 E, F) (B); C-F. Phaenomenella nicoi n. sp.; C, holotype, MNHN-IM-2013-61585 (shell on Fig. 7A-C); D, MNHN-IM-2013-59398 (shell on Fig. 7D); E, MNHN-IM-2013-61639 (shell on Fig. 7E); F, MNHN-IM-2013-61673 (shell on Fig. 7I). Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3652815" httpUri="https://zenodo.org/record/3652815/files/figure.png" pageId="12" pageNumber="43">Figs 5</figureCitation>
C-F; 7; 8)
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MATERIAL EXAMINED. —
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.
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<location id="8EA25E82FFC72024FB7CFEBCFB67C38F" LSID="urn:lsid:plazi:treatment:03D4B94FFFC72039FC32FF15FBF0C697:8EA25E82FFC72024FB7CFEBCFB67C38F" box="[1171,1230,369,393]" country="China" latitude="20.116667" longLatPrecision="1269" longitude="116.36667" municipality="Sea" name="South" pageId="12" pageNumber="43">South</location>
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MNHN- IM-2013-61585 (sequenced,
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A-C);
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off
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, ZHONGSHA 2015; st. CP4132;
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SEQUENCED MATERIAL. —
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lv;
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(
<figureCitation id="134614DCFFC72024FC08FE34FB94C017" box="[999,1085,505,529]" captionStart="FIG" captionStartId="15.[132,143,1895,1913]" captionTargetBox="[131,1454,215,1827]" captionTargetId="figure@15.[131,1454,214,1827]" captionTargetPageId="15" captionText="FIG. 7. — Shells of Phaenomenella nicoi n. sp.: A-C, holotype, MNHN-IM-2013-61585, SL 41.1 mm; D, ZHONGSHA 2015, st. CP4157, sequenced spm, MNHN- IM-2013-59398, SL 37.5 mm;E, ZHONGSHA 2015,st. CP4133,sequenced spm,MNHN-IM-2013-61639, SL 43.2 mm;F, AURORA 2007,st. CP2685,sequenced spm, MNHN-IM-2007-34639, SL 34.7 mm; G, AURORA 2007, st. CP2685, not-sequenced spm, MNHN-IM-2007-34644, SL 38.7 mm; H, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61637, SL 30.6 mm (anatomy studied); I, ZHONGSHA 2015, st. CP4134, sequenced spm, MNHN-IM-2013-61673, SL 42 mm; J, ZHONGSHA 2015,st. CP4133,not-sequenced spm, MNHN-IM-2013-61636, SL 40.2 mm (anatomy studied); K, L, ZHONGSHA 2015, st. CP4132, not-sequenced spm, MNHN-IM-2013-61592, 39.1 mm; L, enlarged protoconch. Scale bars: A-K, 20 cm; L, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3652823" httpUri="https://zenodo.org/record/3652823/files/figure.png" pageId="12" pageNumber="43">Figs 7H</figureCitation>
;
<figureCitation id="134614DCFFC72024FBA2FE34FBF1C017" box="[1101,1112,505,529]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="12" pageNumber="43">8</figureCitation>
E-G),
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,
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off
<collectorName id="26886D8FFFC72024FB32FDD9FAE9C02A" box="[1245,1344,532,556]" pageId="12" pageNumber="43">Dong Sha</collectorName>
, ZHONG- SHA 2015, st. CP4133;
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lv.;
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Fig.
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); st. CP4134;
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lv;
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); st. CP4157;
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,
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<collectingCountry id="F36A48C9FFC72024FCC2FD55FC0BC0B6" box="[813,930,664,688]" name="Philippines" pageId="12" pageNumber="43">Philippines</collectingCountry>
6 1 l•
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; MNHN-IM-2007-34639;
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2007; st. CP2685;
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(
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Fig. 7
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.
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STUDIED MATERIAL. —
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lv;
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, MNHN-IM-2013-61592 (
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);
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off
<collectorName id="26886D8FFFC72024FCC2FCECFC23C13E" box="[813,906,800,825]" pageId="12" pageNumber="43">Dong Sha</collectorName>
, ZHONGSHA 2015; st. CP4132;
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lv;
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(
<figureCitation id="134614DCFFC72024FCDBFC98FCD9C16B" box="[820,880,853,877]" captionStart="FIG" captionStartId="15.[132,143,1895,1913]" captionTargetBox="[131,1454,215,1827]" captionTargetId="figure@15.[131,1454,214,1827]" captionTargetPageId="15" captionText="FIG. 7. — Shells of Phaenomenella nicoi n. sp.: A-C, holotype, MNHN-IM-2013-61585, SL 41.1 mm; D, ZHONGSHA 2015, st. CP4157, sequenced spm, MNHN- IM-2013-59398, SL 37.5 mm;E, ZHONGSHA 2015,st. CP4133,sequenced spm,MNHN-IM-2013-61639, SL 43.2 mm;F, AURORA 2007,st. CP2685,sequenced spm, MNHN-IM-2007-34639, SL 34.7 mm; G, AURORA 2007, st. CP2685, not-sequenced spm, MNHN-IM-2007-34644, SL 38.7 mm; H, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61637, SL 30.6 mm (anatomy studied); I, ZHONGSHA 2015, st. CP4134, sequenced spm, MNHN-IM-2013-61673, SL 42 mm; J, ZHONGSHA 2015,st. CP4133,not-sequenced spm, MNHN-IM-2013-61636, SL 40.2 mm (anatomy studied); K, L, ZHONGSHA 2015, st. CP4132, not-sequenced spm, MNHN-IM-2013-61592, 39.1 mm; L, enlarged protoconch. Scale bars: A-K, 20 cm; L, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3652823" httpUri="https://zenodo.org/record/3652823/files/figure.png" pageId="12" pageNumber="43">Figs 7J</figureCitation>
,
<figureCitation id="134614DCFFC72024FC95FC98FC2DC16B" box="[890,900,853,877]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="12" pageNumber="43">8</figureCitation>
A-D),
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<specimenCode id="DBDBA022FFC72024FBD2FC98FB19C16B" box="[1085,1200,853,877]" pageId="12" pageNumber="43">2013-63029</specimenCode>
,
<collectionCode id="ED6C909CFFC72024FB55FC9BFA9EC16B" box="[1210,1335,854,877]" pageId="12" pageNumber="43">MNHN-IM-</collectionCode>
<specimenCode id="DBDBA022FFC72024FAD9FC98FA00C16B" box="[1334,1449,853,877]" pageId="12" pageNumber="43">2013-63025</specimenCode>
,
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<specimenCode id="DBDBA022FFC72024FC5FFCBDFB80C181" box="[944,1065,879,904]" pageId="12" pageNumber="43">2013-61638</specimenCode>
; st. CP4133;
<geoCoordinate id="EE496E9EFFC72024FB59FCA2FAA5C181" box="[1206,1292,879,903]" degrees="19" direction="north" minutes="59" orientation="latitude" pageId="12" pageNumber="43" precision="925" value="19.983334">19°59N</geoCoordinate>
,
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;
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</quantity>
<specimenCode id="DBDBA022FFC72024FC95FC47FC3DC1A4" box="[890,916,906,930]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="43">
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<specimenCount id="9D7BC3D0FFC72024FC67FC47FC3DC1A4" box="[904,916,906,930]" pageId="12" pageNumber="43">2</specimenCount>
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</specimenCode>
lv,
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<specimenCode id="DBDBA022FFC72024FBDDFC47FB01C1A4" box="[1074,1192,906,930]" pageId="12" pageNumber="43">2013-61671</specimenCode>
,
<collectionCode id="ED6C909CFFC72024FB5CFC47FA9AC1A7" box="[1203,1331,906,929]" pageId="12" pageNumber="43">MNHN-IM-</collectionCode>
<specimenCode id="DBDBA022FFC72024FADDFC47FA01C1A4" box="[1330,1448,906,930]" pageId="12" pageNumber="43">2013-61672</specimenCode>
; st. CP4134;
<geoCoordinate id="EE496E9EFFC72024FC43FC69FBADC1BA" box="[940,1028,932,956]" degrees="19" direction="north" minutes="50" orientation="latitude" pageId="12" pageNumber="43" precision="925" value="19.833334">19°50N</geoCoordinate>
,
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;
<quantity id="4C85A5BCFFC72024FB94FC68FAACC1BA" box="[1147,1285,933,957]" metricMagnitude="3" metricUnit="m" metricValue="1.2029999999999998" metricValueMax="1.278" metricValueMin="1.128" pageId="12" pageNumber="43" unit="m" value="1203.0" valueMax="1278.0" valueMin="1128.0">
<elevation id="0050EF6AFFC72024FB94FC68FAACC1BA" box="[1147,1285,933,957]" metricMagnitude="3" metricUnit="m" metricValue="1.2029999999999998" metricValueMax="1.278" metricValueMin="1.128" pageId="12" pageNumber="43" unit="m" value="1203.0" valueMax="1278.0" valueMin="1128.0">1128-1278 m</elevation>
</quantity>
<specimenCode id="DBDBA022FFC72024FAE2FC68FA85C1BB" box="[1293,1324,933,957]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="43">
<materialsCitation id="3B150204FFC72024FAF0FC68FA85C1BB" ID-GBIF-Occurrence="2573220333" box="[1311,1324,933,957]" collectionCode="MNHN" country="China" elevation="1297" latitude="19.8" location="Helen Shoal" longLatPrecision="1270" longitude="116.48333" pageId="12" pageNumber="43" specimenCode="IM-2013-59397" specimenCount="1">
<specimenCount id="9D7BC3D0FFC72024FAF0FC68FA85C1BB" box="[1311,1324,933,957]" pageId="12" pageNumber="43">1</specimenCount>
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</specimenCode>
lv;
<collectionCode id="ED6C909CFFC72024FABCFC68FA06C1BA" box="[1363,1455,933,956]" pageId="12" pageNumber="43">MNHN-</collectionCode>
<specimenCode id="DBDBA022FFC72024FCC2FC72FC67C1D1" box="[813,974,959,983]" pageId="12" pageNumber="43">IM-2013-59397</specimenCode>
; st. CP4157;
<geoCoordinate id="EE496E9EFFC72024FBB8FC72FB07C1D1" box="[1111,1198,959,983]" degrees="19" direction="north" minutes="48" orientation="latitude" pageId="12" pageNumber="43" precision="925" value="19.8">19°48N</geoCoordinate>
,
<geoCoordinate id="EE496E9EFFC72024FB57FC72FABEC1D1" box="[1208,1303,959,983]" degrees="116" direction="east" minutes="29" orientation="longitude" pageId="12" pageNumber="43" precision="925" value="116.48333">116°29E</geoCoordinate>
;
<quantity id="4C85A5BCFFC72024FACEFC72FA02C1D0" box="[1313,1451,959,983]" metricMagnitude="3" metricUnit="m" metricValue="1.297" metricValueMax="1.389" metricValueMin="1.205" pageId="12" pageNumber="43" unit="m" value="1297.0" valueMax="1389.0" valueMin="1205.0">
<elevation id="0050EF6AFFC72024FACEFC72FA02C1D0" box="[1313,1451,959,983]" metricMagnitude="3" metricUnit="m" metricValue="1.297" metricValueMax="1.389" metricValueMin="1.205" pageId="12" pageNumber="43" unit="m" value="1297.0" valueMax="1389.0" valueMin="1205.0">1205-1389 m</elevation>
</quantity>
, S off
<location id="8EA25E82FFC72024FC89FC14FC4FC1F7" LSID="urn:lsid:plazi:treatment:03D4B94FFFC72039FC32FF15FBF0C697:8EA25E82FFC72024FC89FC14FC4FC1F7" box="[870,998,985,1009]" country="China" latitude="19.8" longLatPrecision="1270" longitude="116.48333" name="Helen Shoal" pageId="12" pageNumber="43">Helen Shoal</location>
<specimenCode id="DBDBA022FFC72024FC00FC17FBA7C1F7" box="[1007,1038,985,1009]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="43">
<materialsCitation id="3B150204FFC72024FBEEFC14FBA7C1F7" ID-GBIF-Occurrence="2573220330" box="[1025,1038,985,1009]" collectionCode="MNHN, MNHN" country="China" elevation="1219" latitude="18.816668" longLatPrecision="1274" longitude="113.96667" pageId="12" pageNumber="43" specimenCode="IM-2013-61815, IM-2013-59663" specimenCount="2">
<specimenCount id="9D7BC3D0FFC72024FBEEFC14FBA7C1F7" box="[1025,1038,985,1009]" pageId="12" pageNumber="43">2</specimenCount>
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lv;
<collectionCode id="ED6C909CFFC72024FBD6FC17FB6BC1F7" box="[1081,1218,986,1009]" pageId="12" pageNumber="43">MNHN-IM-</collectionCode>
<specimenCode id="DBDBA022FFC72024FB2EFC14FAE8C1F7" box="[1217,1345,985,1009]" pageId="12" pageNumber="43">2013-61815</specimenCode>
,
<collectionCode id="ED6C909CFFC72024FABEFC17FA19C1F7" box="[1361,1456,986,1009]" pageId="12" pageNumber="43">MNHN-</collectionCode>
<specimenCode id="DBDBA022FFC72024FCC2FC39FC67C60A" box="[813,974,1012,1036]" pageId="12" pageNumber="43">IM-2013-59663</specimenCode>
; st. CP4141;
<geoCoordinate id="EE496E9EFFC72024FBB8FC39FB04C60A" box="[1111,1197,1012,1036]" degrees="18" direction="north" minutes="49" orientation="latitude" pageId="12" pageNumber="43" precision="925" value="18.816668">18°49N</geoCoordinate>
,
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;
<quantity id="4C85A5BCFFC72024FACFFC39FA02C60D" box="[1312,1451,1012,1036]" metricMagnitude="3" metricUnit="m" metricValue="1.2185" metricValueMax="1.286" metricValueMin="1.151" pageId="12" pageNumber="43" unit="m" value="1218.5" valueMax="1286.0" valueMin="1151.0">
<elevation id="0050EF6AFFC72024FACFFC39FA02C60D" box="[1312,1451,1012,1036]" metricMagnitude="3" metricUnit="m" metricValue="1.2185" metricValueMax="1.286" metricValueMin="1.151" pageId="12" pageNumber="43" unit="m" value="1218.5" valueMax="1286.0" valueMin="1151.0">1151-1286 m</elevation>
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.
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<emphasis id="B909D44BFFC72024FCC2FBC3FC60C620" bold="true" box="[813,969,1038,1062]" pageId="12" pageNumber="43">
<collectingCountry id="F36A48C9FFC72024FCC2FBC3FC31C620" box="[813,920,1038,1062]" name="Philippines" pageId="12" pageNumber="43">Philippines</collectingCountry>
• • l•
</emphasis>
; MNHN-IM-2007-34644 (
<figureCitation id="134614DCFFC72024FB20FBC3FAA9C620" box="[1231,1280,1038,1062]" captionStart="FIG" captionStartId="15.[132,143,1895,1913]" captionTargetBox="[131,1454,215,1827]" captionTargetId="figure@15.[131,1454,214,1827]" captionTargetPageId="15" captionText="FIG. 7. — Shells of Phaenomenella nicoi n. sp.: A-C, holotype, MNHN-IM-2013-61585, SL 41.1 mm; D, ZHONGSHA 2015, st. CP4157, sequenced spm, MNHN- IM-2013-59398, SL 37.5 mm;E, ZHONGSHA 2015,st. CP4133,sequenced spm,MNHN-IM-2013-61639, SL 43.2 mm;F, AURORA 2007,st. CP2685,sequenced spm, MNHN-IM-2007-34639, SL 34.7 mm; G, AURORA 2007, st. CP2685, not-sequenced spm, MNHN-IM-2007-34644, SL 38.7 mm; H, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61637, SL 30.6 mm (anatomy studied); I, ZHONGSHA 2015, st. CP4134, sequenced spm, MNHN-IM-2013-61673, SL 42 mm; J, ZHONGSHA 2015,st. CP4133,not-sequenced spm, MNHN-IM-2013-61636, SL 40.2 mm (anatomy studied); K, L, ZHONGSHA 2015, st. CP4132, not-sequenced spm, MNHN-IM-2013-61592, 39.1 mm; L, enlarged protoconch. Scale bars: A-K, 20 cm; L, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3652823" httpUri="https://zenodo.org/record/3652823/files/figure.png" pageId="12" pageNumber="43">Fig. 7</figureCitation>
);
<collectingRegion id="49B9C6BBFFC72024FAFCFBC3FADDC620" box="[1299,1396,1038,1062]" country="Philippines" name="Aurora" pageId="12" pageNumber="43">AURORA</collectingRegion>
2007; st. CP2685;
<geoCoordinate id="EE496E9EFFC72024FC47FBE4FC56C647" box="[936,1023,1065,1089]" degrees="15" direction="north" minutes="00" orientation="latitude" pageId="12" pageNumber="43" precision="925" value="15.0">15°00N</geoCoordinate>
,
<geoCoordinate id="EE496E9EFFC72024FBE6FBE4FBCEC647" box="[1033,1127,1065,1089]" degrees="123" direction="east" minutes="06" orientation="longitude" pageId="12" pageNumber="43" precision="925" value="123.1">123°06E</geoCoordinate>
;
<quantity id="4C85A5BCFFC72024FB9EFBE4FB52C646" box="[1137,1275,1065,1089]" metricMagnitude="3" metricUnit="m" metricValue="1.2285" metricValueMax="1.302" metricValueMin="1.155" pageId="12" pageNumber="43" unit="m" value="1228.5" valueMax="1302.0" valueMin="1155.0">
<elevation id="0050EF6AFFC72024FB9EFBE4FB52C646" box="[1137,1275,1065,1089]" metricMagnitude="3" metricUnit="m" metricValue="1.2285" metricValueMax="1.302" metricValueMin="1.155" pageId="12" pageNumber="43" unit="m" value="1228.5" valueMax="1302.0" valueMin="1155.0">1155-1302 m</elevation>
</quantity>
, SL
<quantity id="4C85A5BCFFC72024FAC9FBE4FA2DC646" box="[1318,1412,1065,1089]" metricMagnitude="-2" metricUnit="m" metricValue="3.87" pageId="12" pageNumber="43" unit="mm" value="38.7">38.7 mm</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC72024FCC2FB94FBA9C6C7" pageId="12" pageNumber="43" type="etymology">
<paragraph id="8BC20859FFC72024FCC2FB94FBA9C6C7" blockId="12.[813,1456,1113,1217]" pageId="12" pageNumber="43">ETYMOLOGY. — Named in honour of Nicolas Puillandre, associate professor of MNHN, our long term co-author and participant of many expeditions with whom we had a pleasure to share the lab bench for long hours.</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC72024FCC2FB17FAD7C770" pageId="12" pageNumber="43" type="diagnosis">
<paragraph id="8BC20859FFC72024FCC2FB17FAD7C770" blockId="12.[813,1456,1242,1399]" pageId="12" pageNumber="43">
DIAGNOSIS. — Shell medium sized for genus, up to
<quantity id="4C85A5BCFFC72024FAE0FB17FAFFC6F4" box="[1295,1366,1242,1266]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="43" unit="mm" value="42.0">42 mm</quantity>
, fusiform with high spire and attenuated, medium long, and slightly inclined to left siphonal canal. Protoconch large, paucispiral, of about 2 whorls. Spiral sculpture of distinct flat on top spiral cords, about 55 on last whorl. Radula with tricuspate central teeth with rectangular basal part and tricuspate lateral teeth with longest outermost cusp.
</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC72024FCC2FA42FA19C7C4" pageId="12" pageNumber="43" type="distribution">
<paragraph id="8BC20859FFC72024FCC2FA42FA19C7C4" blockId="12.[813,1456,1423,1474]" pageId="12" pageNumber="43">
DISTRIBUTION. — The species is found so far in the South
<collectingCountry id="F36A48C9FFC72024FAA0FA5DFA22C7AE" box="[1359,1419,1424,1448]" name="China" pageId="12" pageNumber="43">China</collectingCountry>
Sea at
<quantity id="4C85A5BCFFC72024FCACFA67FC12C7C4" box="[835,955,1450,1474]" metricMagnitude="3" metricUnit="m" metricValue="1.194" metricValueMax="1.389" metricValueMin="0.999" pageId="12" pageNumber="43" unit="m" value="1194.0" valueMax="1389.0" valueMin="999.0">999-1389 m</quantity>
and
<collectingCountry id="F36A48C9FFC72024FC07FA67FBFDC7C4" box="[1000,1108,1450,1474]" name="Philippines" pageId="12" pageNumber="43">Philippines</collectingCountry>
(in Philippine Sea) at
<quantity id="4C85A5BCFFC72024FACAFA67FA05C7C4" box="[1317,1452,1450,1474]" metricMagnitude="3" metricUnit="m" metricValue="1.2285" metricValueMax="1.302" metricValueMin="1.155" pageId="12" pageNumber="43" unit="m" value="1228.5" valueMax="1302.0" valueMin="1155.0">1155-1302 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFC72039FCC2FA39FBF5C054" lastPageId="17" lastPageNumber="48" pageId="12" pageNumber="43" type="description">
<paragraph id="8BC20859FFC72024FCC2FA39FBE9C408" blockId="12.[813,1456,1524,2030]" box="[813,1088,1524,1550]" pageId="12" pageNumber="43">
DESCRIPTION (
<typeStatus id="54C6B6FBFFC72024FC27FA34FB9EC40B" box="[968,1079,1529,1549]" pageId="12" pageNumber="43" type="holotype">HOLOTYPE</typeStatus>
)
</paragraph>
<paragraph id="8BC20859FFC72024FCC2F9D9FCF6C428" blockId="12.[813,1456,1524,2030]" box="[813,863,1556,1582]" pageId="12" pageNumber="43">
<emphasis id="B909D44BFFC72024FCC2F9D9FCF6C428" box="[813,863,1556,1582]" italics="true" pageId="12" pageNumber="43">Shell</emphasis>
</paragraph>
<paragraph id="8BC20859FFC72039FCC2F9F9FE4DC3F4" blockId="12.[813,1456,1524,2030]" lastBlockId="17.[132,775,217,786]" lastPageId="17" lastPageNumber="48" pageId="12" pageNumber="43">
Shell fusiform with high spire and attenuated siphonal canal (
<figureCitation id="134614DCFFC72024FCD9F999FCDEC468" box="[822,887,1620,1646]" captionStart="FIG" captionStartId="15.[132,143,1895,1913]" captionTargetBox="[131,1454,215,1827]" captionTargetId="figure@15.[131,1454,214,1827]" captionTargetPageId="15" captionText="FIG. 7. — Shells of Phaenomenella nicoi n. sp.: A-C, holotype, MNHN-IM-2013-61585, SL 41.1 mm; D, ZHONGSHA 2015, st. CP4157, sequenced spm, MNHN- IM-2013-59398, SL 37.5 mm;E, ZHONGSHA 2015,st. CP4133,sequenced spm,MNHN-IM-2013-61639, SL 43.2 mm;F, AURORA 2007,st. CP2685,sequenced spm, MNHN-IM-2007-34639, SL 34.7 mm; G, AURORA 2007, st. CP2685, not-sequenced spm, MNHN-IM-2007-34644, SL 38.7 mm; H, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61637, SL 30.6 mm (anatomy studied); I, ZHONGSHA 2015, st. CP4134, sequenced spm, MNHN-IM-2013-61673, SL 42 mm; J, ZHONGSHA 2015,st. CP4133,not-sequenced spm, MNHN-IM-2013-61636, SL 40.2 mm (anatomy studied); K, L, ZHONGSHA 2015, st. CP4132, not-sequenced spm, MNHN-IM-2013-61592, 39.1 mm; L, enlarged protoconch. Scale bars: A-K, 20 cm; L, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3652823" httpUri="https://zenodo.org/record/3652823/files/figure.png" pageId="12" pageNumber="43">Fig. 7</figureCitation>
A-C), fragile, white under periostracum. Siphonal canal medium long, slightly inclined to left and crossing coiling axis. Protoconch paucispiral, of about 2 whorls, partially eroded as well as upper teleoconch whorls, teleoconch whorls 6 ½ in number. [Better preserved protoconch found in specimen MNHN-IM-2013-61592 (
<figureCitation id="134614DCFFC72024FB38F939FA8BC508" box="[1239,1314,1780,1806]" captionStart="FIG" captionStartId="15.[132,143,1895,1913]" captionTargetBox="[131,1454,215,1827]" captionTargetId="figure@15.[131,1454,214,1827]" captionTargetPageId="15" captionText="FIG. 7. — Shells of Phaenomenella nicoi n. sp.: A-C, holotype, MNHN-IM-2013-61585, SL 41.1 mm; D, ZHONGSHA 2015, st. CP4157, sequenced spm, MNHN- IM-2013-59398, SL 37.5 mm;E, ZHONGSHA 2015,st. CP4133,sequenced spm,MNHN-IM-2013-61639, SL 43.2 mm;F, AURORA 2007,st. CP2685,sequenced spm, MNHN-IM-2007-34639, SL 34.7 mm; G, AURORA 2007, st. CP2685, not-sequenced spm, MNHN-IM-2007-34644, SL 38.7 mm; H, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61637, SL 30.6 mm (anatomy studied); I, ZHONGSHA 2015, st. CP4134, sequenced spm, MNHN-IM-2013-61673, SL 42 mm; J, ZHONGSHA 2015,st. CP4133,not-sequenced spm, MNHN-IM-2013-61636, SL 40.2 mm (anatomy studied); K, L, ZHONGSHA 2015, st. CP4132, not-sequenced spm, MNHN-IM-2013-61592, 39.1 mm; L, enlarged protoconch. Scale bars: A-K, 20 cm; L, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3652823" httpUri="https://zenodo.org/record/3652823/files/figure.png" pageId="12" pageNumber="43">Fig. 7L</figureCitation>
): bulbous, of nearly 2 whorls, diameter
<quantity id="4C85A5BCFFC72024FBAFF8DEFB0CC528" box="[1088,1189,1811,1838]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="12" pageNumber="43" unit="mm" value="2.05">2.05 mm</quantity>
, exposed height
<quantity id="4C85A5BCFFC72024FABBF8D9FA05C528" box="[1364,1452,1812,1838]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="43" unit="mm" value="2.1">2.1 mm</quantity>
. Protoconch surface eroded, sculpture not seen. Boundary with teleoconch marked by weak (about 5) orthocline ribs.] Teleoconch whorls convex, weakly angulated at shoulder. Suture distinct, shallowly impressed. Spiral sculpture of distinct flat on top spiral cords, covering entire shell surface. Number of cords increasing from 8 on first teleoconch whorl to 19 on penultimate whorl, on last whorl 55 cords, of which about 20 on canal. Cords differing slightly in width, with most narrow on subsutural ramp, interspaces between cords about 1/3-½ of cords width. Strong axial ribs present on entire shell, suture to suture on uppermost teleoconch whorls, gradually becoming obsolete on subsutural ramp and absent on ramp of last and penultimate whorls. On last whorl ribs distinct only on shoulder, producing its angulated appearance. Number of ribs stable, i.e., 15-16 per whorl.
</paragraph>
<caption id="DF0258D1FFC62025FF6BF861FBADC5ED" ID-DOI="http://doi.org/10.5281/zenodo.3652815" ID-Zenodo-Dep="3652815" httpUri="https://zenodo.org/record/3652815/files/figure.png" pageId="13" pageNumber="44" startId="13.[132,143,1964,1982]" targetBox="[132,1455,215,1835]" targetPageId="13">
<paragraph id="8BC20859FFC62025FF6BF861FBADC5ED" blockId="13.[132,1456,1964,2028]" pageId="13" pageNumber="44">
FIG. 5. — Radulae of
<taxonomicName id="4C7D73DAFFC62025FEDFF861FE19C5B8" authorityName="Fraussen &amp; Hadorn" authorityYear="2006" box="[304,432,1964,1982]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="44" phylum="Mollusca" rank="genus">
<emphasis id="B909D44BFFC62025FEDFF861FE19C5B8" box="[304,432,1964,1982]" italics="true" pageId="13" pageNumber="44">Phaenomenella</emphasis>
</taxonomicName>
spp.
<emphasis id="B909D44BFFC62025FE33F861FE40C5B8" bold="true" box="[476,489,1964,1982]" pageId="13" pageNumber="44">A</emphasis>
,
<emphasis id="B909D44BFFC62025FE1CF861FDA9C5B8" bold="true" box="[499,512,1964,1982]" pageId="13" pageNumber="44">B</emphasis>
,
<taxonomicName id="4C7D73DAFFC62025FDE6F861FD75C5B8" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[521,732,1964,1982]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="44" phylum="Mollusca" rank="species" species="samadiae" status="sp. nov.">
<emphasis id="B909D44BFFC62025FDE6F861FD75C5B8" box="[521,732,1964,1982]" italics="true" pageId="13" pageNumber="44">Phaenomenella samadiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A6930FFC62025FD0EF861FCBBC5B8" box="[737,786,1964,1982]" pageId="13" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
; holotype,MNHN-IM-2013-61617 (shell on Fig. 4A-C) (
<emphasis id="B909D44BFFC62025FB3CF861FB49C5B8" bold="true" box="[1235,1248,1964,1982]" pageId="13" pageNumber="44">A</emphasis>
); MNHN-IM-2013-61674 (shell on Fig. 4 E, F) (
<emphasis id="B909D44BFFC62025FEDCF80EFEE9C5D3" bold="true" box="[307,320,1987,2005]" pageId="13" pageNumber="44">B</emphasis>
);
<emphasis id="B909D44BFFC62025FEA0F80EFEC6C5D3" bold="true" box="[335,367,1987,2005]" pageId="13" pageNumber="44">C -F</emphasis>
.
<taxonomicName id="4C7D73DAFFC62025FE95F80EFD8FC5D3" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[378,550,1987,2005]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="44" phylum="Mollusca" rank="species" species="nicoi" status="sp. nov.">
<emphasis id="B909D44BFFC62025FE95F80EFD8FC5D3" box="[378,550,1987,2005]" italics="true" pageId="13" pageNumber="44">Phaenomenella nicoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A6930FFC62025FDC3F80EFDF7C5D3" box="[556,606,1987,2005]" pageId="13" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B909D44BFFC62025FD8AF80EFDDAC5D3" bold="true" box="[613,627,1987,2005]" pageId="13" pageNumber="44">C</emphasis>
, holotype, MNHN-IM-2013-61585 (shell on Fig. 7A-C);
<emphasis id="B909D44BFFC62025FBAFF80EFBE7C5D3" bold="true" box="[1088,1102,1987,2005]" pageId="13" pageNumber="44">D</emphasis>
, MNHN-IM-2013-59398 (shell on Fig. 7D);
<emphasis id="B909D44BFFC62025FF6BF817FF39C5EA" bold="true" box="[132,144,2010,2028]" pageId="13" pageNumber="44">E</emphasis>
, MNHN-IM-2013-61639 (shell on Fig. 7E);
<emphasis id="B909D44BFFC62025FE1BF817FE56C5EA" bold="true" box="[500,511,2010,2028]" pageId="13" pageNumber="44">F</emphasis>
, MNHN-IM-2013-61673 (shell on Fig. 7I). Scale bars: 100 μm.
</paragraph>
</caption>
<caption id="DF0258D1FFC52026FF6BF861FD6EC5ED" ID-DOI="http://doi.org/10.5281/zenodo.3652819" ID-Zenodo-Dep="3652819" httpUri="https://zenodo.org/record/3652819/files/figure.png" pageId="14" pageNumber="45" startId="14.[132,143,1964,1982]" targetBox="[144,1435,220,1825]" targetPageId="14">
<paragraph id="8BC20859FFC52026FF6BF861FD6EC5ED" blockId="14.[132,1455,1964,2027]" pageId="14" pageNumber="45">
FIG. 6. — Anatomy of
<taxonomicName id="4C7D73DAFFC52026FED5F861FDA7C5B8" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[314,526,1964,1982]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="45" phylum="Mollusca" rank="species" species="samadiae" status="sp. nov.">
<emphasis id="B909D44BFFC52026FED5F861FDA7C5B8" box="[314,526,1964,1982]" italics="true" pageId="14" pageNumber="45">Phaenomenella samadiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A6930FFC52026FDFBF861FDEFC5B8" box="[532,582,1964,1982]" pageId="14" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B909D44BFFC52026FDA2F861FDF3C5B8" bold="true" box="[589,602,1964,1982]" pageId="14" pageNumber="45">A</emphasis>
,
<emphasis id="B909D44BFFC52026FD8BF861FDD8C5B8" bold="true" box="[612,625,1964,1982]" pageId="14" pageNumber="45">B</emphasis>
,
<emphasis id="B909D44BFFC52026FD93F861FD34C5B8" bold="true" box="[636,669,1964,1982]" pageId="14" pageNumber="45">E -G</emphasis>
, holotype, MNHN-IM-2013-61617, female;
<emphasis id="B909D44BFFC52026FBE6F861FBBEC5B8" bold="true" box="[1033,1047,1964,1982]" pageId="14" pageNumber="45">C</emphasis>
,
<emphasis id="B909D44BFFC52026FBCEF861FB86C5B8" bold="true" box="[1057,1071,1964,1982]" pageId="14" pageNumber="45">D</emphasis>
, MNHN-IM-2013-61674, male;
<emphasis id="B909D44BFFC52026FAD7F861FAECC5B8" bold="true" box="[1336,1349,1964,1982]" pageId="14" pageNumber="45">A</emphasis>
, cephalopodium, front view, mantle removed;
<emphasis id="B909D44BFFC52026FE4CF80EFE19C5D3" bold="true" box="[419,432,1987,2005]" pageId="14" pageNumber="45">B</emphasis>
, mantle;
<emphasis id="B909D44BFFC52026FE11F80EFDA5C5D3" bold="true" box="[510,524,1987,2005]" pageId="14" pageNumber="45">C</emphasis>
, uppermost part of penis;
<emphasis id="B909D44BFFC52026FD07F80EFD5FC5D3" bold="true" box="[744,758,1987,2005]" pageId="14" pageNumber="45">D</emphasis>
, stomach, external view;
<emphasis id="B909D44BFFC52026FC24F80EFC7EC5D3" bold="true" box="[971,983,1987,2005]" pageId="14" pageNumber="45">E</emphasis>
, foregut, ventral view;
<emphasis id="B909D44BFFC52026FB7BF80EFB36C5D3" bold="true" box="[1172,1183,1987,2005]" pageId="14" pageNumber="45">F</emphasis>
, foregut, lateral view;
<emphasis id="B909D44BFFC52026FAB9F80EFACDC5D3" bold="true" box="[1366,1380,1987,2005]" pageId="14" pageNumber="45">G</emphasis>
, valve of Leiblein. Abbreviations: see material and methods. Scale bars: 5 mm.
</paragraph>
</caption>
<caption id="DF0258D1FFC42027FF6BF8AAFB90C5ED" ID-DOI="http://doi.org/10.5281/zenodo.3652823" ID-Zenodo-Dep="3652823" httpUri="https://zenodo.org/record/3652823/files/figure.png" pageId="15" pageNumber="46" startId="15.[132,143,1895,1913]" targetBox="[131,1454,215,1827]" targetPageId="15">
<paragraph id="8BC20859FFC42027FF6BF8AAFB90C5ED" blockId="15.[132,1457,1895,2027]" pageId="15" pageNumber="46">
FIG. 7. — Shells of
<taxonomicName id="4C7D73DAFFC42027FECFF8AAFE62C57F" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[288,459,1895,1913]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="46" phylum="Mollusca" rank="species" species="nicoi" status="sp. nov.">
<emphasis id="B909D44BFFC42027FECFF8AAFE62C57F" box="[288,459,1895,1913]" italics="true" pageId="15" pageNumber="46">Phaenomenella nicoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A6930FFC42027FE3FF8AAFDABC57F" box="[464,514,1895,1913]" pageId="15" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B909D44BFFC42027FDE6F8A5FD82C57C" bold="true" box="[521,555,1895,1914]" pageId="15" pageNumber="46">A -C</emphasis>
, holotype, MNHN-IM-2013-61585, SL 41.1 mm;
<emphasis id="B909D44BFFC42027FC2FF8A5FC67C57C" bold="true" box="[960,974,1896,1914]" pageId="15" pageNumber="46">D</emphasis>
, ZHONGSHA 2015, st. CP4157, sequenced spm, MNHN- IM-2013-59398, SL 37.5 mm;
<emphasis id="B909D44BFFC42027FE95F8B3FE2FC596" bold="true" box="[378,390,1918,1936]" pageId="15" pageNumber="46">E</emphasis>
, ZHONGSHA 2015,st. CP4133,sequenced spm,MNHN-IM-2013-61639, SL 43.2 mm;
<emphasis id="B909D44BFFC42027FBB9F8B3FBC8C596" bold="true" box="[1110,1121,1918,1936]" pageId="15" pageNumber="46">F</emphasis>
, AURORA 2007,st. CP2685,sequenced spm, MNHN-IM-2007-34639, SL 34.7 mm;
<emphasis id="B909D44BFFC42027FE01F858FE55C5A1" bold="true" box="[494,508,1941,1959]" pageId="15" pageNumber="46">G</emphasis>
, AURORA 2007, st. CP2685, not-sequenced spm, MNHN-IM-2007-34644, SL 38.7 mm;
<emphasis id="B909D44BFFC42027FB0EF858FB46C5A1" bold="true" box="[1249,1263,1941,1959]" pageId="15" pageNumber="46">H</emphasis>
, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61637, SL 30.6 mm (anatomy studied);
<emphasis id="B909D44BFFC42027FCD3F861FCEBC5B8" bold="true" box="[828,834,1964,1982]" pageId="15" pageNumber="46">I</emphasis>
, ZHONGSHA 2015, st. CP4134, sequenced spm, MNHN-IM-2013-61673, SL 42 mm;
<emphasis id="B909D44BFFC42027FF0CF80FFF44C5D2" bold="true" box="[227,237,1986,2004]" pageId="15" pageNumber="46">J</emphasis>
, ZHONGSHA 2015,st. CP4133,not-sequenced spm, MNHN-IM-2013-61636, SL 40.2 mm (anatomy studied);
<emphasis id="B909D44BFFC42027FB6EF80FFB27C5D2" bold="true" box="[1153,1166,1986,2004]" pageId="15" pageNumber="46">K</emphasis>
,
<emphasis id="B909D44BFFC42027FB77F80FFB0AC5D2" bold="true" box="[1176,1187,1986,2004]" pageId="15" pageNumber="46">L</emphasis>
, ZHONGSHA 2015, st. CP4132, not-sequenced spm, MNHN-IM-2013-61592, 39.1 mm;
<emphasis id="B909D44BFFC42027FDB9F814FDC8C5ED" bold="true" box="[598,609,2009,2027]" pageId="15" pageNumber="46">L</emphasis>
, enlarged protoconch. Scale bars: A-K, 20 cm; L, 1 mm.
</paragraph>
</caption>
<caption id="DF0258D1FFDB2038FF6BF861FBC9C5EA" ID-DOI="http://doi.org/10.5281/zenodo.3652827" ID-Zenodo-Dep="3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="16" pageNumber="47" startId="16.[132,143,1964,1982]" targetBox="[140,1438,230,1880]" targetPageId="16">
<paragraph id="8BC20859FFDB2038FF6BF861FBC9C5EA" blockId="16.[132,1457,1964,2028]" pageId="16" pageNumber="47">
FIG. 8. — Anatomy of
<taxonomicName id="4C7D73DAFFDB2038FED7F861FE4AC5B8" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[312,483,1964,1982]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="47" phylum="Mollusca" rank="species" species="nicoi" status="sp. nov.">
<emphasis id="B909D44BFFDB2038FED7F861FE4AC5B8" box="[312,483,1964,1982]" italics="true" pageId="16" pageNumber="47">Phaenomenella nicoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A6930FFDB2038FE06F861FDB3C5B8" box="[489,538,1964,1982]" pageId="16" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B909D44BFFDB2038FDCEF860FDEAC5B9" bold="true" box="[545,579,1964,1983]" pageId="16" pageNumber="47">A -D</emphasis>
, MNHN-IM-2013-61636, male;
<emphasis id="B909D44BFFDB2038FCA5F860FCC2C5B8" bold="true" box="[842,875,1964,1983]" pageId="16" pageNumber="47">E -G</emphasis>
, MNHN-IM-2013-61637, female:
<emphasis id="B909D44BFFDB2038FB6DF860FB26C5B9" bold="true" box="[1154,1167,1965,1983]" pageId="16" pageNumber="47">A</emphasis>
, soft body removed from the shell;
<emphasis id="B909D44BFFDB2038FF6BF80EFF38C5D3" bold="true" box="[132,145,1987,2005]" pageId="16" pageNumber="47">B</emphasis>
, penis;
<emphasis id="B909D44BFFDB2038FF3DF80EFF49C5D3" bold="true" box="[210,224,1987,2005]" pageId="16" pageNumber="47">C</emphasis>
, proboscis, opened dorsally;
<emphasis id="B909D44BFFDB2038FE3CF80EFE48C5D3" bold="true" box="[467,481,1987,2005]" pageId="16" pageNumber="47">D</emphasis>
, foregut,right lateral view;
<emphasis id="B909D44BFFDB2038FD50F80EFD62C5D3" bold="true" box="[703,715,1987,2005]" pageId="16" pageNumber="47">E</emphasis>
, foregut,right lateral view, right salivary gland displaced;
<emphasis id="B909D44BFFDB2038FB4DF80EFB04C5D3" bold="true" box="[1186,1197,1987,2005]" pageId="16" pageNumber="47">F</emphasis>
, foregut,right lateral view, right salivary gland in original position;
<emphasis id="B909D44BFFDB2038FE71F817FE05C5EA" bold="true" box="[414,428,2010,2028]" pageId="16" pageNumber="47">G</emphasis>
, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm.
</paragraph>
</caption>
<paragraph id="8BC20859FFDA2039FF74FE35FD53C054" blockId="17.[132,775,217,786]" pageId="17" pageNumber="48">Aperture ovate, white inside, angulated posteriorly, outer lip thin, evenly convex, concave at transition to canal. Parietal wall and columella with narrow and thin glossy callus.</paragraph>
<paragraph id="8BC20859FFDA2039FF74FD95FE8CC0B4" blockId="17.[132,775,217,786]" pageId="17" pageNumber="48">Shell covered with very light olive adhering periostracum, forming densely spaced low axial lamellae visible in interspaces between cords.</paragraph>
<paragraph id="8BC20859FFDA2039FF74FD7AFE41C114" blockId="17.[132,775,217,786]" pageId="17" pageNumber="48">
Operculum partially abraded, when intact (
<figureCitation id="134614DCFFDA2039FD88FD7AFD17C0D4" box="[615,702,695,722]" captionStart="FIG" captionStartId="15.[132,143,1895,1913]" captionTargetBox="[131,1454,215,1827]" captionTargetId="figure@15.[131,1454,214,1827]" captionTargetPageId="15" captionText="FIG. 7. — Shells of Phaenomenella nicoi n. sp.: A-C, holotype, MNHN-IM-2013-61585, SL 41.1 mm; D, ZHONGSHA 2015, st. CP4157, sequenced spm, MNHN- IM-2013-59398, SL 37.5 mm;E, ZHONGSHA 2015,st. CP4133,sequenced spm,MNHN-IM-2013-61639, SL 43.2 mm;F, AURORA 2007,st. CP2685,sequenced spm, MNHN-IM-2007-34639, SL 34.7 mm; G, AURORA 2007, st. CP2685, not-sequenced spm, MNHN-IM-2007-34644, SL 38.7 mm; H, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61637, SL 30.6 mm (anatomy studied); I, ZHONGSHA 2015, st. CP4134, sequenced spm, MNHN-IM-2013-61673, SL 42 mm; J, ZHONGSHA 2015,st. CP4133,not-sequenced spm, MNHN-IM-2013-61636, SL 40.2 mm (anatomy studied); K, L, ZHONGSHA 2015, st. CP4132, not-sequenced spm, MNHN-IM-2013-61592, 39.1 mm; L, enlarged protoconch. Scale bars: A-K, 20 cm; L, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3652823" httpUri="https://zenodo.org/record/3652823/files/figure.png" pageId="17" pageNumber="48">Fig. 7 D</figureCitation>
) spanning most of aperture with distinctly turned leftwards terminal nucleus and rounded upper part.
</paragraph>
<paragraph id="8BC20859FFDA2039FF6BFCFAFEFEC157" blockId="17.[132,775,823,1264]" box="[132,343,823,849]" pageId="17" pageNumber="48">
<heading id="D08ABF35FFDA2039FF6BFCFAFEFEC157" box="[132,343,823,849]" fontSize="11" level="5" pageId="17" pageNumber="48" reason="1">
<emphasis id="B909D44BFFDA2039FF6BFCFAFEFEC157" box="[132,343,823,849]" italics="true" pageId="17" pageNumber="48">
Radula (
<figureCitation id="134614DCFFDA2039FF30FCF5FEB5C157" box="[223,284,823,849]" captionStart="FIG" captionStartId="13.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1835]" captionTargetId="figure@13.[132,1455,215,1836]" captionTargetPageId="13" captionText="FIG. 5. — Radulae of Phaenomenella spp.A, B, Phaenomenella samadiae n. sp.; holotype,MNHN-IM-2013-61617 (shell on Fig. 4A-C) (A); MNHN-IM-2013-61674 (shell on Fig. 4 E, F) (B); C-F. Phaenomenella nicoi n. sp.; C, holotype, MNHN-IM-2013-61585 (shell on Fig. 7A-C); D, MNHN-IM-2013-59398 (shell on Fig. 7D); E, MNHN-IM-2013-61639 (shell on Fig. 7E); F, MNHN-IM-2013-61673 (shell on Fig. 7I). Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3652815" httpUri="https://zenodo.org/record/3652815/files/figure.png" pageId="17" pageNumber="48">Fig. 5</figureCitation>
C-F)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC20859FFDA2039FF6BFC9AFE2EC6F6" blockId="17.[132,775,823,1264]" pageId="17" pageNumber="48">
Examined in five specimens, including
<typeStatus id="54C6B6FBFFDA2039FDA8FC9AFD07C177" box="[583,686,855,881]" pageId="17" pageNumber="48" type="holotype">holotype</typeStatus>
. Rather similar in all specimens; central tooth with rectangular basal part with weakly arcuate anterior margin and three medium long triangular broad cusps, central one shorter and narrower than lateral ones. Lateral teeth normally tricuspate with weakly curved basal side, attached to membrane. Outermost cusp recurved, medium long, inner cusp weakly recurved, about 2/3 of outer cusp length. Intermediate cusp shortest, situated slightly closer to inner cusp. In one sequenced specimen (MNHN-IM-2013-61673,
<figureCitation id="134614DCFFDA2039FE57FBBBFDA1C697" box="[440,520,1142,1169]" captionStart="FIG" captionStartId="13.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1835]" captionTargetId="figure@13.[132,1455,215,1836]" captionTargetPageId="13" captionText="FIG. 5. — Radulae of Phaenomenella spp.A, B, Phaenomenella samadiae n. sp.; holotype,MNHN-IM-2013-61617 (shell on Fig. 4A-C) (A); MNHN-IM-2013-61674 (shell on Fig. 4 E, F) (B); C-F. Phaenomenella nicoi n. sp.; C, holotype, MNHN-IM-2013-61585 (shell on Fig. 7A-C); D, MNHN-IM-2013-59398 (shell on Fig. 7D); E, MNHN-IM-2013-61639 (shell on Fig. 7E); F, MNHN-IM-2013-61673 (shell on Fig. 7I). Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3652815" httpUri="https://zenodo.org/record/3652815/files/figure.png" pageId="17" pageNumber="48">Fig. 5F</figureCitation>
) intermediate cusps of the left lateral teeth of the
<taxonomicName id="4C7D73DAFFDA2039FE72FB5BFE49C6B6" box="[413,480,1174,1200]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="17" pageNumber="52" phylum="Bryophyta" rank="genus">radula</taxonomicName>
paired, nearly equal in size, while lateral teeth on right side have broader intermediate cusp subdivided on top.
</paragraph>
<paragraph id="8BC20859FFDA2039FF6BFADBFE2BC729" blockId="17.[132,775,1301,1424]" box="[132,386,1301,1327]" pageId="17" pageNumber="48">
<heading id="D08ABF35FFDA2039FF6BFADBFE2BC729" box="[132,386,1301,1327]" fontSize="11" level="5" pageId="17" pageNumber="48" reason="1">
<emphasis id="B909D44BFFDA2039FF6BFADBFE2BC729" box="[132,386,1301,1327]" italics="true" pageId="17" pageNumber="48">
Measurements (
<typeStatus id="54C6B6FBFFDA2039FECBFAD8FED2C729" box="[292,379,1301,1327]" pageId="17" pageNumber="48" type="holotype">holotype</typeStatus>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC20859FFDA2039FF6BFAF8FD7DC796" blockId="17.[132,775,1301,1424]" pageId="17" pageNumber="48">
Shell length
<quantity id="4C85A5BCFFDA2039FEE3FAF8FEDAC756" box="[268,371,1333,1360]" metricMagnitude="-2" metricUnit="m" metricValue="4.13" pageId="17" pageNumber="48" unit="mm" value="41.3">41.3 mm</quantity>
, last whorl length
<quantity id="4C85A5BCFFDA2039FDD4FAF8FD0BC756" box="[571,674,1333,1360]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="17" pageNumber="48" unit="mm" value="26.9">26.9 mm</quantity>
, aperture length (without canal) 16.0 mm, diameter
<quantity id="4C85A5BCFFDA2039FDBBFA98FD15C776" box="[596,700,1365,1392]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="17" pageNumber="48" unit="mm" value="16.5">16.5 mm</quantity>
. In the largest specimen studied, shell length reaching
<quantity id="4C85A5BCFFDA2039FD92FAB8FD79C796" box="[637,720,1397,1424]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="17" pageNumber="48" unit="mm" value="43.0">43 mm</quantity>
.
</paragraph>
<paragraph id="8BC20859FFDA2039FF6BFA78FF48C7C9" blockId="17.[132,775,1461,1743]" box="[132,225,1461,1487]" pageId="17" pageNumber="48">
<heading id="D08ABF35FFDA2039FF6BFA78FF48C7C9" box="[132,225,1461,1487]" fontSize="11" level="5" pageId="17" pageNumber="48" reason="1">
<emphasis id="B909D44BFFDA2039FF6BFA78FF48C7C9" box="[132,225,1461,1487]" italics="true" pageId="17" pageNumber="48">Anatomy</emphasis>
</heading>
</paragraph>
<paragraph id="8BC20859FFDA2039FF6BFA18FE66C4C9" blockId="17.[132,775,1461,1743]" pageId="17" pageNumber="48">
Two specimens examined: MNHN-IM-2013-61636, male, and MNHN-IM-2013-61637, female, sequenced
<typeStatus id="54C6B6FBFFDA2039FD44FA38FCAEC409" box="[683,775,1525,1551]" pageId="17" pageNumber="48" type="paratype">paratype</typeStatus>
(similar in both studied specimens). Soft body partly extracted from the shell. Head medium large, with two long tentacles and large black eyes on lobes at bases of tentacles. Mantle similar to that of
<taxonomicName id="4C7D73DAFFDA2039FEA7F9B9FDE4C488" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[328,589,1652,1678]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="48" phylum="Mollusca" rank="species" species="samadiae" status="sp. nov.">
<emphasis id="B909D44BFFDA2039FEA7F9B9FDE4C488" box="[328,589,1652,1678]" italics="true" pageId="17" pageNumber="48">Phaenomenella samadiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A6930FFDA2039FDBAF9B8FD3BC489" box="[597,658,1653,1679]" pageId="17" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
Penis flattened, with seminal papilla situated on its top and surrounded by circle fold of skin (
<figureCitation id="134614DCFFDA2039FE9FF979FE17C4C8" box="[368,446,1716,1742]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8B</figureCitation>
).
</paragraph>
<paragraph id="8BC20859FFDA2039FF6BF93EFDA8C5E8" blockId="17.[132,775,1779,2030]" pageId="17" pageNumber="48">
<emphasis id="B909D44BFFDA2039FF6BF93EFEEAC50B" bold="true" box="[132,323,1779,1805]" pageId="17" pageNumber="48">Digestive system.</emphasis>
Proboscis almost completely inverted into rhynchodaeum (
<figureCitation id="134614DCFFDA2039FEAFF8D9FE31C528" box="[320,408,1812,1838]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8D</figureCitation>
, pr). Several bands of proboscis retractors muscles attached to middle part on both sides of rhynchodaeum (
<figureCitation id="134614DCFFDA2039FED9F89EFEDAC568" box="[310,371,1875,1902]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8</figureCitation>
D-F, prr). Buccal mass slightly shorter than retracted proboscis (
<figureCitation id="134614DCFFDA2039FE78F8BEFE40C58B" box="[407,489,1907,1933]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8C</figureCitation>
, bm), attached to its walls by multiple odontophoral retractors (
<figureCitation id="134614DCFFDA2039FDF3F85EFDC7C5AB" box="[540,622,1939,1965]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8C</figureCitation>
, odr). Radula lying in middle of buccal mass and attached to proboscis walls by median retractor (
<figureCitation id="134614DCFFDA2039FE86F81EFE13C5EB" box="[361,442,2003,2029]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8C</figureCitation>
, mrr).
</paragraph>
<paragraph id="8BC20859FFDA2039FCACFF14FBF5C054" blockId="17.[813,1456,217,594]" pageId="17" pageNumber="48">
Anterior oesophagus straight, following along ventral side of proboscis (
<figureCitation id="134614DCFFDA2039FC52FF34FBA3C315" box="[957,1034,249,275]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8E</figureCitation>
, aoe). Valve of Leiblein (
<figureCitation id="134614DCFFDA2039FAFAFF34FA2CC312" box="[1301,1413,249,276]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8D, E</figureCitation>
, vl) medium large, coniform, situated immediately before nerve ring (
<figureCitation id="134614DCFFDA2039FC82FEF4FC6EC355" box="[877,967,313,339]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8D</figureCitation>
, nr). Salivary glands medium-large (about 0.3 proboscis length), fused ventrally beneath nerve ring (
<figureCitation id="134614DCFFDA2039FCD9FEB4FC36C395" box="[822,927,377,403]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8E, F</figureCitation>
, sg), with very thin strongly convoluted salivary ducts following along anterior oesophagus. Gland of Leiblein medium in size (
<figureCitation id="134614DCFFDA2039FC34FE75FBE0C3D5" box="[987,1097,440,467]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8D, E</figureCitation>
, gl), following along posterior oesophagus and anterior aorta. Stomach rather large, spanning about 0.4 whorl (
<figureCitation id="134614DCFFDA2039FC0AFE35FB9CC014" box="[997,1077,504,530]" captionStart="FIG" captionStartId="16.[132,143,1964,1982]" captionTargetBox="[140,1438,230,1880]" captionTargetId="figure@16.[132,1455,216,1953]" captionTargetPageId="16" captionText="FIG. 8. — Anatomy of Phaenomenella nicoi n. sp.: A-D, MNHN-IM-2013-61636, male; E-G, MNHN-IM-2013-61637, female: A, soft body removed from the shell; B, penis; C, proboscis, opened dorsally; D, foregut,right lateral view; E, foregut,right lateral view, right salivary gland displaced; F, foregut,right lateral view, right salivary gland in original position; G, stomach, outer view. Abbreviations: see material and methods. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652827" httpUri="https://zenodo.org/record/3652827/files/figure.png" pageId="17" pageNumber="48">Fig. 8G</figureCitation>
). Posterior mixing area (pma) twice larger than in
<taxonomicName id="4C7D73DAFFDA2039FC2EFDD5FB6BC034" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[961,1218,536,562]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="48" phylum="Mollusca" rank="species" species="samadiae">
<emphasis id="B909D44BFFDA2039FC2EFDD5FB6BC034" box="[961,1218,536,562]" italics="true" pageId="17" pageNumber="48">Phaenomenella samadiae</emphasis>
</taxonomicName>
, posterior oesophagus and intestine medium wide.
</paragraph>
</subSubSection>
<subSubSection id="C3675BD2FFDA2039FCC2FDBAFBF0C697" pageId="17" pageNumber="48" type="discussion">
<paragraph id="8BC20859FFDA2039FCC2FDBAFC27C089" blockId="17.[813,1456,631,1169]" box="[813,910,631,657]" pageId="17" pageNumber="48">REMARKS</paragraph>
<paragraph id="8BC20859FFDA2039FCC2FD55FBF0C697" blockId="17.[813,1456,631,1169]" pageId="17" pageNumber="48">
The new species is highly variable in shell shape. Some of the specimens are much more slender (
<figureCitation id="134614DCFFDA2039FB77FD7AFAB6C0D4" box="[1176,1311,695,722]" captionStart="FIG" captionStartId="15.[132,143,1895,1913]" captionTargetBox="[131,1454,215,1827]" captionTargetId="figure@15.[131,1454,214,1827]" captionTargetPageId="15" captionText="FIG. 7. — Shells of Phaenomenella nicoi n. sp.: A-C, holotype, MNHN-IM-2013-61585, SL 41.1 mm; D, ZHONGSHA 2015, st. CP4157, sequenced spm, MNHN- IM-2013-59398, SL 37.5 mm;E, ZHONGSHA 2015,st. CP4133,sequenced spm,MNHN-IM-2013-61639, SL 43.2 mm;F, AURORA 2007,st. CP2685,sequenced spm, MNHN-IM-2007-34639, SL 34.7 mm; G, AURORA 2007, st. CP2685, not-sequenced spm, MNHN-IM-2007-34644, SL 38.7 mm; H, ZHONGSHA 2015, st. CP4133, sequenced spm, MNHN-IM-2013-61637, SL 30.6 mm (anatomy studied); I, ZHONGSHA 2015, st. CP4134, sequenced spm, MNHN-IM-2013-61673, SL 42 mm; J, ZHONGSHA 2015,st. CP4133,not-sequenced spm, MNHN-IM-2013-61636, SL 40.2 mm (anatomy studied); K, L, ZHONGSHA 2015, st. CP4132, not-sequenced spm, MNHN-IM-2013-61592, 39.1 mm; L, enlarged protoconch. Scale bars: A-K, 20 cm; L, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3652823" httpUri="https://zenodo.org/record/3652823/files/figure.png" pageId="17" pageNumber="48">Fig. 7D, I, G</figureCitation>
) and the axial ribs are either very weak or obsolete. The specimen with no axial ribs was collected at a maximal depth (
<quantity id="4C85A5BCFFDA2039FAE7FD3AFA0BC114" box="[1288,1442,759,786]" metricMagnitude="3" metricUnit="m" metricValue="1.6585" metricValueMax="1.683" metricValueMin="1.634" pageId="17" pageNumber="48" unit="m" value="1658.5" valueMax="1683.0" valueMin="1634.0">1634-1683 m</quantity>
), but there is not clear correlation with depth, since syntopic specimens can have strong or weak axial ribs. Nevertheless the molecular data clearly indicates the conspecifity of “typical” angulated specimens with well-developed ribs and smooth ones. The specimens collected at greater depth have the protoconch and upper teleoconch whorls more eroded or missing. In general shape the new species has some resemblance to
<taxonomicName id="4C7D73DAFFDA2039FCC2FC3AFBCFC617" authority="Fraussen, 2008" authorityName="Fraussen" authorityYear="2008" box="[813,1126,1014,1041]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="48" phylum="Mollusca" rank="species" species="mokenorum">
<emphasis id="B909D44BFFDA2039FCC2FC3AFC17C616" box="[813,958,1014,1040]" italics="true" pageId="17" pageNumber="48">P. mokenorum</emphasis>
<bibRefCitation id="EFEC75A8FFDA2039FC2BFC3AFBCFC617" author="FRAUSSEN K." box="[964,1126,1015,1041]" journalOrPublisher="Veliger" pageId="17" pageNumber="48" pagination="48 - 50" part="50" refId="ref12618" refString="FRAUSSEN K. 2008 - A new Phaenomenella Fraussen &amp; Hadorn, 2006 (Gastropoda: Buccinidae), from the Andaman Sea. Veliger 50: 48 - 50." title="A new Phaenomenella Fraussen &amp; Hadorn, 2006 (Gastropoda: Buccinidae), from the Andaman Sea" type="journal article" year="2008">Fraussen, 2008</bibRefCitation>
</taxonomicName>
from the Andaman Sea, differing in better pronounced axial ribs in later teleoconch whorls and more attenuated narrow siphonal canal. Also
<taxonomicName id="4C7D73DAFFDA2039FA8FFBFAFA06C656" authority="Kantor &amp; Kosyan &amp; Sorokin, 2020" authorityName="Kantor &amp; Kosyan &amp; Sorokin" authorityYear="2020" box="[1376,1455,1079,1104]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="48" phylum="Mollusca" rank="species" species="nicoi" status="sp. nov.">
<emphasis id="B909D44BFFDA2039FA8FFBFAFA06C656" box="[1376,1455,1079,1104]" italics="true" pageId="17" pageNumber="48">P. nicoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A6930FFDA2039FCC2FB9AFCC0C677" box="[813,873,1111,1137]" pageId="17" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
has smaller size (maximal shell length
<quantity id="4C85A5BCFFDA2039FAFAFB9BFACFC677" box="[1301,1382,1110,1137]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="17" pageNumber="48" unit="mm" value="42.0">42 mm</quantity>
versus
<quantity id="4C85A5BCFFDA2039FCC2FBBBFC38C697" box="[813,913,1142,1169]" metricMagnitude="-2" metricUnit="m" metricValue="5.56" pageId="17" pageNumber="48" unit="mm" value="55.6">55.6 mm</quantity>
in
<taxonomicName id="4C7D73DAFFDA2039FC5AFBBAFBE1C696" authorityName="Fraussen" authorityYear="2008" box="[949,1096,1142,1168]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="48" phylum="Mollusca" rank="species" species="mokenorum">
<emphasis id="B909D44BFFDA2039FC5AFBBAFBE1C696" box="[949,1096,1142,1168]" italics="true" pageId="17" pageNumber="48">P. mokenorum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>