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<mods:title id="271C01B23F68448F941AABD27011295D">Systematic revision of the fruit fly genus Carpophthoromyia Austen (Diptera, Tephritidae)</mods:title>
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<paragraph id="8BA6213EFFCBFFDA5629FBE26235FBEE" blockId="20.[264,594,1044,1105]" box="[264,594,1044,1070]" pageId="20" pageNumber="21">
<heading id="D0EE9652FFCBFFDA5629FBE26235FBEE" bold="true" box="[264,594,1044,1070]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<taxonomicName id="4C195ABDFFCBFFDA5629FBE26235FBEE" ID-CoL="RGHX" authority="Bezzi, 1918" authorityName="Bezzi" authorityYear="1918" box="[264,594,1044,1070]" class="Insecta" family="Tephritidae" genus="Carpophthoromyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pseudotritea">
<emphasis id="B96DFD2CFFCBFFDA5629FBE26235FBEE" bold="true" box="[264,594,1044,1070]" pageId="20" pageNumber="21">
<emphasis id="B96DFD2CFFCBFFDA5629FBE2617CFBEE" bold="true" box="[264,283,1044,1070]" italics="true" pageId="20" pageNumber="21">C</emphasis>
. 
<emphasis id="B96DFD2CFFCBFFDA560BFBE261D9FBEE" bold="true" box="[298,446,1044,1070]" italics="true" pageId="20" pageNumber="21">pseudotritea</emphasis>
<bibRefCitation id="EF885CCFFFCBFFDA56E7FBE26235FBEE" author="Bezzi" box="[454,594,1044,1070]" pageId="20" pageNumber="40" refString="Bezzi, M. (1918) Notes on the Ethiopian fruit-flies of the family Trypaneidae, other than Dacus (s. l.), with descriptions of new genera and species (Dipt.) - I. Bulletin of Entomological Research, 8, 215 - 251." type="journal article" year="1918">Bezzi, 1918</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA6213EFFCBFFDA5629FBC16183FB91" blockId="20.[264,594,1044,1105]" box="[264,484,1079,1105]" pageId="20" pageNumber="21">
(
<figureCitation id="13223DBBFFCBFFDA5630FBC1613FFB91" box="[273,344,1079,1105]" captionStart="FIGURES 7 – 9" captionStartId="43.[264,382,1752,1776]" captionTargetBox="[365,1145,325,1668]" captionTargetId="figure@43.[286,1264,298,1711]" captionTargetPageId="43" captionText="FIGURES 7 – 9. Wing. — 7. Carpophthoromyia dimidiata. — 8. C. vittata. — 9. C. pseudotritea." httpUri="https://zenodo.org/record/172783/files/figure.png" pageId="20" pageNumber="21">Fig. 9</figureCitation>
, 
<figureCitation id="13223DBBFFCBFFDA5644FBC161E2FB91" box="[357,389,1079,1105]" captionStart="FIGURES 13 – 22" captionStartId="45.[264,382,1790,1814]" captionTargetBox="[266,1276,321,1790]" captionTargetId="figure@45.[253,1307,305,1790]" captionTargetPageId="45" captionText="FIGURES 13 – 22. Aculeus full length. — 13. Carpophthoromyia flavofasciata. — 14. C. radulata. — 15. C. tritea. — 16. C. interrupta. — 17. C. schoutedeni. — 18. C. dividua. — 19. C. tessmanni. — 20. C. pseudotritea. — 21. C. virgata. — 22. C. vittata." httpUri="https://zenodo.org/record/172785/files/figure.png" pageId="20" pageNumber="21">20</figureCitation>
, 
<figureCitation id="13223DBBFFCBFFDA56B0FBC161D6FB91" box="[401,433,1079,1105]" captionStart="FIGURES 23 – 27" captionStartId="46.[264,382,740,764]" captionTargetBox="[264,1322,317,718]" captionTargetId="figure@46.[253,1354,305,733]" captionTargetPageId="46" captionText="FIGURES 23 – 27. Apical part of aculeus. — 23. Carpophthoromyia dividua. — 24. C. pseudotritea. — 25. C. tessmanni. — 26. C. vittata. — 27. C. vittata (lateral view)." httpUri="https://zenodo.org/record/172786/files/figure.png" pageId="20" pageNumber="21">24</figureCitation>
, 
<figureCitation id="13223DBBFFCBFFDA569CFBC161BCFB91" box="[445,475,1079,1105]" captionStart="FIGURES 34 – 35" captionStartId="47.[264,382,1269,1293]" captionText="FIGURES 34 – 35. Epandrium, posterior view. — 34. Carpophthoromyia dividua. — 35. C. pseudotritea." pageId="20" pageNumber="21">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30372B5FFCBFFDA5629FB71649BFB1D" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="8BA6213EFFCBFFDA5629FB716344FB5D" blockId="20.[264,1276,1158,1246]" box="[264,803,1158,1182]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="AB090610FFCBFFDA5629FB716344FB5D" box="[264,803,1158,1182]" pageId="20" pageNumber="21">
<taxonomicName id="4C195ABDFFCBFFDA5629FB716378FB5D" ID-CoL="RGHX" authority="Bezzi, 1918: 225" authorityName="Bezzi" authorityPageNumber="225" authorityYear="1918" box="[264,799,1158,1182]" class="Insecta" family="Tephritidae" genus="Carpophthoromyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pseudotritea">
<emphasis id="B96DFD2CFFCBFFDA5629FB716238FB5E" box="[264,607,1159,1182]" italics="true" pageId="20" pageNumber="21">Carpophthoromyia pseudotritea</emphasis>
<treatmentCitation id="0AB8072FFFCBFFDA5547FB706378FB5D" author="Bezzi" box="[614,799,1158,1181]" page="225" pageId="20" pageNumber="21" year="1918">
<bibRefCitation id="EF885CCFFFCBFFDA5547FB70628EFB5D" author="Bezzi" box="[614,745,1158,1181]" pageId="20" pageNumber="40" refString="Bezzi, M. (1918) Notes on the Ethiopian fruit-flies of the family Trypaneidae, other than Dacus (s. l.), with descriptions of new genera and species (Dipt.) - I. Bulletin of Entomological Research, 8, 215 - 251." type="journal article" year="1918">Bezzi, 1918</bibRefCitation>
: 225
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA6213EFFCBFFDA5629FB516306FB7D" blockId="20.[264,1276,1158,1246]" box="[264,865,1190,1214]" pageId="20" pageNumber="21">
<taxonomicName id="4C195ABDFFCBFFDA5629FB51633BFB7D" ID-CoL="RGHX" authority="Cogan &amp; Munro 1980" authorityName="Cogan &amp; Munro" authorityYear="1980" box="[264,860,1190,1214]" class="Insecta" family="Tephritidae" genus="Carpophthoromyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pseudotritea">
<emphasis id="B96DFD2CFFCBFFDA5629FB516238FB7E" box="[264,607,1191,1214]" italics="true" pageId="20" pageNumber="21">Carpophthoromyia pseudotritea</emphasis>
: Cogan &amp; Munro 1980
</taxonomicName>
.
</paragraph>
<paragraph id="8BA6213EFFCBFFDA5629FB31649BFB1D" blockId="20.[264,1276,1158,1246]" box="[264,1276,1222,1246]" pageId="20" pageNumber="21">
<taxonomicName id="4C195ABDFFCBFFDA5629FB316248FB1D" authority="Bezzi 1913" authorityName="Bezzi" authorityYear="1913" box="[264,559,1222,1246]" class="Insecta" family="Tephritidae" genus="Ceratitis" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tritea">
<emphasis id="B96DFD2CFFCBFFDA5629FB3161C1FB1E" box="[264,422,1223,1246]" italics="true" pageId="20" pageNumber="21">Ceratitis tritea</emphasis>
: 
<bibRefCitation id="EF885CCFFFCBFFDA5695FB306248FB1D" author="Bezzi" box="[436,559,1222,1245]" pageId="20" pageNumber="40" refString="Bezzi, M. (1913) Altre Ceratitis africane allevate dal Prof. F. Silvestri. Bollettino del Laboratorio di Zoologia Generale e Agraria della R. Scuola Superiore d'Agricoltura in Portici, 7, 19 - 26." type="journal article" year="1913">Bezzi 1913</bibRefCitation>
</taxonomicName>
, 
<bibRefCitation id="EF885CCFFFCBFFDA551BFB3062B9FB1D" author="Enderlein" box="[570,734,1222,1245]" pageId="20" pageNumber="40" refString="Enderlein, G. (1920) Zur Kenntnis tropischer Frucht-Bohrfliegen. Zoologische Jahrbucher Abteilung fur Systematik, Geografie und Biologie der Tiere, 43, 336 - 360." type="journal article" year="1920">Enderlein 1920</bibRefCitation>
(misident.), Cogan &amp; Munro 1980 (p.p. misident.)
</paragraph>
</subSubSection>
<subSubSection id="C30372B5FFCBFFD95629FAF963ECFE3E" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="8BA6213EFFCBFFDA5629FAF9611AFAE9" blockId="20.[264,1323,1295,1565]" box="[264,381,1295,1321]" pageId="20" pageNumber="21">Diagnosis</paragraph>
<paragraph id="8BA6213EFFCBFFDA5619FAC16342F9D9" blockId="20.[264,1323,1295,1565]" pageId="20" pageNumber="21">Arista distinctly plumose; frons with longitudinal brown band for entire length; distance between posterior frontal and anterior orbital distinctly shorter than distance between anterior and posterior orbital; three frontals; scutum without transverse band; postpronotum white; scutellum with three apical brown spots, only visible in ventral view; anterior margin of wing with one indentation near junction of vein C with apical part of vein R1; S­band and inverted V­band separate.</paragraph>
<paragraph id="8BA6213EFFCBFFDA5629F9A461F6F9AC" blockId="20.[264,1324,1618,1884]" box="[264,401,1618,1644]" pageId="20" pageNumber="21">Description</paragraph>
<paragraph id="8BA6213EFFCBFFDB5619F98C622EFE9E" blockId="20.[264,1324,1618,1884]" lastBlockId="21.[264,1325,284,1196]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">Head. Antennal segments dark yellow to brown. Arista distinctly plumose; longest rays longer than width of first flagellomere. Frons white to yellow, longitudinal yellowbrown band for entire length from ocellar triangle to antennal base, equal to width between anterior orbitals. Three frontals placed on oblique line, with anterior frontal at least 3 times as far from the inner eye margin than posterior frontal; two orbitals, posterior one shorter than anterior one but well developed. Distance between posterior frontal and anterior orbital distinctly shorter than distance between anterior and posterior orbital. Face white to yellow, gena darker brown.</paragraph>
<paragraph id="8BA6213EFFCAFFDB5619FE9A62ECFDB6" blockId="21.[264,1325,284,1196]" pageId="21" pageNumber="22">Thorax. Scutum shining black­brown, along transverse suture often yellow­brown, sometimes with faint yellow fascia; black setulae, without transverse bands of silvery setulae. Postpronotum white. Anepisternum with white to yellow band with lower margin reaching to lower fourth of posterior margin; with pale setulae, lower fourth black setulae sometimes more extensively so along posterior margin, two anepisternals. Katatergite and anatergite both white. Scutellum white, ventrally with 3 brown apical spots, not visible in dorsal view. Subscutellum black.</paragraph>
<paragraph id="8BA6213EFFCAFFDB5619FD726287FD06" blockId="21.[264,1325,284,1196]" pageId="21" pageNumber="22">Legs brown, tibia, tarsal segments and extreme apex of front femur yellow, at most basal margin of tibiae slightly darkened.</paragraph>
<paragraph id="8BA6213EFFCAFFDB5619FD226263FC54" blockId="21.[264,1325,284,1196]" pageId="21" pageNumber="22">
Wing (
<figureCitation id="13223DBBFFCAFFDB56A9FD2261B7FD2E" box="[392,464,724,750]" captionStart="FIGURES 7 – 9" captionStartId="43.[264,382,1752,1776]" captionTargetBox="[365,1145,325,1668]" captionTargetId="figure@43.[286,1264,298,1711]" captionTargetPageId="43" captionText="FIGURES 7 – 9. Wing. — 7. Carpophthoromyia dimidiata. — 8. C. vittata. — 9. C. pseudotritea." httpUri="https://zenodo.org/record/172783/files/figure.png" pageId="21" pageNumber="22">Fig. 9</figureCitation>
). Hyaline indentation near junction of vein C with apical part of vein R1, reaching vein R4+5, sometimes continuing slightly beyond vein. S­band and inverted Vband not fused; rarely V­band less strongly developed near subapical tooth, almost divided in two parts. S­band with small subapical tooth. Crossvein DM­Cu straight or slightly sinuous. R­M ratio 1.40.
</paragraph>
<paragraph id="8BA6213EFFCAFFDB5619FC546438FB44" blockId="21.[264,1325,284,1196]" pageId="21" pageNumber="22">
Abdomen. Shining black­brown, tergite 4 with median yellow spot and/or silvery microtrichosity posteriorly; with black setulae. Spermatheca ovoid in apical part, base slender. Female terminalia, oviscape shorter than abdomen; shining black­brown. Aculeus orange, flattened, about 5 times longer than wide (
<figureCitation id="13223DBBFFCAFFDB5448FBEC63A5FBF4" box="[873,962,1050,1076]" captionStart="FIGURES 13 – 22" captionStartId="45.[264,382,1790,1814]" captionTargetBox="[266,1276,321,1790]" captionTargetId="figure@45.[253,1307,305,1790]" captionTargetPageId="45" captionText="FIGURES 13 – 22. Aculeus full length. — 13. Carpophthoromyia flavofasciata. — 14. C. radulata. — 15. C. tritea. — 16. C. interrupta. — 17. C. schoutedeni. — 18. C. dividua. — 19. C. tessmanni. — 20. C. pseudotritea. — 21. C. virgata. — 22. C. vittata." httpUri="https://zenodo.org/record/172785/files/figure.png" pageId="21" pageNumber="22">Fig. 20</figureCitation>
); tip simply pointed (
<figureCitation id="13223DBBFFCAFFDB53E3FBEC657CFBF4" box="[1218,1307,1050,1076]" captionStart="FIGURES 23 – 27" captionStartId="46.[264,382,740,764]" captionTargetBox="[264,1322,317,718]" captionTargetId="figure@46.[253,1354,305,733]" captionTargetPageId="46" captionText="FIGURES 23 – 27. Apical part of aculeus. — 23. Carpophthoromyia dividua. — 24. C. pseudotritea. — 25. C. tessmanni. — 26. C. vittata. — 27. C. vittata (lateral view)." httpUri="https://zenodo.org/record/172786/files/figure.png" pageId="21" pageNumber="22">Fig. 24</figureCitation>
). Male terminalia (
<figureCitation id="13223DBBFFCAFFDB56F7FBB46249FB9C" box="[470,558,1090,1116]" captionStart="FIGURES 34 – 35" captionStartId="47.[264,382,1269,1293]" captionText="FIGURES 34 – 35. Epandrium, posterior view. — 34. Carpophthoromyia dividua. — 35. C. pseudotritea." pageId="21" pageNumber="22">Fig. 35</figureCitation>
), epandrium rounded in posterior view; posterior lobe of lateral surstylus elongated and almost straight; medial surstylus at apex pointed.
</paragraph>
<paragraph id="8BA6213EFFCAFFDB5619FB646436FB6C" blockId="21.[264,1325,284,1196]" box="[312,1105,1170,1196]" pageId="21" pageNumber="22">Body length: 4.87 (4.00­6.00)mm; wing length 4.93 (4.00­5.89)mm</paragraph>
<paragraph id="8BA6213EFFCAFFDB5629FB146182FB3C" blockId="21.[264,1323,1250,1876]" box="[264,485,1250,1276]" pageId="21" pageNumber="22">Material examined</paragraph>
<paragraph id="8BA6213EFFCAFFDB5619FAFC62A5FA5C" blockId="21.[264,1323,1250,1876]" pageId="21" pageNumber="22">
<typeStatus id="54A29F9CFFCAFFDB5619FAFC6115FAE4" box="[312,370,1290,1316]" pageId="21" pageNumber="22">Type</typeStatus>
material: 
<typeStatus id="54A29F9CFFCAFFDB56D5FAFC620EFAE4" box="[500,617,1290,1316]" pageId="21" pageNumber="22" type="lectotype">Lectotype</typeStatus>
Ψ (herewith designated): 
<collectingCountry id="F30E61AEFFCAFFDB5491FAFC647AFAE4" box="[944,1053,1290,1316]" name="Ghana" pageId="21" pageNumber="22">GHANA</collectingCountry>
, Aburi, 
<date id="FFA707FEFFCAFFDB53A7FAFC6567FAE4" box="[1158,1280,1290,1316]" pageId="21" pageNumber="22" value="1911-05-10">10.V.1911</date>
, L. Armstrong, “on fallen calabash” (BMNH). 
<typeStatus id="54A29F9CFFCAFFDB542CFAC463D6FA8C" box="[781,945,1330,1356]" pageId="21" pageNumber="22" type="paralectotype">Paralectotype</typeStatus>
: 1Ψ, same locality as 
<typeStatus id="54A29F9CFFCAFFDB5399FAC46540FA8C" box="[1208,1319,1330,1356]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
, 
<date id="FFA707FEFFCAFFDB5629FAAC6114FAB4" box="[264,371,1370,1396]" pageId="21" pageNumber="22" value="1911-05-08">8.V.1911</date>
, L. Armstrong, “on fallen calabash” (BMNH). 
<collectingCountry id="F30E61AEFFCAFFDB5323FAAC64E7FAB4" box="[1026,1152,1370,1396]" name="Nigeria" pageId="21" pageNumber="22">NIGERIA</collectingCountry>
: 1Ψ, Agege, 
<date id="FFA707FEFFCAFFDB5629FA7461EAFA5C" box="[264,397,1410,1436]" pageId="21" pageNumber="22" value="1914-04-17">17.IV.1914</date>
, W.A. Lamborn (BMNH).
</paragraph>
<paragraph id="8BA6213EFFCAFFD85619FA5C63E5FBBE" blockId="21.[264,1323,1250,1876]" lastBlockId="22.[264,1324,283,1150]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
Other material: 
<collectingCountry id="F30E61AEFFCAFFDB56D3FA5C6237FA04" box="[498,592,1450,1476]" name="Benin" pageId="21" pageNumber="22">BENIN</collectingCountry>
: 1Ψ, Attogon, 
<date id="FFA707FEFFCAFFDB55DAFA5C6315FA04" box="[763,882,1450,1476]" pageId="21" pageNumber="22" value="2000-03-03">3.III.2000</date>
, “sweep netting”, G. Goergen (IITA); 1Ψ, Lana, 
<date id="FFA707FEFFCAFFDB56A5FA24626BFA2C" box="[388,524,1490,1516]" pageId="21" pageNumber="22" value="1997-11-16">16.XI.1997</date>
, “sweep netting forest”, G. Goergen (IITA). 
<collectingCountry id="F30E61AEFFCAFFDB5336FA2464A3FA2C" box="[1047,1220,1490,1516]" name="Cameroon" pageId="21" pageNumber="22">CAMEROON</collectingCountry>
: Befang, Rt N11, 60km N Bamenda, 
<quantity id="4CE18CDBFFCAFFDB557FFA0C62C3F9D4" box="[606,676,1530,1556]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="22" unit="m" value="500.0">500m</quantity>
, 
<date id="FFA707FEFFCAFFDB5593FA0C635DF9D4" box="[690,826,1530,1556]" pageId="21" pageNumber="22" value="1987-11-24">24.XI.1987</date>
, 2ɗɗ, Ψ. Kaplan; 3ɗɗ 1Ψ, A. Freidberg (both 
<collectingCountry id="F30E61AEFFCAFFDB5673F9D461F6F9FC" box="[338,401,1570,1596]" name="China" pageId="21" pageNumber="22">TAU</collectingCountry>
); 5ɗɗ 5ΨΨ, nr Douala, 
<date id="FFA707FEFFCAFFDB55E0F9D463EEF9FC" box="[705,905,1570,1596]" pageId="21" pageNumber="22" value="1936-05-23" valueMax="1936-06-17" valueMin="1936-05-23">23.V­17.VI.1936</date>
, Van Zwaluwenburg &amp; McGough (USNM); 1Ψ, Lolodorf, L. Conradt (BMNH); 1Ψ, rtN9, 20km E Sangmelina, 
<date id="FFA707FEFFCAFFDB538CF9BC6541F9A4" box="[1197,1318,1610,1636]" pageId="21" pageNumber="22" value="1987-11-07">7.XI.1987</date>
, A. Freidberg (
<collectingCountry id="F30E61AEFFCAFFDB568FF9846188F94C" box="[430,495,1650,1676]" name="China" pageId="21" pageNumber="22">TAU</collectingCountry>
); 2ɗɗ, nr Yaoundé, 
<date id="FFA707FEFFCAFFDB55FEF98463FDF94C" box="[735,922,1650,1676]" pageId="21" pageNumber="22" value="1936-05-29" valueMax="1936-06-08" valueMin="1936-05-29">29.V­8.VI.1936</date>
, Van Zwaluwenburg &amp; McGough (USNM). 
<collectingCountry id="F30E61AEFFCAFFDB56A4F96C635DF974" box="[389,826,1690,1716]" name="Central African Republic" pageId="21" pageNumber="22">CENTRAL AFRICAN REPUBLIC</collectingCountry>
?: 1ɗ, “S.O. 
<collectingCountry id="F30E61AEFFCAFFDB54FAF96C6429F974" box="[987,1102,1690,1716]" name="Cameroon" pageId="21" pageNumber="22">Kamerun</collectingCountry>
”, Uam Gebiet, nr Bosum, 
<date id="FFA707FEFFCAFFDB564EF93461BCF91C" box="[367,475,1730,1756]" pageId="21" pageNumber="22" value="1914-05-02">2.V.1914</date>
, G. Tessmann (MNHU). 
<collectingCountry id="F30E61AEFFCAFFDB5437F9346319F91C" box="[790,894,1730,1756]" name="Republic of the Congo" pageId="21" pageNumber="22">CONGO</collectingCountry>
(D.R.): Eala, 1Ψ, 
<date id="FFA707FEFFCAFFDB534BF9346497F91C" box="[1130,1264,1730,1756]" pageId="21" pageNumber="22" value="1936-04-16">16.IV.1936</date>
; 1Ψ, 
<date id="FFA707FEFFCAFFDB5629F91C61EAF8C4" box="[264,397,1770,1796]" pageId="21" pageNumber="22" value="1936-04-24">24.IV.1936</date>
; 1ɗ, 
<date id="FFA707FEFFCAFFDB56F7F91C623CF8C4" box="[470,603,1770,1796]" pageId="21" pageNumber="22" value="1936-04-25">25.IV.1936</date>
; 1ɗ, 
<date id="FFA707FEFFCAFFDB5585F91C6343F8C4" box="[676,804,1770,1796]" pageId="21" pageNumber="22" value="1936-05">May 1936</date>
, all J. Ghesquière (KBIN); 1ɗ, 
<collectingCountry id="F30E61AEFFCAFFDB539AF91C6540F8C4" box="[1211,1319,1770,1796]" name="Ecuador" pageId="21" pageNumber="22">Equateur</collectingCountry>
, Bokuma, 
<date id="FFA707FEFFCAFFDB56A1F8E4619CF8EC" box="[384,507,1810,1836]" pageId="21" pageNumber="22" value="1952-07">July 1952</date>
, R.P. Lootens (KMMA); 1ɗ, Flandria, 
<date id="FFA707FEFFCAFFDB54CAF8E46414F8EC" box="[1003,1139,1810,1836]" pageId="21" pageNumber="22" value="1935-09-10">10.IX.1935</date>
, J. Ghesquière (KBIN); 1ɗ, Haut­Lopori, May­June.1927, J. Ghesquière (KMMA); 1ɗ, Kanda­Kanda, 1930, Hissette (KMMA); Kasai, Ngombe, 1ɗ, 
<date id="FFA707FEFFC9FFD85419FEEA63A6FEF6" box="[824,961,284,310]" pageId="22" pageNumber="23" value="1921-11-16">16.XI.1921</date>
, H. Schouteden (KMMA); 1ɗ 
<date id="FFA707FEFFC9FFD85629FEB261F7FE9E" box="[264,400,324,350]" pageId="22" pageNumber="23" value="1921-11-17">17.XI.1921</date>
, H. Schouteden (KBIN); 1Ψ, Kivu, Bassin Liadi, Lutunguru, Ratendele, “for. mont.”, 
<date id="FFA707FEFFC9FFD8564EFE9A618BFE46" box="[367,492,364,390]" pageId="22" pageNumber="23" value="1950-02-17">17.II.1950</date>
, G. Marlier (KMMA); 1ɗ, Kivu, Kissenyi, 1924, Van Saceghem (KMMA) 1ɗ, Kunungu, 
<date id="FFA707FEFFC9FFD85515FE6262CDFE6E" box="[564,682,404,430]" pageId="22" pageNumber="23" value="1921-04-08">8.IV.1921</date>
, H. Schouteden (KMMA); 1ɗ, Likete, 
<date id="FFA707FEFFC9FFD853A3FE62656DFE6E" box="[1154,1290,404,430]" pageId="22" pageNumber="23" value="1936-06-16">16.VI.1936</date>
, J. Ghesquière (KBIN); 1Ψ, Luebo, Kamalembi, 
<date id="FFA707FEFFC9FFD85402FE4A63CCFE16" box="[803,939,444,470]" pageId="22" pageNumber="23" value="1921-09-17">17.IX.1921</date>
, H. Schouteden (KBIN); 1ɗ 1Ψ, Lulonga, Lokolenge, 
<date id="FFA707FEFFC9FFD85525FE126219FE3E" box="[516,638,484,510]" pageId="22" pageNumber="23" value="1927-06">June 1927</date>
, J. Ghesquière (KMMA); Parc National de Garamba, mission H. De Saeger, 1ɗ II/fc/3, 
<date id="FFA707FEFFC9FFD85596FDFA632EFDE6" box="[695,841,524,550]" pageId="22" pageNumber="23" value="1951-07-16">16.VII.1951</date>
, H. De Saeger (2102); 1Ψ, II/hd/9, 
<date id="FFA707FEFFC9FFD85629FDC261C4FD8E" box="[264,419,564,590]" pageId="22" pageNumber="23" value="1951-08-21">21.VIII.1951</date>
, H. De Saeger (2297); 1Ψ, II/fd/9, 
<date id="FFA707FEFFC9FFD85478FDC2638CFD8E" box="[857,1003,564,590]" pageId="22" pageNumber="23" value="1952-07-16">16.VII.1952</date>
, H. De Saeger (3805), all (KMMA); 1ɗ, Uele, Buta, 
<date id="FFA707FEFFC9FFD8555BFDAA6352FDB6" box="[634,821,604,630]" pageId="22" pageNumber="23" value="1938-02">February 1938</date>
, J. Vrydagh (KMMA); 1Ψ, Tshikapa, 
<date id="FFA707FEFFC9FFD85629FD7261E8FD5E" box="[264,399,644,670]" pageId="22" pageNumber="23" value="1921-11-21">21.XI.1921</date>
, H. Schouteden (KMMA). 
<collectingCountry id="F30E61AEFFC9FFD855C9FD726444FD5E" box="[744,1059,644,670]" name="Equatorial Guinea" pageId="22" pageNumber="23">EQUATORIAL GUINEA</collectingCountry>
: 1ɗ, Uelleburg, VI­ 
<date id="FFA707FEFFC9FFD85629FD5A6119FD06" box="[264,382,684,710]" pageId="22" pageNumber="23" value="1908-08">VIII.1908</date>
, G. Tessmann (MNHU). 
<collectingCountry id="F30E61AEFFC9FFD85589FD5A6309FD06" box="[680,878,684,710]" name="Ivory Coast" pageId="22" pageNumber="23">IVORY COAST</collectingCountry>
: 1ɗ, Banco Nat. Park, N. Abidjan, S. side 
<geoCoordinate id="EE2D47F9FFC9FFD8561EFD2261C3FD2E" box="[319,420,724,750]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="5.366667">05°22’N</geoCoordinate>
<geoCoordinate id="EE2D47F9FFC9FFD8568DFD226270FD2E" box="[428,535,724,750]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-4.05">04°03’W</geoCoordinate>
, 
<date id="FFA707FEFFC9FFD85505FD2262A8FD2E" box="[548,719,724,750]" pageId="22" pageNumber="23" value="1989-04-23" valueMax="1989-04-27" valueMin="1989-04-23">23­27.IV.1989</date>
, “edges of wide track in forest”, J. Londt (NMSA); 1ɗ, Man, Bethani, 
<geoCoordinate id="EE2D47F9FFC9FFD856D3FD0A622FFCD6" box="[498,584,764,790]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="7.4">7°24’N</geoCoordinate>
<geoCoordinate id="EE2D47F9FFC9FFD85572FD0A62D7FCD6" box="[595,688,764,790]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-7.55">7°33’W</geoCoordinate>
, 
<date id="FFA707FEFFC9FFD855E1FD0A6349FCD6" box="[704,814,764,790]" pageId="22" pageNumber="23" value="1998-02-08">8.II.1998</date>
; 1Ψ, SWW Man, Idessa, 
<geoCoordinate id="EE2D47F9FFC9FFD85346FD0A64D9FCD6" box="[1127,1214,764,790]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="7.35">7°21’N</geoCoordinate>
<geoCoordinate id="EE2D47F9FFC9FFD853E8FD0A6541FCD6" box="[1225,1318,764,790]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-7.6">7°36’W</geoCoordinate>
, 
<date id="FFA707FEFFC9FFD85629FCD261E2FCFE" box="[264,389,804,830]" pageId="22" pageNumber="23" value="1998-02-10">10.II.1998</date>
, both C. Kassebeer &amp; S. Hilger (
<collectingCountry id="F30E61AEFFC9FFD85412FCD26313FCFE" box="[819,884,804,830]" name="China" pageId="22" pageNumber="23">TAU</collectingCountry>
). 
<collectingCountry id="F30E61AEFFC9FFD854ACFCD26462FCFE" box="[909,1029,804,830]" name="Liberia" pageId="22" pageNumber="23">LIBERIA</collectingCountry>
: 1ɗ, 
<quantity id="4CE18CDBFFC9FFD8536DFCD264EFFCFE" box="[1100,1160,804,830]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="22" pageNumber="23" unit="mi" value="20.0">20mi</quantity>
S Voinjama, 
<date id="FFA707FEFFC9FFD85629FCBA61C4FCA6" box="[264,419,844,870]" pageId="22" pageNumber="23" value="1966-08-13">13.VIII.1966</date>
, E.S. Ross &amp; K. Lorenzen (CAS). 
<collectingCountry id="F30E61AEFFC9FFD8541BFCBA63D0FCA6" box="[826,951,844,870]" name="Nigeria" pageId="22" pageNumber="23">NIGERIA</collectingCountry>
: Ilaro Forest, 1ɗ, 
<date id="FFA707FEFFC9FFD853A9FCBA656BFCA6" box="[1160,1292,844,870]" pageId="22" pageNumber="23" value="1974-04-21">21.IV.1974</date>
, J. Riley (BMNH); 1Ψ, 
<date id="FFA707FEFFC9FFD85527FC82620CFC4E" box="[518,619,884,910]" pageId="22" pageNumber="23" value="1974-01-05">5.I.1974</date>
, M.A. Cornes (BMNH); 2ɗɗ, Forest Mamu, 
<date id="FFA707FEFFC9FFD853BDFC82656FFC4E" box="[1180,1288,884,910]" pageId="22" pageNumber="23" value="1961-05-06">6.V.1961</date>
, J. Riley (BMNH); 1ɗ, Ibadan, 
<date id="FFA707FEFFC9FFD8554CFC6A6295FC76" box="[621,754,924,950]" pageId="22" pageNumber="23" value="1922-06-16">16.VI.1922</date>
(BMNH); 1Ψ, Ilesha, 
<date id="FFA707FEFFC9FFD8532EFC6A641CFC76" box="[1039,1147,924,950]" pageId="22" pageNumber="23" value="1911-02-03">3.II.1911</date>
, T.F.G. Mayer (BMNH); Olokemeji, 1ɗ, 
<date id="FFA707FEFFC9FFD85564FC326362FC1E" box="[581,773,964,990]" pageId="22" pageNumber="23" value="1912-12">December 1912</date>
; 2ɗɗ, 
<date id="FFA707FEFFC9FFD85478FC3263A1FC1E" box="[857,966,964,990]" pageId="22" pageNumber="23" value="1936-05-06">6.V.1936</date>
, “swept”, Van Zwaluwenburg &amp; McGough (USNM); 4ɗɗ, “Bridwell collection” (USNM). 
<collectingCountry id="F30E61AEFFC9FFD8533AFC1A6416FBC6" box="[1051,1137,1004,1030]" name="Togo" pageId="22" pageNumber="23">TOGO</collectingCountry>
: 1ɗ, Sodo, 
<date id="FFA707FEFFC9FFD85233FC1A611CFBEE" pageId="22" pageNumber="23" value="1982-01-02" valueMax="1982-01-21" valueMin="1982-01-02">2­ 21.I.1982</date>
, G.J. Steck (
<collectingCountry id="F30E61AEFFC9FFD8552DFBE2622AFBEE" box="[524,589,1044,1070]" name="China" pageId="22" pageNumber="23">TAU</collectingCountry>
). 
<collectingCountry id="F30E61AEFFC9FFD85541FBE26283FBEE" box="[608,740,1044,1070]" name="Uganda" pageId="22" pageNumber="23">UGANDA</collectingCountry>
: 1Ψ, N.W. Madi, W. Nile, Metu Hills, 
<date id="FFA707FEFFC9FFD8538CFBE26541FBEE" box="[1197,1318,1044,1070]" pageId="22" pageNumber="23" value="1954-06-05">5.VI.1954</date>
, van Someren (BMNH); 1ɗ, Masaka, Katera Forest, 
<date id="FFA707FEFFC9FFD85453FBCA63B1FB96" box="[882,982,1084,1110]" pageId="22" pageNumber="23" value="1955-09">IX.1955</date>
, van Someren (BMNH); 1Ψ, Budongo Forest, 
<date id="FFA707FEFFC9FFD856F3FB92623EFBBE" box="[466,601,1124,1150]" pageId="22" pageNumber="23" value="1938-02-07" valueMax="1938-02-08" valueMin="1938-02-07">7­8.II.1938</date>
, F.W. Edwards (BMNH).
</paragraph>
<paragraph id="8BA6213EFFC9FFD85629FB4261F2FB0E" blockId="22.[264,1324,1204,1310]" box="[264,405,1204,1230]" pageId="22" pageNumber="23">Distribution</paragraph>
<paragraph id="8BA6213EFFC9FFD85619FB2A637FFADE" blockId="22.[264,1324,1204,1310]" pageId="22" pageNumber="23">
<collectingCountry id="F30E61AEFFC9FFD85619FB2A61EFFB36" box="[312,392,1244,1270]" name="Ghana" pageId="22" pageNumber="23">Ghana</collectingCountry>
, 
<collectingCountry id="F30E61AEFFC9FFD856BAFB2A627FFB36" box="[411,536,1244,1270]" name="Cameroon" pageId="22" pageNumber="23">Cameroon</collectingCountry>
, 
<collectingCountry id="F30E61AEFFC9FFD8550DFB2A630EFB36" box="[556,873,1244,1270]" name="Central African Republic" pageId="22" pageNumber="23">Central African Republic</collectingCountry>
?, 
<collectingCountry id="F30E61AEFFC9FFD854A9FB2A63B1FB36" box="[904,982,1244,1270]" name="Republic of the Congo" pageId="22" pageNumber="23">Congo</collectingCountry>
(D.R.), 
<collectingCountry id="F30E61AEFFC9FFD85366FB2A654FFB36" box="[1095,1320,1244,1270]" name="Equatorial Guinea" pageId="22" pageNumber="23">Equatorial Guinea</collectingCountry>
, 
<collectingCountry id="F30E61AEFFC9FFD85629FAF261F2FADE" box="[264,405,1284,1310]" name="Ivory Coast" pageId="22" pageNumber="23">Ivory Coast</collectingCountry>
, 
<collectingCountry id="F30E61AEFFC9FFD85681FAF2619FFADE" box="[416,504,1284,1310]" name="Liberia" pageId="22" pageNumber="23">Liberia</collectingCountry>
, 
<collectingCountry id="F30E61AEFFC9FFD85525FAF26238FADE" box="[516,607,1284,1310]" name="Nigeria" pageId="22" pageNumber="23">Nigeria</collectingCountry>
, 
<collectingCountry id="F30E61AEFFC9FFD8554AFAF262CDFADE" box="[619,682,1284,1310]" name="Togo" pageId="22" pageNumber="23">Togo</collectingCountry>
, 
<collectingCountry id="F30E61AEFFC9FFD85594FAF26373FADE" box="[693,788,1284,1310]" name="Uganda" pageId="22" pageNumber="23">Uganda</collectingCountry>
.
</paragraph>
<paragraph id="8BA6213EFFC9FFD85629FAA2611DFAAE" blockId="22.[264,1323,1364,1510]" box="[264,378,1364,1390]" pageId="22" pageNumber="23">Hostplant</paragraph>
<paragraph id="8BA6213EFFC9FFD85619FA8A637FFA26" blockId="22.[264,1323,1364,1510]" pageId="22" pageNumber="23">
Material identified as 
<taxonomicName id="4C195ABDFFC9FFD8551BFA8A62FBFA55" box="[570,668,1404,1429]" class="Insecta" family="Tephritidae" genus="Ceratitis" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tritea">
<emphasis id="B96DFD2CFFC9FFD8551BFA8A62FBFA55" box="[570,668,1404,1429]" italics="true" pageId="22" pageNumber="23">C. tritea</emphasis>
</taxonomicName>
, but belonging to this species (according to the original description by 
<bibRefCitation id="EF885CCFFFC9FFD856E3FA52620BFA7E" author="Bezzi" box="[450,620,1444,1470]" pageId="22" pageNumber="40" refString="Bezzi, M. (1918) Notes on the Ethiopian fruit-flies of the family Trypaneidae, other than Dacus (s. l.), with descriptions of new genera and species (Dipt.) - I. Bulletin of Entomological Research, 8, 215 - 251." type="journal article" year="1918">Bezzi (1918))</bibRefCitation>
, from Olokomeji by Silvestri, was reared from fruits of 
<taxonomicName id="4C195ABDFFC9FFD85629FA3A637FFA26" authority="Baill." authorityName="Baill." box="[264,792,1484,1510]" class="Magnoliopsida" family="Icacinaceae" genus="Pyrenacantha" kingdom="Plantae" order="Icacinales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="vogeliana">
<emphasis id="B96DFD2CFFC9FFD85629FA3A6241FA25" box="[264,550,1484,1509]" italics="true" pageId="22" pageNumber="23">Pyrenacantha vogeliana</emphasis>
Baill. (Icacinaceae)
</taxonomicName>
</paragraph>
<paragraph id="8BA6213EFFC9FFD85629F9EA61E0F9F6" blockId="22.[264,1324,1564,1870]" box="[264,391,1564,1590]" pageId="22" pageNumber="23">Comments</paragraph>
<paragraph id="8BA6213EFFC9FFD95619F9B261BBFE46" blockId="22.[264,1324,1564,1870]" lastBlockId="23.[264,1324,284,510]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
Specimens originally identified as 
<taxonomicName id="4C195ABDFFC9FFD855F6F9B263E8F99D" box="[727,911,1604,1630]" class="Insecta" family="Tephritidae" genus="Carpophthoromyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pseudotritea">
<emphasis id="B96DFD2CFFC9FFD855F6F9B2628DF99D" box="[727,746,1604,1629]" italics="true" pageId="22" pageNumber="23">C</emphasis>
. 
<emphasis id="B96DFD2CFFC9FFD855DDF9B263E8F99D" box="[764,911,1604,1629]" italics="true" pageId="22" pageNumber="23">pseudotritea</emphasis>
</taxonomicName>
proved to belong to a complex of different species. They are all characterized by the presence of a brown medial band running from the ocellar triangle downwards to the antennal base, a character state absent in other 
<taxonomicName id="4C195ABDFFC9FFD8566DF94A624BF915" box="[332,556,1724,1749]" class="Insecta" family="Tephritidae" genus="Carpophthoromyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B96DFD2CFFC9FFD8566DF94A624BF915" box="[332,556,1724,1749]" italics="true" pageId="22" pageNumber="23">Carpophthoromyia</emphasis>
</taxonomicName>
. The species can only be differentiated from each other by slight differences in the position of the frontal and orbital setae, the width of the medial band, and the male and female genitalia. However, since the material for most of the sibling species is limited, and for two of the species the male sex is unknown, the intraspecific variation with regard to these characteristics is still largely unknown. Caution has to be taken in order to identify the species concerned correctly. The genitalia of those known sexes are hereby illustrated.
</paragraph>
<paragraph id="8BA6213EFFC8FFD95619FE6263ECFE3E" blockId="23.[264,1324,284,510]" pageId="23" pageNumber="24">
One specimen in the MNHU collection is labelled as originating from SE 
<collectingCountry id="F30E61AEFFC8FFD95388FE626540FE6E" box="[1193,1319,404,430]" name="Cameroon" pageId="23" pageNumber="24">Cameroon</collectingCountry>
. However, the other specifications on the label “Uam Gebiet” [Uam river territory] and “Bosum”, are situated in the 
<collectingCountry id="F30E61AEFFC8FFD9557BFE1263E0FE3E" box="[602,903,484,510]" name="Central African Republic" pageId="23" pageNumber="24">Central African Republic</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>