treatments-xml/data/03/80/87/038087FEFFC5FFDBF691F8BF3B6CC776.xml

<document id="1D486A56FEC3885C8BFB2D869A124CE3" ID-DOI="10.11646/zootaxa.3861.3.2" ID-GBIF-Dataset="7beda7dd-c67d-44f3-9e62-cf4500db236c" ID-ISSN="1175-5326" ID-Zenodo-Dep="252285" ID-ZooBank="0F6451EC-F2A1-4EC8-8C86-CB0A4EA73AB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460910737130" checkinUser="plazi" docAuthor="Grishin, Nick V." docDate="2014" docId="038087FEFFC5FFDBF691F8BF3B6CC776" docLanguage="en" docName="zt03861p248.pdf" docOrigin="Zootaxa 3861 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clito littera Mabille 1877" docType="treatment" docVersion="9" lastPageNumber="237" masterDocId="FFB9FF86FFC1FFDDF606FF8E3F3BC30B" masterDocTitle="Two new species of Clito from South America and a revision of the Clito littera group (Lepidoptera: Hesperiidae: Pyrginae)" masterLastPageNumber="248" masterPageNumber="231" pageNumber="235" updateTime="1698361186773" updateUser="plazi">
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<mods:title id="A7CD95103BD0A73B405FBCFB7B7B608B">Two new species of Clito from South America and a revision of the Clito littera group (Lepidoptera: Hesperiidae: Pyrginae)</mods:title>
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<treatment id="038087FEFFC5FFDBF691F8BF3B6CC776" ID-DOI="http://doi.org/10.5281/zenodo.6140304" ID-GBIF-Taxon="119588491" ID-Zenodo-Dep="6140304" LSID="urn:lsid:plazi:treatment:038087FEFFC5FFDBF691F8BF3B6CC776" httpUri="http://treatment.plazi.org/id/038087FEFFC5FFDBF691F8BF3B6CC776" lastPageId="6" lastPageNumber="237" pageId="4" pageNumber="235">
<subSubSection id="C3336563FFC5FFD9F691F8BF3D10C466" pageId="4" pageNumber="235" type="nomenclature">
<paragraph id="8B9636E8FFC5FFD9F691F8BF3DC1C447" blockId="4.[151,762,1841,1901]" box="[151,762,1841,1868]" pageId="4" pageNumber="235">
<heading id="D0DE8184FFC5FFD9F691F8BF3DC1C447" bold="true" box="[151,762,1841,1868]" fontSize="11" level="1" pageId="4" pageNumber="235" reason="1">
<emphasis id="B95DEAFAFFC5FFD9F691F8BF3DC1C447" bold="true" box="[151,762,1841,1868]" pageId="4" pageNumber="235">
<taxonomicName id="4C294D6BFFC5FFD9F691F8BF3EDDC447" ID-CoL="W89D" authority="Mabille, 1877" authorityName="Mabille" authorityYear="1877" box="[151,486,1841,1868]" class="Insecta" family="Hesperiidae" genus="Clito" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC5FFD9F691F8BF3E1AC440" bold="true" box="[151,289,1841,1867]" italics="true" pageId="4" pageNumber="235">Clito littera</emphasis>
(
<bibRefCitation id="EFB84B19FFC5FFD9F736F8BC3EE5C447" author="Mabille" box="[304,478,1842,1868]" pageId="4" pageNumber="235" refString="Mabille, P. (1877) Diagnoses de lepidopteres nouveaux du groupe des hesperides. Petites Nouvelles entomologiques, 2, 165 - 166." type="journal article" year="1877">Mabille, 1877</bibRefCitation>
)
</taxonomicName>
, 
<typeStatus id="5492884AFFC5FFD9F7F5F8BC3D58C447" box="[499,611,1842,1868]" pageId="4" pageNumber="235" type="lectotype">lectotype</typeStatus>
designation
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9636E8FFC5FFD9F691F8DA3D10C466" blockId="4.[151,762,1841,1901]" box="[151,555,1876,1901]" pageId="4" pageNumber="235">
(
<figureCitation id="13122A6DFFC5FFD9F699F8DA3E22C466" box="[159,281,1876,1901]" captionStart="FIGURES 9 – 16" captionStartId="3.[151,264,1731,1753]" captionTargetBox="[154,1433,193,1707]" captionTargetId="figure@3.[152,1435,193,1709]" captionTargetPageId="3" captionText="FIGURES 9 – 16. Type specimens of Clito from the C. littera group. 9 – 10. C. littera lectotype, designated here, &quot; Brasil &quot;, P. Mabille collection 1923, R. Oberthür Coll., Brit. Mus. 1927 - 3, specimen No. BMNH (E) # 982948 (genitalia Fig. 26 a, g); 11 – 12. C. anda [holo] type, French Guiana, Collect. C. Bar, R. Oberthür Coll., Brit. Mus. 1931 - 136, specimen No. BMNH (E) # 982947; (genitalia Fig. 26 m, n, o); 13 – 14. C. anda [para] type, French Guiana: Cayenne, R. Oberthür Coll., Brit. Mus. 1931 - 136, specimen No. BMNH (E) # 1054238; 15 – 16. C. nebulosa holotype, Bolivia, Rio Songo, 750 m, Coll. Fassl (genitalia Fig. 26 d). All specimens are males. Dorsal and ventral surfaces are shown on odd- and even-numbered figures, respectively. Labels are shown in-line with the specimen images. Labels are reduced 2.5 times compared to specimens: small scale bar below C. littera labels refers to labels, and larger scale bars refer to specimens. 15 – 16 is in SMF collection and is photographed by Ernst Brockmann, other specimens are in BMNH collection and images are copyright (©) Trustees of the Natural History Museum, London (used with permission)." httpUri="https://zenodo.org/record/252287/files/figure.png" pageId="4" pageNumber="235">Figs. 9–10</figureCitation>
, 
<figureCitation id="13122A6DFFC5FFD9F723F8DA3E78C466" box="[293,323,1876,1901]" captionStart="FIGURE 26" captionStartId="12.[151,250,1216,1238]" captionTargetBox="[202,1373,193,1183]" captionTargetId="figure@12.[202,1373,193,1183]" captionTargetPageId="12" captionText="FIGURE 26. Male genitalia of Clito littera group species. a, g. C. littera lectotype, designated here, &quot; Brasil &quot;, P. Mabille collection 1923, R. Oberthür Coll., Brit. Mus. 1927 - 3, specimen No. BMNH (E) # 982948 [BMNH], in situ on a specimen, scales brushed off: a. various views rotating from right posterolateral through posterior to left posterolateral; g. ventrolateral (specimen Figs. 9 – 10); b, c. C. litteroides holotype, Venezuela: Aragua State, Rancho Grande [Biological Station, approximate GPS coordinates 10 ° 21 ' N 67 ° 41 ' W], elevation 1100 m, 4 - Jun- 1985, leg. S. S. Nicolay, genitalia vial NVG 130614 - 42 [USNM]: b. section of complete genitalia in posterolateral view; c. complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 7 – 8); d. C. nebulosa holotype, ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Mielke &amp; Schröder (1994) (specimen Figs. 15 – 16); e, f. C. trinidadensis holotype, Trinidad, Morne Catherine, 24 - Mar- 1982, leg. M. J. W. Cock, specimen No. BMNH (E) # 982949 [BMNH], dorsal and lateral views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Cock (1998) (specimen Figs. 17 – 18); h. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall &amp; M. A. Solis [USNM], complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 5 – 6); i. C. jonkersi holotype, Surinam, Domburg, 8 - Mar- 1975, leg. E. H. Jonkers [RMNH], lateral internal view of left valva and lateral view of uncus and gnathos, illustration from de Jong (1983) (specimen Figs. 19 – 20); j. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall &amp; M. A. Solis [USNM], right valva and uncus in situ on a specimen, scales brushed off (specimen Figs. 3 – 4); k. C. palotschka holotype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, 4 - Mar- 2001, leg. D. H. Ahrenholz [USNM], right valva in situ on a specimen, scales brushed off (specimen Figs. 1 – 2); l. C. anda, Brazil: Rondônia, Cacaulândia municipality, Fazenda Rancho Grande, genitalia No. GTA 4377, illustration from Austin (2000), ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis; m – o. C. anda [holo] type, French Guiana, Collect. C. Bar, R. Oberthür Coll., Brit. Mus. 1931 – 136, specimen No. BMNH (E) # 982947 [BMNH] (specimen Figs. 11 – 12), left valva on m. and Evans's sketches: n. pinned in the collection [BMNH]; o. published, and more stylized, version (Evans 1953), lateral internal view of left valva, lateral and ventral views of uncus, gnathos and distal half of penis. Species name is given below each image, holotypes and paratypes are labeled as HT and PT. Gray &quot; F &quot; indicates mirror image (left-right inverted). Images a, f, g, m – o. are copyright (©) Trustees of the Natural History Museum, London (used with permission)." httpUri="https://zenodo.org/record/252291/files/figure.png" pageId="4" pageNumber="235">26</figureCitation>
a, g, 27 part, 28 part)
</paragraph>
</subSubSection>
<subSubSection id="C3336563FFC5FFDBF691F8123B6CC776" lastPageId="6" lastPageNumber="237" pageId="4" pageNumber="235" type="reference_group">
<paragraph id="8B9636E8FFC5FFD8F691F8123B85C08B" blockId="4.[151,1436,1948,2009]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="236" pageId="4" pageNumber="235">
<taxonomicName id="4C294D6BFFC5FFD9F691F8123E52C4BE" ID-CoL="98HWJ" box="[151,361,1948,1973]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC5FFD9F691F8123E52C4BE" box="[151,361,1948,1973]" italics="true" pageId="4" pageNumber="235">Telemiades littera</emphasis>
</taxonomicName>
has been described by 
<bibRefCitation id="EFB84B19FFC5FFD9F4A4F8123C6EC4BE" author="Mabille" box="[674,853,1948,1973]" pageId="4" pageNumber="235" refString="Mabille, P. (1877) Diagnoses de lepidopteres nouveaux du groupe des hesperides. Petites Nouvelles entomologiques, 2, 165 - 166." type="journal article" year="1877">Mabille (1877)</bibRefCitation>
from the unstated number of specimens from &quot;Apurimacensi valle et 
<collectingCountry id="F33E7678FFC5FFD9F7A4F84E3EC0C4D2" box="[418,507,1984,2009]" name="Peru" pageId="4" pageNumber="235">Peruvia</collectingCountry>
&quot; in the Mabille collection, of which one possible 
<typeStatus id="5492884AFFC5FFD9F228F84F3BBEC4D2" box="[1070,1157,1985,2009]" pageId="4" pageNumber="235" type="syntype">syntype</typeStatus>
is located in the BMNH collection. A search in ZMHB, where a significant fraction of Mabille 
<typeStatus id="5492884AFFC4FFD8F5C7FF163CC7C3BB" box="[961,1020,152,176]" pageId="5" pageNumber="236">types</typeStatus>
resides, did not reveal specimens of this species. This sole 
<typeStatus id="5492884AFFC4FFD8F796FF333EDCC3DE" box="[400,487,189,213]" pageId="5" pageNumber="236" type="syntype">syntype</typeStatus>
in BMNH agrees with the original description. According to the label, it comes from the Mabille collection and is curated as a 
<typeStatus id="5492884AFFC4FFD8F46DFF6E3DA0C3F3" box="[619,667,224,248]" pageId="5" pageNumber="236">type</typeStatus>
of &quot; 
<taxonomicName id="4C294D6BFFC4FFD8F4CDFF513C29C3F3" ID-CoL="98HWJ" box="[715,786,223,248]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">littera</taxonomicName>
&quot; (
<figureCitation id="13122A6DFFC4FFD8F52CFF513C9DC3F3" box="[810,934,223,248]" captionStart="FIGURES 9 – 16" captionStartId="3.[151,264,1731,1753]" captionTargetBox="[154,1433,193,1707]" captionTargetId="figure@3.[152,1435,193,1709]" captionTargetPageId="3" captionText="FIGURES 9 – 16. Type specimens of Clito from the C. littera group. 9 – 10. C. littera lectotype, designated here, &quot; Brasil &quot;, P. Mabille collection 1923, R. Oberthür Coll., Brit. Mus. 1927 - 3, specimen No. BMNH (E) # 982948 (genitalia Fig. 26 a, g); 11 – 12. C. anda [holo] type, French Guiana, Collect. C. Bar, R. Oberthür Coll., Brit. Mus. 1931 - 136, specimen No. BMNH (E) # 982947; (genitalia Fig. 26 m, n, o); 13 – 14. C. anda [para] type, French Guiana: Cayenne, R. Oberthür Coll., Brit. Mus. 1931 - 136, specimen No. BMNH (E) # 1054238; 15 – 16. C. nebulosa holotype, Bolivia, Rio Songo, 750 m, Coll. Fassl (genitalia Fig. 26 d). All specimens are males. Dorsal and ventral surfaces are shown on odd- and even-numbered figures, respectively. Labels are shown in-line with the specimen images. Labels are reduced 2.5 times compared to specimens: small scale bar below C. littera labels refers to labels, and larger scale bars refer to specimens. 15 – 16 is in SMF collection and is photographed by Ernst Brockmann, other specimens are in BMNH collection and images are copyright (©) Trustees of the Natural History Museum, London (used with permission)." httpUri="https://zenodo.org/record/252287/files/figure.png" pageId="5" pageNumber="236">Figs. 9–10</figureCitation>
). Handwriting on the label written in red ink and on two green labels matches that of Mabille. One of the green labels is a locality label. Contrary to the text in the description, it says &quot; 
<collectingCountry id="F33E7678FFC4FFD8F7A9FEA93EC1C24B" box="[431,506,295,320]" name="Brazil" pageId="5" pageNumber="236">Brasil</collectingCountry>
&quot;. Several possibilities may explain this discrepancy. First this specimen is not a 
<typeStatus id="5492884AFFC4FFD8F691FEC33FC9C26E" box="[151,242,333,357]" pageId="5" pageNumber="236" type="syntype">syntype</typeStatus>
. Second the 
<typeStatus id="5492884AFFC4FFD8F780FEC33EE6C26E" box="[390,477,333,357]" pageId="5" pageNumber="236" type="syntype">syntype</typeStatus>
is mislabeled and the Mabille description is accurate about the location. Third, the label on the 
<typeStatus id="5492884AFFC4FFD8F734FEFE3EB2C283" box="[306,393,368,392]" pageId="5" pageNumber="236" type="syntype">syntype</typeStatus>
is accurate and the Mabille description is in error. It is mostly likely that the Mabille description is in error and this specimen is a 
<typeStatus id="5492884AFFC4FFD8F48AFE1B3DD8C2A6" box="[652,739,405,429]" pageId="5" pageNumber="236" type="syntype">syntype</typeStatus>
originating from 
<collectingCountry id="F33E7678FFC4FFD8F5ACFE1A3CC9C2A6" box="[938,1010,404,429]" name="Brazil" pageId="5" pageNumber="236">Brazil</collectingCountry>
. The locality from the description can be translated as &quot;Apurimac valley and 
<collectingCountry id="F33E7678FFC4FFD8F449FE363DB3C2DB" box="[591,648,440,464]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
&quot;. The usage of &quot;and&quot; may indicate that there could have been at least two specimens. However, the Apurimac river is in 
<collectingCountry id="F33E7678FFC4FFD8F4D4FE533C32C2FF" box="[722,777,477,500]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
. Moreover, in a later publication (
<bibRefCitation id="EFB84B19FFC4FFD8F280FE523FF9C113" author="Mabille" pageId="5" pageNumber="236" refString="Mabille, P. &amp; Boullet E. (1912) Essai de revision de la famille des hesperides (suite). Annales des Sciences naturelles (Zoologie), Series 9, 16 (1 / 4), 1 - 159, pl. 1 - 2." type="journal article" year="1912">Mabille &amp; Boullet 1912: 121</bibRefCitation>
) only one specimen from &quot;Am. Mérid.&quot; (i.e., South 
<collectingCountry id="F33E7678FFC4FFD8F509FE713C4AC113" box="[783,881,511,536]" name="United States of America" pageId="5" pageNumber="236">America</collectingCountry>
) is mentioned, in the Mabille collection. While it is possible that the original description (
<bibRefCitation id="EFB84B19FFC4FFD8F451FDAA3DCBC137" author="Mabille" box="[599,752,548,573]" pageId="5" pageNumber="236" refString="Mabille, P. (1877) Diagnoses de lepidopteres nouveaux du groupe des hesperides. Petites Nouvelles entomologiques, 2, 165 - 166." type="journal article" year="1877">Mabille 1877</bibRefCitation>
) was written off specimens different from this one, 
<taxonomicName id="4C294D6BFFC4FFD8F331FDAB3AA0C136" ID-CoL="98HWJ" box="[1335,1435,548,573]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F331FDAB3AA0C136" box="[1335,1435,548,573]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
is very rare in collections and this sole specimen agrees very well with the detailed original description. It is possible that &quot;et&quot; in the original locality was meant to indicate a phrase &quot;that is&quot; rather than a conjunction &quot;and&quot;, and this specimen was the only one to base the description upon, or the locality statement is simply in error. Comparative analysis of the three Mabille labels suggests (John V. Calhoun, pers. comm.) that the label in red ink might have been the first one, and the green labels were added later. The red ink label is very similar to the other name labels used by Mabille. The name &quot; 
<taxonomicName id="4C294D6BFFC4FFD8F460FD733D89C01F" ID-CoL="98HWJ" box="[614,690,765,788]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="247" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F460FD733D89C01F" box="[614,690,765,788]" italics="true" pageId="5" pageNumber="236">Littera</emphasis>
</taxonomicName>
&quot; on the green label is overwritten with the letters &quot; 
<emphasis id="B95DEAFAFFC4FFD8F2E8FD733A32C01F" box="[1262,1289,765,788]" italics="true" pageId="5" pageNumber="236">Vu</emphasis>
&quot; (French for &quot;seen&quot;? beginning of a different name?). The style of &quot; 
<emphasis id="B95DEAFAFFC4FFD8F50BFCAF3C25C033" box="[781,798,801,824]" italics="true" pageId="5" pageNumber="236">V</emphasis>
&quot; matches well unique Mabille &quot; 
<emphasis id="B95DEAFAFFC4FFD8F294FCAF3B98C033" box="[1170,1187,801,824]" italics="true" pageId="5" pageNumber="236">V</emphasis>
&quot; on other labels. The &quot; 
<collectingCountry id="F33E7678FFC4FFD8F6A6FCCA3FD3C056" box="[160,232,836,861]" name="Brazil" pageId="5" pageNumber="236">Brasil</collectingCountry>
&quot; locality label is written on the same green paper as the identification label and was probably added at the same time. Either Mabille forgot the locality given in the description, or corrected that locality.
</paragraph>
<paragraph id="8B9636E8FFC4FFD8F6C1FC033CA4C697" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="236">
In addition to 
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<emphasis id="B95DEAFAFFC4FFD8F76AFC033EEFC0AE" box="[364,468,908,933]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
, Mabille
</taxonomicName>
described four other species in the same work (
<bibRefCitation id="EFB84B19FFC4FFD8F26CFC023A32C0AF" author="Mabille" box="[1130,1289,908,933]" pageId="5" pageNumber="236" refString="Mabille, P. (1877) Diagnoses de lepidopteres nouveaux du groupe des hesperides. Petites Nouvelles entomologiques, 2, 165 - 166." type="journal article" year="1877">Mabille 1877</bibRefCitation>
): 
<taxonomicName id="4C294D6BFFC4FFD8F319FC023E32C0C3" ID-CoL="555TB" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="lurideolus">
<emphasis id="B95DEAFAFFC4FFD8F319FC023E32C0C3" italics="true" pageId="5" pageNumber="236">Telemiades lurideolus</emphasis>
</taxonomicName>
, currently treated as a subjective synonym of 
<taxonomicName id="4C294D6BFFC4FFD8F547FC213B5DC0C3" ID-CoL="555SV" authority="Stoll" authorityName="Stoll" box="[833,1126,943,968]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="avitus">
<emphasis id="B95DEAFAFFC4FFD8F547FC213B35C0C3" box="[833,1038,943,968]" italics="true" pageId="5" pageNumber="236">Telemiades avitus</emphasis>
(Stoll)
</taxonomicName>
; 
<taxonomicName id="4C294D6BFFC4FFD8F27EFC213A69C0C3" ID-CoL="4BT8D" box="[1144,1362,943,968]" class="Insecta" family="Hesperiidae" genus="Pachyneuria" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="B95DEAFAFFC4FFD8F27EFC213A69C0C3" box="[1144,1362,943,968]" italics="true" pageId="5" pageNumber="236">Pachyneuria inops</emphasis>
</taxonomicName>
; 
<taxonomicName id="4C294D6BFFC4FFD8F361FC3F3E1DC0E6" ID-CoL="5D2XG" class="Insecta" family="Hesperiidae" genus="Zera" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="hyacinthinus">
<emphasis id="B95DEAFAFFC4FFD8F361FC3F3E1DC0E6" italics="true" pageId="5" pageNumber="236">Zera hyacinthinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C294D6BFFC4FFD8F759FC5A3DB1C0E6" ID-CoL="QFQC" box="[351,650,980,1005]" class="Insecta" family="Hesperiidae" genus="Camptopleura" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="theramenes">
<emphasis id="B95DEAFAFFC4FFD8F759FC5A3DB1C0E6" box="[351,650,980,1005]" italics="true" pageId="5" pageNumber="236">Camptopleura theramenes</emphasis>
</taxonomicName>
. Possible 
<typeStatus id="5492884AFFC4FFD8F4F9FC5B3C5AC0E6" box="[767,865,981,1005]" pageId="5" pageNumber="236" type="syntype">syntypes</typeStatus>
for all these taxa are extant in several collections, but locality labels for neither of them agree fully with their descriptions. While no locality is given in the description for 
<emphasis id="B95DEAFAFFC4FFD8F742FB933EA1C73F" box="[324,410,1053,1076]" italics="true" pageId="5" pageNumber="236">P. i n op s</emphasis>
, 
<taxonomicName id="4C294D6BFFC4FFD8F7ABFB933D62C73E" ID-CoL="5D2XG" box="[429,601,1052,1077]" class="Insecta" family="Hesperiidae" genus="Zera" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="hyacinthinus">
<emphasis id="B95DEAFAFFC4FFD8F7ABFB933D62C73E" box="[429,601,1052,1077]" italics="true" pageId="5" pageNumber="236">P. hyacinthinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C294D6BFFC4FFD8F494FB933C0EC73E" ID-CoL="QFQC" box="[658,821,1052,1077]" class="Insecta" family="Hesperiidae" genus="Camptopleura" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="theramenes">
<emphasis id="B95DEAFAFFC4FFD8F494FB933C0EC73E" box="[658,821,1052,1077]" italics="true" pageId="5" pageNumber="236">C. theramenes</emphasis>
</taxonomicName>
, labels on likely 
<typeStatus id="5492884AFFC4FFD8F5FCFB933B67C73E" box="[1018,1116,1053,1077]" pageId="5" pageNumber="236" type="syntype">syntypes</typeStatus>
state: &quot; 
<collectingCountry id="F33E7678FFC4FFD8F2B7FB923BC2C73E" box="[1201,1273,1052,1077]" name="Brazil" pageId="5" pageNumber="236">Brésil</collectingCountry>
&quot; (and &quot; 
<collectingCountry id="F33E7678FFC4FFD8F34BFB923AAFC73F" box="[1357,1428,1052,1076]" name="Peru" pageId="5" pageNumber="236">Perou</collectingCountry>
&quot; on a possible 
<typeStatus id="5492884AFFC4FFD8F73FFBCE3EABC753" box="[313,400,1088,1112]" pageId="5" pageNumber="236" type="syntype">syntype</typeStatus>
from IRSN), &quot;Chiriquí&quot; (on a 
<typeStatus id="5492884AFFC4FFD8F4FEFBCE3C74C753" box="[760,847,1088,1112]" pageId="5" pageNumber="236" type="syntype">syntype</typeStatus>
in ZMHB, but the BMNH 
<typeStatus id="5492884AFFC4FFD8F294FBCE3BD2C753" box="[1170,1257,1088,1112]" pageId="5" pageNumber="236" type="syntype">syntype</typeStatus>
has no locality label) and &quot;Bogota, 
<collectingCountry id="F33E7678FFC4FFD8F77AFBEA3ECBC776" box="[380,496,1124,1149]" name="Colombia" pageId="5" pageNumber="236">Colombia</collectingCountry>
&quot;, respectively. For 
<taxonomicName id="4C294D6BFFC4FFD8F4CFFBEB3C63C776" ID-CoL="555TB" box="[713,856,1124,1149]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="lurideolus">
<emphasis id="B95DEAFAFFC4FFD8F4CFFBEB3C63C776" box="[713,856,1124,1149]" italics="true" pageId="5" pageNumber="236">T. lurideolus</emphasis>
</taxonomicName>
, the description states &quot; 
<collectingCountry id="F33E7678FFC4FFD8F261FBEA3BFEC776" box="[1127,1221,1124,1149]" name="Brazil" pageId="5" pageNumber="236">Brasilia</collectingCountry>
&quot;, but the specimen label gives &quot; 
<collectingCountry id="F33E7678FFC4FFD8F719FB093EADC7AB" box="[287,406,1159,1184]" name="Colombia" pageId="5" pageNumber="236">Colombia</collectingCountry>
&quot;. Most of these localities from the labels have been used as deduced 
<typeStatus id="5492884AFFC4FFD8F29EFB073BF3C7AA" box="[1176,1224,1161,1185]" pageId="5" pageNumber="236">type</typeStatus>
localities for these taxa (
<bibRefCitation id="EFB84B19FFC4FFD8F6D0FB223E50C7CF" author="Mielke" box="[214,363,1196,1221]" pageId="5" pageNumber="236" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B." type="book" year="2004">Mielke 2004</bibRefCitation>
, 
<bibRefCitation id="EFB84B19FFC4FFD8F77EFB223E8EC7CE" author="Mielke" box="[376,437,1196,1221]" pageId="5" pageNumber="236" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera). Sociedade Brasileira de Zoologia, Curitiba, Parana, Brazil, xiii + 1536 pp." type="book" year="2005">2005</bibRefCitation>
). Interestingly, the sequence in which these taxa were described was 
<taxonomicName id="4C294D6BFFC4FFD8F2C9FB223A7AC7CE" ID-CoL="555TB" box="[1231,1345,1196,1221]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="lurideolus">
<emphasis id="B95DEAFAFFC4FFD8F2C9FB223A7AC7CE" box="[1231,1345,1196,1221]" italics="true" pageId="5" pageNumber="236">lurideolus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C294D6BFFC4FFD8F356FB223AAFC7CE" ID-CoL="98HWJ" box="[1360,1428,1196,1221]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F356FB223AAFC7CE" box="[1360,1428,1196,1221]" italics="true" pageId="5" pageNumber="236">littera</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C294D6BFFC4FFD8F691FB5F3FEFC7E3" ID-CoL="4BT8D" box="[151,212,1233,1256]" class="Insecta" family="Hesperiidae" genus="Pachyneuria" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="B95DEAFAFFC4FFD8F691FB5F3FEFC7E3" box="[151,212,1233,1256]" italics="true" pageId="5" pageNumber="236">inops</emphasis>
</taxonomicName>
, etc. Localities on their labels are: 
<collectingCountry id="F33E7678FFC4FFD8F473FB413DDDC7E3" box="[629,742,1231,1256]" name="Colombia" pageId="5" pageNumber="236">Colombia</collectingCountry>
, 
<collectingCountry id="F33E7678FFC4FFD8F4F3FB413C07C7E3" box="[757,828,1231,1256]" name="Brazil" pageId="5" pageNumber="236">Brazil</collectingCountry>
, 
<collectingCountry id="F33E7678FFC4FFD8F54DFB5F3C45C7E3" box="[843,894,1233,1256]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
or 
<collectingCountry id="F33E7678FFC4FFD8F5ACFB413CCAC7E3" box="[938,1009,1231,1256]" name="Brazil" pageId="5" pageNumber="236">Brazil</collectingCountry>
, but localities in the description are: 
<collectingCountry id="F33E7678FFC4FFD8F691FB7A3FE5C606" box="[151,222,1268,1293]" name="Brazil" pageId="5" pageNumber="236">Brazil</collectingCountry>
, 
<collectingCountry id="F33E7678FFC4FFD8F6ECFB7B3E1BC607" box="[234,288,1269,1292]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
, no locality given. It is not inconceivable that a shift by one in the localities and taxa has occurred in the descriptions. If this shift is corrected, locality for 
<taxonomicName id="4C294D6BFFC4FFD8F4E2FA963C72C63B" ID-CoL="98HWJ" box="[740,841,1303,1328]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F4E2FA963C72C63B" box="[740,841,1303,1328]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
becomes 
<collectingCountry id="F33E7678FFC4FFD8F5BEFA993CC4C63B" box="[952,1023,1303,1328]" name="Brazil" pageId="5" pageNumber="236">Brazil</collectingCountry>
, and 
<taxonomicName id="4C294D6BFFC4FFD8F23FFA973BAAC63B" ID-CoL="4BT8D" box="[1081,1169,1305,1328]" class="Insecta" family="Hesperiidae" genus="Pachyneuria" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="inops">
<emphasis id="B95DEAFAFFC4FFD8F23FFA973BAAC63B" box="[1081,1169,1305,1328]" italics="true" pageId="5" pageNumber="236">P. inops</emphasis>
</taxonomicName>
, for which two possible 
<typeStatus id="5492884AFFC4FFD8F691FAB33FC2C65E" box="[151,249,1341,1365]" pageId="5" pageNumber="236" type="syntype">syntypes</typeStatus>
(in IRSN and BMNH) are extant, will get the current 
<taxonomicName id="4C294D6BFFC4FFD8F555FAB33C83C65E" ID-CoL="98HWJ" box="[851,952,1340,1365]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F555FAB33C83C65E" box="[851,952,1340,1365]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
description locality &quot;Apurimacensi valle et 
<collectingCountry id="F33E7678FFC4FFD8F691FAD13FC8C673" box="[151,243,1375,1400]" name="Peru" pageId="5" pageNumber="236">Peruvia</collectingCountry>
&quot;. Notably, specimens matching 
<taxonomicName id="4C294D6BFFC4FFD8F45CFAEE3D84C673" ID-CoL="98HWJ" box="[602,703,1375,1400]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F45CFAEE3D84C673" box="[602,703,1375,1400]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
description are not known from 
<collectingCountry id="F33E7678FFC4FFD8F235FAEF3B52C673" box="[1075,1129,1377,1400]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
, but have been collected in 
<collectingCountry id="F33E7678FFC4FFD8F691FA0A3FDBC696" box="[151,224,1412,1437]" name="Brazil" pageId="5" pageNumber="236">Brazil</collectingCountry>
: Rio de Janeiro (e.g., Gávea, Petrópolis) [OM-DZUP, MGCL].
</paragraph>
<paragraph id="8B9636E8FFC4FFD8F6C1FA263E71C4D6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="236">
One additional curiosity is that 
<bibRefCitation id="EFB84B19FFC4FFD8F432FA293DEEC6CB" author="Draudt" box="[564,725,1447,1472]" pageId="5" pageNumber="236" refString="Draudt, M. W. K. (1922) B. Grypocera, breitkopfige Tagfalter. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Vol. 5. Alfred Kernen, Stuttgart, pp. 857 - 872, pls. 160 - 191." type="book chapter" year="1922">Draudt (1922)</bibRefCitation>
attempted to illustrate dorsal and ventral aspects of 
<taxonomicName id="4C294D6BFFC4FFD8F332FA263AA0C6CB" ID-CoL="98HWJ" box="[1332,1435,1447,1472]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F332FA263AA0C6CB" box="[1332,1435,1447,1472]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB84B19FFC4FFD8F698FA423EA4C6EE" author="Draudt" box="[158,415,1484,1509]" pageId="5" pageNumber="236" refString="Draudt, M. W. K. (1922) B. Grypocera, breitkopfige Tagfalter. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Vol. 5. Alfred Kernen, Stuttgart, pp. 857 - 872, pls. 160 - 191." type="book chapter" year="1922" yearSuffix="a">Draudt 1922: pl. 169a</bibRefCitation>
). Interestingly, these illustrations not only disagree with the characters from the 
<taxonomicName id="4C294D6BFFC4FFD8F333FA433AA0C6EE" ID-CoL="98HWJ" box="[1333,1435,1484,1509]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F333FA433AA0C6EE" box="[1333,1435,1484,1509]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
original description, but also do not agree with the text about 
<taxonomicName id="4C294D6BFFC4FFD8F556FA7E3C8DC503" ID-CoL="98HWJ" box="[848,950,1519,1544]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F556FA7E3C8DC503" box="[848,950,1519,1544]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
that they were supposed to illustrate in the publication itself (
<bibRefCitation id="EFB84B19FFC4FFD8F769F99A3D51C526" author="Draudt" box="[367,618,1556,1581]" pageId="5" pageNumber="236" refString="Draudt, M. W. K. (1922) B. Grypocera, breitkopfige Tagfalter. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Vol. 5. Alfred Kernen, Stuttgart, pp. 857 - 872, pls. 160 - 191." type="book chapter" year="1922" yearSuffix="p">Draudt 1922: p. 869</bibRefCitation>
). 
<bibRefCitation id="EFB84B19FFC4FFD8F485F99B3CBAC526" author="Evans" box="[643,897,1556,1581]" pageId="5" pageNumber="236" refString="Evans, W. H. (1953) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part III (Groups E, F, G) Pyrginae. Section 2. British Museum (Natural History), London, v + 178 pp., pls. 26 - 53." type="book chapter" year="1953" yearSuffix="p">Evans (1953: p. 153)</bibRefCitation>
noticed a problem and corrected the Draudt identification of illustrations to 
<taxonomicName id="4C294D6BFFC4FFD8F7FAF9B93C9DC55B" ID-CoL="555TF" authority="Hewitson, 1867" authorityName="Hewitson" authorityYear="1867" box="[508,934,1591,1616]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="penidas">
<emphasis id="B95DEAFAFFC4FFD8F7FAF9B93DECC55B" box="[508,727,1591,1616]" italics="true" pageId="5" pageNumber="236">Telemiades penidas</emphasis>
(Hewitson, 1867)
</taxonomicName>
, which is likely to be correct. Evans himself illustrated genitalia of 
<taxonomicName id="4C294D6BFFC4FFD8F794F9D33ECCC57E" ID-CoL="98HWJ" box="[402,503,1628,1653]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F794F9D33ECCC57E" box="[402,503,1628,1653]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
with a sketch (1953: Plate 44: E.52. 1). These sketches, showing ventral and lateral views of the side of uncus, gnathos, tegumen and penis and left valva in medial view were drawn by Evans from crude preparation of genitalia that he glued to carton cards. However, genitalia of 
<taxonomicName id="4C294D6BFFC4FFD8F237F92B3BADC5B6" ID-CoL="98HWJ" box="[1073,1174,1700,1725]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F237F92B3BADC5B6" box="[1073,1174,1700,1725]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
<typeStatus id="5492884AFFC4FFD8F298F92B3BCEC5B6" box="[1182,1269,1701,1725]" pageId="5" pageNumber="236" type="syntype">syntype</typeStatus>
were intact on the specimen, only the scales around genitalia were brushed off to reveal the distal ends of the valvae and uncus (
<figureCitation id="13122A6DFFC4FFD8F699F9623FC5C40E" box="[159,254,1772,1797]" captionStart="FIGURE 26" captionStartId="12.[151,250,1216,1238]" captionTargetBox="[202,1373,193,1183]" captionTargetId="figure@12.[202,1373,193,1183]" captionTargetPageId="12" captionText="FIGURE 26. Male genitalia of Clito littera group species. a, g. C. littera lectotype, designated here, &quot; Brasil &quot;, P. Mabille collection 1923, R. Oberthür Coll., Brit. Mus. 1927 - 3, specimen No. BMNH (E) # 982948 [BMNH], in situ on a specimen, scales brushed off: a. various views rotating from right posterolateral through posterior to left posterolateral; g. ventrolateral (specimen Figs. 9 – 10); b, c. C. litteroides holotype, Venezuela: Aragua State, Rancho Grande [Biological Station, approximate GPS coordinates 10 ° 21 ' N 67 ° 41 ' W], elevation 1100 m, 4 - Jun- 1985, leg. S. S. Nicolay, genitalia vial NVG 130614 - 42 [USNM]: b. section of complete genitalia in posterolateral view; c. complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 7 – 8); d. C. nebulosa holotype, ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Mielke &amp; Schröder (1994) (specimen Figs. 15 – 16); e, f. C. trinidadensis holotype, Trinidad, Morne Catherine, 24 - Mar- 1982, leg. M. J. W. Cock, specimen No. BMNH (E) # 982949 [BMNH], dorsal and lateral views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Cock (1998) (specimen Figs. 17 – 18); h. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall &amp; M. A. Solis [USNM], complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 5 – 6); i. C. jonkersi holotype, Surinam, Domburg, 8 - Mar- 1975, leg. E. H. Jonkers [RMNH], lateral internal view of left valva and lateral view of uncus and gnathos, illustration from de Jong (1983) (specimen Figs. 19 – 20); j. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall &amp; M. A. Solis [USNM], right valva and uncus in situ on a specimen, scales brushed off (specimen Figs. 3 – 4); k. C. palotschka holotype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, 4 - Mar- 2001, leg. D. H. Ahrenholz [USNM], right valva in situ on a specimen, scales brushed off (specimen Figs. 1 – 2); l. C. anda, Brazil: Rondônia, Cacaulândia municipality, Fazenda Rancho Grande, genitalia No. GTA 4377, illustration from Austin (2000), ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis; m – o. C. anda [holo] type, French Guiana, Collect. C. Bar, R. Oberthür Coll., Brit. Mus. 1931 – 136, specimen No. BMNH (E) # 982947 [BMNH] (specimen Figs. 11 – 12), left valva on m. and Evans's sketches: n. pinned in the collection [BMNH]; o. published, and more stylized, version (Evans 1953), lateral internal view of left valva, lateral and ventral views of uncus, gnathos and distal half of penis. Species name is given below each image, holotypes and paratypes are labeled as HT and PT. Gray &quot; F &quot; indicates mirror image (left-right inverted). Images a, f, g, m – o. are copyright (©) Trustees of the Natural History Museum, London (used with permission)." httpUri="https://zenodo.org/record/252291/files/figure.png" pageId="5" pageNumber="236">Figs. 26</figureCitation>
a, g). Apparently, Evans's sketch that shows the base of internal surface of a valva could not have been drawn from this specimen. No other nominal 
<taxonomicName id="4C294D6BFFC4FFD8F4ACF89E3C29C423" ID-CoL="98HWJ" box="[682,786,1807,1832]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F4ACF89E3C29C423" box="[682,786,1807,1832]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
specimens were mentioned by 
<bibRefCitation id="EFB84B19FFC4FFD8F287F89E3A24C423" author="Evans" box="[1153,1311,1807,1832]" pageId="5" pageNumber="236" refString="Evans, W. H. (1953) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part III (Groups E, F, G) Pyrginae. Section 2. British Museum (Natural History), London, v + 178 pp., pls. 26 - 53." type="book chapter" year="1953">Evans (1953)</bibRefCitation>
: &quot;Only the 
<typeStatus id="5492884AFFC4FFD8F691F8BA3FF1C447" box="[151,202,1844,1868]" pageId="5" pageNumber="236">type</typeStatus>
&quot; specimen was in the collection. On the other hand, genitalia of 
<taxonomicName id="4C294D6BFFC4FFD8F5AFF8BB3BD9C446" authority="Evans, 1953" authorityName="Evans" authorityYear="1953" box="[937,1250,1844,1869]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="subSpecies" species="littera" subSpecies="anda">
<emphasis id="B95DEAFAFFC4FFD8F5AFF8BB3B74C446" box="[937,1103,1844,1869]" italics="true" pageId="5" pageNumber="236">C. littera anda</emphasis>
<bibRefCitation id="EFB84B19FFC4FFD8F251F8BB3BD9C446" author="Evans" box="[1111,1250,1844,1869]" pageId="5" pageNumber="236" refString="Evans, W. H. (1953) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part III (Groups E, F, G) Pyrginae. Section 2. British Museum (Natural History), London, v + 178 pp., pls. 26 - 53." type="book chapter" year="1953">Evans, 1953</bibRefCitation>
</taxonomicName>
[holo] 
<typeStatus id="5492884AFFC4FFD8F32AF8BA3A64C446" box="[1324,1375,1844,1869]" pageId="5" pageNumber="236">type</typeStatus>
have been prepared (
<figureCitation id="13122A6DFFC4FFD8F743F8D93E80C47B" box="[325,443,1879,1904]" captionStart="FIGURES 9 – 16" captionStartId="3.[151,264,1731,1753]" captionTargetBox="[154,1433,193,1707]" captionTargetId="figure@3.[152,1435,193,1709]" captionTargetPageId="3" captionText="FIGURES 9 – 16. Type specimens of Clito from the C. littera group. 9 – 10. C. littera lectotype, designated here, &quot; Brasil &quot;, P. Mabille collection 1923, R. Oberthür Coll., Brit. Mus. 1927 - 3, specimen No. BMNH (E) # 982948 (genitalia Fig. 26 a, g); 11 – 12. C. anda [holo] type, French Guiana, Collect. C. Bar, R. Oberthür Coll., Brit. Mus. 1931 - 136, specimen No. BMNH (E) # 982947; (genitalia Fig. 26 m, n, o); 13 – 14. C. anda [para] type, French Guiana: Cayenne, R. Oberthür Coll., Brit. Mus. 1931 - 136, specimen No. BMNH (E) # 1054238; 15 – 16. C. nebulosa holotype, Bolivia, Rio Songo, 750 m, Coll. Fassl (genitalia Fig. 26 d). All specimens are males. Dorsal and ventral surfaces are shown on odd- and even-numbered figures, respectively. Labels are shown in-line with the specimen images. Labels are reduced 2.5 times compared to specimens: small scale bar below C. littera labels refers to labels, and larger scale bars refer to specimens. 15 – 16 is in SMF collection and is photographed by Ernst Brockmann, other specimens are in BMNH collection and images are copyright (©) Trustees of the Natural History Museum, London (used with permission)." httpUri="https://zenodo.org/record/252287/files/figure.png" pageId="5" pageNumber="236">Fig. 11-12</figureCitation>
, valva on 
<figureCitation id="13122A6DFFC4FFD8F429F8D93DBBC47B" box="[559,640,1879,1904]" captionStart="FIGURE 26" captionStartId="12.[151,250,1216,1238]" captionTargetBox="[202,1373,193,1183]" captionTargetId="figure@12.[202,1373,193,1183]" captionTargetPageId="12" captionText="FIGURE 26. Male genitalia of Clito littera group species. a, g. C. littera lectotype, designated here, &quot; Brasil &quot;, P. Mabille collection 1923, R. Oberthür Coll., Brit. Mus. 1927 - 3, specimen No. BMNH (E) # 982948 [BMNH], in situ on a specimen, scales brushed off: a. various views rotating from right posterolateral through posterior to left posterolateral; g. ventrolateral (specimen Figs. 9 – 10); b, c. C. litteroides holotype, Venezuela: Aragua State, Rancho Grande [Biological Station, approximate GPS coordinates 10 ° 21 ' N 67 ° 41 ' W], elevation 1100 m, 4 - Jun- 1985, leg. S. S. Nicolay, genitalia vial NVG 130614 - 42 [USNM]: b. section of complete genitalia in posterolateral view; c. complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 7 – 8); d. C. nebulosa holotype, ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Mielke &amp; Schröder (1994) (specimen Figs. 15 – 16); e, f. C. trinidadensis holotype, Trinidad, Morne Catherine, 24 - Mar- 1982, leg. M. J. W. Cock, specimen No. BMNH (E) # 982949 [BMNH], dorsal and lateral views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Cock (1998) (specimen Figs. 17 – 18); h. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall &amp; M. A. Solis [USNM], complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 5 – 6); i. C. jonkersi holotype, Surinam, Domburg, 8 - Mar- 1975, leg. E. H. Jonkers [RMNH], lateral internal view of left valva and lateral view of uncus and gnathos, illustration from de Jong (1983) (specimen Figs. 19 – 20); j. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall &amp; M. A. Solis [USNM], right valva and uncus in situ on a specimen, scales brushed off (specimen Figs. 3 – 4); k. C. palotschka holotype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, 4 - Mar- 2001, leg. D. H. Ahrenholz [USNM], right valva in situ on a specimen, scales brushed off (specimen Figs. 1 – 2); l. C. anda, Brazil: Rondônia, Cacaulândia municipality, Fazenda Rancho Grande, genitalia No. GTA 4377, illustration from Austin (2000), ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis; m – o. C. anda [holo] type, French Guiana, Collect. C. Bar, R. Oberthür Coll., Brit. Mus. 1931 – 136, specimen No. BMNH (E) # 982947 [BMNH] (specimen Figs. 11 – 12), left valva on m. and Evans's sketches: n. pinned in the collection [BMNH]; o. published, and more stylized, version (Evans 1953), lateral internal view of left valva, lateral and ventral views of uncus, gnathos and distal half of penis. Species name is given below each image, holotypes and paratypes are labeled as HT and PT. Gray &quot; F &quot; indicates mirror image (left-right inverted). Images a, f, g, m – o. are copyright (©) Trustees of the Natural History Museum, London (used with permission)." httpUri="https://zenodo.org/record/252291/files/figure.png" pageId="5" pageNumber="236">
Fig. 
<quantity id="4CD19B0DFFC4FFD8F465F8D93DBBC47B" box="[611,640,1879,1904]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="236" unit="m" value="26.0">26</quantity>
</figureCitation>
m, unfortunately both valvae now damaged). Evans's sketch must refer to 
<taxonomicName id="4C294D6BFFC4FFD8F6B3F8F33E66C49E" box="[181,349,1916,1941]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="subSpecies" species="littera" subSpecies="anda">
<emphasis id="B95DEAFAFFC4FFD8F6B3F8F33E66C49E" box="[181,349,1916,1941]" italics="true" pageId="5" pageNumber="236">C. littera anda</emphasis>
</taxonomicName>
<typeStatus id="5492884AFFC4FFD8F763F8F23EF1C49E" box="[357,458,1916,1941]" pageId="5" pageNumber="236" type="holotype">holotype</typeStatus>
, rather than to nominal 
<taxonomicName id="4C294D6BFFC4FFD8F4DBF8F33C78C49E" ID-CoL="98HWJ" box="[733,835,1916,1941]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC4FFD8F4DBF8F33C78C49E" box="[733,835,1916,1941]" italics="true" pageId="5" pageNumber="236">C. littera</emphasis>
</taxonomicName>
, because: (1) Evans mentions no 
<taxonomicName id="4C294D6BFFC4FFD8F2C1F8F33E05C4B3" authority="Draudt, 1924" authorityName="Draudt" authorityYear="1924" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="subSpecies" species="littera" subSpecies="nebulosa">
<emphasis id="B95DEAFAFFC4FFD8F2C1F8F33AA0C49E" box="[1223,1435,1916,1941]" italics="true" pageId="5" pageNumber="236">C. littera nebulosa</emphasis>
(
<bibRefCitation id="EFB84B19FFC4FFD8F698F82E3E0DC4B3" author="Draudt" box="[158,310,1952,1977]" pageId="5" pageNumber="236" refString="Draudt, M. W. K. (1924) Nachtrage: Lycaenidae-Nachtrage: Hesperidae. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Vol. 5. Alfred Kernen, Stuttgart, pp. 1043 - 1048." type="book" year="1924">Draudt, 1924</bibRefCitation>
)
</taxonomicName>
specimens, (2) the 
<taxonomicName id="4C294D6BFFC4FFD8F41AF82F3DFAC4B2" box="[540,705,1952,1977]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="subSpecies" species="littera" subSpecies="anda">
<emphasis id="B95DEAFAFFC4FFD8F41AF82F3DFAC4B2" box="[540,705,1952,1977]" italics="true" pageId="5" pageNumber="236">C. littera anda</emphasis>
</taxonomicName>
<typeStatus id="5492884AFFC4FFD8F4CFF82F3DC2C4B2" box="[713,761,1953,1977]" pageId="5" pageNumber="236">type</typeStatus>
series consists of two specimens and (3) 
<taxonomicName id="4C294D6BFFC4FFD8F2C9F82E3A3CC4B2" ID-CoL="9BTRG" box="[1231,1287,1952,1977]" class="Insecta" family="Hesperiidae" genus="Clito" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="anda">
<emphasis id="B95DEAFAFFC4FFD8F2C9F82E3A3CC4B2" box="[1231,1287,1952,1977]" italics="true" pageId="5" pageNumber="236">anda</emphasis>
</taxonomicName>
<typeStatus id="5492884AFFC4FFD8F308F82F3A55C4B2" box="[1294,1390,1953,1977]" pageId="5" pageNumber="236" type="paratype">paratype</typeStatus>
has intact abdomen.
</paragraph>
<paragraph id="8B9636E8FFC7FFDBF6C1FF193C07C113" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="237">
Therefore, neither facies, nor genitalia of nominal 
<taxonomicName id="4C294D6BFFC7FFDBF506FF163C5EC3BB" ID-CoL="98HWJ" box="[768,869,151,176]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF506FF163C5EC3BB" box="[768,869,151,176]" italics="true" pageId="6" pageNumber="237">C. littera</emphasis>
</taxonomicName>
have been illustrated in publications. However, in the BMNH collection drawer, next to the 
<taxonomicName id="4C294D6BFFC7FFDBF47EFF333DE4C3DE" ID-CoL="98HWJ" box="[632,735,188,213]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF47EFF333DE4C3DE" box="[632,735,188,213]" italics="true" pageId="6" pageNumber="237">C. littera</emphasis>
</taxonomicName>
possible 
<typeStatus id="5492884AFFC7FFDBF54BFF333C9CC3DE" box="[845,935,189,213]" pageId="6" pageNumber="237" type="syntype">syntype</typeStatus>
, there is an excellent drawing of both wing surfaces (
<figureCitation id="13122A6DFFC7FFDBF705FF513E6DC3F3" box="[259,342,223,248]" captionStart="FIGURES 17 – 23" captionStartId="4.[151,264,1463,1485]" captionTargetBox="[154,1434,195,1433]" captionTargetId="figure@4.[152,1435,193,1442]" captionTargetPageId="4" captionText="FIGURES 17 – 23. Type specimens and illustrations of Clito from the C. littera group. 17 – 18. C. trinidadensis holotype, Trinidad, Morne Catherine, 24 - Mar- 1982, leg. M. J. W. Cock, specimen No. BMNH (E) # 982949 [BMNH] (genitalia shown in Figs. 26 e, f); 19 – 20. C. jonkersi holotype, Surinam, Domburg, 8 - Mar- 1975, leg. E. H. Jonkers [RMNH] (genitalia Fig. 26 i); 21. C. littera, unpublished illustration, likely of the lectotype, dorsal—left, ventral—right, pinned in the collection next to the specimen [BMNH]; 22. C. nebulosa original illustration of the holotype from Draudt (1924) dorsal side, half-specimen image is shown for easy comparison on 23. (and Figs. 15 – 16). Dorsal and ventral surfaces are shown on odd- and even-numbered figures, respectively, except illustrations. Labels are shown between the specimen images. Labels are reduced 2.5 times compared to specimens: small scale bar below C. trinidadensis locality labels refers to labels, and larger scale bars refer to specimens. Photographs 19 – 20 are by Bernard Hermier, images 17 – 18 and 21 are copyright (©) Trustees of the Natural History Museum, London (used with permission)." httpUri="https://zenodo.org/record/252288/files/figure.png" pageId="6" pageNumber="237">Fig. 21</figureCitation>
), mostly likely of this specimen. John V. Calhoun (pers. comm.) suggested that this unpublished drawing might be by Jules Culot (
<date id="FF971028FFC7FFDBF431FE8A3D81C216" box="[567,698,260,285]" pageId="6" pageNumber="237" value="1861" valueMax="1933">1861–1933</date>
), who illustrated many species in &quot;Etudes de lépidoptérologie comparée&quot; by Charles Oberthür. The Oberthürs frequently pinning often uncolored cut-outs of Culot’s published figures with the associated specimens in their collection. It is possible that Charles Oberthür thought about publishing a 
<taxonomicName id="4C294D6BFFC7FFDBF72DFEFE3EAAC283" ID-CoL="98HWJ" box="[299,401,367,392]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF72DFEFE3EAAC283" box="[299,401,367,392]" italics="true" pageId="6" pageNumber="237">C. littera</emphasis>
</taxonomicName>
figure, but never got beyond this original drawing before his death in 1924, only one year after his brother, René, received Mabille’s collection. The drawing might have been pinned near the specimen, where it remained until René gave this part of the collection to BMNH in 1927. The inked name below the drawing does not seem to match the handwriting of Charles Oberthür, but it could have been written by René, Culot, or someone else at a later date (John V. Calhoun, pers. comm.).
</paragraph>
<paragraph id="8B9636E8FFC7FFDBF6C1FDAA3B6CC776" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="237">
While we cannot completely exclude the possibility that this sole 
<taxonomicName id="4C294D6BFFC7FFDBF5D3FDAB3B04C136" ID-CoL="98HWJ" box="[981,1087,548,573]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF5D3FDAB3B04C136" box="[981,1087,548,573]" italics="true" pageId="6" pageNumber="237">C. littera</emphasis>
</taxonomicName>
specimen in BMNH is not a 
<typeStatus id="5492884AFFC7FFDBF691FDC63FCAC16B" box="[151,241,584,608]" pageId="6" pageNumber="237" type="syntype">syntype</typeStatus>
, the evidence presented above suggests that it is likely to be one. This specimen in the Natural History Museum (BMNH) bearing eight labels: circular, yellow on one side, handprinted: / 890 /, white, covered in glue on the other side; round, white with red round on one side, printed: / 
<typeStatus id="5492884AFFC7FFDBF569FD1E3C9DC1A3" box="[879,934,656,680]" pageId="6" pageNumber="237">Type</typeStatus>
/ inside the red circle; rectangular, asparagus green, handwritten: / 
<collectingCountry id="F33E7678FFC7FFDBF788FD3A3EEAC1C6" box="[398,465,692,717]" name="Brazil" pageId="6" pageNumber="237">Brasil</collectingCountry>
/; rectangular, asparagus green, handwritten: / 
<taxonomicName id="4C294D6BFFC7FFDBF5F7FD3A3B07C1C6" ID-CoL="98HWJ" box="[1009,1084,692,717]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="247" phylum="Arthropoda" rank="species" species="littera">Littera</taxonomicName>
P. Mab. /; rectangular, white, handwritten in red ink: / 
<taxonomicName id="4C294D6BFFC7FFDBF7C1FD593D33C1FB" ID-CoL="98HWJ" box="[455,520,727,752]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">littera</taxonomicName>
/ Mab. /; almost square, white, printed: / Ex musaeo / P. Mabille / 1923 /; rectangular, white, printed: / Ex Oberthür Coll. / Brit. Mus. 1927-3. /; rectangular, printed: / BMNH(E) #1982948 / is hereby designated as the 
<typeStatus id="5492884AFFC7FFDBF7CDFC913D09C033" box="[459,562,799,824]" pageId="6" pageNumber="237" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="4C294D6BFFC7FFDBF451FC913C1BC033" ID-CoL="98HWJ" box="[599,800,799,824]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF451FC913C1BC033" box="[599,800,799,824]" italics="true" pageId="6" pageNumber="237">Telemiades littera</emphasis>
</taxonomicName>
. The following label will be added to the specimen after publication of this study: / 
<typeStatus id="5492884AFFC7FFDBF7CFFCCA3D52C057" box="[457,617,836,860]" pageId="6" pageNumber="237" type="lectotype">LECTOTYPE</typeStatus>
/ 
<taxonomicName id="4C294D6BFFC7FFDBF487FCCA3C71C056" ID-CoL="98HWJ" box="[641,842,836,861]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF487FCCA3C71C056" box="[641,842,836,861]" italics="true" pageId="6" pageNumber="237">Telemiades littera</emphasis>
</taxonomicName>
/ 
<bibRefCitation id="EFB84B19FFC7FFDBF565FCCA3B3FC057" author="Mabille" box="[867,1028,836,861]" pageId="6" pageNumber="237" refString="Mabille, P. (1877) Diagnoses de lepidopteres nouveaux du groupe des hesperides. Petites Nouvelles entomologiques, 2, 165 - 166." type="journal article" year="1877">Mabille, 1877</bibRefCitation>
. Pet. Nouv. Ent. 2: 165 / designated by 
<bibRefCitation id="EFB84B19FFC7FFDBF6BCFCE93E6FC08B" author="Grishin" box="[186,340,871,896]" pageId="6" pageNumber="237" refString="Warren, A. D., Davis, K. J., Stangeland, E. M., Pelham, J. P. &amp; Grishin, N. V. (2014) Illustrated Lists of American Butterflies. Available from: http: // www. butterfliesofamerica. com / (accessed 15 March 2014)" type="journal article" year="2014">Grishin, 2014</bibRefCitation>
/. This specimen is illustrated in 
<figureCitation id="13122A6DFFC7FFDBF4CEFCE93C5CC08B" box="[712,871,871,896]" captionStart="FIGURES 9 – 16" captionStartId="3.[151,264,1731,1753]" captionTargetBox="[154,1433,193,1707]" captionTargetId="figure@3.[152,1435,193,1709]" captionTargetPageId="3" captionText="FIGURES 9 – 16. Type specimens of Clito from the C. littera group. 9 – 10. C. littera lectotype, designated here, &quot; Brasil &quot;, P. Mabille collection 1923, R. Oberthür Coll., Brit. Mus. 1927 - 3, specimen No. BMNH (E) # 982948 (genitalia Fig. 26 a, g); 11 – 12. C. anda [holo] type, French Guiana, Collect. C. Bar, R. Oberthür Coll., Brit. Mus. 1931 - 136, specimen No. BMNH (E) # 982947; (genitalia Fig. 26 m, n, o); 13 – 14. C. anda [para] type, French Guiana: Cayenne, R. Oberthür Coll., Brit. Mus. 1931 - 136, specimen No. BMNH (E) # 1054238; 15 – 16. C. nebulosa holotype, Bolivia, Rio Songo, 750 m, Coll. Fassl (genitalia Fig. 26 d). All specimens are males. Dorsal and ventral surfaces are shown on odd- and even-numbered figures, respectively. Labels are shown in-line with the specimen images. Labels are reduced 2.5 times compared to specimens: small scale bar below C. littera labels refers to labels, and larger scale bars refer to specimens. 15 – 16 is in SMF collection and is photographed by Ernst Brockmann, other specimens are in BMNH collection and images are copyright (©) Trustees of the Natural History Museum, London (used with permission)." httpUri="https://zenodo.org/record/252287/files/figure.png" pageId="6" pageNumber="237">Figs. 9 and 10</figureCitation>
and its genitalia 
<emphasis id="B95DEAFAFFC7FFDBF220FCE73B57C08B" box="[1062,1132,873,896]" italics="true" pageId="6" pageNumber="237">in situ</emphasis>
in 
<figureCitation id="13122A6DFFC7FFDBF289FCE93BD0C08B" box="[1167,1259,871,896]" captionStart="FIGURE 26" captionStartId="12.[151,250,1216,1238]" captionTargetBox="[202,1373,193,1183]" captionTargetId="figure@12.[202,1373,193,1183]" captionTargetPageId="12" captionText="FIGURE 26. Male genitalia of Clito littera group species. a, g. C. littera lectotype, designated here, &quot; Brasil &quot;, P. Mabille collection 1923, R. Oberthür Coll., Brit. Mus. 1927 - 3, specimen No. BMNH (E) # 982948 [BMNH], in situ on a specimen, scales brushed off: a. various views rotating from right posterolateral through posterior to left posterolateral; g. ventrolateral (specimen Figs. 9 – 10); b, c. C. litteroides holotype, Venezuela: Aragua State, Rancho Grande [Biological Station, approximate GPS coordinates 10 ° 21 ' N 67 ° 41 ' W], elevation 1100 m, 4 - Jun- 1985, leg. S. S. Nicolay, genitalia vial NVG 130614 - 42 [USNM]: b. section of complete genitalia in posterolateral view; c. complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 7 – 8); d. C. nebulosa holotype, ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Mielke &amp; Schröder (1994) (specimen Figs. 15 – 16); e, f. C. trinidadensis holotype, Trinidad, Morne Catherine, 24 - Mar- 1982, leg. M. J. W. Cock, specimen No. BMNH (E) # 982949 [BMNH], dorsal and lateral views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Cock (1998) (specimen Figs. 17 – 18); h. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall &amp; M. A. Solis [USNM], complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 5 – 6); i. C. jonkersi holotype, Surinam, Domburg, 8 - Mar- 1975, leg. E. H. Jonkers [RMNH], lateral internal view of left valva and lateral view of uncus and gnathos, illustration from de Jong (1983) (specimen Figs. 19 – 20); j. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall &amp; M. A. Solis [USNM], right valva and uncus in situ on a specimen, scales brushed off (specimen Figs. 3 – 4); k. C. palotschka holotype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, 4 - Mar- 2001, leg. D. H. Ahrenholz [USNM], right valva in situ on a specimen, scales brushed off (specimen Figs. 1 – 2); l. C. anda, Brazil: Rondônia, Cacaulândia municipality, Fazenda Rancho Grande, genitalia No. GTA 4377, illustration from Austin (2000), ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis; m – o. C. anda [holo] type, French Guiana, Collect. C. Bar, R. Oberthür Coll., Brit. Mus. 1931 – 136, specimen No. BMNH (E) # 982947 [BMNH] (specimen Figs. 11 – 12), left valva on m. and Evans's sketches: n. pinned in the collection [BMNH]; o. published, and more stylized, version (Evans 1953), lateral internal view of left valva, lateral and ventral views of uncus, gnathos and distal half of penis. Species name is given below each image, holotypes and paratypes are labeled as HT and PT. Gray &quot; F &quot; indicates mirror image (left-right inverted). Images a, f, g, m – o. are copyright (©) Trustees of the Natural History Museum, London (used with permission)." httpUri="https://zenodo.org/record/252291/files/figure.png" pageId="6" pageNumber="237">Figs. 26</figureCitation>
a, g. It is a male and is recognized by two splits on right hindwing, from margin to discal band, at Rs vein and near tornus, left antenna missing, scales brushed off genitalia. This specimen seems to be the only 
<typeStatus id="5492884AFFC7FFDBF264FC3E3B82C0C3" box="[1122,1209,944,968]" pageId="6" pageNumber="237" type="syntype">syntype</typeStatus>
in collections. The 
<typeStatus id="5492884AFFC7FFDBF691FC5A3FC5C0E6" box="[151,254,980,1005]" pageId="6" pageNumber="237" type="lectotype">lectotype</typeStatus>
is designated to ensure nomenclatural stability and to preserve current usage of the name, which is necessary taking into account that a new species very close to 
<taxonomicName id="4C294D6BFFC7FFDBF55DFC763CFBC71B" ID-CoL="98HWJ" box="[859,960,1015,1040]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF55DFC763CFBC71B" box="[859,960,1015,1040]" italics="true" pageId="6" pageNumber="237">C. littera</emphasis>
</taxonomicName>
is described below, and taxa treated as 
<taxonomicName id="4C294D6BFFC7FFDBF384FC763FE0C73E" ID-CoL="98HWJ" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF384FC763FE0C73E" italics="true" pageId="6" pageNumber="237">C. littera</emphasis>
</taxonomicName>
subspecies are likely to be distinct biological species. The 
<typeStatus id="5492884AFFC7FFDBF597FB933CFAC73E" box="[913,961,1053,1077]" pageId="6" pageNumber="237">type</typeStatus>
locality of 
<taxonomicName id="4C294D6BFFC7FFDBF24DFB933B88C73E" ID-CoL="98HWJ" box="[1099,1203,1052,1077]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF24DFB933B88C73E" box="[1099,1203,1052,1077]" italics="true" pageId="6" pageNumber="237">C. littera</emphasis>
</taxonomicName>
is &quot; 
<collectingCountry id="F33E7678FFC7FFDBF2E2FB923A14C73E" box="[1252,1327,1052,1077]" name="Brazil" pageId="6" pageNumber="237">Brazil</collectingCountry>
&quot;, and the locality specified in the original publication (
<collectingCountry id="F33E7678FFC7FFDBF493FBCE3DF7C753" box="[661,716,1088,1112]" name="Peru" pageId="6" pageNumber="237">Peru</collectingCountry>
) must be erroneous. Since 
<taxonomicName id="4C294D6BFFC7FFDBF206FBCE3B5DC753" ID-CoL="98HWJ" box="[1024,1126,1087,1112]" class="Insecta" family="Hesperiidae" genus="Telemiades" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="littera">
<emphasis id="B95DEAFAFFC7FFDBF206FBCE3B5DC753" box="[1024,1126,1087,1112]" italics="true" pageId="6" pageNumber="237">C. littera</emphasis>
</taxonomicName>
specimens are known only from Rio de Janeiro State, it is likely that the 
<typeStatus id="5492884AFFC7FFDBF492FBEA3DC0C776" box="[660,763,1124,1149]" pageId="6" pageNumber="237" type="lectotype">lectotype</typeStatus>
originated in Southeast 
<collectingCountry id="F33E7678FFC7FFDBF20DFBEA3B68C776" box="[1035,1107,1124,1149]" name="Brazil" pageId="6" pageNumber="237">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>