216 lines
25 KiB
XML
216 lines
25 KiB
XML
<document id="FF137BDCB30EE916C4D92896687BC73A" ID-CLB-Dataset="28391" ID-DOI="10.11646/zootaxa.4565.2.9" ID-GBIF-Dataset="7ca3a729-11e2-4c6c-9f55-9ecd6b13a262" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589961" ID-ZooBank="C9C6BCBD-D262-47F5-953F-D241FEBDA5DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552292382976" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V., Bukejs, Andris & Telnov, Dmitry" docDate="2019" docId="038087AC904AFFDFFF76AB38FC47B086" docLanguage="en" docName="zootaxa.4565.2.9.pdf" docOrigin="Zootaxa 4565 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gonialaena groehni Nabozhenko & Bukejs & Telnov 2019, sp. nov." docType="treatment" docUuid="7437512D-27AB-48AB-B532-402EBA6B81D7" docUuidSource="ZooBank" docVersion="9" lastPageNumber="257" masterDocId="FFB9FFD4904BFFDBFFE1AC07FFCDB443" masterDocTitle="Gonialaenini, a new tribe of Lagriinae (Coleoptera: Tenebrionidae) from Eocene Baltic Amber" masterLastPageNumber="260" masterPageNumber="253" pageNumber="254" updateTime="1698701512721" updateUser="ExternalLinkService">
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<mods:title id="61C77DF81002EABB4D2D9902EB2C52FA">Gonialaenini, a new tribe of Lagriinae (Coleoptera: Tenebrionidae) from Eocene Baltic Amber</mods:title>
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<mods:namePart id="1B567033683E64351FA39624103EF69C">Nabozhenko, Maxim V.</mods:namePart>
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<mods:namePart id="896E0736072320993223366C6EBA1B82">Bukejs, Andris</mods:namePart>
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<mods:namePart id="1760230D7EDC3792963CC0EA0FD542D7">Telnov, Dmitry</mods:namePart>
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<mods:title id="D248C7F413F0D583869CA47C9A69002B">Zootaxa</mods:title>
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<mods:date id="C8D3CD4A1C795C90458895016DB30FFF">2019</mods:date>
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<subSubSection id="C3336531904AFFDAFF76AB38FC46B338" pageId="1" pageNumber="254" type="nomenclature">
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<paragraph id="8B9636BA904AFFDAFF76AB38FE22B319" blockId="1.[151,907,1855,1951]" box="[151,495,1855,1882]" pageId="1" pageNumber="254">
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<heading id="D0DE81D6904AFFDAFF76AB38FE22B319" bold="true" box="[151,495,1855,1882]" fontSize="11" level="1" pageId="1" pageNumber="254" reason="1">
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<emphasis id="B95DEAA8904AFFDAFF76AB38FE22B319" bold="true" box="[151,495,1855,1882]" pageId="1" pageNumber="254">
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<taxonomicName id="4C294D39904AFFDAFF76AB38FE47B31A" authority="Nabozhenko & Bukejs & Telnov, 2019" authorityName="Nabozhenko & Bukejs & Telnov" authorityYear="2019" box="[151,394,1855,1881]" class="Insecta" family="Tenebrionidae" genus="Gonialaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="254" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
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<emphasis id="B95DEAA8904AFFDAFF76AB38FE47B31A" bold="true" box="[151,394,1855,1881]" italics="true" pageId="1" pageNumber="254">Gonialaena groehni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26E57D3904AFFDAFE70AB47FE22B319" box="[401,495,1856,1882]" pageId="1" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9636BA904AFFDAFF76AB65FC46B338" blockId="1.[151,907,1855,1951]" box="[151,907,1890,1915]" pageId="1" pageNumber="254">
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http://zoobank.org/
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336531904AFFDAFF76AB81FEC6B3DC" box="[151,267,1926,1951]" pageId="1" pageNumber="254" type="description">
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<paragraph id="8B9636BA904AFFDAFF76AB81FEC6B3DC" blockId="1.[151,907,1855,1951]" box="[151,267,1926,1951]" pageId="1" pageNumber="254">
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(
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<figureCitation id="13122A3F904AFFDAFF7EAB81FECFB3DC" box="[159,258,1926,1951]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–7" captionStart-2="FIGURES 8–9" captionStartId-0="2.[151,264,1839,1862]" captionStartId-1="3.[151,264,1939,1962]" captionStartId-2="6.[151,264,1874,1897]" captionTargetBox-0="[151,1435,620,1826]" captionTargetBox-1="[150,1436,301,1931]" captionTargetBox-2="[188,1398,280,1838]" captionTargetId-0="figure@2.[151,1435,620,1826]" captionTargetId-1="figure@3.[150,1436,298,1931]" captionTargetId-2="figure@6.[188,1398,280,1838]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionText-0="FIGURES 1–2. Gonialaena groehni gen. et sp. nov., holotype: 1—habitus, dorsal view; 2—habitus, ventral view. Scale bars represent 2 mm." captionText-1="FIGURES 3–7. Gonialaena groehni gen. et sp. nov.: holotype: 3—right antenna, dorsal view; 4—left antenna, ventral view; 5—elytral pubescence in apical one-third; 6—right protibial, dorsal view; 7—apical part of ovipositor. Abbreviations: a, bantennomere 1, c—outer apical spines of tibia, d—lobe VI of coxite, e—gonostyli. Not reproduced to the same scale." captionText-2="FIGURES 8–9. Gonialaena groehni gen. et sp. nov.: holotype: 8—details of forebody, dorsal view; 9—details of abdomen and metathorax. Scale bars represent 1 mm." httpUri-0="https://zenodo.org/record/2589963/files/figure.png" httpUri-1="https://zenodo.org/record/2589965/files/figure.png" httpUri-2="https://zenodo.org/record/2589967/files/figure.png" pageId="1" pageNumber="254">Figs 1–9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336531904AFFD9FF76ABC9FA9BB5CB" lastPageId="2" lastPageNumber="255" pageId="1" pageNumber="254" type="materials_examined">
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<paragraph id="8B9636BA904AFFD9FF76ABC9FAFDB503" blockId="1.[151,1436,1998,2023]" lastBlockId="2.[151,1437,151,573]" lastPageId="2" lastPageNumber="255" pageId="1" pageNumber="254">
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<emphasis id="B95DEAA8904AFFDAFF76ABC9FE85B3A4" bold="true" box="[151,328,1998,2023]" pageId="1" pageNumber="254">Type material.</emphasis>
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<typeStatus id="54928818904AFFDAFEB3ABC9FE77B3A4" box="[338,442,1998,2023]" pageId="1" pageNumber="254" type="holotype">Holotype</typeStatus>
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adult
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<specimenCount id="9D2FFD33904AFFDAFDE6ABC8FDD3B3A5" box="[519,542,1999,2022]" pageId="1" pageNumber="254" type="female">♀</specimenCount>
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: “GPIH 4959, coll. Gröhn 8589”, deposited at the Center of Natural History (CeNak) (formerly Geological-Paleontological Institute and Museum, GPIH) of the University of
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<collectingRegion id="49EDF8589049FFD9FACDAC90FA55B4F3" box="[1324,1432,151,176]" country="Germany" name="Hamburg" pageId="2" pageNumber="255">Hamburg</collectingRegion>
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,
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<collectingCountry id="F33E762A9049FFD9FF76ACBBFECCB497" box="[151,257,188,212]" name="Germany" pageId="2" pageNumber="255">Germany</collectingCountry>
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. Rather complete beetle inclusion (apical tarsomeres of left mesotarsus missing) with partially exposed genitalia, included in transparent yellow piece of amber with dimensions 55×33×
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<quantity id="4CD19B5F9049FFD9FBC9ACD8FBB4B4B4" box="[1064,1145,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="255" unit="mm" value="13.0">13 mm</quantity>
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. Syninclusions: one small indet.
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<taxonomicName id="4C294D399049FFD9FF00AD03FDDBB55F" authority="(Hymenoptera)" baseAuthorityName="Hymenoptera" box="[225,534,260,285]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="family">Formicidae (Hymenoptera)</taxonomicName>
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specimen, one indet. Nematocera (Diptera) specimen, one strongly damaged specimen of indet. Insecta, a few stellate
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<taxonomicName id="4C294D399049FFD9FD85AD2FFD01B503" authorityName="" authorityYear="1829" box="[612,716,296,320]" class="Magnoliopsida" family="Fagaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="255" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
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trichomes, and a few small pieces of organic material.
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</paragraph>
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<paragraph id="8B9636BA9049FFD9FF26AD4BFA41B527" blockId="2.[151,1437,151,573]" box="[199,1420,332,357]" pageId="2" pageNumber="255">
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<emphasis id="B95DEAA89049FFD9FF26AD4BFE98B526" bold="true" box="[199,341,332,357]" pageId="2" pageNumber="255">
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<typeStatus id="549288189049FFD9FF26AD4BFECFB526" box="[199,258,332,357]" pageId="2" pageNumber="255">Type</typeStatus>
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strata.
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</emphasis>
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Baltic amber, mid-Eocene to Upper Eocene, Prussian Formation, Priabonian, ca. 37.2–33.9 Ma.
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</paragraph>
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<paragraph id="8B9636BA9049FFD9FF26AD68FA9BB5CB" blockId="2.[151,1437,151,573]" box="[199,1366,367,392]" pageId="2" pageNumber="255">
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<materialsCitation id="3B413CE79049FFD9FF26AD68FA9FB5CB" ID-GBIF-Occurrence="2012823424" box="[199,1362,367,392]" country="Russia" location="Baltic Sea" municipality="Yantarny" pageId="2" pageNumber="255" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
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<emphasis id="B95DEAA89049FFD9FF26AD68FEA8B5CB" bold="true" box="[199,357,367,392]" pageId="2" pageNumber="255">
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<typeStatus id="549288189049FFD9FF26AD68FECFB5CB" box="[199,258,367,392]" pageId="2" pageNumber="255">Type</typeStatus>
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locality.
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</emphasis>
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<location id="8EF660619049FFD9FE8DAD68FE12B5CB" LSID="urn:lsid:plazi:treatment:038087AC904AFFDFFF76AB38FC47B086:8EF660619049FFD9FE8DAD68FE12B5CB" box="[364,479,367,392]" country="Russia" municipality="Yantarny" name="Baltic Sea" pageId="2" pageNumber="255" stateProvince="Kaliningrad">Baltic Sea</location>
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coast,
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<collectingMunicipality id="6BF2ACC09049FFD9FDCCAD77FD5FB5CB" box="[557,658,368,392]" pageId="2" pageNumber="255">Yantarny</collectingMunicipality>
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settlement (formerly Palmnicken),
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<collectingRegion id="49EDF8589049FFD9FBC0AD68FB6AB5CB" box="[1057,1191,367,392]" country="Russia" name="Kaliningrad" pageId="2" pageNumber="255">Kaliningrad</collectingRegion>
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region,
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<collectingCountry id="F33E762A9049FFD9FAE2AD68FA9FB5CB" box="[1283,1362,367,392]" name="Russia" pageId="2" pageNumber="255">Russia</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33365319049FFD9FF26AD93FE9CB593" pageId="2" pageNumber="255" type="etymology">
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<paragraph id="8B9636BA9049FFD9FF26AD93FE9CB593" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="255">
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<emphasis id="B95DEAA89049FFD9FF26AD93FE83B5EE" bold="true" box="[199,334,404,429]" pageId="2" pageNumber="255">Etymology.</emphasis>
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The species is named in honour of Carsten Gröhn (Glinde,
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<collectingCountry id="F33E762A9049FFD9FC0CAD93FB9AB5EF" box="[1005,1111,404,428]" name="Germany" pageId="2" pageNumber="255">Germany</collectingCountry>
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) who provided the specimen for examination.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33365319049FFD9FF26ADDBFBAEB5B6" box="[199,1123,476,501]" pageId="2" pageNumber="255" type="diagnosis">
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<paragraph id="8B9636BA9049FFD9FF26ADDBFBAEB5B6" blockId="2.[151,1437,151,573]" box="[199,1123,476,501]" pageId="2" pageNumber="255">
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<emphasis id="B95DEAA89049FFD9FF26ADDBFE03B5B6" bold="true" box="[199,462,476,501]" pageId="2" pageNumber="255">Differential diagnosis.</emphasis>
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See comparison of the genus
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<taxonomicName id="4C294D399049FFD9FCFCADDBFC6CB5B7" authority="Nabozhenko & Bukejs & Telnov, 2019" authorityName="Nabozhenko & Bukejs & Telnov" authorityYear="2019" box="[797,929,476,500]" class="Insecta" family="Tenebrionidae" genus="Gonialaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B95DEAA89049FFD9FCFCADDBFC6CB5B7" box="[797,929,476,500]" italics="true" pageId="2" pageNumber="255">Gonialaena</emphasis>
|
||
</taxonomicName>
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||
<emphasis id="B95DEAA89049FFD9FC49ADDBFBDCB5B6" bold="true" box="[936,1041,476,501]" pageId="2" pageNumber="255">
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||
<taxonomicNameLabel id="A26E57D39049FFD9FC49ADDBFBDCB5B6" box="[936,1041,476,501]" pageId="2" pageNumber="255" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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above.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33365319049FFDFFF26ADF8FC47B086" lastPageId="4" lastPageNumber="257" pageId="2" pageNumber="255" type="description">
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||
<paragraph id="8B9636BA9049FFD9FF26ADF8FF2BB67E" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="255">
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<emphasis id="B95DEAA89049FFD9FF26ADF8FE94B65B" bold="true" box="[199,345,511,536]" pageId="2" pageNumber="255">Description.</emphasis>
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Total body length about
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<quantity id="4CD19B5F9049FFD9FD64AE07FD2FB654" box="[645,738,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="255" unit="mm" value="9.1">9.1 mm</quantity>
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; body elongated, slightly convex dorsally; unicolorous dark brown.
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</paragraph>
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<caption id="DF5666329049FFD9FF76AB28FEF6B326" httpUri="https://zenodo.org/record/2589963/files/figure.png" pageId="2" pageNumber="255" startId="2.[151,264,1839,1862]" targetBox="[151,1435,620,1826]" targetPageId="2">
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<paragraph id="8B9636BA9049FFD9FF76AB28FEF6B326" blockId="2.[151,1435,1839,1893]" pageId="2" pageNumber="255">
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<emphasis id="B95DEAA89049FFD9FF76AB28FEF6B306" bold="true" box="[151,315,1839,1862]" pageId="2" pageNumber="255">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="4C294D399049FFD9FEA3AB37FDC2B305" authority="Nabozhenko & Bukejs & Telnov, 2019" authorityName="Nabozhenko & Bukejs & Telnov" authorityYear="2019" box="[322,527,1840,1862]" class="Insecta" family="Tenebrionidae" genus="Gonialaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="species" species="groehni" status="gen. et sp. nov.">
|
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<emphasis id="B95DEAA89049FFD9FEA3AB37FDC2B305" box="[322,527,1840,1862]" italics="true" pageId="2" pageNumber="255">Gonialaena groehni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26E57D39049FFD9FDF7AB37FD78B306" box="[534,693,1840,1862]" pageId="2" pageNumber="255" rank="species">
|
||
<emphasis id="B95DEAA89049FFD9FDF7AB37FD8EB306" bold="true" box="[534,579,1840,1861]" pageId="2" pageNumber="255">gen.</emphasis>
|
||
et
|
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<emphasis id="B95DEAA89049FFD9FD85AB37FD78B306" bold="true" box="[612,693,1840,1861]" pageId="2" pageNumber="255">sp. nov.</emphasis>
|
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</taxonomicNameLabel>
|
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, holotype: 1—habitus, dorsal view; 2—habitus, ventral view. Scale bars represent 2 mm.
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</paragraph>
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</caption>
|
||
<paragraph id="8B9636BA9049FFD8FF26AB95FE72B4BB" blockId="2.[151,1436,1937,2034]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="256" pageId="2" pageNumber="255">
|
||
<emphasis id="B95DEAA89049FFD9FF26AB95FEC9B3E9" box="[199,260,1938,1962]" italics="true" pageId="2" pageNumber="255">Head</emphasis>
|
||
prognathous; apical maxillary palpomere securiform; eyes large, transverse, oval, slightly prominent; antennae of 11 antennomeres, filiform, moderately long, reaching basal quarter of elytra, evenly covered with sparse, erect to semierect simple trichoid setae; scape markedly wider than pedicel; pedicel short, about 0.6× as long as antennomere 3, antennomeres 3–10 elongate, subequal in length, antennomeres 8–10 slightly dilated apically, antennomere 11 spindle-shaped, 1.7× as long as wide; relative length ratios of antennomeres 1– 11 equal to?-5-9-9-9-9-9-
|
||
<date id="FF97107A9048FFD8FEB4ACD8FE77B4BB" box="[341,442,223,248]" pageId="3" pageNumber="256" value="2010-09-09" valueMax="2010-09-09" valueMin="2010-09-09">9-9-9-10</date>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF5666329048FFD8FF76AB94FAF6B3A4" httpUri="https://zenodo.org/record/2589965/files/figure.png" pageId="3" pageNumber="256" startId="3.[151,264,1939,1962]" targetBox="[150,1436,301,1931]" targetPageId="3">
|
||
<paragraph id="8B9636BA9048FFD8FF76AB94FAF6B3A4" blockId="3.[151,1436,1939,2023]" pageId="3" pageNumber="256">
|
||
<emphasis id="B95DEAA89048FFD8FF76AB94FEF9B3EA" bold="true" box="[151,308,1939,1962]" pageId="3" pageNumber="256">FIGURES 3–7</emphasis>
|
||
.
|
||
<taxonomicName id="4C294D399048FFD8FEA2AB93FDC2B3E9" authority="Nabozhenko & Bukejs & Telnov, 2019" authorityName="Nabozhenko & Bukejs & Telnov" authorityYear="2019" box="[323,527,1940,1962]" class="Insecta" family="Tenebrionidae" genus="Gonialaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="groehni" status="gen. et sp. nov.">
|
||
<emphasis id="B95DEAA89048FFD8FEA2AB93FDC2B3E9" box="[323,527,1940,1962]" italics="true" pageId="3" pageNumber="256">Gonialaena groehni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26E57D39048FFD8FDF6AB93FD78B3EA" box="[535,693,1940,1962]" pageId="3" pageNumber="256" rank="species">
|
||
<emphasis id="B95DEAA89048FFD8FDF6AB93FD8EB3EA" bold="true" box="[535,579,1940,1961]" pageId="3" pageNumber="256">gen.</emphasis>
|
||
et
|
||
<emphasis id="B95DEAA89048FFD8FD85AB93FD78B3EA" bold="true" box="[612,693,1940,1961]" pageId="3" pageNumber="256">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: holotype: 3—right antenna, dorsal view; 4—left antenna, ventral view; 5—elytral pubescence in apical one-third; 6—right protibial, dorsal view; 7—apical part of ovipositor. Abbreviations: a, bantennomere 1, c—outer apical spines of tibia, d—lobe VI of coxite, e—gonostyli. Not reproduced to the same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9636BA904FFFDFFF26AC9FFBEBB5EE" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="257">
|
||
<emphasis id="B95DEAA8904FFFDFFF26AC9FFE8CB4F3" box="[199,321,152,176]" italics="true" pageId="4" pageNumber="257">Prothorax.</emphasis>
|
||
Pronotum nearly circular, 1.1× as wide as long, widest medially. Lateral margins widely rounded, emarginate at base; anterior margin rounded; posterior margin nearly straight; anterior and lateral margins not beaded, posterior margin apparently narrowly beaded; anterior angles obsolete, vaguely expressed, posterior angles distinct, orthogonal; disc flattened, without impressions; pronotal punctation minute and dense, punctures sparser on disc than laterally; disc sparsely covered with dark erect setae of various lengths (longer setae are sparser); prothoracic hypomera finely sparsely punctate and covered with short erect setae; prosternum covered with suberect setae, before procoxal cavities much longer than one diameter of a procoxal cavity; prosternal process slightly convex, not protruding, narrow, about 0.3× as wide as procoxal diameter.
|
||
</paragraph>
|
||
<paragraph id="8B9636BA904FFFDFFF26ADBFFDABB6EB" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="257">
|
||
<emphasis id="B95DEAA8904FFFDFFF26ADBFFE98B593" box="[199,341,440,464]" italics="true" pageId="4" pageNumber="257">Pterothorax.</emphasis>
|
||
Elytra elongate-oval, weakly convex dorsally, with apex prolonged, nearly mucronate; humeri rounded; each elytron with 10 indistinct striae of round coarse punctures; interstriae slightly convex, with fine punctation; sparsely covered with two
|
||
<typeStatus id="54928818904FFFDFFD83AE07FD50B65B" box="[610,669,512,536]" pageId="4" pageNumber="257">types</typeStatus>
|
||
of setae: longer erect and shorter suberect ones. Epipleura wide anteriorly, gradually narrowing posteriorly, reaching elytral apex, finely densely punctate. Prohypomera, meso- and metaventrite with delicate, moderately sparse punctation; mes- and metepisternum with small and distinctly denser punctures; covered with short, fine semierect setae. Metaventrite with shallow median impression in basal half and with longitudinal sulcus posteromedially.
|
||
</paragraph>
|
||
<paragraph id="8B9636BA904FFFDFFF26AEB3FE58B71E" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="257">
|
||
<emphasis id="B95DEAA8904FFFDFFF26AEB3FEFCB68F" box="[199,305,692,716]" italics="true" pageId="4" pageNumber="257">Abdomen</emphasis>
|
||
finely and sparsely punctured (ventrites 4 and 5 with denser punctation), covered with short, suberect setae; narrow intersegmental membranes present between 3–5 abdominal ventrites; ventrite 5 with apical margin rounded; relative length ratios of ventrites 1–5 equal to
|
||
<date id="FF97107A904FFFDFFCCFAEFBFC61B757" box="[814,940,764,788]" pageId="4" pageNumber="257" value="2011-08-20" valueMax="2011-08-14" valueMin="2011-08-20">20-14-11-8</date>
|
||
-13 (medially). Ovipositor of lagrioid
|
||
<typeStatus id="54928818904FFFDFFA8DAEFAFA51B756" box="[1388,1436,765,789]" pageId="4" pageNumber="257">type</typeStatus>
|
||
(
|
||
<bibRefCitation id="EFB84B4B904FFFDFFF7FAF18FE04B77B" author="Tschinkel, W. R. & Doyen, J. T." box="[158,457,799,824]" pageId="4" pageNumber="257" pagination="321 - 368" refId="ref4436" refString="Tschinkel, W. R. & Doyen, J. T. (1980) Comparative anatomy of the defensive glands, ovipositors and female tubes of tenebrionid beetles (Coleoptera). International journal of insect morphology and embryology, 9, 321 - 368. https: // doi. org / 10.1016 / 0020 - 7322 (80) 90009 - 4" type="journal article" year="1980">Tschinkel & Doyen 1980</bibRefCitation>
|
||
): with long cylindrical finger-like lobe 4 of the coxite and terminal position of long cylindrical gonostyles.
|
||
</paragraph>
|
||
<paragraph id="8B9636BA904FFFDFFF26AF6FFC47B086" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="257">
|
||
Legs long, slender, covered with sparse setae of two
|
||
<typeStatus id="54928818904FFFDFFCC6AF6FFCAFB7C3" box="[807,866,872,896]" pageId="4" pageNumber="257">types</typeStatus>
|
||
and delicate punctation. Procoxal cavities closed. Mesotrochantine exposed. Procoxae globose; mesocoxae oval, longitudinal; metacoxae narrowly oval, transverse; all coxae distinctly not contiguous. Femora unarmed, without teeth, profemora slightly clavate, meso- and metafemora nearly parallel-sided, metafomora distinctly longer than mesofemora. Tibiae almost straight, nearly parallel-sided, narrow, slightly shorter than femora, with two short spines apically, covered with two
|
||
<typeStatus id="54928818904FFFDFFB1CAFFFFAF5B053" box="[1277,1336,1016,1040]" pageId="4" pageNumber="257">types</typeStatus>
|
||
of setae: oblique short dense erect ones and 6-8 very long erect setae on outer margin. Tarsi long, but shorter than tibiae, with dense setation on plantar surface; penultimate tarsomere not bilobed or widened, insertion of terminal tarsomere is central according to the face of slightly cupuliform penultimate tarsomere; metatarsus about 1.4× as long as protarsus; relative length ratios of protarsomeres 1–5 equal to 8-
|
||
<date id="FF97107A904FFFDFFBEBA88FFBA2B0E3" box="[1034,1135,1160,1184]" pageId="4" pageNumber="257" value="2011-04-04" valueMax="2011-04-04" valueMin="2011-04-04">4-4-4-11</date>
|
||
; relative length ratios of metatarsomeres 1–4 equal to
|
||
<date id="FF97107A904FFFDFFE3EA8ABFDBCB086" box="[479,625,1196,1221]" pageId="4" pageNumber="257" value="1920-11-10" valueMax="1920-11-15" valueMin="1920-11-10">20-11-10-15.</date>
|
||
Claws thin, free, simple.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |