519 lines
97 KiB
XML
519 lines
97 KiB
XML
<document id="B8C986E93EC52C657056AABBA48D9FED" ID-DOI="10.5281/zenodo.12996779" ID-Zenodo-Dep="12996779" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="luisschmitz" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1721870935690" checkinUser="luisschmitz" docAuthor="Gerry Cassis & Loren Vanags" docDate="2006" docId="0380878FFFA0FFBEFC2CFCDBC0E1FB84" docLanguage="en" docName="Denisia.19.275-398.pdf.imf" docOrigin="Denisia 19" docTitle="Coleotichus WHITE 1839" docType="treatment" docVersion="11" lastPageNumber="310" masterDocId="FFB9FFF7FFBFFF9DFFC4FF8CC239FFED" masterDocTitle="Jewel Bugs of Australia (Insecta, Heteroptera, Scutelleridae)" masterLastPageNumber="398" masterPageNumber="275" pageNumber="306" updateTime="1725646846613" updateUser="ExternalLinkService">
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<mods:mods id="ACCFC9E3E2D1DFD4BB6316CB3B418E12" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="99FBA9FBCB7A2C8170A2F613A0FBEE9E">
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<mods:title id="E8851787C0C339CBF862A146853D2639">Jewel Bugs of Australia (Insecta, Heteroptera, Scutelleridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="1D5C34B2E72B3C1D81FCCD99C640FD18" type="personal">
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<mods:roleTerm id="EE12EE6C89EF8DEA17C0674D9AE9835F">Author</mods:roleTerm>
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<mods:namePart id="D6BF2786780890CD56ED2588B6B07E58">Gerry Cassis</mods:namePart>
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<mods:name id="24CEF70683420F1416B2550F01F88F55" type="personal">
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<mods:roleTerm id="1D3815B4FD860CA8A0B079EFEF9B5560">Author</mods:roleTerm>
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<mods:namePart id="5CEFC7543FC9472778BD06DDFED536A3">Loren Vanags</mods:namePart>
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<mods:typeOfResource id="B0F6A6A119BC45ED7C523791F664C94F">text</mods:typeOfResource>
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<mods:titleInfo id="895A84773C9190BDD96B00ACD6077472">
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<mods:title id="AA7C3249AFDD3709F0440E4F2221A1CD">Denisia</mods:title>
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</mods:titleInfo>
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<mods:part id="78B58AA6E5A274C57363BD91749A3DE4">
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<mods:date id="0CB4E34168711B4C931A8D6A1C0B20FA">2006</mods:date>
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<mods:detail id="0C4D8D912A0762C1528AAFD3EB5E1A06" type="volume">
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<mods:number id="CD0041D4F57D6E77C4DF25C89350F10F">19</mods:number>
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<mods:start id="BC8012ED00089BD96C63BBA46FB78455">275</mods:start>
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<mods:end id="BC52E96F22CFA3C7A9E2704B238475AC">398</mods:end>
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<mods:classification id="C30AB61118D2FA79AFBB5053400F47EF">journal article</mods:classification>
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<mods:identifier id="68B1F6B3DD8081E9030EB7AC8F5ED4A8" type="DOI">10.5281/zenodo.12996779</mods:identifier>
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<mods:identifier id="B546124B4F2725875D8CA1101A2491EC" type="Zenodo-Dep">12996779</mods:identifier>
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<subSubSection id="C3336512FFA0FF82FC2CFCDBC694FC62" pageId="31" pageNumber="306" type="nomenclature">
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<paragraph id="8B963699FFA0FF82FC2CFCDBC694FC62" blockId="31.[1000,1435,110,1917]" pageId="31" pageNumber="306">
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<taxonomicName id="4C294D1AFFA0FF82FC2CFCDBC6D6FC82" authority="WHITE 1839" authorityName="WHITE" authorityYear="1839" box="[1000,1263,855,879]" class="Insecta" family="Scutelleridae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="306" phylum="Arthropoda" rank="genus">
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<emphasis id="B95DEA8BFFA0FF82FC2CFCDBC652FC82" box="[1000,1131,855,879]" italics="true" pageId="31" pageNumber="306">Coleotichus</emphasis>
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<bibRefCitation id="EFB84B68FFA0FF82FBB7FCDBC6D6FC82" author="WHITE A." box="[1139,1263,855,879]" journalOrPublisher="Mag. Nat. Hist." pageId="31" pageNumber="306" pagination="537 - 543" part="2" refId="ref89958" refString="WHITE A. (1839): Descriptions of two Hemipterous insects. - Mag. Nat. Hist. (2) 3: 537 - 543." title="Descriptions of two Hemipterous insects" type="journal article" year="1839">
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<smallCapsWord id="8D70A045FFA0FF82FBB7FCDBC688FC83" baselines="873,873" box="[1139,1201,855,879]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="White" pageId="31" pageNumber="306">WHITE</smallCapsWord>
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1839
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</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="13122A1CFFA0FF82FC2AFCFBC609FC62" box="[1006,1072,887,911]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="31" pageNumber="306">Figs 3</figureCitation>
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c-e, 6, 7, 8)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336512FFA0FF82FC2CFC2CC6D2FB04" pageId="31" pageNumber="306" type="reference_group">
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<paragraph id="8B963699FFA0FF82FC2CFC2CC6D2FB04" blockId="31.[1000,1435,110,1917]" pageId="31" pageNumber="306">
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<taxonomicName id="4C294D1AFFA0FF82FC2CFC2CC6DCFC58" authority="WHITE 1839" authorityName="WHITE" authorityYear="1839" box="[1000,1253,928,949]" class="Insecta" family="Scutelleridae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="306" phylum="Arthropoda" rank="genus">
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<emphasis id="B95DEA8BFFA0FF82FC2CFC2CC671FC58" box="[1000,1096,928,949]" italics="true" pageId="31" pageNumber="306">Coleotichus</emphasis>
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<bibRefCitation id="EFB84B68FFA0FF82FB89FC2CC6DCFC58" author="WHITE A." box="[1101,1253,928,949]" journalOrPublisher="Mag. Nat. Hist." pageId="31" pageNumber="306" pagination="537 - 543" part="2" refId="ref89958" refString="WHITE A. (1839): Descriptions of two Hemipterous insects. - Mag. Nat. Hist. (2) 3: 537 - 543." title="Descriptions of two Hemipterous insects" type="journal article" year="1839">
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<smallCapsWord id="8D70A045FFA0FF82FB89FC2CC6B0FC5E" baselines="944,943" box="[1101,1161,928,949]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="White" pageId="31" pageNumber="306">WHITE</smallCapsWord>
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1839: 541
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</bibRefCitation>
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</taxonomicName>
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(nov. gen.);
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<bibRefCitation id="EFB84B68FFA0FF82FA91FC2CC610FC39" author="DALLAS W. S." journalOrPublisher="British Museum, London" pageId="31" pageNumber="306" pagination="1 - 368" refId="ref83529" refString="DALLAS W. S. (1851): List of specimens of Hemipterous insects in the collection of the British Museum. - British Museum, London: 1 - 368, pls I- XI." title="List of specimens of Hemipterous insects in the collection of the British Museum" type="book" year="1851">
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<smallCapsWord id="8D70A045FFA0FF82FA91FC2CC7A0FC5E" baselines="944,943" box="[1365,1433,928,949]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Dallas" pageId="31" pageNumber="306">DALLAS</smallCapsWord>
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1851: 5
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</bibRefCitation>
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(description);
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<bibRefCitation id="EFB84B68FFA0FF82FB6FFC33C710FC39" author="STAL C." box="[1195,1321,959,980]" journalOrPublisher="Noestedtiana, Holmiae" pageId="31" pageNumber="306" pagination="1 - 256" refId="ref88808" refString="STAL C. (1865): Hemiptera Africana. Vol. 1. - Noestedtiana, Holmiae: iv + 1 - 256." title="Hemiptera Africana. Vol. 1." type="book chapter" year="1865">
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<smallCapsWord id="8D70A045FFA0FF82FB6FFC33C6E1FC3E" baselines="975,975" box="[1195,1240,959,980]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Stal" pageId="31" pageNumber="306">STÅL</smallCapsWord>
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1865: 35
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</bibRefCitation>
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(key);
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<bibRefCitation id="EFB84B68FFA0FF82FAA3FC33C601FC1E" author="MAYR G." journalOrPublisher="Karl Gerold's Sohn, Wien" pageId="31" pageNumber="306" pagination="1 - 204" refId="ref87428" refString="MAYR G. (1866): Hemiptera. - In: Reise der Osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf- Urbair. Zoologischer Theil. Zweiter Band. I. Abtheilung. B. - Karl Gerold's Sohn, Wien: 1 - 204, pls I-V." title="Hemiptera" type="book chapter" volumeTitle="Reise der Osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf- Urbair. Zoologischer Theil. Zweiter Band. I. Abtheilung. B." year="1866">
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<smallCapsWord id="8D70A045FFA0FF82FAA3FC33C7A3FC3F" baselines="975,974" box="[1383,1434,959,980]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Mayr" pageId="31" pageNumber="306">MAYR</smallCapsWord>
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1866: 13
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</bibRefCitation>
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(diagnosis);
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<bibRefCitation id="EFB84B68FFA0FF82FB74FC52C712FC1E" author="STAL C." box="[1200,1323,990,1011]" journalOrPublisher="Kongl. Svenska Vetensk. - Akad. Handl." pageId="31" pageNumber="306" pagination="1 - 163" part="11" refId="ref89047" refString="STAL C. (1873): Enumeratio Hemipterorum. Bidrag till en forteckning ofver aller hittills kanda Hemiptera, jemte systrmatiska meddelanden. 3. - Kongl. Svenska Vetensk. - Akad. Handl. 11 (2): 1 - 163." title="Enumeratio Hemipterorum. Bidrag till en forteckning ofver aller hittills kanda Hemiptera, jemte systrmatiska meddelanden. 3." type="journal article" year="1873">
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<smallCapsWord id="8D70A045FFA0FF82FB74FC52C6E4FC1F" baselines="1006,1006" box="[1200,1245,990,1011]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Stal" pageId="31" pageNumber="306">STÅL</smallCapsWord>
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1873: 4
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</bibRefCitation>
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(diagnosis);
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<bibRefCitation id="EFB84B68FFA0FF82FC2CFC70C718FBFC" author="LETHIERRY L. & G. SEVERIN" box="[1000,1313,1020,1041]" journalOrPublisher="F Hayez, Bruxelles" pageId="31" pageNumber="306" pagination="1 - 286" refId="ref86627" refString="LETHIERRY L. & G. SEVERIN (1893): Catalogue General des Hemipteres. Tome I. Heteropteres Pentatomidae. - F Hayez, Bruxelles: xi + 1 - 286." title="Catalogue General des Hemipteres. Tome I. Heteropteres Pentatomidae" type="book chapter" year="1893">
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<smallCapsWord id="8D70A045FFA0FF82FC2CFC70C67AFBFD" baselines="1036,1036" box="[1000,1091,1020,1041]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Lethierry" pageId="31" pageNumber="306">LETHIERRY</smallCapsWord>
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&
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<smallCapsWord id="8D70A045FFA0FF82FBB0FC70C682FBFD" baselines="1036,1036" box="[1140,1211,1020,1041]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Severin" pageId="31" pageNumber="306">SEVERIN</smallCapsWord>
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1893: 15
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</bibRefCitation>
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(catalogue);
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<bibRefCitation id="EFB84B68FFA0FF82FC2CFB97C69EFBDD" author="SCHOUTEDEN H." bookContentInfo="5 pls." box="[1000,1191,1051,1072]" journalOrPublisher="Genera Insectorum" pageId="31" pageNumber="306" pagination="1 - 98" part="24" refId="ref87964" refString="SCHOUTEDEN H. (1904): Heteroptera. Fam. Pentatomidae. Subfam. Scutellerinae. - Genera Insectorum 24: 1 - 98, 5 pls." title="Heteroptera. Fam. Pentatomidae. Subfam. Scutellerinae" type="book chapter" year="1904">
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<smallCapsWord id="8D70A045FFA0FF82FC2CFB97C665FBC3" baselines="1067,1066" box="[1000,1116,1051,1072]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Schouteden" pageId="31" pageNumber="306">SCHOUTEDEN</smallCapsWord>
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1904: 4
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</bibRefCitation>
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, 5 (description);
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<bibRefCitation id="EFB84B68FFA0FF82FA80FB97C67AFBA2" author="KIRKALDY G. W." editor="SHARP D." journalOrPublisher="University Press, Cambridge" pageId="31" pageNumber="306" pagination="93 - 174" refId="ref85896" refString="KIRKALDY G. W. (1902): Hemiptera. - In: SHARP D. (Ed.), Fauna Hawaiiensis Volume III Part II. - University Press, Cambridge: 93 - 174, pls IV-V." title="Hemiptera" type="book chapter" volumeTitle="Fauna Hawaiiensis Volume III Part II." year="1902">
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<smallCapsWord id="8D70A045FFA0FF82FA80FB97C7A3FBC3" baselines="1067,1066" box="[1348,1434,1051,1072]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Kirkaldy" pageId="31" pageNumber="306">KIRKALDY</smallCapsWord>
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1902: 172
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</bibRefCitation>
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(Hawaii);
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<bibRefCitation id="EFB84B68FFA0FF82FB6CFBB6C776FBA2" author="SCHOUTEDEN H." box="[1192,1359,1082,1103]" journalOrPublisher="Ann. Mus. Nat. Hung." pageId="31" pageNumber="306" pagination="317 - 361" part="3" refId="ref87993" refString="SCHOUTEDEN H. (1905): Monographie du genre Coleotichus. - Ann. Mus. Nat. Hung. 3: 317 - 361." title="Monographie du genre Coleotichus" type="journal article" year="1905">
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<smallCapsWord id="8D70A045FFA0FF82FB6CFBB6C722FBA0" baselines="1098,1097" box="[1192,1307,1082,1103]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Schouteden" pageId="31" pageNumber="306">SCHOUTEDEN</smallCapsWord>
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1905
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</bibRefCitation>
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(monograph);
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<bibRefCitation id="EFB84B68FFA0FF82FBF4FBD5C6DFFB83" author="KIRKALDY G. W." box="[1072,1254,1113,1134]" journalOrPublisher="Felix Dames, Berlin" pageId="31" pageNumber="306" pagination="1 - 392" refId="ref86002" refString="KIRKALDY G. W. (1909): Catalogue of the Hemiptera (Heteroptera) Vol. 1. Cimicidae. - Felix Dames, Berlin: xl, 1 - 392." title="Catalogue of the Hemiptera (Heteroptera) Vol. 1. Cimicidae" type="book chapter" year="1909">
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<smallCapsWord id="8D70A045FFA0FF82FBF4FBD5C6BCFB81" baselines="1129,1128" box="[1072,1157,1113,1134]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Kirkaldy" pageId="31" pageNumber="306">KIRKALDY</smallCapsWord>
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1909: 313
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</bibRefCitation>
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(catalogue);
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<bibRefCitation id="EFB84B68FFA0FF82FA9BFBD5C60CFB60" author="GROSS G. F." journalOrPublisher="A. B. James, Adelaide" pageId="31" pageNumber="306" pagination="1 - 250" refId="ref84925" refString="GROSS G. F. (1975): Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1. A. B. James, Adelaide: 1 - 250, 3 colour pls." title="Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1" type="book chapter" year="1975">
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<smallCapsWord id="8D70A045FFA0FF82FA9BFBD5C7A3FB81" baselines="1129,1128" box="[1375,1434,1113,1134]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gross" pageId="31" pageNumber="306">GROSS</smallCapsWord>
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1975: 80
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</bibRefCitation>
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(description);
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<bibRefCitation id="EFB84B68FFA0FF82FB7CFBF4C758FB60" author="DISTANT W. L." box="[1208,1377,1144,1165]" journalOrPublisher="Ann. Mag. Nat. Hist." pageId="31" pageNumber="306" pagination="143 - 164" part="9" refId="ref83802" refString="DISTANT W. L. (1920): Rhynchota from New Caledonia. - Ann. Mag. Nat. Hist. (9) 6: 143 - 164." title="Rhynchota from New Caledonia" type="journal article" year="1920">
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<smallCapsWord id="8D70A045FFA0FF82FB7CFBF4C73DFB66" baselines="1160,1159" box="[1208,1284,1144,1165]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Distant" pageId="31" pageNumber="306">DISTANT</smallCapsWord>
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1920: 143
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</bibRefCitation>
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(catalogue);
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<bibRefCitation id="EFB84B68FFA0FF82FBEAFB1AC6F8FB46" author="GROSS G. F." box="[1070,1217,1174,1195]" journalOrPublisher="A. B. James, Adelaide" pageId="31" pageNumber="306" pagination="1 - 250" refId="ref84925" refString="GROSS G. F. (1975): Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1. A. B. James, Adelaide: 1 - 250, 3 colour pls." title="Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1" type="book chapter" year="1975">
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<smallCapsWord id="8D70A045FFA0FF82FBEAFB1AC650FB44" baselines="1190,1189" box="[1070,1129,1174,1195]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gross" pageId="31" pageNumber="306">GROSS</smallCapsWord>
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1975: 80
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</bibRefCitation>
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(description);
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<bibRefCitation id="EFB84B68FFA0FF82FA8EFB1AC6F7FB27" author="MCDONALD F. J. D. & G. CASSIS" journalOrPublisher="Aust. J. Zool." pageId="31" pageNumber="306" pagination="537 - 572" part="32" refId="ref87212" refString="MCDONALD F. J. D. & G. CASSIS (1984): Revision of the Australian Scutelleridae LEACH (Hemiptera). - Aust. J. Zool. 32: 537 - 572." title="Revision of the Australian Scutelleridae LEACH (Hemiptera)" type="journal article" year="1984">
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<smallCapsWord id="8D70A045FFA0FF82FA8EFB1AC753FB44" baselines="1190,1189" box="[1354,1386,1174,1195]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Mc" pageId="31" pageNumber="306">
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<smallCapsWord id="8D70A045FFA0FF82FAADFB1AC630FB25" baselines="1190,1189,1190,1220" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Donald" pageId="31" pageNumber="306">MCDONALD</smallCapsWord>
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</smallCapsWord>
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&
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<smallCapsWord id="8D70A045FFA0FF82FBE9FB39C653FB25" baselines="1221,1220" box="[1069,1130,1205,1226]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Cassis" pageId="31" pageNumber="306">CASSIS</smallCapsWord>
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1984: 538
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</bibRefCitation>
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(genitalia);
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<bibRefCitation id="EFB84B68FFA0FF82FA86FB39C6B8FB04" author="CASSIS G. & G. F. GROSS" editor="HOUSTON W. W. K. & G. V. MAYNARD" journalOrPublisher="CSIRO Publishing, Melbourne: xiv" pageId="31" pageNumber="306" pagination="1 - 732" refId="ref83117" refString="CASSIS G. & G. F. GROSS (2002): Hemiptera: Heteroptera (Pentatomomorpha). - In: HOUSTON W. W. K. & G. V. MAYNARD (Eds), Zoological Catalogue of Australia. Vol. 27.3 B. CSIRO Publishing, Melbourne: xiv + 1 - 732." title="Hemiptera: Heteroptera (Pentatomomorpha)" type="book chapter" volumeTitle="Zoological Catalogue of Australia. Vol. 27.3 B" year="2002">
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&
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<smallCapsWord id="8D70A045FFA0FF82FC2CFB58C61AFB0A" baselines="1252,1251" box="[1000,1059,1236,1257]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gross" pageId="31" pageNumber="306">GROSS</smallCapsWord>
|
|
2002: 583
|
|
</bibRefCitation>
|
|
(catalogue)
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3336512FFA0FF82FC2CFB7FC679FACB" pageId="31" pageNumber="306" type="type_taxon">
|
|
<paragraph id="8B963699FFA0FF82FC2CFB7FC679FACB" blockId="31.[1000,1435,110,1917]" pageId="31" pageNumber="306">
|
|
Type species:
|
|
<taxonomicName id="4C294D1AFFA0FF82FBA2FB7FC745FAE5" authority="FABRICIUS 1787" authorityName="FABRICIUS" authorityYear="1787" box="[1126,1404,1267,1288]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="306" phylum="Arthropoda" rank="species" species="costatus">
|
|
<emphasis id="B95DEA8BFFA0FF82FBA2FB7FC6DFFAE5" box="[1126,1254,1267,1288]" italics="true" pageId="31" pageNumber="306">Cimex costatus</emphasis>
|
|
<bibRefCitation id="EFB84B68FFA0FF82FB34FB7FC745FAE5" author="FABRICIUS J. C." box="[1264,1404,1267,1288]" journalOrPublisher="C. G. Proft, Hafnia" pageId="31" pageNumber="306" pagination="1 - 348" refId="ref84316" refString="FABRICIUS J. C. (1787): Mantissa insectorum sistens eorum species nuper detectas adjectis characteribus, genericis, differentiis specificis, emendationibus, observationibus. Vol. 2. - C. G. Proft, Hafnia: iii-xx + 1 - 348." title="Mantissa insectorum sistens eorum species nuper detectas adjectis characteribus, genericis, differentiis specificis, emendationibus, observationibus. Vol. 2." type="book chapter" year="1787">
|
|
<smallCapsWord id="8D70A045FFA0FF82FB34FB7FC771FAEB" baselines="1283,1282" box="[1264,1352,1267,1288]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Fabricius" pageId="31" pageNumber="306">FABRICIUS</smallCapsWord>
|
|
1787
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
by monotypy
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3336512FFA0FF82FBCAFACCC760F8E7" pageId="31" pageNumber="306" type="diagnosis">
|
|
<paragraph id="8B963699FFA0FF82FBCAFACCC760F8E7" blockId="31.[1000,1435,110,1917]" pageId="31" pageNumber="306">
|
|
Diagnosis:
|
|
<taxonomicName id="4C294D1AFFA0FF82FBBAFACCC6D1FABA" box="[1150,1256,1344,1367]" class="Insecta" family="Scutelleridae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="306" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFFA0FF82FBBAFACCC6D1FABA" box="[1150,1256,1344,1367]" italics="true" pageId="31" pageNumber="306">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
is recognised by the following combination of characters: elongate-ovoid (
|
|
<figureCitation id="13122A1CFFA0FF82FB42FA0FC686FA77" box="[1158,1215,1411,1434]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="31" pageNumber="306">Figs 3</figureCitation>
|
|
c-e); lateral margins of jugae rounded (
|
|
<figureCitation id="13122A1CFFA0FF82FB46FA28C6F3FA56" box="[1154,1226,1444,1467]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="31" pageNumber="306">Fig. 7b</figureCitation>
|
|
); posterior margin of pronotum strongly convex (
|
|
<figureCitation id="13122A1CFFA0FF82FB39FA4AC700FA30" box="[1277,1337,1478,1501]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="31" pageNumber="306">Figs 3</figureCitation>
|
|
c-e), covering anterior margin of scutellum; metathoracic peritreme mostly rectilinear, sulcate, with apex anteriorly arcuate (
|
|
<figureCitation id="13122A1CFFA0FF82FAD5F9A6C762F9AC" box="[1297,1371,1578,1601]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="31" pageNumber="306">Fig. 7d</figureCitation>
|
|
); thoracic sterna strongly keel-like, pro- and mesosternal keels overlapping (
|
|
<figureCitation id="13122A1CFFA0FF82FAD2F9E1C762F969" box="[1302,1371,1645,1668]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="31" pageNumber="306">Fig. 7c</figureCitation>
|
|
); CAI absent; CAII symmetrical or asymmetrical, bifid (
|
|
<figureCitation id="13122A1CFFA0FF82FBE5F93CC6BAF92A" box="[1057,1155,1712,1735]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="31" pageNumber="306">Figs 8c, d</figureCitation>
|
|
), with rounded or digitiform lobal sclerites; CAIII separated or fused (
|
|
<figureCitation id="13122A1CFFA0FF82FAB0F95DC63DF8E7" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="31" pageNumber="306">Fig. 8d</figureCitation>
|
|
); and, spermathecal reservoir oval.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3336512FFA0FFBFFBCAF8AFC2F2F94B" lastPageId="34" lastPageNumber="309" pageId="31" pageNumber="306" type="description">
|
|
<paragraph id="8B963699FFA0FFBFFBCAF8AFC2F2F94B" blockId="31.[1000,1435,110,1917]" lastBlockId="34.[154,588,110,1917]" lastPageId="34" lastPageNumber="309" pageId="31" pageNumber="306">
|
|
Description: Body elongate-ovoid (
|
|
<figureCitation id="13122A1CFFA0FF82FAB5F8AFC1CAF8B1" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="31" pageNumber="306">Figs 3</figureCitation>
|
|
c-e); moderately-sized to large species; moderately convex dorsally and ventrally; wing tip extending beyond scutellum; moderately to densely punctate; pale to dark brown, sometimes with red or green iridescent highlighting (e.g.
|
|
<taxonomicName id="4C294D1AFF9FFFBDFE46FA7AC2F9F9C2" class="Insecta" family="Scutelleridae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="307" phylum="Arthropoda" rank="species" species="blackburniae">
|
|
<emphasis id="B95DEA8BFF9FFFBDFE46FA7AC2F9F9C2" italics="true" pageId="32" pageNumber="307">Coleotichus blackbur-niae</emphasis>
|
|
</taxonomicName>
|
|
; Hawaii), rarely with patterned darker brown markings. Head: triangular (
|
|
<figureCitation id="13122A1CFF9FFFBDFE3FF9B5C00EF9BD" box="[507,567,1593,1616]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="32" pageNumber="307">Figs 3</figureCitation>
|
|
ce, 7a), moderately convex; clypeus broadly rounded at apex and barely surpassing juga (
|
|
<figureCitation id="13122A1CFF9FFFBDFF65F911C2DEF959" box="[161,231,1693,1716]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="32" pageNumber="307">Fig. 7a</figureCitation>
|
|
); jugal margins rounded (
|
|
<figureCitation id="13122A1CFF9FFFBDFE19F911C027F959" box="[477,542,1693,1716]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="32" pageNumber="307">Fig 7b</figureCitation>
|
|
); lorae strongly demarcated, margins carinate (
|
|
<figureCitation id="13122A1CFF9FFFBDFF65F96CC2D5F91A" box="[161,236,1760,1783]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="32" pageNumber="307">Fig. 7b</figureCitation>
|
|
); bucculae narrow, lateral margins weakly arcuate (
|
|
<figureCitation id="13122A1CFF9FFFBDFE8CF88EC3AEF8F4" box="[328,407,1794,1817]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="32" pageNumber="307">Fig. 7c</figureCitation>
|
|
). Antennae: AI-AII(a) shortest segments, subequal; AIII-AIV longest segments, subequal or AIV slightly longer. Labium: reaching apex of metasternum (
|
|
<figureCitation id="13122A1CFF9FFFBDFCC6FA1EC174FA44" box="[770,845,1426,1449]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="32" pageNumber="307">Fig. 7c</figureCitation>
|
|
) to abdominal SIV; LII usually longest segment. Pronotum: large, shield-like, strongly convex (
|
|
<figureCitation id="13122A1CFF9FFFBDFC0BFA59C635FA01" box="[975,1036,1493,1516]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="32" pageNumber="307">Figs 3</figureCitation>
|
|
ce); anterior margin concave; anterolateral margins rectilinear, carinate; humeral angles rounded to weakly angulate; posterior region of disc greatly expanded, posterior margin strongly convex, overlapping anterior margin of scutellum (
|
|
<figureCitation id="13122A1CFF9FFFBDFC87F911C1B9F959" box="[835,896,1693,1716]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="32" pageNumber="307">Figs 3</figureCitation>
|
|
c-e). Scutellum: elongate, strongly convex, U-shaped, most often strongly tapered posteriorly (
|
|
<figureCitation id="13122A1CFF9FFFBDFC0AF96CC635F91A" box="[974,1036,1760,1783]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="32" pageNumber="307">Figs 3</figureCitation>
|
|
ce); basal aspects of corium and clavus, and connexiva III-VII exposed (
|
|
<figureCitation id="13122A1CFF9FFFBDFC45F8AFC183F8D7" box="[897,954,1827,1850]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="32" pageNumber="307">Fig. 3</figureCitation>
|
|
c-e). Thoracic pleura: external efferent system of metathoracic glands well-developed, raised (
|
|
<figureCitation id="13122A1CFF9DFFBFFF65FFE2C2D0FF68" box="[161,233,110,133]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="34" pageNumber="309">Fig. 7c</figureCitation>
|
|
); evaporative areas occupying most of metepisternum, and extending to mesepimeron (
|
|
<figureCitation id="13122A1CFF9DFFBFFEEEFF3CC3B5FF2A" box="[298,396,176,199]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="34" pageNumber="309">Figs 7c, d</figureCitation>
|
|
); ostiole moderately-sized (
|
|
<figureCitation id="13122A1CFF9DFFBFFF34FF5EC300FF04" box="[240,313,210,233]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="34" pageNumber="309">Fig. 7d</figureCitation>
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); peritreme elongate, mostly rectilinear, with apex weakly curved at apex, gutter-like, medially sulcate (
|
|
<figureCitation id="13122A1CFF9DFFBFFE32FE98C079FEC6" box="[502,576,276,299]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="34" pageNumber="309">Fig. 7d</figureCitation>
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). Thoracic sterna: lateral margins strongly sulcate, prosternal and mesosternal keels overlapping (
|
|
<figureCitation id="13122A1CFF9DFFBFFEDFFEF5C358FE7D" box="[283,353,377,400]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="34" pageNumber="309">Fig. 7c</figureCitation>
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), with latter contiguous with explanate anterior margin of proepisternum, rounded (
|
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<figureCitation id="13122A1CFF9DFFBFFE8EFE37C3A9FE3F" box="[330,400,443,466]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="34" pageNumber="309">Fig. 7c</figureCitation>
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); metasternal keels thickened (
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<figureCitation id="13122A1CFF9DFFBFFED6FE51C358FE19" box="[274,353,477,500]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="34" pageNumber="309">Fig. 7c</figureCitation>
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). Male Genitalia: pygophore (
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<figureCitation id="13122A1CFF9DFFBFFF32FE72C369FDF8" box="[246,336,510,533]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="34" pageNumber="309">Figs 7e, f</figureCitation>
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,
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<figureCitation id="13122A1CFF9DFFBFFE9EFE72C34BFDF8" box="[346,370,510,533]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">8a</figureCitation>
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) large, subquadrate to suboval, ventral margin convex; genital opening dorsal in orientation, broad, suboval, lateral margins densely setose; parameres (
|
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<figureCitation id="13122A1CFF9DFFBFFF23FD08C30FFD76" box="[231,310,644,667]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Fig. 8b</figureCitation>
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), columnar stem, strongly hook-shaped crown, sometimes base of crown with flange; aedeagus (
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<figureCitation id="13122A1CFF9DFFBFFE15FD4BC007FD33" box="[465,574,711,734]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Figs 8c, d</figureCitation>
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), asymmetrical (
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<figureCitation id="13122A1CFF9DFFBFFEEBFD64C3A0FD12" box="[303,409,744,767]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Fig. 8c, d</figureCitation>
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) or symmetrical; phallotheca squat (
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|
<figureCitation id="13122A1CFF9DFFBFFEA9FC85C386FCCD" box="[365,447,777,800]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Fig. 8c</figureCitation>
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), sclerotized, without processes; ejaculatory apparatus moderately developed (
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<figureCitation id="13122A1CFF9DFFBFFE51FCC0C3DEFC8E" box="[405,487,844,867]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Fig. 8c</figureCitation>
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); ventral conducting canal with 8-10 paired convolutions (
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<figureCitation id="13122A1CFF9DFFBFFF18FC03C31FFC4B" box="[220,294,911,934]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Fig. 8c</figureCitation>
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); ejaculatory reservoir short, oval (
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<figureCitation id="13122A1CFF9DFFBFFF17FC3CC325FC2A" box="[211,284,944,967]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Fig. 8c</figureCitation>
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); vesica short to elongate, tapered towards apex, strongly arcuate; conjunctival appendages (
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<figureCitation id="13122A1CFF9DFFBFFEBEFC7FC3D8FBE7" box="[378,481,1011,1034]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Figs 8c, d</figureCitation>
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): CAI absent; CAII asymmetrical (
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<figureCitation id="13122A1CFF9DFFBFFE52FB98C3CFFBC6" box="[406,502,1044,1067]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Figs 8c, d</figureCitation>
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) or symmetrical, greatly enlarged, mostly membraneous, distally bifurcate, CAII(L) with short (acute) or large (sickle-shaped) lobal sclerite, CAII(M) with short acute lobal sclerite, or blunt bifid lobal sclerites; CAIII sclerotized (
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<figureCitation id="13122A1CFF9DFFBFFF29FB51C36BFB19" box="[237,338,1245,1268]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Fig. 8c, d</figureCitation>
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) or membraneous, fused medially or separated, flange-shaped or Sshaped, sometimes bifid. Female Terminalia: ventrally oriented, co-planar; paratergites VIII large, broad, subelliptoid, posterior margin rounded; paratergites IX narrowly elliptoid, medial margins truncate; gonocoxae I transverse, undivided, posterior margin concave, medial margins elevated. Spermatheca: short to moderately-sized proximal fecundation canal; spermathecal reservoir narrowly dilated elliptoid, membraneous; pump with dorsal and proximal flanges; bulb oval.
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</paragraph>
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</subSubSection>
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<caption id="DF566611FF9FFFBDFB3FFFE7C78FFD7E" ID-DOI="http://doi.org/10.5281/zenodo.13713977" ID-Zenodo-Dep="13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="32" pageNumber="307" startId="32.[1275,1311,107,126]" targetBox="[151,1257,106,1409]" targetPageId="32" targetType="figure">
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<paragraph id="8B963699FF9FFFBDFB3FFFE7C78FFD7E" blockId="32.[1275,1526,107,659]" pageId="32" pageNumber="307">
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<emphasis id="B95DEA8BFF9FFFBDFB3FFFE7C70EFF93" bold="true" box="[1275,1335,107,126]" pageId="32" pageNumber="307">Fig. 7</emphasis>
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: Scanning electron micrographs of key characters of
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<taxonomicName id="4C294D1AFF9FFFBDFA46FF2CC775FF23" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="307" phylum="Arthropoda" rank="species" species="costatus">
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<emphasis id="B95DEA8BFF9FFFBDFA46FF2CC775FF23" italics="true" pageId="32" pageNumber="307">Coleotichus costatus</emphasis>
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</taxonomicName>
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(
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<emphasis id="B95DEA8BFF9FFFBDFA9DFF37C75FFF23" bold="true" box="[1369,1382,187,206]" pageId="32" pageNumber="307">a</emphasis>
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) Head, dorsal view (
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<emphasis id="B95DEA8BFF9FFFBDFAF2FF5AC77DFF04" bold="true" box="[1334,1348,214,233]" pageId="32" pageNumber="307">b</emphasis>
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) Head, lateral view (
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<emphasis id="B95DEA8BFF9FFFBDFAF2FF7CC778FEEE" bold="true" box="[1334,1345,240,259]" pageId="32" pageNumber="307">c</emphasis>
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) Head and thorax, ventral view (
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<emphasis id="B95DEA8BFF9FFFBDFA15FE87C7E6FEF3" bold="true" box="[1489,1503,267,286]" pageId="32" pageNumber="307">d</emphasis>
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) External efferent system of metathoracic glands, ventral view (
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<emphasis id="B95DEA8BFF9FFFBDFA47FED7C7A9FE83" bold="true" box="[1411,1424,347,366]" pageId="32" pageNumber="307">e</emphasis>
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) male pygophore, ventral view (
|
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<emphasis id="B95DEA8BFF9FFFBDFAC6FE1CC732FE4E" bold="true" box="[1282,1291,400,419]" pageId="32" pageNumber="307">f</emphasis>
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) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm.
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</paragraph>
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</caption>
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|
<caption id="DF566611FF9EFFBCFFF8F9C7C2AEF86E" ID-DOI="http://doi.org/10.5281/zenodo.13713979" ID-Zenodo-Dep="13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="33" pageNumber="308" startId="33.[60,96,1611,1630]" subCaptionStartIDs="33.[118,260,1664,1683]" subCaptionStarts="Abbr" targetBox="[60,1436,117,1923]" targetPageId="33" targetType="figure">
|
|
<paragraph id="8B963699FF9EFFBCFFF8F9C7C2AEF86E" blockId="33.[60,882,1611,1923]" pageId="33" pageNumber="308">
|
|
<emphasis id="B95DEA8BFF9EFFBCFFF8F9C7C241F9B3" bold="true" box="[60,120,1611,1630]" pageId="33" pageNumber="308">Fig. 8</emphasis>
|
|
: Male genitalia of
|
|
<taxonomicName id="4C294D1AFF9EFFBCFEF3F9C7C03BF9B3" box="[311,514,1611,1630]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="308" phylum="Arthropoda" rank="species" species="costatus">
|
|
<emphasis id="B95DEA8BFF9EFFBCFEF3F9C7C03BF9B3" box="[311,514,1611,1630]" italics="true" pageId="33" pageNumber="308">Coleotichus costatus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis id="B95DEA8BFF9EFFBCFDCBF9C7C025F9B3" bold="true" box="[527,540,1611,1630]" pageId="33" pageNumber="308">a</emphasis>
|
|
) pygophore, dorsal view (
|
|
<emphasis id="B95DEA8BFF9EFFBCFF80F9E9C26BF995" bold="true" box="[68,82,1637,1656]" pageId="33" pageNumber="308">b</emphasis>
|
|
) paramere, lateral view (
|
|
<emphasis id="B95DEA8BFF9EFFBCFE8AF9E9C360F995" bold="true" box="[334,345,1637,1656]" pageId="33" pageNumber="308">c</emphasis>
|
|
) aedeagus, lateral view (
|
|
<emphasis id="B95DEA8BFF9EFFBCFD90F9E9C05BF995" bold="true" box="[596,610,1637,1656]" pageId="33" pageNumber="308">d</emphasis>
|
|
) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection id="C3336512FF9DFFBFFF04F933C185FDB4" pageId="34" pageNumber="309" type="distribution">
|
|
<paragraph id="8B963699FF9DFFBFFF04F933C185FDB4" blockId="34.[154,588,110,1917]" lastBlockId="34.[622,1056,110,1917]" pageId="34" pageNumber="309">
|
|
Diversity and distribution: The highest area of species diversity in
|
|
<taxonomicName id="4C294D1AFF9DFFBFFE68F96DC02FF915" box="[428,534,1761,1784]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFE68F96DC02FF915" box="[428,534,1761,1784]" italics="true" pageId="34" pageNumber="309">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
is in the Australian zoogeographic region, with seven of the thirteen described species. A number of the species are widespread, with all three Australian species, found also in New Guinea, or surrounding islands, with
|
|
<taxonomicName id="4C294D1AFF9DFFBFFBC2FFE2C087FF4B" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="excellens">
|
|
<emphasis id="B95DEA8BFF9DFFBFFBC2FFE2C087FF4B" italics="true" pageId="34" pageNumber="309">C. excellens</emphasis>
|
|
</taxonomicName>
|
|
now known from as far north as Taiwan (see below). The genus is also represented in the Oriental region, with two species known from Indonesia (Borneo and Sumatra).
|
|
<taxonomicName id="4C294D1AFF9DFFBFFD1DFE99C17AFEC1" box="[729,835,277,300]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFD1DFE99C17AFEC1" box="[729,835,277,300]" italics="true" pageId="34" pageNumber="309">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
is also represented in Pacific islands and archipelagos, distant from Australia, including Samoa, Hawaii and the Marquesas. The most spectacular species of
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDAAFE16C0E1FE5C" box="[622,728,410,433]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDAAFE16C0E1FE5C" box="[622,728,410,433]" italics="true" pageId="34" pageNumber="309">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C294D1AFF9DFFBFFD2EFE16C1B8FE5C" box="[746,897,410,433]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="blackburniae">
|
|
<emphasis id="B95DEA8BFF9DFFBFFD2EFE16C1B8FE5C" box="[746,897,410,433]" italics="true" pageId="34" pageNumber="309">C. blackburniae</emphasis>
|
|
</taxonomicName>
|
|
, is endemic to the Hawaiian Islands, where it breeds on koa, a native species of
|
|
<taxonomicName id="4C294D1AFF9DFFBFFCA5FE51C199FE19" box="[865,928,477,500]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="34" pageNumber="309" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFCA5FE51C199FE19" box="[865,928,477,500]" italics="true" pageId="34" pageNumber="309">Acacia</emphasis>
|
|
</taxonomicName>
|
|
, where both plant and insect are threatened, the jewel bug by green vegetable bug biological control agents (
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD27FDCEC104FDB5" baselines="595,596" box="[739,829,578,601]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Johnson" pageId="34" pageNumber="309">JOHNSON</smallCapsWord>
|
|
et al. 2005).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3336512FF9DFFBFFDABFDFEC626FACB" pageId="34" pageNumber="309" type="taxon_list">
|
|
<paragraph id="8B963699FF9DFFBFFDABFDFEC626FACB" blockId="34.[622,1056,110,1917]" pageId="34" pageNumber="309">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFDFEC124FD64" bold="true" box="[623,797,626,649]" pageId="34" pageNumber="309">Included species:</emphasis>
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFD18C619FD46" authority="VAN DUZEE 1932 Marquesas" authorityName="VAN DUZEE" authorityYear="1932" box="[623,1056,660,683]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="adamsoni">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFD18C0DCFD46" box="[623,741,660,683]" italics="true" pageId="34" pageNumber="309">C. adamsoni</emphasis>
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD28FD18C121FD47" baselines="677,678" box="[748,792,660,683]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Van" pageId="34" pageNumber="309">VAN</smallCapsWord>
|
|
<smallCapsWord id="8D70A045FF9DFFBFFCDBFD18C164FD44" baselines="677,677" box="[799,861,660,683]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Duzee" pageId="34" pageNumber="309">DUZEE</smallCapsWord>
|
|
1932 Marquesas
|
|
</taxonomicName>
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDAAFD39C18BFD21" authority="(MONTROUZIER 1858)" baseAuthorityName="MONTROUZIER" baseAuthorityYear="1858" box="[622,946,693,716]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="artensis">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDAAFD39C0EDFD21" box="[622,724,693,716]" italics="true" pageId="34" pageNumber="309">C. artensis</emphasis>
|
|
(
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFD26FD39C192FD21" author="MONTROUZIER P." box="[738,939,693,716]" journalOrPublisher="Ann. Soc. Linn. Lyon" pageId="34" pageNumber="309" pagination="243 - 260" part="2" refId="ref87667" refString="MONTROUZIER P. (1858): Description de quelque Hemipteres de la Nouvelle-Caledonie. - Ann. Soc. Linn. Lyon (2) 5: 243 - 260." title="Description de quelque Hemipteres de la Nouvelle-Caledonie" type="journal article" year="1858">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD26FD39C148FD26" baselines="710,711" box="[738,881,693,716]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Montrouzier" pageId="34" pageNumber="309">MONTROUZIER</smallCapsWord>
|
|
1858
|
|
</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
Australia, New Caledonia, Fiji, Vanuatu, Samoa, Aru Island
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFC96C163FCDC" authority="TAEUBER 1929" authorityName="TAEUBER" authorityYear="1929" box="[623,858,794,817]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="bakeri">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFC96C0FFFCDC" box="[623,710,794,817]" italics="true" pageId="34" pageNumber="309">C. bakeri</emphasis>
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFD09FC96C163FCDC" author="TAEUBER H. W." box="[717,858,794,817]" journalOrPublisher="Konowia" pageId="34" pageNumber="309" pagination="194 - 223" part="8" refId="ref89214" refString="TAEUBER H. W. (1929): Beitrage zur Kenntnis der Heteropteren-Fauna der Philippinen. - Konowia 8 (2): 194 - 223." title="Beitrage zur Kenntnis der Heteropteren-Fauna der Philippinen" type="journal article" year="1929">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD09FC96C11DFCDD" baselines="811,812" box="[717,804,794,817]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Taeuber" pageId="34" pageNumber="309">TAEUBER</smallCapsWord>
|
|
1929
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
Philippines
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFCB7C14FFCBF" authority="SCHOUTEDEN 1905" authorityName="SCHOUTEDEN" authorityYear="1905" box="[623,886,827,850]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="biroi">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFCB7C081FCBF" box="[623,696,827,850]" italics="true" pageId="34" pageNumber="309">C. biroi</emphasis>
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFD7AFCB7C14FFCBF" author="SCHOUTEDEN H." box="[702,886,827,850]" journalOrPublisher="Ann. Mus. Nat. Hung." pageId="34" pageNumber="309" pagination="317 - 361" part="3" refId="ref87993" refString="SCHOUTEDEN H. (1905): Monographie du genre Coleotichus. - Ann. Mus. Nat. Hung. 3: 317 - 361." title="Monographie du genre Coleotichus" type="journal article" year="1905">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD7AFCB7C106FCBC" baselines="844,845" box="[702,831,827,850]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Schouteden" pageId="34" pageNumber="309">SCHOUTEDEN</smallCapsWord>
|
|
1905
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
New Guinea
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFCD1C626FC99" authority="WHITE 1881 Hawaii" authorityName="WHITE" authorityYear="1881" box="[623,1055,861,884]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="blackburniae">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFCD1C13AFC99" box="[623,771,861,884]" italics="true" pageId="34" pageNumber="309">C. blackburniae</emphasis>
|
|
<smallCapsWord id="8D70A045FF9DFFBFFCCEFCD1C175FC9F" baselines="878,878" box="[778,844,861,884]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="White" pageId="34" pageNumber="309">WHITE</smallCapsWord>
|
|
1881 Hawaii
|
|
</taxonomicName>
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFCF2C1AFFC78" authority="SCHOUTEDEN 1905" authorityName="SCHOUTEDEN" authorityYear="1905" box="[623,918,894,917]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="breddini">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFCF2C0E1FC78" box="[623,728,894,917]" italics="true" pageId="34" pageNumber="309">C. breddini</emphasis>
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFD1BFCF2C1AFFC78" author="SCHOUTEDEN H." box="[735,918,894,917]" journalOrPublisher="Ann. Mus. Nat. Hung." pageId="34" pageNumber="309" pagination="317 - 361" part="3" refId="ref87993" refString="SCHOUTEDEN H. (1905): Monographie du genre Coleotichus. - Ann. Mus. Nat. Hung. 3: 317 - 361." title="Monographie du genre Coleotichus" type="journal article" year="1905">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD1BFCF2C166FC79" baselines="911,912" box="[735,863,894,917]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Schouteden" pageId="34" pageNumber="309">SCHOUTEDEN</smallCapsWord>
|
|
1905
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
Micronesia
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFC13C1B3FC5B" authority="SCHOUTEDEN 1905" authorityName="SCHOUTEDEN" authorityYear="1905" box="[623,906,927,950]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="bulowi">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFC13C0F5FC5B" box="[623,716,927,950]" italics="true" pageId="34" pageNumber="309">C. bulowi</emphasis>
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFD16FC13C1B3FC5B" author="SCHOUTEDEN H." box="[722,906,927,950]" journalOrPublisher="Ann. Mus. Nat. Hung." pageId="34" pageNumber="309" pagination="317 - 361" part="3" refId="ref87993" refString="SCHOUTEDEN H. (1905): Monographie du genre Coleotichus. - Ann. Mus. Nat. Hung. 3: 317 - 361." title="Monographie du genre Coleotichus" type="journal article" year="1905">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD16FC13C16AFC5B" baselines="944,946" box="[722,851,927,950]" lowerCaseFontSize="10" mainFontSize="10" normCase="title" normString="Schouteden" pageId="34" pageNumber="309">SCHOUTEDEN</smallCapsWord>
|
|
1905
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
Samoa
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDAAFC4DC1B5FC35" authority="(FABRICIUS 1787)" baseAuthorityName="FABRICIUS" baseAuthorityYear="1787" box="[622,908,961,984]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="costatus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDAAFC4DC0E1FC35" box="[622,728,961,984]" italics="true" pageId="34" pageNumber="309">C. costatus</emphasis>
|
|
(
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFD22FC4DC1BCFC35" author="FABRICIUS J. C." box="[742,901,961,984]" journalOrPublisher="C. G. Proft, Hafnia" pageId="34" pageNumber="309" pagination="1 - 348" refId="ref84316" refString="FABRICIUS J. C. (1787): Mantissa insectorum sistens eorum species nuper detectas adjectis characteribus, genericis, differentiis specificis, emendationibus, observationibus. Vol. 2. - C. G. Proft, Hafnia: iii-xx + 1 - 348." title="Mantissa insectorum sistens eorum species nuper detectas adjectis characteribus, genericis, differentiis specificis, emendationibus, observationibus. Vol. 2." type="book chapter" year="1787">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD22FC4DC172FC3A" baselines="978,979" box="[742,843,961,984]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabricius" pageId="34" pageNumber="309">FABRICIUS</smallCapsWord>
|
|
1787
|
|
</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
Australia, New Caledonia, Tonga
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFB88C154FBF6" authority="WALKER 1867" authorityName="WALKER" authorityYear="1867" box="[623,877,1028,1051]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="excellens">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFB88C0E6FBF6" box="[623,735,1028,1051]" italics="true" pageId="34" pageNumber="309">C. excellens</emphasis>
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFD22FB88C154FBF6" author="WALKER F." box="[742,877,1028,1051]" journalOrPublisher="E. Newman, London" pageId="34" pageNumber="309" pagination="1 - 240" refId="ref89733" refString="WALKER F. (1867): Catalogue of the specimens of Heteropterous Hemiptera in the collection of the British Museum. Part I. Scutata. - E. Newman, London: 1 - 240." title="Catalogue of the specimens of Heteropterous Hemiptera in the collection of the British Museum. Part I. Scutata." type="book chapter" year="1867">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD22FB88C10EFBF7" baselines="1045,1046" box="[742,823,1028,1051]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Walker" pageId="34" pageNumber="309">WALKER</smallCapsWord>
|
|
1867
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
Australia, Fiji, New Guinea, New Caledonia, Samoa, Philippines, Taiwan, Indonesia (Irian Jaya)
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFB06C1AAFB4C" authority="VOLLENHOVEN 1863" authorityName="VOLLENHOVEN" authorityYear="1863" box="[623,915,1162,1185]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="fuscus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFB06C0F1FB4C" box="[623,712,1162,1185]" italics="true" pageId="34" pageNumber="309">C. fuscus</emphasis>
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFD0AFB06C1AAFB4C" author="VOLLENHOVEN S. C. S. VAN" box="[718,915,1162,1185]" journalOrPublisher="Martinus Nijhoff, La Haye" pageId="34" pageNumber="309" pagination="1 - 60" refId="ref89654" refString="VOLLENHOVEN S. C. S. VAN (1863): Essai d'une Faune Entomologique de l'archipel Indo-neerlandais. Premiere Monographie: Famille des Scutellerides. - Martinus Nijhoff, La Haye: 1 - 60." title="Essai d'une Faune Entomologique de l'archipel Indo-neerlandais. Premiere Monographie: Famille des Scutellerides" type="book chapter" year="1863">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD0AFB06C165FB4D" baselines="1179,1180" box="[718,860,1162,1185]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Vollenhoven" pageId="34" pageNumber="309">VOLLENHOVEN</smallCapsWord>
|
|
1863
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
Ambon, Ceram
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFB40C1B7FB0E" authority="USINGER 1946" authorityName="USINGER" authorityYear="1946" box="[623,910,1228,1251]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="marianensis">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFB40C0C2FB0E" box="[623,763,1228,1251]" italics="true" pageId="34" pageNumber="309">C. marianensis</emphasis>
|
|
<smallCapsWord id="8D70A045FF9DFFBFFCC6FB40C161FB0E" baselines="1245,1247" box="[770,856,1228,1251]" lowerCaseFontSize="10" mainFontSize="10" normCase="title" normString="Usinger" pageId="34" pageNumber="309">USINGER</smallCapsWord>
|
|
1946
|
|
</taxonomicName>
|
|
Guam
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDAAFB62C626FAE8" authority="BERGROTH 1915" authorityName="BERGROTH" authorityYear="1915" box="[622,1055,1262,1285]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="ornamentifer">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDAAFB62C13CFAE8" box="[622,773,1262,1285]" italics="true" pageId="34" pageNumber="309">C. ornamentifer</emphasis>
|
|
<smallCapsWord id="8D70A045FF9DFFBFFCCFFB62C14AFAE9" baselines="1279,1280" box="[779,883,1262,1285]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Bergroth" pageId="34" pageNumber="309">BERGROTH</smallCapsWord>
|
|
1915 Borneo
|
|
</taxonomicName>
|
|
<taxonomicName id="4C294D1AFF9DFFBFFDABFA83C1B2FACB" authority="BREDDIN 1900" authorityName="BREDDIN" authorityYear="1900" box="[623,907,1295,1318]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="sumatranus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFDABFA83C0C3FACB" box="[623,762,1295,1318]" italics="true" pageId="34" pageNumber="309">C. sumatranus</emphasis>
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFCC4FA83C1B2FACB" author="BREDDIN G." box="[768,907,1295,1318]" journalOrPublisher="Abh. Senckenb. Naturforsch. Ges." pageId="34" pageNumber="309" pagination="139 - 202" part="25" refId="ref82705" refString="BREDDIN G. (1900): Hemiptera gesammelt von Professor Kukenthal im Malayischen Archipel. - Abh. Senckenb. Naturforsch. Ges. 25: 139 - 202." title="Hemiptera gesammelt von Professor Kukenthal im Malayischen Archipel" type="journal article" year="1900">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFCC4FA83C16DFAC8" baselines="1312,1313" box="[768,852,1295,1318]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Breddin" pageId="34" pageNumber="309">BREDDIN</smallCapsWord>
|
|
1900
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
Sumatra
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3336512FF9DFFBEFD50FAC3C0E1FB84" lastPageId="35" lastPageNumber="310" pageId="34" pageNumber="309" type="discussion">
|
|
<paragraph id="8B963699FF9DFFBFFD50FAC3C162F890" blockId="34.[622,1056,110,1917]" pageId="34" pageNumber="309">
|
|
Remarks:
|
|
<taxonomicName id="4C294D1AFF9DFFBFFCC4FAC3C153FA8B" box="[768,874,1359,1382]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFCC4FAC3C153FA8B" box="[768,874,1359,1382]" italics="true" pageId="34" pageNumber="309">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
is a distinctive genus of
|
|
<taxonomicName id="4C294D1AFF9DFFBFFD00FAFDC10AFA65" box="[708,819,1393,1416]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="300" phylum="Arthropoda" rank="subFamily" subFamily="Elvisurinae">Elvisurinae</taxonomicName>
|
|
, having greatly enlarged thoracic sternal keels, with the pro- and mesothoracic keels overlapping (
|
|
<figureCitation id="13122A1CFF9DFFBFFC00FA3FC62CFA27" box="[964,1045,1459,1482]" captionStart="Fig" captionStartId="32.[1275,1311,107,126]" captionTargetBox="[151,1257,106,1409]" captionTargetId="figure-280@32.[150,1258,105,1410]" captionTargetPageId="32" captionText="Fig. 7: Scanning electron micrographs of key characters of Coleotichus costatus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Ms(k) = mesosternal keel; Pe = peritreme; Pe(k) = proepisternal keel, ventral view; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713977" httpUri="https://zenodo.org/record/13713977/files/figure.png" pageId="34" pageNumber="309">Fig. 7d</figureCitation>
|
|
). The genus shares some similarities with the nominotypical genus
|
|
<taxonomicName id="4C294D1AFF9DFFBFFC9FFA7AC19FF9E0" box="[859,934,1526,1549]" class="Insecta" genus="Elvisura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFC9FFA7AC19FF9E0" box="[859,934,1526,1549]" italics="true" pageId="34" pageNumber="309">Elvisura</emphasis>
|
|
</taxonomicName>
|
|
, with the pronotum greatly expanded posteriorly, covering the anterior margin of the scutellum (
|
|
<figureCitation id="13122A1CFF9DFFBFFDB1F9D7C08EF99F" box="[629,695,1627,1650]" captionStart="Fig" captionStartId="26.[620,656,1714,1733]" captionTargetBox="[620,1517,114,1920]" captionTargetId="figure-375@26.[619,1531,105,1925]" captionTargetPageId="26" captionText="Fig. 3: Habitus of Australian Elvisurinae species (a) Austrotichus rugosus, Oi (b) Solenotichus circuliferus, Oi (c) Coleotichus artensis, Oi (d) Coleotichus costatus, Oi (e) Coleotichus excellens, Oi. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13713967" httpUri="https://zenodo.org/record/13713967/files/figure.png" pageId="34" pageNumber="309">Figs 3</figureCitation>
|
|
c-e). The male genitalia resemble those of
|
|
<taxonomicName id="4C294D1AFF9DFFBFFD01F9F0C129F97E" box="[709,784,1660,1683]" class="Insecta" genus="Elvisura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFD01F9F0C129F97E" box="[709,784,1660,1683]" italics="true" pageId="34" pageNumber="309">Elvisura</emphasis>
|
|
</taxonomicName>
|
|
, in the following ways: the CAII have lateral and medial lobes with lobal sclerites, and the CAIII are sclerotized and fused medially (as in
|
|
<taxonomicName id="4C294D1AFF9DFFBFFC9CF96CC1FEF91A" baseAuthorityName="FABRICIUS" baseAuthorityYear="1787" box="[856,967,1760,1783]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="costatus">
|
|
<emphasis id="B95DEA8BFF9DFFBFFC9CF96CC1FEF91A" box="[856,967,1760,1783]" italics="true" pageId="34" pageNumber="309">C. costatus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C294D1AFF9DFFBFFBC2F96CC08BF8F4" baseAuthorityName="MONTROUZIER" baseAuthorityYear="1858" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="artensis">
|
|
<emphasis id="B95DEA8BFF9DFFBFFBC2F96CC08BF8F4" italics="true" pageId="34" pageNumber="309">C. artensis</emphasis>
|
|
</taxonomicName>
|
|
) (e.g.,
|
|
<figureCitation id="13122A1CFF9DFFBFFCC5F88EC15FF8F4" box="[769,870,1794,1817]" captionStart="Fig" captionStartId="33.[60,96,1611,1630]" captionTargetBox="[60,1436,117,1923]" captionTargetId="figure-15@33.[215,1436,105,1912]" captionTargetPageId="33" captionText="Fig. 8: Male genitalia of Coleotichus costatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L)L = left second conjunctival appendage, lateral branch; CAII(L)R = right second conjunctival appendage, lateral branch; CAII(M)L = left second conjunctival appendage, medial branch; CAII(M)R = right second conjunctival appendage, lateral branch; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13713979" httpUri="https://zenodo.org/record/13713979/files/figure.png" pageId="34" pageNumber="309">Figs 8c, d</figureCitation>
|
|
).
|
|
<bibRefCitation id="EFB84B68FF9DFFBFFCB9F88EC626F8F4" author="LESTON D." box="[893,1055,1794,1817]" journalOrPublisher="Ann. Sth Afr." pageId="34" pageNumber="309" pagination="46 - 60" part="51" refId="ref86586" refString="LESTON D. (1953 b): Notes on the Ethiopian Pentatomidae. IV. A new genus and species from the Cape Province. - Ann. Sth Afr. 51 (2): 46 - 60." title="Notes on the Ethiopian Pentatomidae. IV. A new genus and species from the Cape Province" type="journal article" year="1953">
|
|
<smallCapsWord id="8D70A045FF9DFFBFFCB9F88EC1FFF8F5" baselines="1811,1812" box="[893,966,1794,1817]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Leston" pageId="34" pageNumber="309">LESTON</smallCapsWord>
|
|
(1953b)
|
|
</bibRefCitation>
|
|
described the male genitalia of
|
|
<taxonomicName id="4C294D1AFF9DFFBFFC61F8AFC0D5F8B6" authority="SPINOLA" authorityName="SPINOLA" class="Insecta" genus="Elvisura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="309" phylum="Arthropoda" rank="species" species="irrorata">
|
|
<emphasis id="B95DEA8BFF9DFFBFFC61F8AFC0AAF8B1" italics="true" pageId="34" pageNumber="309">Elvisura irro-rata</emphasis>
|
|
<smallCapsWord id="8D70A045FF9DFFBFFD5EF8C9C0D5F8B6" baselines="1878,1879" box="[666,748,1861,1884]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Spinola" pageId="34" pageNumber="309">SPINOLA</smallCapsWord>
|
|
</taxonomicName>
|
|
, which serve as a basis for this comparative hypothesis.
|
|
</paragraph>
|
|
<paragraph id="8B963699FF9CFFBEFDFEFFE2C10DFDB3" blockId="35.[532,966,110,1918]" pageId="35" pageNumber="310">
|
|
<bibRefCitation id="EFB84B68FF9CFFBEFDFEFFE2C0F0FF68" author="WHITE A." box="[570,713,110,133]" journalOrPublisher="Mag. Nat. Hist." pageId="35" pageNumber="310" pagination="537 - 543" part="2" refId="ref89958" refString="WHITE A. (1839): Descriptions of two Hemipterous insects. - Mag. Nat. Hist. (2) 3: 537 - 543." title="Descriptions of two Hemipterous insects" type="journal article" year="1839">
|
|
<smallCapsWord id="8D70A045FF9CFFBEFDFEFFE2C045FF6E" baselines="127,127" box="[570,636,110,133]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="White" pageId="35" pageNumber="310">WHITE</smallCapsWord>
|
|
(1839)
|
|
</bibRefCitation>
|
|
described
|
|
<taxonomicName id="4C294D1AFF9CFFBEFCF3FFE2C19BFF68" box="[823,930,110,133]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="310" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9CFFBEFCF3FFE2C19BFF68" box="[823,930,110,133]" italics="true" pageId="35" pageNumber="310">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
on the basis of the Australian species,
|
|
<emphasis id="B95DEA8BFF9CFFBEFCABFF03C001FF25" italics="true" pageId="35" pageNumber="310">
|
|
<taxonomicName id="4C294D1AFF9CFFBEFCABFF03C00DFF25" baseAuthorityName="FABRICIUS" baseAuthorityYear="1787" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="310" phylum="Arthropoda" rank="species" species="costatus">C. costa-tus</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
<bibRefCitation id="EFB84B68FF9CFFBEFD81FF3DC123FF25" author="SCHOUTEDEN H." bookContentInfo="5 pls." box="[581,794,177,200]" journalOrPublisher="Genera Insectorum" pageId="35" pageNumber="310" pagination="1 - 98" part="24" refId="ref87964" refString="SCHOUTEDEN H. (1904): Heteroptera. Fam. Pentatomidae. Subfam. Scutellerinae. - Genera Insectorum 24: 1 - 98, 5 pls." title="Heteroptera. Fam. Pentatomidae. Subfam. Scutellerinae" type="book chapter" year="1904">
|
|
<smallCapsWord id="8D70A045FF9CFFBEFD81FF3DC0FFFF2A" baselines="194,195" box="[581,710,177,200]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Schouteden" pageId="35" pageNumber="310">SCHOUTEDEN</smallCapsWord>
|
|
(1904)
|
|
</bibRefCitation>
|
|
, in redescribing the genus, established three subgenera (
|
|
<taxonomicName id="4C294D1AFF9CFFBEFDD9FF79C04EFEC3" authority=", EpicoleoticHus and ParacoleoticHus" authorityName="EpicoleoticHus and ParacoleoticHus" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="310" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9CFFBEFDD9FF79C0B1FEE1" box="[541,648,245,268]" italics="true" pageId="35" pageNumber="310">Coleotichus</emphasis>
|
|
,
|
|
<emphasis id="B95DEA8BFF9CFFBEFD69FF79C116FEE1" box="[685,815,245,268]" italics="true" pageId="35" pageNumber="310">Epicoleotichus</emphasis>
|
|
and
|
|
<emphasis id="B95DEA8BFF9CFFBEFC56FF79C04EFEC3" italics="true" pageId="35" pageNumber="310">Para-coleotichus</emphasis>
|
|
</taxonomicName>
|
|
) on the basis of the prosternal shape, declivity of the male pygophore, and the shape of the female genital plates.
|
|
<bibRefCitation id="EFB84B68FF9CFFBEFDD0FEF0C09BFE7E" author="GROSS G. F." box="[532,674,380,403]" journalOrPublisher="A. B. James, Adelaide" pageId="35" pageNumber="310" pagination="1 - 250" refId="ref84925" refString="GROSS G. F. (1975): Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1. A. B. James, Adelaide: 1 - 250, 3 colour pls." title="Plant-feeding and other bugs (Hemiptera) of South Australia. Heteroptera - Part 1" type="book chapter" year="1975">
|
|
<smallCapsWord id="8D70A045FF9CFFBEFDD0FEF0C06FFE7F" baselines="397,398" box="[532,598,380,403]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Gross" pageId="35" pageNumber="310">GROSS</smallCapsWord>
|
|
(1975)
|
|
</bibRefCitation>
|
|
redescribed
|
|
<taxonomicName id="4C294D1AFF9CFFBEFCE2FEF0C1A8FE7E" box="[806,913,380,403]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="310" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9CFFBEFCE2FEF0C1A8FE7E" box="[806,913,380,403]" italics="true" pageId="35" pageNumber="310">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
, and used these subgeneric categories, giving credence to the shape of the prosternum in relation to the eyes. Our observations indicate that differences in these characters are more indicative of species relationships, and not diagnostic at the genus level.
|
|
</paragraph>
|
|
<paragraph id="8B963699FF9CFFBEFDFEFDF4C0E1FB84" blockId="35.[532,966,110,1918]" pageId="35" pageNumber="310">
|
|
<bibRefCitation id="EFB84B68FF9CFFBEFDFEFDF4C152FD62" author="MCDONALD F. J. D. & G. CASSIS" box="[570,875,632,655]" journalOrPublisher="Aust. J. Zool." pageId="35" pageNumber="310" pagination="537 - 572" part="32" refId="ref87212" refString="MCDONALD F. J. D. & G. CASSIS (1984): Revision of the Australian Scutelleridae LEACH (Hemiptera). - Aust. J. Zool. 32: 537 - 572." title="Revision of the Australian Scutelleridae LEACH (Hemiptera)" type="journal article" year="1984">MCDONALD & CASSIS (1984)</bibRefCitation>
|
|
added a description of the male genitalia for
|
|
<taxonomicName id="4C294D1AFF9CFFBEFDD0FD30C046FD3E" box="[532,639,700,723]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="310" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9CFFBEFDD0FD30C046FD3E" box="[532,639,700,723]" italics="true" pageId="35" pageNumber="310">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
, and did not adhere to subgeneric categories, without any explicit explanation. Pending a more detailed comparative morphological investigation and phylogenetic analysis of
|
|
<taxonomicName id="4C294D1AFF9CFFBEFD1CFCCFC17AFCB7" box="[728,835,835,858]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="310" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9CFFBEFD1CFCCFC17AFCB7" box="[728,835,835,858]" italics="true" pageId="35" pageNumber="310">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
, the sisterspecies relationships are uncertain, and we know of no new evidence that supports the use of subgeneric categories. Aside from the Australian species, the male and female genitalia of the extralimital species of
|
|
<taxonomicName id="4C294D1AFF9CFFBEFDD0FB82C046FBC8" box="[532,639,1038,1061]" class="Insecta" family="Pentatomidae" genus="Coleotichus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="310" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95DEA8BFF9CFFBEFDD0FB82C046FBC8" box="[532,639,1038,1061]" italics="true" pageId="35" pageNumber="310">Coleotichus</emphasis>
|
|
</taxonomicName>
|
|
have not been documented; a necessary prerequisite for any further classificatory hypotheses.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |