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<document id="849A9072B2A5641DB8D410F477587A5C" ID-DOI="10.1080/00222933.2017.1324050" ID-ISSN="1464-5262" ID-Zenodo-Dep="5181888" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628698566541" checkinUser="felipe" docAuthor="Schmitter-Soto, Juan J." docDate="2017" docId="03806F39C918FFB6FE58FC6BD751FCFC" docLanguage="en" docName="JNatHist.51.23-24.1331-1424.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 51 (21 - 24)" docSource="http://dx.doi.org/10.1080/00222933.2017.1324050" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Astyanax tamiahua Schmitter-Soto 2017, sp. nov." docType="treatment" docVersion="4" lastPageNumber="1411" masterDocId="FFB91741C957FFE7FFBEFFB5D316FFA7" masterDocTitle="A revision of Astyanax (Characiformes: Characidae) in Central and North America, with the description of nine new species" masterLastPageNumber="1424" masterPageNumber="1331" pageNumber="1409" updateTime="1699050015946" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B1952ACB4EDB4B476B748CC086B8884D">A revision of Astyanax (Characiformes: Characidae) in Central and North America, with the description of nine new species</mods:title>
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<mods:namePart id="987ED09FB1235D5EB2905EE1165E489F">Schmitter-Soto, Juan J.</mods:namePart>
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<mods:date id="F1E5C0B948B23018B7EC3AF3A9C01B3D">2017</mods:date>
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<mods:number id="53B2768351FDE393B660A3EFD4A30825">2017-07-03</mods:number>
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<treatment id="03806F39C918FFB6FE58FC6BD751FCFC" ID-DOI="http://doi.org/10.5281/zenodo.5191164" ID-Zenodo-Dep="5191164" LSID="urn:lsid:plazi:treatment:03806F39C918FFB6FE58FC6BD751FCFC" httpUri="http://treatment.plazi.org/id/03806F39C918FFB6FE58FC6BD751FCFC" lastPageId="81" lastPageNumber="1411" pageId="79" pageNumber="1409">
<subSubSection id="C3338DA4C918FFA8FE58FC6BD02BFC5F" box="[486,829,990,1016]" pageId="79" pageNumber="1409" type="nomenclature">
<paragraph id="8B96DE2FC918FFA8FE58FC6BD02BFC5F" blockId="79.[486,829,990,1051]" box="[486,829,990,1016]" pageId="79" pageNumber="1409">
<heading id="D0DE6943C918FFA8FE58FC6BD02BFC5F" box="[486,829,990,1016]" centered="true" fontSize="11" level="2" pageId="79" pageNumber="1409" reason="5">
<taxonomicName id="4C29A5ACC918FFA8FE58FC6BD1C1FC5F" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[486,727,990,1016]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="tamiahua" status="sp. nov.">
<emphasis id="B95D023DC918FFA8FE58FC6BD1C1FC5F" bold="true" box="[486,727,990,1016]" italics="true" pageId="79" pageNumber="1409">Astyanax tamiahua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EBF46C918FFA8FD5FFC6AD02BFC5F" box="[737,829,991,1016]" pageId="79" pageNumber="1409" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3338DA4C918FFA8FD88FBB7D196FBC3" pageId="79" pageNumber="1409" type="description">
<paragraph id="8B96DE2FC918FFA8FD88FBB7D1FBFBBC" blockId="79.[486,829,990,1051]" box="[566,749,1026,1051]" pageId="79" pageNumber="1409">
(
<figureCitation id="1312C2AAC918FFA8FD80FBB7D1ACFBBC" box="[574,698,1026,1051]" captionStart="Figure 12" captionStartId="15.[160,225,1445,1468]" captionTargetBox="[166,1150,742,1409]" captionTargetId="figure-57@15.[163,1153,738,1414]" captionTargetPageId="15" captionText="Figure 12. Map of Mexico, showing distribution localities of Astyanax acatlanensis sp. nov., A. argentatus, A. mexicanus, A. rioverde sp. nov., A. salvatoris sp. nov., A. tamiahua sp. nov. and A. tehuacanensis sp. nov. A larger symbol denotes the type locality (ambiguous for A. mexicanus; see text)." figureDoi="http://doi.org/10.5281/zenodo.5181913" httpUri="https://zenodo.org/record/5181913/files/figure.png" pageId="79" pageNumber="1409">Figures 12</figureCitation>
,
<figureCitation id="1312C2AAC918FFA8FD77FBB7D1F0FBBC" box="[713,742,1026,1051]" captionStart="Figure 46" captionStartId="80.[162,227,1728,1751]" captionTargetBox="[164,1154,1289,1697]" captionTargetId="figure-378@80.[164,1154,1289,1697]" captionTargetPageId="80" captionText="Figure 46. Astyanax tamiahua sp. nov., holotype, UMMZ 250294." figureDoi="http://doi.org/10.5281/zenodo.5181967" httpUri="https://zenodo.org/record/5181967/files/figure.png" pageId="79" pageNumber="1409">46</figureCitation>
)
</paragraph>
<paragraph id="8B96DE2FC918FFA8FF1EFBF9D196FBC3" blockId="79.[160,640,1100,1124]" box="[160,640,1100,1124]" pageId="79" pageNumber="1409">
<taxonomicName id="4C29A5ACC918FFA8FF1EFBF9D215FBC3" authorityName="Baird and Girard" authorityYear="1854" box="[160,259,1100,1124]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="genus">
<emphasis id="B95D023DC918FFA8FF1EFBF9D215FBC3" box="[160,259,1100,1124]" italics="true" pageId="79" pageNumber="1409">Astyanax</emphasis>
</taxonomicName>
Tamiahua,
<bibRefCitation id="EFB8A3DEC918FFA8FE2DFBF9D16CFBC3" author="Schmitter-Soto JJ" box="[403,634,1100,1124]" pageId="79" pageNumber="1409" pagination="101 - 130" refId="ref49172" refString="Schmitter-Soto JJ. 2016. A phylogeny of Astyanax (Characiformes: Characidae) in Central and North America. Zootaxa. 4109: 101 - 130." type="journal article" year="2016">Schmitter-Soto 2016</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3338DA4C918FFA8FF1EFB24D71AFADD" pageId="79" pageNumber="1409" type="diagnosis">
<paragraph id="8B96DE2FC918FFA8FF1EFB24D202FB0D" blockId="79.[160,1156,1169,1402]" box="[160,276,1169,1194]" pageId="79" pageNumber="1409">
<emphasis id="B95D023DC918FFA8FF1EFB24D202FB0D" bold="true" box="[160,276,1169,1194]" italics="true" pageId="79" pageNumber="1409">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B96DE2FC918FFA8FF1EFB01D71AFADD" blockId="79.[160,1156,1169,1402]" pageId="79" pageNumber="1409">
Diagnosed from other
<taxonomicName id="4C29A5ACC918FFA8FE23FB01D140FB6B" box="[413,598,1204,1228]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B95D023DC918FFA8FE23FB01D2E8FB6B" box="[413,510,1204,1228]" italics="true" pageId="79" pageNumber="1409">Astyanax</emphasis>
species
</taxonomicName>
in
<collectingRegion id="49ED10CDC918FFA8FDC7FB01D1CEFB6B" box="[633,728,1204,1228]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="79" pageNumber="1409">Veracruz</collectingRegion>
and northern
<collectingCountry id="F33E9EBFC918FFA8FCC4FB01D0DFFB6B" box="[890,969,1204,1228]" name="Mexico" pageId="79" pageNumber="1409">Mexico</collectingCountry>
as follows: body deeper, mean 35.9% SL (mean 33.9% SL in
<taxonomicName id="4C29A5ACC918FFA8FD3EFB62D006FB48" authorityName="Baird and Girard" authorityYear="1854" box="[640,784,1239,1263]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="B95D023DC918FFA8FD3EFB62D006FB48" box="[640,784,1239,1263]" italics="true" pageId="79" pageNumber="1409">A. argentatus</emphasis>
</taxonomicName>
); head longer, more than 26% SL (26% or less in
<taxonomicName id="4C29A5ACC918FFA8FEF1FB4FD2E6FAB5" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[335,496,1274,1298]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="acatlanensis" status="sp. nov.">
<emphasis id="B95D023DC918FFA8FEF1FB4FD2E6FAB5" box="[335,496,1274,1298]" italics="true" pageId="79" pageNumber="1409">A. acatlanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EBF46C918FFA8FE47FB4FD144FAB5" box="[505,594,1274,1298]" pageId="79" pageNumber="1409" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C29A5ACC918FFA8FDE0FB4FD1F8FAB5" authorityName="Baird and Girard" authorityYear="1854" box="[606,750,1274,1298]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="B95D023DC918FFA8FDE0FB4FD1F8FAB5" box="[606,750,1274,1298]" italics="true" pageId="79" pageNumber="1409">A. argentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C29A5ACC918FFA8FC94FB4FD08BFAB5" authority="Schmitter-Soto, 2017" authorityName="Lozano-Vilano and Schmitter-Soto" authorityYear="2017" box="[810,925,1274,1298]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="rioverde" status="sp. nov.">
<emphasis id="B95D023DC918FFA8FC94FB4FD08BFAB5" box="[810,925,1274,1298]" italics="true" pageId="79" pageNumber="1409">A. rioverde</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EBF46C918FFA8FC18FB4FD716FAB5" box="[934,1024,1274,1298]" pageId="79" pageNumber="1409" rank="species">sp. nov.</taxonomicNameLabel>
); supraoccipital in lateral view, slightly concave (angled in
<emphasis id="B95D023DC918FFA8FD09FAA9D039FA93" box="[695,815,1308,1332]" italics="true" pageId="79" pageNumber="1409">
A.
<emphasis id="B95D023DC918FFA8FD6AFAA9D1F7FA93" bold="true" box="[724,737,1308,1332]" italics="true" pageId="79" pageNumber="1409">fi</emphasis>
nitimus
</emphasis>
); infraorbital II, triangular with an angled base (with a long convex base in
<taxonomicName id="4C29A5ACC918FFA8FD22FA8AD03BFAF0" authorityName="Baird and Girard" authorityYear="1854" box="[668,813,1343,1367]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="B95D023DC918FFA8FD22FA8AD03BFAF0" box="[668,813,1343,1367]" italics="true" pageId="79" pageNumber="1409">A. argentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C29A5ACC918FFA8FCD7FA8AD0E1FAF0" authorityName="? Regan" authorityYear="1908" box="[873,1015,1343,1367]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="B95D023DC918FFA8FCD7FA8AD0E1FAF0" box="[873,1015,1343,1367]" italics="true" pageId="79" pageNumber="1409">A. mexicanus</emphasis>
</taxonomicName>
); first dorsal pterygiophore, rostral edge angled (curved in
<taxonomicName id="4C29A5ACC918FFA8FD18FAD7D020FADD" authorityName="Baird and Girard" authorityYear="1854" box="[678,822,1378,1402]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="argentatus">
<emphasis id="B95D023DC918FFA8FD18FAD7D020FADD" box="[678,822,1378,1402]" italics="true" pageId="79" pageNumber="1409">A. argentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C29A5ACC918FFA8FCCCFAD7D0E9FADD" authorityName="? Regan" authorityYear="1908" box="[882,1023,1378,1402]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="B95D023DC918FFA8FCCCFAD7D0E9FADD" box="[882,1023,1378,1402]" italics="true" pageId="79" pageNumber="1409">A. mexicanus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3338DA4C918FFB7FF1EFA13D2FCFB34" lastPageId="80" lastPageNumber="1410" pageId="79" pageNumber="1409" type="description">
<paragraph id="8B96DE2FC918FFA8FF1EFA13D230FA18" blockId="79.[160,1157,1446,1714]" box="[160,294,1446,1471]" pageId="79" pageNumber="1409">
<emphasis id="B95D023DC918FFA8FF1EFA13D230FA18" bold="true" box="[160,294,1446,1471]" italics="true" pageId="79" pageNumber="1409">Description</emphasis>
</paragraph>
<paragraph id="8B96DE2FC918FFA8FF1EFA7FD793FA45" blockId="79.[160,1157,1446,1714]" box="[160,1157,1482,1506]" pageId="79" pageNumber="1409">
A species of
<taxonomicName id="4C29A5ACC918FFA8FE92FA7FD299FA45" authorityName="Baird and Girard" authorityYear="1854" box="[300,399,1482,1506]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="genus">
<emphasis id="B95D023DC918FFA8FE92FA7FD299FA45" box="[300,399,1482,1506]" italics="true" pageId="79" pageNumber="1409">Astyanax</emphasis>
</taxonomicName>
, subgenus
<taxonomicName id="4C29A5ACC918FFA8FDB1FA7FD164FA45" authorityName="Baird and Girard" authorityYear="1854" box="[527,626,1482,1506]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="79" pageNumber="1409" phylum="Chordata" rank="genus">
<emphasis id="B95D023DC918FFA8FDB1FA7FD164FA45" box="[527,626,1482,1506]" italics="true" pageId="79" pageNumber="1409">Astyanax</emphasis>
</taxonomicName>
(i.e. with a complete predorsal series of scales).
</paragraph>
<paragraph id="8B96DE2FC918FFA8FF7EFA59D06CF9ED" blockId="79.[160,1157,1446,1714]" pageId="79" pageNumber="1409">Head profile approximately straight at nape; snout rounded; lower and upper lips even, mouth terminal. Pectoral fins usually reach posteriorly to pelvic fin origin; anal and dorsal fins may overlap vertically. Lobes of caudal fin, subequal.</paragraph>
<paragraph id="8B96DE2FC918FFB7FF7EF9E1D103FEA9" blockId="79.[160,1157,1446,1714]" lastBlockId="80.[160,1156,142,1171]" lastPageId="80" lastPageNumber="1410" pageId="79" pageNumber="1409">
D. 1011; A. 2025, modally 22; pect. 1011. Procurrent rays on dorsal caudal fin, 9 or fewer. Gill rakers on first arch, 1720, modally 19; on lower limb, 1112. Scales in lateral line, 3034, modally 32; predorsal scales, 912, modally 10; scale rows from lateral line to base of first dorsal-fin ray, 7; to base of pelvic fin, 5; to base of pectoral fin, modally 4, as few as 3; circumpeduncular scales, 1518, modally 16. A short, single scale row on anal fin base. Nuptial tubercles may be bifid. Total vertebrae 3133, 1819 caudal. Detailed frequencies are given in
<tableCitation id="C6ABEB94C907FFB7FE05FF43D119FEA9" box="[443,527,246,270]" captionStart="Table 3" captionStartId="16.[160,216,142,165]" captionText="Table 3. Meristics of Central and North American species in the genus Astyanax. Numbers are" pageId="80" pageNumber="1410">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8B96DE2FC907FFB7FF7EFEACD1A1FED1" blockId="80.[160,1156,142,1171]" pageId="80" pageNumber="1410">
Largest examined specimen,
<quantity id="4CD173CAC907FFB7FDA4FEACD194FE96" box="[538,642,281,305]" metricMagnitude="-2" metricUnit="m" metricValue="6.27" pageId="80" pageNumber="1410" unit="mm" value="62.7">62.7 mm</quantity>
SL. Body depth, 3338% SL. Head length, 2628% SL; orbital diameter, 3037% HL; interorbital distance, 8.211.1% SL, mean 9.3 (further morphometric data appear in
<tableCitation id="C6ABEB94C907FFB7FDE9FEEBD1BDFED1" box="[599,683,350,374]" captionStart="Table 4" captionStartId="19.[160,216,137,160]" captionText="Table 4. Selected morphometric proportions of Central and North American species in the genus Astyanax (abbreviated by first letters of specific epithet). Specimens measured are those marked dig in Appendix. All measurements given as percentage of standard length. Data are means; whole range in parentheses." pageId="80" pageNumber="1410">Table 4</tableCitation>
).
</paragraph>
<paragraph id="8B96DE2FC907FFB7FF7EFE34D281FBE9" blockId="80.[160,1156,142,1171]" pageId="80" pageNumber="1410">Anterior fontanel short, convex-sided. Supraoccipital process in dorsal view, long, narrow-based, slightly concave in lateral view. Vomer rostrally slightly concave. Arms of premaxilla, subequal; 04 teeth. Highest tooth on dentary, first or third; posterior teeth, abruptly smaller. Articular, dorsal arm, angled. Maxillary teeth, 13. Quadrate, dorsal process not expanded. Metapterygoid, rostral arm longer than ventral. Infraorbital II, a triangle with an angled base; infraorbital III, inferoposteriorly semicircular; infraorbital IV, square. Urohyal rostral end turned up, its ventrorostral edge convex, its ventral apex about equidistant between rostral and caudal end; ceratohyal rostral vertices angled. Epibranchial III, distal segment of the main body, straight. Lower pharyngeal teeth plate, single; its caudal side concave. Dorsal side of hyomandibular, convex. Opercle, sides of dorsal half, parallel; caudoventrally straight-convex, caudodorsally concave; ventral tip, sharp. Interopercular posterior edge, straight-convex, with a spine. Preopercular ventral rim, straight; posteroventral edge, curved; 2 canals at angle. Five predorsal bony elements; rostral edge of first pterygiophore angled. Coracoid with 2 interdigitations in suture to cleithrum, a concave caudal edge. Caudad process of postcleithrum, with parallel sides and convex end. Proximal edge of pelvic bone, convex. Postanal element, short. Caudal tip of scaphium, spiny; its dorsoposterior edge, slightly concave. Neural spines under dorsal fin, straight. Sixth caudal vertebra from tail, with a haemal spine displaced caudad. Rostral edge of largest hypuric plate, concave. Epuric plate on last neural spine, convex-edged.</paragraph>
<paragraph id="8B96DE2FC907FFB7FF7EFBECD2FCFB34" blockId="80.[160,1156,142,1171]" pageId="80" pageNumber="1410">Humeral spot, squarish. Pigment on anal fin, concentrated distally. Caudal spot, both on peduncle and on fin rays.</paragraph>
</subSubSection>
<caption id="DF568EA7C907FFB7FF1CF975D026F970" ID-DOI="http://doi.org/10.5281/zenodo.5181967" ID-Zenodo-Dep="5181967" box="[162,816,1728,1751]" httpUri="https://zenodo.org/record/5181967/files/figure.png" pageId="80" pageNumber="1410" startId="80.[162,227,1728,1751]" targetBox="[164,1154,1289,1697]" targetPageId="80">
<paragraph id="8B96DE2FC907FFB7FF1CF975D026F970" blockId="80.[162,816,1728,1751]" box="[162,816,1728,1751]" pageId="80" pageNumber="1410">
<emphasis id="B95D023DC907FFB7FF1CF975D21CF970" bold="true" box="[162,266,1728,1751]" pageId="80" pageNumber="1410">Figure 46.</emphasis>
<taxonomicName id="4C29A5ACC907FFB7FEADF975D2D9F970" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[275,463,1728,1751]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="80" pageNumber="1410" phylum="Chordata" rank="species" species="tamiahua" status="sp. nov.">
<emphasis id="B95D023DC907FFB7FEADF975D2D9F970" box="[275,463,1728,1751]" italics="true" pageId="80" pageNumber="1410">Astyanax tamiahua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EBF46C907FFB7FE69F975D13EF970" box="[471,552,1728,1751]" pageId="80" pageNumber="1410" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, UMMZ 250294.
</paragraph>
</caption>
<subSubSection id="C3338DA4C906FFB6FF1EFF38D172FE96" pageId="81" pageNumber="1411" type="materials_examined">
<paragraph id="8B96DE2FC906FFB6FF1EFF38D2E2FF01" blockId="81.[160,1156,141,305]" box="[160,500,141,166]" pageId="81" pageNumber="1411">
<emphasis id="B95D023DC906FFB6FF1EFF38D2E2FF01" bold="true" box="[160,500,141,166]" italics="true" pageId="81" pageNumber="1411">Type material and depositor</emphasis>
</paragraph>
<paragraph id="8B96DE2FC906FFB6FF1EFF04D172FE96" blockId="81.[160,1156,141,305]" pageId="81" pageNumber="1411">
<materialsCitation id="3B41D472C906FFB6FF1EFF04D157FF4B" collectingDate="1930-04" collectionCode="UMMZ" country="Mexico" location="Rio Cucharas" municipality="Tamiahua" pageId="81" pageNumber="1411" specimenCode="UMMZ 250294" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<typeStatus id="5492608DC906FFB6FF1EFF04D211FF6E" box="[160,263,177,201]" pageId="81" pageNumber="1411" type="holotype">Holotype</typeStatus>
<specimenCode id="DB8F7654C906FFB6FEAEFF04D2ACFF6E" box="[272,442,177,201]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="81" pageNumber="1411" type="University or college">UMMZ 250294</specimenCode>
,
<quantity id="4CD173CAC906FFB6FE78FF04D13CFF6E" box="[454,554,177,201]" metricMagnitude="-2" metricUnit="m" metricValue="5.42" pageId="81" pageNumber="1411" unit="mm" value="54.2">54.2 mm</quantity>
SL,
<location id="8EF688F4C906FFB6FDE3FF04D1F9FF6E" LSID="urn:lsid:plazi:treatment:03806F39C918FFB6FE58FC6BD751FCFC:8EF688F4C906FFB6FDE3FF04D1F9FF6E" box="[605,751,177,201]" country="Mexico" municipality="Tamiahua" name="Rio Cucharas" pageId="81" pageNumber="1411" stateProvince="Veracruz de Ignacio de la Llave">Río Cucharas</location>
near
<collectingMunicipality id="6BF24455C906FFB6FC8AFF04D0B3FF6E" box="[820,933,177,201]" pageId="81" pageNumber="1411">Tamiahua</collectingMunicipality>
, northern
<collectingRegion id="49ED10CDC906FFB6FBA3FF04D796FF6E" box="[1053,1152,177,201]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="81" pageNumber="1411">Veracruz</collectingRegion>
,
<collectingCountry id="F33E9EBFC906FFB6FF1EFF61D3E5FF4B" box="[160,243,212,236]" name="Mexico" pageId="81" pageNumber="1411">Mexico</collectingCountry>
, coll.
<date id="FF97F8EFC906FFB6FE82FF61D2A3FF4B" box="[316,437,212,236]" pageId="81" pageNumber="1411" value="1930-04">
<collectingDate id="EFD30107C906FFB6FE82FF61D2A3FF4B" box="[316,437,212,236]" pageId="81" pageNumber="1411" value="1930-04">April 1930</collectingDate>
</date>
(
<figureCitation id="1312C2AAC906FFB6FE77FF61D121FF4B" box="[457,567,212,236]" captionStart="Figure 46" captionStartId="80.[162,227,1728,1751]" captionTargetBox="[164,1154,1289,1697]" captionTargetId="figure-378@80.[164,1154,1289,1697]" captionTargetPageId="80" captionText="Figure 46. Astyanax tamiahua sp. nov., holotype, UMMZ 250294." figureDoi="http://doi.org/10.5281/zenodo.5181967" httpUri="https://zenodo.org/record/5181967/files/figure.png" pageId="81" pageNumber="1411">Figure 46</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3B41D472C906FFB6FDECFF61D2F6FEA9" collectingDate="1930-04" collectionCode="UMMZ" country="Mexico" location="Rio Cucharas" municipality="Tamiahua" pageId="81" pageNumber="1411" specimenCode="UMMZ 97362" specimenCount="103" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<typeStatus id="5492608DC906FFB6FDECFF61D1D4FF4B" box="[594,706,212,236]" pageId="81" pageNumber="1411" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="DB8F7654C906FFB6FD6FFF61D07BFF4B" box="[721,877,212,236]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="81" pageNumber="1411" type="University or college">UMMZ 97362</specimenCode>
(
<specimenCount id="9D2F15A6C906FFB6FC3FFF61D725FF4B" box="[897,1075,212,236]" pageId="81" pageNumber="1411" type="generic">103 specimens</specimenCount>
), same collection data as holotype
</materialsCitation>
.
<materialsCitation id="3B41D472C906FFB6FE51FF43D281FE96" collectionCode="UMMZ" country="Mexico" location="Rio Nautla" municipality="Ixtacocoa" pageId="81" pageNumber="1411" specimenCode="UMMZ 167489" specimenCount="100" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<specimenCode id="DB8F7654C906FFB6FE51FF43D181FEA9" box="[495,663,246,270]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="81" pageNumber="1411" type="University or college">UMMZ 167489</specimenCode>
(
<specimenCount id="9D2F15A6C906FFB6FD17FF43D04CFEA9" box="[681,858,246,270]" pageId="81" pageNumber="1411" type="generic">100 specimens</specimenCount>
),
<location id="8EF688F4C906FFB6FCD0FF43D0F0FEA9" LSID="urn:lsid:plazi:treatment:03806F39C918FFB6FE58FC6BD751FCFC:8EF688F4C906FFB6FCD0FF43D0F0FEA9" box="[878,998,246,270]" country="Mexico" municipality="Ixtacocoa" name="Rio Nautla" pageId="81" pageNumber="1411" stateProvince="Veracruz de Ignacio de la Llave">Río Nautla</location>
at
<collectingMunicipality id="6BF24455C906FFB6FBADFF43D796FEA9" box="[1043,1152,246,270]" pageId="81" pageNumber="1411">Ixtacocoa</collectingMunicipality>
,
<collectingRegion id="49ED10CDC906FFB6FF1EFEACD215FE96" box="[160,259,281,305]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="81" pageNumber="1411">Veracruz</collectingRegion>
,
<collectingCountry id="F33E9EBFC906FFB6FEB1FEACD274FE96" box="[271,354,281,305]" name="Mexico" pageId="81" pageNumber="1411">Mexico</collectingCountry>
, coll
</materialsCitation>
.
<date id="FF97F8EFC906FFB6FE1DFEACD176FE96" box="[419,608,281,305]" pageId="81" pageNumber="1411" value="1951-09">September 1951</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3338DA4C906FFB6FF1EFEE8D1E3FE3E" box="[160,757,349,409]" pageId="81" pageNumber="1411" type="etymology">
<paragraph id="8B96DE2FC906FFB6FF1EFEE8D20BFED1" blockId="81.[160,757,349,409]" box="[160,285,349,374]" pageId="81" pageNumber="1411">
<emphasis id="B95D023DC906FFB6FF1EFEE8D20BFED1" bold="true" box="[160,285,349,374]" italics="true" pageId="81" pageNumber="1411">Etymology</emphasis>
</paragraph>
<paragraph id="8B96DE2FC906FFB6FF1EFE34D1E3FE3E" blockId="81.[160,757,349,409]" box="[160,757,385,409]" pageId="81" pageNumber="1411">
Named after the
<typeStatus id="5492608DC906FFB6FEDAFE34D280FE3E" box="[356,406,385,409]" pageId="81" pageNumber="1411">type</typeStatus>
locality. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3338DA4C906FFB6FF1EFE70D14AFDCE" pageId="81" pageNumber="1411" type="distribution">
<paragraph id="8B96DE2FC906FFB6FF1EFE70D23AFE79" blockId="81.[160,833,453,513]" box="[160,300,453,478]" pageId="81" pageNumber="1411">
<emphasis id="B95D023DC906FFB6FF1EFE70D23AFE79" bold="true" box="[160,300,453,478]" italics="true" pageId="81" pageNumber="1411">Distribution</emphasis>
</paragraph>
<paragraph id="8B96DE2FC906FFB6FF1EFE5CD057FDA6" blockId="81.[160,833,453,513]" box="[160,833,489,513]" pageId="81" pageNumber="1411">
Tamiahua to Nautla, northern
<collectingRegion id="49ED10CDC906FFB6FE44FE5CD14BFDA6" box="[506,605,489,513]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="81" pageNumber="1411">Veracruz</collectingRegion>
,
<collectingCountry id="F33E9EBFC906FFB6FDD7FE5CD1AFFDA6" box="[617,697,489,513]" name="Mexico" pageId="81" pageNumber="1411">Mexico</collectingCountry>
(
<figureCitation id="1312C2AAC906FFB6FD77FE5CD022FDA6" box="[713,820,489,513]" captionStart="Figure 12" captionStartId="15.[160,225,1445,1468]" captionTargetBox="[166,1150,742,1409]" captionTargetId="figure-57@15.[163,1153,738,1414]" captionTargetPageId="15" captionText="Figure 12. Map of Mexico, showing distribution localities of Astyanax acatlanensis sp. nov., A. argentatus, A. mexicanus, A. rioverde sp. nov., A. salvatoris sp. nov., A. tamiahua sp. nov. and A. tehuacanensis sp. nov. A larger symbol denotes the type locality (ambiguous for A. mexicanus; see text)." figureDoi="http://doi.org/10.5281/zenodo.5181913" httpUri="https://zenodo.org/record/5181913/files/figure.png" pageId="81" pageNumber="1411">Figure 12</figureCitation>
).
</paragraph>
<paragraph id="8B96DE2FC906FFB6FF1EFD98D2C0FDE1" blockId="81.[160,604,557,617]" box="[160,470,557,582]" pageId="81" pageNumber="1411">
<emphasis id="B95D023DC906FFB6FF1EFD98D2C0FDE1" bold="true" box="[160,470,557,582]" italics="true" pageId="81" pageNumber="1411">Proposed common names</emphasis>
</paragraph>
<paragraph id="8B96DE2FC906FFB6FF1EFDE4D14AFDCE" blockId="81.[160,604,557,617]" box="[160,604,593,617]" pageId="81" pageNumber="1411">Tamiahua tetra, sardinita de Tamiahua.</paragraph>
</subSubSection>
<subSubSection id="C3338DA4C906FFB6FF1EFD20D751FCFC" pageId="81" pageNumber="1411" type="discussion">
<paragraph id="8B96DE2FC906FFB6FF1EFD20D210FD09" blockId="81.[160,1155,661,859]" box="[160,262,661,686]" pageId="81" pageNumber="1411">
<emphasis id="B95D023DC906FFB6FF1EFD20D210FD09" bold="true" box="[160,262,661,686]" italics="true" pageId="81" pageNumber="1411">Remarks</emphasis>
</paragraph>
<paragraph id="8B96DE2FC906FFB6FF1EFD0CD026FCB1" blockId="81.[160,1155,661,859]" pageId="81" pageNumber="1411">
The southern population at Nautla may reach 25 anal-fin rays and 34 scales on lateral line, whereas the
<typeStatus id="5492608DC906FFB6FED1FD6ED2B7FD54" box="[367,417,731,755]" pageId="81" pageNumber="1411">type</typeStatus>
population near Tamiahua does not exceed 23 and 33, respectively; pectoral fins are also longer in the northern range.
</paragraph>
<paragraph id="8B96DE2FC906FFB6FF7EFC94D751FCFC" blockId="81.[160,1155,661,859]" pageId="81" pageNumber="1411">
The species, represented by both populations, is a part of clade Ia of
<bibRefCitation id="EFB8A3DEC906FFB6FC5EFC94D23EFCFC" author="Ornelas-Garcia CP &amp; Dominguez-Dominguez O &amp; Doadrio I" pageId="81" pageNumber="1411" pagination="340" refId="ref48693" refString="Ornelas-Garcia CP, Dominguez-Dominguez O, Doadrio I. 2008. Evolutionary history of the fish genus Astyanax Baird and Girard 1854 (Actinopterygii, Characidae) in Mesoamerica reveals multiple morphological homoplasies. BMC Evol Biol. 8: 340." type="journal article" year="2008">Ornelas-García et al. (2008)</bibRefCitation>
, in a subgroup different from from that of the inland northern forms.
</paragraph>
</subSubSection>
</treatment>
</document>