treatments-xml/data/07/2D/87/072D87D2FFCDFF8841DFFD3AE082FDD7.xml
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<document id="7E77971D5795E0067948CE246DB76F9B" ID-CLB-Dataset="29242" ID-DOI="10.11646/zootaxa.4496.1.42" ID-GBIF-Dataset="895f6990-113a-4193-a279-7dfbcd8be850" ID-ISSN="1175-5326" ID-Zenodo-Dep="1447025" ID-ZooBank="F0E7B997-D3E2-45F9-9DB8-1CE60196FEC3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538748284745" checkinUser="plazi" docAuthor="Bartz, Marie L. C., James, Samuel W., Steffen, Gerusa P. K., Antoniolli, Zaida, Steffen, Ricardo B. &amp; Brown, George G." docDate="2018" docId="072D87D2FFCDFF8841DFFD3AE082FDD7" docLanguage="en" docName="zootaxa.4496.1.42.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glossoscolex GlossosColex, n. sp." docType="treatment" docVersion="7" lastPageNumber="553" masterDocId="FB14FFAAFFC9FF8D4148FFEDE35AFFEA" masterDocTitle="New species-group taxa of Glossoscolex (Clitellata: Glossoscolecidae) from Rio Grande do Sul, Brazil" masterLastPageNumber="561" masterPageNumber="548" pageNumber="552" updateTime="1698566898224" updateUser="ExternalLinkService">
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<mods:title id="F40C33EFD864B2C80A3767A0712168AD">New species-group taxa of Glossoscolex (Clitellata: Glossoscolecidae) from Rio Grande do Sul, Brazil</mods:title>
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<mods:namePart id="03A3444646FEFA18B6083453E2D16E44">Bartz, Marie L. C.</mods:namePart>
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<mods:namePart id="DD05EF71EA0BE3EEB894F6641CC1317F">James, Samuel W.</mods:namePart>
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<mods:namePart id="E6C9635D2FEEC1B2C393A1749A415C10">Steffen, Gerusa P. K.</mods:namePart>
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<mods:namePart id="6CCF6F91CEDFC73FBFB405DD77D2E802">Antoniolli, Zaida</mods:namePart>
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<mods:namePart id="E363DA14F520AF96D86451B92D051D8E">Steffen, Ricardo B.</mods:namePart>
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<mods:namePart id="076ECD4C1E3FA28C349A3ADC3B52E1ED">Brown, George G.</mods:namePart>
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<mods:title id="5B9CDEE30D48DB83DCA151DD51F7D726">Zootaxa</mods:title>
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<mods:date id="FEC2392FF6439803CD0BBBCE2B77B7E3">2018</mods:date>
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<emphasis id="BDF0EAD6FFCDFF8941DFFD3AE74EFD18" bold="true" box="[151,1044,727,754]" pageId="4" pageNumber="552">
<taxonomicName id="48844D47FFCDFF8941DFFD3AE2B0FD1B" authority="(GlossosColex)" authorityName="GlossosColex" box="[151,490,727,753]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="genus">
<emphasis id="BDF0EAD6FFCDFF8941DFFD3AE2B0FD1B" bold="true" box="[151,490,727,753]" italics="true" pageId="4" pageNumber="552">GlossosColex (GlossosColex)</emphasis>
</taxonomicName>
<taxonomicName id="48844D47FFCDFF8940BAFD3AE09EFD18" authority="Bartz &amp; James" authorityName="Bartz &amp; James &amp; Steffen &amp; Antoniolli &amp; Steffen &amp; Brown" authorityYear="2018" box="[498,964,727,754]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="subSpecies" species="riograndensis" status="sp. nov." subSpecies="pollulus">
<emphasis id="BDF0EAD6FFCDFF8940BAFD3AE05BFD1B" bold="true" box="[498,769,727,753]" italics="true" pageId="4" pageNumber="552">riograndensis pollulus</emphasis>
Bartz &amp; James
</taxonomicName>
,
<taxonomicNameLabel id="A6C357ADFFCDFF894299FD35E74EFD18" box="[977,1044,728,754]" pageId="4" pageNumber="552" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="17BF2A41FFCDFF8941D6FD17E26AFCF9" box="[158,304,762,787]" captionStart="FIGURE 1" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[307,1259,233,1868]" captionTargetId="figure@3.[190,1406,193,1912]" captionTargetPageId="3" captionText="FIGURE 1. Glossoscolex (Glossoscolex) riograndensis n. sp.. A, B. Glossoscolex (Glossoscolex) riograndensis riograndensis. C, D. Glossoscolex (Glossoscolex) riograndensis nativus n. ssp. E, F. Glossoscolex (Glossoscolex) riograndensis pollulus n. ssp. FM = female pOres; MP = male pOres; NF = nephridial pOres." httpUri="https://zenodo.org/record/1447027/files/figure.png" pageId="4" pageNumber="552">Figures 1E,F</figureCitation>
,
<figureCitation id="17BF2A41FFCDFF894075FD16E223FCF9" box="[317,377,763,787]" captionStart="FIGURE 2" captionStartId="6.[151,250,1324,1346]" captionTargetBox="[151,1436,339,1302]" captionTargetId="figure@6.[151,1436,339,1302]" captionTargetPageId="6" captionText="FIGURE 2. Sperm-filled nephridia Of Glossoscolex (Glossoscolex) riograndensis n. sp., histOlOgical sectiOns. A, B. Glossoscolex (G.) riograndensis riograndensis n. sp., segment xii (A) and xiii (B). C, D. Glossoscolex (G.) riograndensis pollulus n. ssp., segments Vi (C) and Vii (D)." httpUri="https://zenodo.org/record/1447029/files/figure.png" pageId="4" pageNumber="552">2C,D</figureCitation>
)
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<typeStatus id="503F8866FFCDFF8941DFFCAFE25CFCB1" box="[151,262,834,859]" pageId="4" pageNumber="552" type="holotype">Holotype</typeStatus>
.
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<collectionCode id="E995AE01FFCDFF89405FFCAEE23CFCB1" box="[279,358,835,859]" pageId="4" pageNumber="552">COFM</collectionCode>
<specimenCode id="DF229EBFFFCDFF89403BFCAFE2A9FCB1" box="[371,499,834,859]" collectionCode="BRRS" pageId="4" pageNumber="552">BRRS0045</specimenCode>
one adult, fragment of natiVe forest,
<location id="8A5B601FFFCDFF8942F2FCAEE711FCB1" LSID="urn:lsid:plazi:treatment:072D87D2FFCDFF8841DFFD3AE082FDD7:8A5B601FFFCDFF8942F2FCAEE711FCB1" box="[954,1099,834,859]" country="Brazil" latitude="-29.66247" longLatPrecision="1" longitude="-53.91399" name="Santa Maria" pageId="4" pageNumber="552" stateProvince="Rio Grande do Sul">Santa Maria</location>
,
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Rio
<collectingRegion id="4D40F826FFCDFF8945D8FCAFE61AFCB1" box="[1168,1344,834,859]" inferredBySuffix="true" pageId="4" pageNumber="552">Grande do Sul</collectingRegion>
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,
<collectingCountry id="F7937654FFCDFF894418FCAFE6CDFCB1" box="[1360,1431,834,859]" name="Brazil" pageId="4" pageNumber="552">Brazil</collectingCountry>
,
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<geoCoordinate id="EAB05003FFCDFF894074FC8BE2BBFC95" box="[316,481,870,895]" direction="west" orientation="longitude" pageId="4" pageNumber="552" precision="1" value="-53.91399">53°54'50,36''O</geoCoordinate>
,
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.
<collectingDate id="EB7EE9ECFFCDFF894329FC8BE07AFC95" box="[609,800,870,895]" pageId="4" pageNumber="552" value="2009-10-26">
<date id="FB3A1004FFCDFF894329FC8BE07AFC95" box="[609,800,870,895]" pageId="4" pageNumber="552" value="2009-10-26">26 October 2009</date>
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,
<collectorName id="22715312FFCDFF894263FC8AE093FC95" box="[811,969,870,895]" pageId="4" pageNumber="552">G.P.K. Steffen</collectorName>
&amp;
<collectorName id="22715312FFCDFF8942A4FC8AE72DFC95" box="[1004,1143,870,895]" pageId="4" pageNumber="552">R.B. Steffen</collectorName>
colls.
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</paragraph>
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<typeStatus id="503F8866FFCDFF89418FFC67E26CFC49" box="[199,310,906,931]" pageId="4" pageNumber="552" type="paratype">Paratype</typeStatus>
.
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<collectionCode id="E995AE01FFCDFF894000FC66E147FC49" box="[328,541,906,931]" pageId="4" pageNumber="552">A—COFM BRRS</collectionCode>
0046 one adult. Same locality and collection data as
<typeStatus id="503F8866FFCDFF8945EFFC67E657FC49" box="[1191,1293,906,931]" pageId="4" pageNumber="552" type="holotype">holotype</typeStatus>
.
<collectionCode id="E995AE01FFCDFF894456FC66E384FC2D" pageId="4" pageNumber="552">B—COFM BRRS</collectionCode>
0047 one subadult. Same locality and collection data as
<typeStatus id="503F8866FFCDFF89421BFC43E0E3FC2D" box="[851,953,942,967]" pageId="4" pageNumber="552" type="holotype">holotype</typeStatus>
.
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<paragraph id="8F3B36C4FFCDFF89418FFC3FE0AFFBE5" blockId="4.[151,1437,834,2011]" pageId="4" pageNumber="552">
<materialsCitation id="3FEC3C99FFCDFF89418FFC3FE0AFFBE5" ID-GBIF-Occurrence="1914265845" collectingDate="2009-10-26" collectionCode="COFM" collectorName="M. P. Pontelli" country="Brazil" elevation="276" latitude="-29.66247" location="Other" longLatPrecision="1" longitude="-53.91399" municipality="Santa Maria" pageId="4" pageNumber="552" specimenCode="BRRS0048" specimenCount="1" stateProvince="Rio Grande do Sul">
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<location id="8A5B601FFFCDFF89418FFC3FE255FC01" LSID="urn:lsid:plazi:treatment:072D87D2FFCDFF8841DFFD3AE082FDD7:8A5B601FFFCDFF89418FFC3FE255FC01" box="[199,271,978,1003]" country="Brazil" latitude="-29.66247" longLatPrecision="1" longitude="-53.91399" municipality="Santa Maria" name="Other" pageId="4" pageNumber="552" stateProvince="Rio Grande do Sul">Other</location>
material.
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<collectionCode id="E995AE01FFCDFF8940C2FC3EE283FC01" box="[394,473,979,1003]" pageId="4" pageNumber="552">COFM</collectionCode>
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adult, fragment of natiVe forest,
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,
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Rio
<collectingRegion id="4D40F826FFCDFF8945D6FC3FE61EFC01" box="[1182,1348,978,1003]" inferredBySuffix="true" pageId="4" pageNumber="552">Grande do Sul</collectingRegion>
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,
<collectingCountry id="F7937654FFCDFF894418FC3FE6CDFC01" box="[1360,1431,978,1003]" name="Brazil" pageId="4" pageNumber="552">Brazil</collectingCountry>
,
<geoCoordinate id="EAB05003FFCDFF8941DFFC1AE26FFBE5" box="[151,309,1015,1039]" direction="south" orientation="latitude" pageId="4" pageNumber="552" precision="1" value="-29.66247">29°39'44,89''S</geoCoordinate>
<geoCoordinate id="EAB05003FFCDFF894074FC1BE2BBFBE5" box="[316,481,1014,1039]" direction="west" orientation="longitude" pageId="4" pageNumber="552" precision="1" value="-53.91399">53°54'50,36''O</geoCoordinate>
,
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.
<collectingDate id="EB7EE9ECFFCDFF894329FC1BE07AFBE5" box="[609,800,1014,1039]" pageId="4" pageNumber="552" value="2009-10-26">
<date id="FB3A1004FFCDFF894329FC1BE07AFBE5" box="[609,800,1014,1039]" pageId="4" pageNumber="552" value="2009-10-26">26 October 2009</date>
</collectingDate>
,
<collectorName id="22715312FFCDFF894263FC1AE0E7FBE5" box="[811,957,1014,1039]" pageId="4" pageNumber="552">M.P. Pontelli</collectorName>
coll.
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</paragraph>
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<subSubSection id="C79E654FFFCDFF89418FFBF7E014FBD9" box="[199,846,1050,1075]" pageId="4" pageNumber="552" type="etymology">
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<emphasis id="BDF0EAD6FFCDFF89418FFBF7E214FBD9" bold="true" box="[199,334,1050,1075]" pageId="4" pageNumber="552">Etymology.</emphasis>
The species is named from Latin for “small”.
</paragraph>
</subSubSection>
<subSubSection id="C79E654FFFCDFF89418FFBD3E759F885" pageId="4" pageNumber="552" type="description">
<paragraph id="8F3B36C4FFCDFF89418FFBD3E0C2FAC5" blockId="4.[151,1437,834,2011]" pageId="4" pageNumber="552">
<emphasis id="BDF0EAD6FFCDFF89418FFBD3E203FBBD" bold="true" box="[199,345,1086,1111]" pageId="4" pageNumber="552">Description.</emphasis>
Dimensions:
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<quantity id="487C9B21FFCDFF89432CFBD2E1E9FBBC" box="[612,691,1087,1111]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="552" unit="mm" value="31.0">31 mm</quantity>
by
<quantity id="487C9B21FFCDFF894396FBD3E06EFBBC" box="[734,820,1086,1111]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="552" unit="mm" value="3.5">3.5 mm</quantity>
at x,
<quantity id="487C9B21FFCDFF89423BFBD2E093FBBC" box="[883,969,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="552" unit="mm" value="3.8">3.8 mm</quantity>
at clitellum,
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at xl, 134 segments;
<typeStatus id="503F8866FFCDFF8941DFFB8EE258FB91" box="[151,258,1123,1147]" pageId="4" pageNumber="552" type="paratype">paratypes</typeStatus>
A
<quantity id="487C9B21FFCDFF89406BFB8EE22BFB90" box="[291,369,1123,1147]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="552" unit="mm" value="33.0">33 mm</quantity>
by 3.0 mm at x,
<quantity id="487C9B21FFCDFF894363FB8EE1DAFB90" box="[555,640,1123,1147]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="552" unit="mm" value="3.7">3.7 mm</quantity>
at clitellum,
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at xl, 139 segments;
<typeStatus id="503F8866FFCDFF894519FB8EE7EBFB91" box="[1105,1201,1123,1147]" pageId="4" pageNumber="552" type="paratype">paratype</typeStatus>
B
<quantity id="487C9B21FFCDFF894599FB8FE645FB90" box="[1233,1311,1122,1147]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="552" unit="mm" value="36.0">36 mm</quantity>
by
<quantity id="487C9B21FFCDFF894400FB8FE6C7FB90" box="[1352,1437,1122,1147]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="552" unit="mm" value="3.5">3.5 mm</quantity>
at x, 4.0 mm at clitellum,
<quantity id="487C9B21FFCDFF894084FB6AE17CFB74" box="[460,550,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="552" unit="mm" value="3.7">3.7 mm</quantity>
at xl, 116 segments. Body cylindrical. Nontype specimen
<quantity id="487C9B21FFCDFF89459BFB6AE67FFB74" box="[1235,1317,1159,1183]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="552" unit="mm" value="28.0">28 mm</quantity>
long, 105 segments. Setae closely paired throughout; setal formula aa:ab:bc:cd:dd = 8.0:0.2:1.0:0.1:8.9. Setae commence in xi, not Well-Visible before clitellum. Prostomium prolobous. Unpigmented, Whitish. OVipores post setal, 3.0 mm apart, in line With setae a. Male pores 2.0 mm apart on xVii Within paired round eleVations (like a mound); clitellum saddle, xVxxiii (
<figureCitation id="17BF2A41FFCDFF894011FAFBE29FFAC5" box="[345,453,1302,1327]" captionStart="FIGURE 1" captionStartId="3.[151,250,1936,1958]" captionTargetBox="[307,1259,233,1868]" captionTargetId="figure@3.[190,1406,193,1912]" captionTargetPageId="3" captionText="FIGURE 1. Glossoscolex (Glossoscolex) riograndensis n. sp.. A, B. Glossoscolex (Glossoscolex) riograndensis riograndensis. C, D. Glossoscolex (Glossoscolex) riograndensis nativus n. ssp. E, F. Glossoscolex (Glossoscolex) riograndensis pollulus n. ssp. FM = female pOres; MP = male pOres; NF = nephridial pOres." httpUri="https://zenodo.org/record/1447027/files/figure.png" pageId="4" pageNumber="552">Fig. 1E,F</figureCitation>
). Nephropores after clitellum in line of b.
</paragraph>
<paragraph id="8F3B36C4FFCDFF89418FFAD6E658F9ED" blockId="4.[151,1437,834,2011]" pageId="4" pageNumber="552">
Septa
<date id="FB3A1004FFCDFF894044FAD7E221FAB9" box="[268,379,1338,1363]" pageId="4" pageNumber="552" value="2011-07-06" valueMax="2011-10-06" valueMin="2011-07-06">6/710/11</date>
equally thick and muscular. Alimentary canal With barrel shaped gizzard in Vi; esophagus With high cheVron-patterned lamellae Viixi, ValVular in xii, intestinal origin xiV; typhlosole origin xiV, in xiVxxVi zigzag With Ventral edge bent oVer to form pockets, after xxVii simple WaVy lamina. Calciferous glands paired xii, small heart shaped, pedunculated, composite-tubular
<typeStatus id="503F8866FFCDFF8943B0FA4AE076FA55" box="[760,812,1447,1471]" pageId="4" pageNumber="552">type</typeStatus>
; blood Vessels to gland include large branch of dorsal Vessel to approximate center of each gland. Holonephric, Vesiculate; ducts to body Wall near leVel of b; some nephridia in the anterior segments (Vi and Vii) Were full of sperm (iridescent material) (
<figureCitation id="17BF2A41FFCDFF89452CFA03E7ACF9ED" box="[1124,1270,1518,1543]" captionStart="FIGURE 2" captionStartId="6.[151,250,1324,1346]" captionTargetBox="[151,1436,339,1302]" captionTargetId="figure@6.[151,1436,339,1302]" captionTargetPageId="6" captionText="FIGURE 2. Sperm-filled nephridia Of Glossoscolex (Glossoscolex) riograndensis n. sp., histOlOgical sectiOns. A, B. Glossoscolex (G.) riograndensis riograndensis n. sp., segment xii (A) and xiii (B). C, D. Glossoscolex (G.) riograndensis pollulus n. ssp., segments Vi (C) and Vii (D)." httpUri="https://zenodo.org/record/1447029/files/figure.png" pageId="4" pageNumber="552">Figure 2C,D</figureCitation>
).
</paragraph>
<paragraph id="8F3B36C4FFCDFF89418FF9FFE071F999" blockId="4.[151,1437,834,2011]" pageId="4" pageNumber="552">Vascular system With Ventral trunk, single dorsal trunk, lateral Vessels in Viiix, esophageal hearts in xxi, last pair (xi) free. Extra-esophageal Vessel from pharyngeal glands, along Ventral-lateral face of gizzard, esophagus back to calciferous glands; supraesophageal Vessel in xxi.</paragraph>
<paragraph id="8F3B36C4FFCDFF89418FF993E759F885" blockId="4.[151,1437,834,2011]" pageId="4" pageNumber="552">
OVaries, oVarian funnels free in xiii. Male sexual system metandric, testes and funnels in xi united Ventrally in a sac shifted posteriorly to xii; seminal Vesicles tubular With lobulated edges passing through xiixiii ranging posteriorly along intestine farther than xxxVxxxix; Vasa deferentia long, looped from seminal Vesicles in xii, form dense zig-zag on body Wall in route to middle-anterior face of muscular round copulatory bulbs (
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diameter); bulbs extend oVer xVixViii. Copulatory bulbs With thin muscular outer layer, dense, delicate corrugated glandular inner surface With small lumen leading to male pore at approximate center of bulb connection to body Wall; transVerse muscle bands crossing oVer bulbs in the anterior and posterior face.
</paragraph>
</subSubSection>
<subSubSection id="C79E654FFFCDFF88418FF897E082FDD7" lastPageId="5" lastPageNumber="553" pageId="4" pageNumber="552" type="discussion">
<paragraph id="8F3B36C4FFCDFF88418FF897E63AFF3F" blockId="4.[151,1437,834,2011]" lastBlockId="5.[151,1436,151,573]" lastPageId="5" lastPageNumber="553" pageId="4" pageNumber="552">
<emphasis id="BDF0EAD6FFCDFF89418FF897E261F879" bold="true" box="[199,315,1914,1939]" pageId="4" pageNumber="552">Remarks.</emphasis>
<taxonomicName id="48844D47FFCDFF89400CF896E058F879" authorityName="Bartz &amp; James &amp; Steffen &amp; Antoniolli &amp; Steffen &amp; Brown" authorityYear="2018" box="[324,770,1915,1939]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="pollulus">
<emphasis id="BDF0EAD6FFCDFF89400CF896E058F879" box="[324,770,1915,1939]" italics="true" pageId="4" pageNumber="552">Glossoscolex (Glossoscolex) r. pollulus</emphasis>
</taxonomicName>
is close to
<taxonomicName id="48844D47FFCDFF8942C1F896E6C6F879" authority="Zicsi &amp; Czusdi, 1999" authorityName="Zicsi &amp; Czusdi" authorityYear="1999" box="[905,1436,1914,1939]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="species" species="minor" subGenus="Glossoscolex">
<emphasis id="BDF0EAD6FFCDFF8942C1F896E7C7F878" box="[905,1181,1915,1939]" italics="true" pageId="4" pageNumber="552">Glossoscolex (G.) minor</emphasis>
<bibRefCitation id="EB154B35FFCDFF8945EEF897E6C6F879" box="[1190,1436,1914,1939]" pageId="4" pageNumber="552" refId="ref9066" refString="Zicsi, A. &amp; CsuZdi, C. (1999) Neue und bekannte RegenWurmer aus Verschiedenen Teilen Sudamerikas. Senckenbergiana Biologica, 78 (1 / 2), 123 - 134." type="journal article">Zicsi &amp; Czusdi, 1999</bibRefCitation>
</taxonomicName>
considering the small body size (comp.
<tableCitation id="C206037FFFCDFF894319F873E1F8F85D" box="[593,674,1950,1975]" captionStart="TABLE 1" captionStartId="10.[147,228,591,614]" captionText="TABLE 1. Comparison of characters of the new species and subspecies of Glossoscolex (Glossoscolex) with the type species of the subgenus and with further similar species that have been compared in the remarks sections of the new taxa described." pageId="4" pageNumber="552">Table 1</tableCitation>
). The differences betWeen
<taxonomicName id="48844D47FFCDFF894299F872E7D5F85C" authorityName="Bartz &amp; James &amp; Steffen &amp; Antoniolli &amp; Steffen &amp; Brown" authorityYear="2018" box="[977,1167,1950,1974]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="pollulus">
<emphasis id="BDF0EAD6FFCDFF894299F872E7D5F85C" box="[977,1167,1950,1974]" italics="true" pageId="4" pageNumber="552">G. (G.) r. pollulus</emphasis>
</taxonomicName>
and
<taxonomicName id="48844D47FFCDFF89458CF872E60CF85C" authorityName="Zicsi &amp; Czusdi" authorityYear="1999" box="[1220,1366,1951,1974]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="552" phylum="Annelida" rank="species" species="minor" subGenus="Glossoscolex">
<emphasis id="BDF0EAD6FFCDFF89458CF872E60CF85C" box="[1220,1366,1951,1974]" italics="true" pageId="4" pageNumber="552">G. (G.) minor</emphasis>
</taxonomicName>
are as folloWs, With the characteristics of the latter in parentheses: number of segments 116139 (198231), clitellum saddle (annular), clitellum extension xVxxii (xiVxxiii), testis sacs U shaped sac in xixii (unpaired in xi), seminal Vesicles big tubular With lobulated edges until xxxVxxxix (band until l), copulatory bulbs round (elongated).
</paragraph>
<paragraph id="8F3B36C4FFCCFF88418FFF0DE3B5FDF2" blockId="5.[151,1436,151,573]" pageId="5" pageNumber="553">
<taxonomicName id="48844D47FFCCFF88418FFF0DE2D7FF12" authorityName="Bartz &amp; James &amp; Steffen &amp; Antoniolli &amp; Steffen &amp; Brown" authorityYear="2018" box="[199,397,224,248]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="pollulus">
<emphasis id="BDF0EAD6FFCCFF88418FFF0DE2D7FF12" box="[199,397,224,248]" italics="true" pageId="5" pageNumber="553">G. (G.) r. pollulus</emphasis>
</taxonomicName>
is morphologically Very similar to
<taxonomicName id="48844D47FFCCFF88426CFF0DE734FF12" authority=", With" authorityName="With" box="[804,1134,223,248]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="riograndensis">
<emphasis id="BDF0EAD6FFCCFF88426CFF0DE776FF12" box="[804,1068,224,248]" italics="true" pageId="5" pageNumber="553">G. (G.) r. riograndensis</emphasis>
, With
</taxonomicName>
the main difference being body length, Which is smaller in
<taxonomicName id="48844D47FFCCFF884356FEE9E1B0FEF6" authorityName="Bartz &amp; James &amp; Steffen &amp; Antoniolli &amp; Steffen &amp; Brown" authorityYear="2018" box="[542,746,260,284]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="pollulus">
<emphasis id="BDF0EAD6FFCCFF884356FEE9E1B0FEF6" box="[542,746,260,284]" italics="true" pageId="5" pageNumber="553">G. (G.) r. pollulus</emphasis>
</taxonomicName>
(
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) than in
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<emphasis id="BDF0EAD6FFCCFF8842A4FEE9E7A1FEF6" box="[1004,1275,260,284]" italics="true" pageId="5" pageNumber="553">G. (G.) r. riograndensis</emphasis>
</taxonomicName>
(
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). Furthermore, the shape of the copulatory bulbs in
<taxonomicName id="48844D47FFCCFF884386FEC5E0CAFEAA" authorityName="Bartz &amp; James &amp; Steffen &amp; Antoniolli &amp; Steffen &amp; Brown" authorityYear="2018" box="[718,912,296,320]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="pollulus">
<emphasis id="BDF0EAD6FFCCFF884386FEC5E0CAFEAA" box="[718,912,296,320]" italics="true" pageId="5" pageNumber="553">G. (G.) r. pollulus</emphasis>
</taxonomicName>
is round, While in
<taxonomicName id="48844D47FFCCFF88452EFEC5E630FEAA" box="[1126,1386,296,320]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="riograndensis">
<emphasis id="BDF0EAD6FFCCFF88452EFEC5E630FEAA" box="[1126,1386,296,320]" italics="true" pageId="5" pageNumber="553">G. (G.) r. riograndensis</emphasis>
</taxonomicName>
it is round to oVal (
<tableCitation id="C206037FFFCCFF884008FEA1E2CFFE8E" box="[320,405,332,357]" captionStart="TABLE 1" captionStartId="10.[147,228,591,614]" captionText="TABLE 1. Comparison of characters of the new species and subspecies of Glossoscolex (Glossoscolex) with the type species of the subgenus and with further similar species that have been compared in the remarks sections of the new taxa described." pageId="5" pageNumber="553">Table 1</tableCitation>
). Both subspecies haVe sperm storage in the nephridia, but in
<taxonomicName id="48844D47FFCCFF884529FEA1E67CFE8E" authorityName="Bartz &amp; James &amp; Steffen &amp; Antoniolli &amp; Steffen &amp; Brown" authorityYear="2018" box="[1121,1318,332,356]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="pollulus">
<emphasis id="BDF0EAD6FFCCFF884529FEA1E67CFE8E" box="[1121,1318,332,356]" italics="true" pageId="5" pageNumber="553">G. (G.) r. pollulus</emphasis>
</taxonomicName>
it is more anterior segments (Vi and Vii) than in
<taxonomicName id="48844D47FFCCFF884303FE9DE00EFE62" box="[587,852,368,392]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="riograndensis">
<emphasis id="BDF0EAD6FFCCFF884303FE9DE00EFE62" box="[587,852,368,392]" italics="true" pageId="5" pageNumber="553">G. (G.) r. riograndensis</emphasis>
</taxonomicName>
in xii, xiii. We deemed these differences not to be enough for separation at species leVel, and furthermore, We did not haVe DNA tissue material from
<taxonomicName id="48844D47FFCCFF88447CFE79E3ABFE3A" authorityName="Bartz &amp; James &amp; Steffen &amp; Antoniolli &amp; Steffen &amp; Brown" authorityYear="2018" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="pollulus">
<emphasis id="BDF0EAD6FFCCFF88447CFE79E3ABFE3A" italics="true" pageId="5" pageNumber="553">G. (G.) r. pollulus</emphasis>
</taxonomicName>
to compare With
<taxonomicName id="48844D47FFCCFF88408AFE55E197FE3A" box="[450,717,440,464]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="riograndensis">
<emphasis id="BDF0EAD6FFCCFF88408AFE55E197FE3A" box="[450,717,440,464]" italics="true" pageId="5" pageNumber="553">G. (G.) r. riograndensis</emphasis>
</taxonomicName>
. Further sampling and both morphological and DNA analysis should help reVeal possible differences betWeen these tWo subspecies, and if they correspond or not to different species.
</paragraph>
<paragraph id="8F3B36C4FFCCFF88418FFDC8E082FDD7" blockId="5.[151,1436,151,573]" box="[199,984,548,573]" pageId="5" pageNumber="553">
For comparison of
<taxonomicName id="48844D47FFCCFF8840D4FDC9E1CCFDD7" authority="With" authorityName="With" box="[412,662,548,573]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="pollulus">
<emphasis id="BDF0EAD6FFCCFF8840D4FDC9E106FDD6" box="[412,604,548,572]" italics="true" pageId="5" pageNumber="553">G. (G.) r. pollulus</emphasis>
With
</taxonomicName>
<taxonomicName id="48844D47FFCCFF8843D4FDC9E00EFDD6" authorityName="Bartz &amp; James &amp; Steffen &amp; Antoniolli &amp; Steffen &amp; Brown" authorityYear="2018" box="[668,852,548,572]" class="Clitellata" family="Glossoscolecidae" genus="Glossoscolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="553" phylum="Annelida" rank="subSpecies" species="riograndensis" subGenus="Glossoscolex" subSpecies="nativus">
<emphasis id="BDF0EAD6FFCCFF8843D4FDC9E1F7FDD1" box="[668,685,548,571]" italics="true" pageId="5" pageNumber="553">G</emphasis>
. (
<emphasis id="BDF0EAD6FFCCFF88438CFDC9E18FFDD1" box="[708,725,548,571]" italics="true" pageId="5" pageNumber="553">G</emphasis>
.)
<emphasis id="BDF0EAD6FFCCFF8843A4FDC8E1ACFDD6" box="[748,758,549,572]" italics="true" pageId="5" pageNumber="553">r</emphasis>
.
<emphasis id="BDF0EAD6FFCCFF88424CFDC8E00EFDD6" box="[772,852,549,572]" italics="true" pageId="5" pageNumber="553">nativus</emphasis>
</taxonomicName>
, see beloW.
</paragraph>
</subSubSection>
</treatment>
</document>