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<mods:title id="4B41A959ED90898C996D871722194636">Taxonomy of the sand bubbler crabs Scopimera globosa De Haan, 1835, and S. tuberculata Stimpson, 1858 (Crustacea: Decapoda: Dotillidae) in East Asia, with description of a new species from the Ryukyus, Japan</mods:title>
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<treatment id="C14B3265FF94ED174F956E11FCA7FD39" ID-DOI="http://doi.org/10.5281/zenodo.5618516" ID-GBIF-Taxon="125137934" ID-Zenodo-Dep="5618516" LSID="urn:lsid:plazi:treatment:C14B3265FF94ED174F956E11FCA7FD39" httpUri="http://treatment.plazi.org/id/C14B3265FF94ED174F956E11FCA7FD39" lastPageId="4" lastPageNumber="46" pageId="2" pageNumber="45">
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<paragraph id="495D8373FF94ED114F956E11FDFCFC9C" blockId="2.[151,569,792,852]" box="[151,569,792,818]" pageId="2" pageNumber="45">
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<taxonomicName id="8EE2F8F0FF94ED114F956E11FDFCFC9C" ID-CoL="4VVFK" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[151,569,792,818]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF94ED114F956E11FDFCFC9C" bold="true" box="[151,569,792,818]" pageId="2" pageNumber="45">
<emphasis id="7B965F61FF94ED114F956E11FEB9FC9C" bold="true" box="[151,380,792,818]" italics="true" pageId="2" pageNumber="45">Scopimera globosa</emphasis>
De Haan, 1835
</emphasis>
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</paragraph>
<paragraph id="495D8373FF94ED114F956E33FDD2FCFA" blockId="2.[151,569,792,852]" box="[151,535,826,852]" pageId="2" pageNumber="45">
(
<figureCitation id="D1D99FF6FF94ED114F9D6E33FF34FCFA" box="[159,241,826,852]" captionStart="FIGURE 1" captionStartId="3.[151,256,934,958]" captionTargetBox="[280,1300,202,901]" captionTargetId="figure@3.[265,1321,155,923]" captionTargetPageId="3" captionText="FIGURE 1. Scopimera globosa De Haan, 1835: dorsal view of carapace (a) and suborbital ridge and pterygostomian region (b); S. intermedia Balss, 1934: dorsal view of carapace (c) and suborbital ridge and pterygostomian region (d); S. ryukyuensis sp. nov.: dorsal view of carapace (e) and suborbital ridge and pterygostomian region (f). a, c, e: note arrow highlighting relative size of extra-orbital angle and solid line indicating direction of ridge behind the extra-orbital angle (which is diagnostic between S. ryukyuensis sp. nov. and S. globosa). b, d, f: arrow noting raised branchial region viewed from front in S. ryukyuensis sp. nov. (e) and relative conspicuousness of sub-orbital ridge: that of S. ryukyuensis sp. nov. being most pronounced." httpUri="https://zenodo.org/record/205329/files/figure.png" pageId="2" pageNumber="45">Figs. 1</figureCitation>
a, b,
<figureCitation id="D1D99FF6FF94ED114E2B6E33FEACFCFA" box="[297,361,826,852]" captionStart="FIGURE 2" captionStartId="4.[151,256,1383,1407]" captionTargetBox="[280,1309,721,1345]" captionTargetId="figure@4.[265,1321,677,1362]" captionTargetPageId="4" captionText="FIGURE 2. Scopimera globosa De Haan, 1835: chela (a) and male abdomen (b); S. intermedia Balss, 1934: chela (c) and male abdomen (d); S. ryukyuensis sp. nov.: chela (e) and male abdomen (f). a, c, e: arrow pointing tooth at inner margin of movable finger: that of S. intermedia Balss, 1934, being most rudimentary. b, d, f: large arrows showing dimensions of the sixth abdominal somite and small arrow indicating the convexity of distal border of the fourth somite." httpUri="https://zenodo.org/record/205330/files/figure.png" pageId="2" pageNumber="45">2a, b,</figureCitation>
<figureCitation id="D1D99FF6FF94ED114E736E33FE6FFCFA" box="[369,426,826,852]" captionStart="FIGURE 3" captionStartId="6.[151,255,1641,1665]" captionTargetBox="[156,1430,197,1610]" captionTargetId="figure@6.[151,1436,194,1619]" captionTargetPageId="6" captionText="FIGURE 3. Scopimera globosa De Haan, 1835: G 1 entire view (a), tip (b) and magnification of setae (c); S. intermedia Balss, 1934: G 1 entire view (d), tip (e) and magnification of setae (f); S. ryukyuensis sp. nov.: G 1 entire view (g), tip (h) and tip observed under light microscope (i). Scale bars in µm." httpUri="https://zenodo.org/record/205331/files/figure.png" pageId="2" pageNumber="45">3ac</figureCitation>
,
<figureCitation id="D1D99FF6FF94ED114EB56E33FE32FCFA" box="[439,503,826,852]" captionStart="FIGURE 4" captionStartId="8.[151,255,1858,1882]" captionTargetBox="[183,1394,200,1816]" captionTargetId="figure@8.[178,1403,194,1825]" captionTargetPageId="8" captionText="FIGURE 4. Photographs in life (a e). Scopimera globosa De Haan, 1835: male frontal view (a) and dorsal view (b); S. intermedia Balss, 1934: male frontal view (c) and dorsal view (d); and S. ryukyuensis sp. nov.: male frontal view (e) and close-up of external maxillipeds from preserved specimens, showing ring-like marking on merus (f)." httpUri="https://zenodo.org/record/205332/files/figure.png" pageId="2" pageNumber="45">4a, b,</figureCitation>
<figureCitation id="D1D99FF6FF94ED114EFC6E33FDC8FCFA" box="[510,525,826,852]" captionStart="FIGURE 6" captionStartId="10.[151,257,1519,1543]" captionTargetBox="[171,1403,1002,1466]" captionTargetId="figure@10.[151,1436,903,1498]" captionTargetPageId="10" captionText="FIGURE 6. Dorsal view of the neotype of Scopimera tuberculata Stimpson, 1858. A male (cw 9.9 mm, cl 7.6 mm) (CBM-ZC 4195) deposited in Natural History Museum and Institute, Chiba, Japan." httpUri="https://zenodo.org/record/205334/files/figure.png" pageId="2" pageNumber="45">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="01F8D0F8FF94ED114F956E8CFD1CFB32" pageId="2" pageNumber="45" type="reference_group">
<paragraph id="495D8373FF94ED114F956E8CFC6FFC32" blockId="2.[151,1437,901,1180]" box="[151,938,901,924]" pageId="2" pageNumber="45">
<treatmentCitation id="C843A562FF94ED114F956E8CFD62FC32" box="[151,679,901,924]" pageId="2" pageNumber="45">
<taxonomicName id="8EE2F8F0FF94ED114F956E8CFD62FC32" authority="De Haan 1835: 53" authorityName="De Haan" authorityPageNumber="53" authorityYear="1835" box="[151,679,901,924]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="globosa" subGenus="Scopimera">
<emphasis id="7B965F61FF94ED114F956E8CFF33FC32" box="[151,246,901,924]" italics="true" pageId="2" pageNumber="45">Ocypode</emphasis>
(
<emphasis id="7B965F61FF94ED114E076E8CFEB2FC32" box="[261,375,901,924]" italics="true" pageId="2" pageNumber="45">Scopimera</emphasis>
)
<emphasis id="7B965F61FF94ED114E846E8CFE1EFC32" box="[390,475,901,924]" italics="true" pageId="2" pageNumber="45">globosa</emphasis>
De Haan 1835: 53
</taxonomicName>
</treatmentCitation>
, pl. 11 (3, 3a, 3b), pl. C.
</paragraph>
<paragraph id="495D8373FF94ED114F956EACFEE0FBD2" blockId="2.[151,1437,901,1180]" pageId="2" pageNumber="45">
<treatmentCitationGroup id="69F2A45DFF94ED114F956EACFEE0FBD2" pageId="2" pageNumber="45">
<treatmentCitation id="C843A562FF94ED114F956EACFD07FC12" box="[151,706,933,956]" pageId="2" pageNumber="45">
<taxonomicName id="8EE2F8F0FF94ED114F956EACFEA6FC12" ID-CoL="4VVFK" box="[151,355,933,956]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF94ED114F956EACFEA6FC12" box="[151,355,933,956]" italics="true" pageId="2" pageNumber="45">Scopimera globosa</emphasis>
</taxonomicName>
— H.
<bibRefCitation id="2D73FE82FF94ED114EA96EACFD48FC12" author="Milne Edwards, H." box="[427,653,933,956]" journalOrPublisher="Annales des Sciences naturelles" pageId="2" pageNumber="45" pagination="109 - 166" part="3" refString="Milne Edwards, H. (1852) Observations sur les affinites zoologiques et la classification naturelle des Crustaces. Annales des Sciences naturelles, (3) 18, 109 - 166." title="Observations sur les affinites zoologiques et la classification naturelle des Crustaces" type="journal article" year="1852">Milne Edwards 1852</bibRefCitation>
: 153
</treatmentCitation>
.
<bibRefCitation id="2D73FE82FF94ED114DE26EACFCBBFC12" author="Ortmann, A." box="[736,894,933,956]" journalOrPublisher="Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere" pageId="2" pageNumber="45" pagination="683 - 772" part="7" refString="Ortmann, A. (1894) Die Decapoden - Krebse des Strassburger Museums, mit besonderer Berucksichtigung der van Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. VIII. Theil. Abtheilung: Brachyura (Brachyura genuine Boas) III. Unterabtheilung: Cancroidea, 2. Section: Cancrinea, 2. Gruppe: Catametopa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7, 683 - 772, pl. 23." title="Die Decapoden - Krebse des Strassburger Museums, mit besonderer Berucksichtigung der van Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. VIII. Theil. Abtheilung: Brachyura (Brachyura genuine Boas) III. Unterabtheilung: Cancroidea, 2. Section: Cancrinea, 2. Gruppe: Catametopa" type="journal article" year="1894">Ortmann 1894</bibRefCitation>
: 747.
<bibRefCitation id="2D73FE82FF94ED114CD36EACFB98FC12" author="Koelbel, K." box="[977,1117,933,956]" journalOrPublisher="Wissenschaftliche Ergebnisse der Reise des Grafen Bela Szechenyi in Ostasien" pageId="2" pageNumber="45" pagination="709 - 718" part="2" refString="Koelbel, K. (1897) Beschreibung der Krebse. Wissenschaftliche Ergebnisse der Reise des Grafen Bela Szechenyi in Ostasien, 2, 709 - 718." title="Beschreibung der Krebse" type="journal article" year="1897">Koebel 1897</bibRefCitation>
: 713.
<bibRefCitation id="2D73FE82FF94ED114BB26EACFAE9FC12" author="Parisi, B." box="[1200,1324,933,956]" journalOrPublisher="Atti della Societa Italiana de Scienze Naturali" pageId="2" pageNumber="45" pagination="5 - 42" part="57" refString="Parisi, B. (1918) I Decapodi giapponesi del Museo di Milano. VI. Catometopa e Paguridea. Atti della Societa Italiana de Scienze Naturali, 57, 5 - 42." title="I Decapodi giapponesi del Museo di Milano. VI. Catometopa e Paguridea" type="journal article" year="1918">Parisi 1918</bibRefCitation>
: 97, fig. 2.
<bibRefCitation id="2D73FE82FF94ED114FD66ECCFE97FC72" author="Tesch, J. J." box="[212,338,965,988]" journalOrPublisher="Grapsidae and Gecarcinidae. Sigoba Expeditie Monographie" pageId="2" pageNumber="45" pagination="1 - 148" part="39" refString="Tesch, J. J. (1918) The Brachyura of the Sigoba Expedition. Hymenosomidae, Ocypodidae, Grapsidae and Gecarcinidae. Sigoba Expeditie Monographie, 39 c, 1 - 148." title="The Brachyura of the Sigoba Expedition. Hymenosomidae, Ocypodidae" type="journal article" year="1918">Tesch 1918</bibRefCitation>
: 46, pl. 3(3).
<bibRefCitation id="2D73FE82FF94ED114EFA6ECCFDBCFC72" author="Kemp, S." box="[504,633,965,988]" journalOrPublisher="Records of the Indian Museum" pageId="2" pageNumber="45" pagination="48" part="16" refString="Kemp, S. (1919) Notes on Crustacea Decapoda in the Indian Museum. XII. Scopimerinae. Records of the Indian Museum, 16, 305 - 3 48." title="Notes on Crustacea Decapoda in the Indian Museum. XII. Scopimerinae" type="journal article" year="1919">Kemp 1919</bibRefCitation>
: 312, pl. 12(2).
<bibRefCitation id="2D73FE82FF94ED114C316ECCFC00FC72" author="Yokoya, Y." box="[819,965,965,988]" journalOrPublisher="Science Reports of the Tohoku Imperial University, 4 th Series (Biology)" pageId="2" pageNumber="45" pagination="757 - 784" part="3" refString="Yokoya, Y. (1928) Report of the biological survey of Mutsu Bay. 10. Brachyura and crab-shaped Anomura. Science Reports of the Tohoku Imperial University, 4 th Series (Biology), 3, 757 - 784." title="Report of the biological survey of Mutsu Bay. 10. Brachyura and crab-shaped Anomura" type="journal article" year="1928">Yokoya 1928</bibRefCitation>
: 779.
<bibRefCitation id="2D73FE82FF94ED114B196ECCFB54FC72" author="Shen, C. - J." box="[1051,1169,965,988]" journalOrPublisher="Zoologia Sinica, Series A" pageId="2" pageNumber="45" pagination="1 - 300" part="9" refString="Shen, C. - J. (1932) The brachyuran Crustacea of North China. Zoologia Sinica, Series A, 9, 1 - 300." title="The brachyuran Crustacea of North China" type="journal article" year="1932">Shen 1932</bibRefCitation>
: 253, pl. 10(5), text-figs. 155, 157.
<bibRefCitation id="2D73FE82FF94ED114E456EECFE07FC52" author="Sakai, T." box="[327,450,997,1020]" journalOrPublisher="Science Reports of the Tokyo Bunrika Daigaku, Section B" pageId="2" pageNumber="45" pagination="281 - 330" part="1" refString="Sakai, T. (1934) Brachyura from the coast of Kyushu, Japan. Science Reports of the Tokyo Bunrika Daigaku, Section B, 1 (25), 281 - 330." title="Brachyura from the coast of Kyushu, Japan" type="journal article" year="1934">Sakai 1934</bibRefCitation>
: 320; 1935: 214, pl. 59(4); 1939: 636, pl. 72(4); 1965: 191, pl. 91(1); 1976: 621, pl. 212(2); 1982: 714 (in part).
<bibRefCitation id="2D73FE82FF94ED114EB0690CFD85FBB2" author="Kamita, T." bookContentInfo="289 pp." box="[434,576,1029,1052]" journalOrPublisher="The Fisheries Society of Chosen, Keijo, Korea" pageId="2" pageNumber="45" refString="Kamita, T. (1941) Studies of the Decapod Crustaceans of Chosen. Pt. 1. Crabs. The Fisheries Society of Chosen, Keijo, Korea, 289 pp. (in Japanese)" title="Studies of the Decapod Crustaceans of Chosen. Pt. 1. Crabs" type="book" year="1941">Kamita 1941</bibRefCitation>
: 175, text-fig. 96.
<bibRefCitation id="2D73FE82FF94ED114C1B690CFBF0FBB2" author="Nishimura, S. &amp; Suzuki, K." bookContentInfo="196 pp." box="[793,1077,1029,1052]" journalOrPublisher="Hoikusha, Osaka" pageId="2" pageNumber="45" refString="Nishimura, S. &amp; Suzuki, K. (1971) Common Seashore Animals of Japan in Color (Kaigan Dobutsu). Hoikusha, Osaka, 196 pp. (in Japanese)" title="Common Seashore Animals of Japan in Color (Kaigan Dobutsu)" type="book" year="1971">Nishimura &amp; Suzuki 1971</bibRefCitation>
: 121, pl. 44(9).
<bibRefCitation id="2D73FE82FF94ED114BF1690CFAA6FBB2" author="Kim, H. S." box="[1267,1379,1029,1052]" journalOrPublisher="Seoul, pp. 589 - 682. (in Korean)" pageId="2" pageNumber="45" refString="Kim, H. S. (1973) Anomura-Brachyura. In: Illustrated Encyclopedia of Fauna &amp; Flora of Korea. Vol. 14. Samhwa, Seoul, pp. 589 - 682. (in Korean)" title="Anomura-Brachyura" type="book" volumeTitle="Illustrated Encyclopedia of Fauna &amp; Flora of Korea. Vol. 14. Samhwa" year="1973">Kim 1973</bibRefCitation>
: 641.
<bibRefCitation id="2D73FE82FF94ED114FD8692CFEA3FB92" author="Takeda, M." box="[218,358,1061,1084]" editor="Utinomi, H." journalOrPublisher="in Japanese" pageId="2" pageNumber="45" pagination="188 - 331" refString="Takeda, M. (1975) Brachyura. In: Utinomi, H. (Ed.), The Aquatic Lower Animals of Japan. Gakken, Tokyo, pp. 188 - 331. (in Japanese)" title="Brachyura" type="book chapter" volumeTitle="The Aquatic Lower Animals of Japan. Gakken" year="1975">Takeda 1975</bibRefCitation>
: 144, 243 (in part); 1982: 212; 1994: 260.
<bibRefCitation id="2D73FE82FF94ED114C39692CFC09FB92" author="Miyake, S." bookContentInfo="277 pp." box="[827,972,1061,1084]" journalOrPublisher="Hoikusha, Osaka" pageId="2" pageNumber="45" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in Color. II Brachyura (Crabs). Hoikusha, Osaka, 277 pp. (in Japanese)" title="Japanese Crustacean Decapods and Stomatopods in Color. II Brachyura (Crabs)" type="book" year="1983">Miyake 1983</bibRefCitation>
: 168 (in part).
<bibRefCitation id="2D73FE82FF94ED114B79692CFAFCFB92" author="Dai, A. - Y. &amp; Yang, S. - L." bookContentInfo="682 pp." box="[1147,1337,1061,1084]" journalOrPublisher="Beijing" pageId="2" pageNumber="45" refString="Dai, A. - Y. &amp; Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing, 682 pp." title="Crabs of the China Seas. China Ocean Press" type="book" year="1991">Dai &amp; Yang 1991</bibRefCitation>
: 496, fig. 253(2), pl. 63(5) (in part).
<bibRefCitation id="2D73FE82FF94ED114EF9694CFCCBFBF2" author="Yamaguchi, T. &amp; Baba, K." box="[507,782,1093,1116]" editor="Yamaguchi, T." journalOrPublisher="Tokyo" pageId="2" pageNumber="45" pagination="145 - 570" refString="Yamaguchi, T. &amp; Baba, K. (1993) Crustacean specimens collected in Japan by Ph. F. von Siebold and H. Burger and held by the Nationaal Natuurhistorisch Museum in Leiden and other museums. In: Yamaguchi, T. (Ed.), Ph. F. von Siebold and Natural History of Japan: Crustacea, The Carcinological Society of Japan, Tokyo, pp. 145 - 570." title="Crustacean specimens collected in Japan by Ph. F. von Siebold and H. Burger and held by the Nationaal Natuurhistorisch Museum in Leiden and other museums" type="book chapter" volumeTitle="Ph. F. von Siebold and Natural History of Japan: Crustacea, The Carcinological Society of Japan" year="1993">
<collectingRegion id="8B264D91FF94ED114EF9694CFDB6FBF2" box="[507,627,1093,1116]" country="Japan" name="Yamaguchi" pageId="2" pageNumber="45">Yamaguchi</collectingRegion>
&amp; Baba 1993
</bibRefCitation>
: 506, fig. 195.
<bibRefCitation id="2D73FE82FF94ED114CC4694CFBB1FBF2" author="Minemizu, R." bookContentInfo="344 pp." box="[966,1140,1093,1116]" journalOrPublisher="Tokyo" pageId="2" pageNumber="45" refString="Minemizu, R. (2000) Marine Decapod and Stomatopod Crustaceans Mainly from Japan. Bunnichi-Sogo Shuppan, Tokyo, 344 pp. (in Japanese)" title="Marine Decapod and Stomatopod Crustaceans Mainly from Japan. Bunnichi-Sogo Shuppan" type="book" year="2000">Minemizu 2000</bibRefCitation>
: 309.
<bibRefCitation id="2D73FE82FF94ED114BC8694CFAACFBF2" author="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F." box="[1226,1385,1093,1116]" journalOrPublisher="The Raffles Bulletin of Zoology Supplement" pageId="2" pageNumber="45" pagination="1 - 286" part="17" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of the extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." title="Systema Brachyurorum: Part I. An annotated checklist of the extant brachyuran crabs of the world" type="journal article" year="2008">
Ng
<emphasis id="7B965F61FF94ED114BF0694CFAE7FBF2" box="[1266,1314,1093,1116]" italics="true" pageId="2" pageNumber="45">et al</emphasis>
. 2008
</bibRefCitation>
: 235 (in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="495D8373FF94ED114F95698CFD1CFB32" blockId="2.[151,1437,901,1180]" box="[151,729,1157,1180]" pageId="2" pageNumber="45">
<treatmentCitationGroup id="69F2A45DFF94ED114F95698CFD1CFB32" box="[151,729,1157,1180]" pageId="2" pageNumber="45">
<treatmentCitation id="C843A562FF94ED114F95698CFD9EFB32" box="[151,603,1157,1180]" pageId="2" pageNumber="45">
<taxonomicName id="8EE2F8F0FF94ED114F95698CFD9EFB32" ID-CoL="4VVFY" authority="Stimpson 1858: 98" authorityName="Stimpson" authorityPageNumber="98" authorityYear="1858" box="[151,603,1157,1180]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="7B965F61FF94ED114F95698CFE4CFB32" box="[151,393,1157,1180]" italics="true" pageId="2" pageNumber="45">Scopimera tuberculata</emphasis>
<bibRefCitation id="2D73FE82FF94ED114E92698CFDF1FB32" author="Stimpson, W." box="[400,564,1157,1180]" journalOrPublisher="Pars 5. Proceedings of the Academia of Natural Science, Philadelphia" pageId="2" pageNumber="45" pagination="93 - 110" part="10" refString="Stimpson, W. (1858) Crustacea Ocypodoidea. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit, Pars 5. Proceedings of the Academia of Natural Science, Philadelphia, 10, 93 - 110." title="Crustacea Ocypodoidea. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit" type="journal article" year="1858">Stimpson 1858</bibRefCitation>
: 98
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="C843A562FF94ED114D65698CFD10FB32" box="[615,725,1157,1180]" pageId="2" pageNumber="45">1907: 102</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="01F8D0F8FF94ED114F9569C3FE44FA44" pageId="2" pageNumber="45" type="materials_examined">
<paragraph id="495D8373FF94ED114F9569C3FF3DFAA4" blockId="2.[151,1437,1226,2036]" pageId="2" pageNumber="45">
<emphasis id="7B965F61FF94ED114F9569C3FE4BFB4A" bold="true" box="[151,398,1226,1252]" pageId="2" pageNumber="45">Material examined</emphasis>
.
<typeStatus id="96593DD1FF94ED114E9D69C3FDDCFB4A" box="[415,537,1226,1252]" pageId="2" pageNumber="45" type="lectotype">Lectotype</typeStatus>
and
<typeStatus id="96593DD1FF94ED114D5569C3FD03FB4A" box="[599,710,1226,1252]" pageId="2" pageNumber="45" type="paratype">paratype</typeStatus>
:
<specimenCount id="5FE448FAFF94ED114DD069C3FCF4FB4A" box="[722,817,1226,1252]" pageId="2" pageNumber="45" type="male">2 males</specimenCount>
(
<collectionCode id="2FF31BB6FF94ED114C4069C3FC5AFB4A" box="[834,927,1226,1252]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="45">RMNH</collectionCode>
282),
<collectingCountry id="31F5C3E3FF94ED114CF269C3FBFCFB4A" box="[1008,1081,1226,1252]" name="Japan" pageId="2" pageNumber="45">Japan</collectingCountry>
,
<date id="3D5CA5B3FF94ED114B4469C3FB17FB4A" box="[1094,1234,1226,1252]" pageId="2" pageNumber="45" value="1823" valueMax="1829">18231829</date>
, coll.
<collectingCountry id="31F5C3E3FF94ED114A1D69C3FA86FB4A" box="[1311,1347,1226,1252]" name="Philippines" pageId="2" pageNumber="45">Ph</collectingCountry>
. F. von Siebold.
</paragraph>
<paragraph id="495D8373FF94ED114FC6681CFE44FA44" blockId="2.[151,1437,1226,2036]" pageId="2" pageNumber="45">
<emphasis id="7B965F61FF94ED114FC6681CFDE9FA81" bold="true" box="[196,556,1301,1327]" pageId="2" pageNumber="45">
Additional material.
<collectingCountry id="31F5C3E3FF94ED114ED1681CFDE3FA81" box="[467,550,1301,1327]" name="Japan" pageId="2" pageNumber="45">Japan</collectingCountry>
:
</emphasis>
<specimenCount id="5FE448FAFF94ED114D36681CFD6FFA81" box="[564,682,1301,1327]" pageId="2" pageNumber="45" type="female">7 females</specimenCount>
(NSMT-Cr-3729), Usu Bay,
<collectingRegion id="8B264D91FF94ED114B0C681CFB4DFA81" box="[1038,1160,1301,1327]" country="Japan" name="Hokkaido" pageId="2" pageNumber="45">Hokkaido</collectingRegion>
,
<date id="3D5CA5B3FF94ED114B97681CFAEBFA81" box="[1173,1326,1301,1327]" pageId="2" pageNumber="45" value="1934-07-27">27 Jul. 1934</date>
, coll. M. Iwasa;
<specimenCount id="5FE448FAFF94ED114FED6833FE95FAFA" box="[239,336,1338,1364]" pageId="2" pageNumber="45" type="male">2 males</specimenCount>
,
<specimenCount id="5FE448FAFF94ED114E5C6833FE16FAFA" box="[350,467,1338,1364]" pageId="2" pageNumber="45" type="female">2 females</specimenCount>
(NSMT-Cr-3739), Funabashi,
<collectingRegion id="8B264D91FF94ED114C4C6833FC5FFAFA" box="[846,922,1338,1364]" country="Japan" name="Chiba" pageId="2" pageNumber="45">Chiba</collectingRegion>
,
<date id="3D5CA5B3FF94ED114CA56833FB89FAFA" box="[935,1100,1338,1364]" pageId="2" pageNumber="45" value="1957-04-17">17 Apr. 1957</date>
;
<specimenCount id="5FE448FAFF94ED114B5B6833FB7DFAFA" box="[1113,1208,1338,1364]" pageId="2" pageNumber="45" type="male">3 males</specimenCount>
(NSMT-Cr-3741), Moiwa, Sopporo;
<specimenCount id="5FE448FAFF94ED114E716869FE03FAD4" box="[371,454,1376,1402]" pageId="2" pageNumber="45" type="male">1 male</specimenCount>
(NSMT-Cr-6504),
<collectingRegion id="8B264D91FF94ED114DB06869FD3AFAD4" box="[690,767,1376,1402]" country="Japan" name="Tokyo" pageId="2" pageNumber="45">Tokyo</collectingRegion>
Bay,
<date id="3D5CA5B3FF94ED114C446869FC18FAD4" box="[838,989,1376,1402]" pageId="2" pageNumber="45" value="1977-03-07">7 Mar. 1977</date>
, coll. M. Suzuki;
<specimenCount id="5FE448FAFF94ED114BBA6869FADFFAD4" box="[1208,1306,1376,1402]" pageId="2" pageNumber="45" type="male">2 males</specimenCount>
,
<specimenCount id="5FE448FAFF94ED114A256869FA59FAD4" box="[1319,1436,1376,1402]" pageId="2" pageNumber="45" type="female">2 females</specimenCount>
(NSMT-Cr-6797), Ena Bay,
<collectingRegion id="8B264D91FF94ED114EE7688CFDA1FA31" box="[485,612,1413,1439]" country="Japan" name="Kanagawa" pageId="2" pageNumber="45">Kanagawa</collectingRegion>
,
<date id="3D5CA5B3FF94ED114D72688CFCCFFA31" box="[624,778,1413,1439]" pageId="2" pageNumber="45" value="1980-06-15">15 Jun. 1980</date>
, coll. M. Takeda;
<specimenCount id="5FE448FAFF94ED114CDD688CFB94FA31" box="[991,1105,1413,1439]" pageId="2" pageNumber="45" type="female">5 females</specimenCount>
(CBM-ZC-7979), Usu Bay,
<collectingRegion id="8B264D91FF94ED114F9568A3FECBFA6A" box="[151,270,1450,1476]" country="Japan" name="Hokkaido" pageId="2" pageNumber="45">Hokkaido</collectingRegion>
,
<date id="3D5CA5B3FF94ED114E1A68A3FE7EFA6A" box="[280,443,1450,1476]" pageId="2" pageNumber="45" value="2004-08-18">18 Aug. 2004</date>
, coll. T. Komai;
<specimenCount id="5FE448FAFF94ED114D7E68A3FD22FA6A" box="[636,743,1450,1476]" pageId="2" pageNumber="45" type="male">10 males</specimenCount>
,
<specimenCount id="5FE448FAFF94ED114DF168A3FCA6FA6A" box="[755,867,1450,1476]" pageId="2" pageNumber="45" type="female">2 females</specimenCount>
(CEL-CRU-2008-02),
<collectingRegion id="8B264D91FF94ED114B7668A3FB3CFA6A" box="[1140,1273,1450,1476]" country="Japan" name="Wakayama" pageId="2" pageNumber="45">Wakayama</collectingRegion>
,
<date id="3D5CA5B3FF94ED114A0668A3FA52FA6A" box="[1284,1431,1450,1476]" pageId="2" pageNumber="45" value="2008-08-07">7 Aug. 2008</date>
, coll. B. K. K. Chan.
</paragraph>
</subSubSection>
<subSubSection id="01F8D0F8FF94ED114FC668FCFCCAF90A" pageId="2" pageNumber="45" type="diagnosis">
<paragraph id="495D8373FF94ED114FC668FCFCCAF90A" blockId="2.[151,1437,1226,2036]" pageId="2" pageNumber="45">
<emphasis id="7B965F61FF94ED114FC668FCFEF9F9A1" bold="true" box="[196,316,1525,1551]" pageId="2" pageNumber="45">Diagnosis</emphasis>
. External maxillipeds merus slightly shorter than or subequal to ischium; external orbital angle blunt, crests behind external orbital angles diverging posteriorly (
<figureCitation id="D1D99FF6FF94ED114C906B13FC25F99A" box="[914,992,1562,1588]" captionStart="FIGURE 1" captionStartId="3.[151,256,934,958]" captionTargetBox="[280,1300,202,901]" captionTargetId="figure@3.[265,1321,155,923]" captionTargetPageId="3" captionText="FIGURE 1. Scopimera globosa De Haan, 1835: dorsal view of carapace (a) and suborbital ridge and pterygostomian region (b); S. intermedia Balss, 1934: dorsal view of carapace (c) and suborbital ridge and pterygostomian region (d); S. ryukyuensis sp. nov.: dorsal view of carapace (e) and suborbital ridge and pterygostomian region (f). a, c, e: note arrow highlighting relative size of extra-orbital angle and solid line indicating direction of ridge behind the extra-orbital angle (which is diagnostic between S. ryukyuensis sp. nov. and S. globosa). b, d, f: arrow noting raised branchial region viewed from front in S. ryukyuensis sp. nov. (e) and relative conspicuousness of sub-orbital ridge: that of S. ryukyuensis sp. nov. being most pronounced." httpUri="https://zenodo.org/record/205329/files/figure.png" pageId="2" pageNumber="45">Figs. 1</figureCitation>
a, 6); cheliped length more than twice of carapace length in mature males, carpus elongated; tooth on inner margin of movable finger triangular (
<figureCitation id="D1D99FF6FF94ED114A6F6B49FF6DF9D1" captionStart="FIGURE 2" captionStartId="4.[151,256,1383,1407]" captionTargetBox="[280,1309,721,1345]" captionTargetId="figure@4.[265,1321,677,1362]" captionTargetPageId="4" captionText="FIGURE 2. Scopimera globosa De Haan, 1835: chela (a) and male abdomen (b); S. intermedia Balss, 1934: chela (c) and male abdomen (d); S. ryukyuensis sp. nov.: chela (e) and male abdomen (f). a, c, e: arrow pointing tooth at inner margin of movable finger: that of S. intermedia Balss, 1934, being most rudimentary. b, d, f: large arrows showing dimensions of the sixth abdominal somite and small arrow indicating the convexity of distal border of the fourth somite." httpUri="https://zenodo.org/record/205330/files/figure.png" pageId="2" pageNumber="45">Fig. 2</figureCitation>
a); male sixth abdominal somite approximately as broad as long (
<figureCitation id="D1D99FF6FF94ED114CC66B6CFBCAF9D1" box="[964,1039,1637,1663]" captionStart="FIGURE 2" captionStartId="4.[151,256,1383,1407]" captionTargetBox="[280,1309,721,1345]" captionTargetId="figure@4.[265,1321,677,1362]" captionTargetPageId="4" captionText="FIGURE 2. Scopimera globosa De Haan, 1835: chela (a) and male abdomen (b); S. intermedia Balss, 1934: chela (c) and male abdomen (d); S. ryukyuensis sp. nov.: chela (e) and male abdomen (f). a, c, e: arrow pointing tooth at inner margin of movable finger: that of S. intermedia Balss, 1934, being most rudimentary. b, d, f: large arrows showing dimensions of the sixth abdominal somite and small arrow indicating the convexity of distal border of the fourth somite." httpUri="https://zenodo.org/record/205330/files/figure.png" pageId="2" pageNumber="45">Fig. 2</figureCitation>
b); G1 slender, tip rounded, with inward radiating setae of similar length (
<figureCitation id="D1D99FF6FF94ED114D706B83FD7DF90A" box="[626,696,1674,1700]" captionStart="FIGURE 3" captionStartId="6.[151,255,1641,1665]" captionTargetBox="[156,1430,197,1610]" captionTargetId="figure@6.[151,1436,194,1619]" captionTargetPageId="6" captionText="FIGURE 3. Scopimera globosa De Haan, 1835: G 1 entire view (a), tip (b) and magnification of setae (c); S. intermedia Balss, 1934: G 1 entire view (d), tip (e) and magnification of setae (f); S. ryukyuensis sp. nov.: G 1 entire view (g), tip (h) and tip observed under light microscope (i). Scale bars in µm." httpUri="https://zenodo.org/record/205331/files/figure.png" pageId="2" pageNumber="45">Fig. 3</figureCitation>
a, b, c).
</paragraph>
</subSubSection>
<subSubSection id="01F8D0F8FF94ED104FC66BB9FD9BF95E" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45" type="description">
<paragraph id="495D8373FF94ED114FC66BB9FCA5F85A" blockId="2.[151,1437,1226,2036]" pageId="2" pageNumber="45">
<emphasis id="7B965F61FF94ED114FC66BB9FE99F964" bold="true" box="[196,348,1712,1738]" pageId="2" pageNumber="45">Description.</emphasis>
Carapace inflated, slightly broader than long, regions indistinct, surface covered by rounded tubercles, being densest on branchial regions, smooth on cardiac and intestinal regions; posterior margin longer than width between both external orbital angles (
<figureCitation id="D1D99FF6FF94ED114C3F6BF3FC4BF8BA" box="[829,910,1786,1812]" captionStart="FIGURE 1" captionStartId="3.[151,256,934,958]" captionTargetBox="[280,1300,202,901]" captionTargetId="figure@3.[265,1321,155,923]" captionTargetPageId="3" captionText="FIGURE 1. Scopimera globosa De Haan, 1835: dorsal view of carapace (a) and suborbital ridge and pterygostomian region (b); S. intermedia Balss, 1934: dorsal view of carapace (c) and suborbital ridge and pterygostomian region (d); S. ryukyuensis sp. nov.: dorsal view of carapace (e) and suborbital ridge and pterygostomian region (f). a, c, e: note arrow highlighting relative size of extra-orbital angle and solid line indicating direction of ridge behind the extra-orbital angle (which is diagnostic between S. ryukyuensis sp. nov. and S. globosa). b, d, f: arrow noting raised branchial region viewed from front in S. ryukyuensis sp. nov. (e) and relative conspicuousness of sub-orbital ridge: that of S. ryukyuensis sp. nov. being most pronounced." httpUri="https://zenodo.org/record/205329/files/figure.png" pageId="2" pageNumber="45">Figs. 1</figureCitation>
a, 6); suborbital ridge lined with at least 20 equal-sized small rounded granules (
<figureCitation id="D1D99FF6FF94ED114D476A29FD4DF894" box="[581,648,1824,1850]" captionStart="FIGURE 1" captionStartId="3.[151,256,934,958]" captionTargetBox="[280,1300,202,901]" captionTargetId="figure@3.[265,1321,155,923]" captionTargetPageId="3" captionText="FIGURE 1. Scopimera globosa De Haan, 1835: dorsal view of carapace (a) and suborbital ridge and pterygostomian region (b); S. intermedia Balss, 1934: dorsal view of carapace (c) and suborbital ridge and pterygostomian region (d); S. ryukyuensis sp. nov.: dorsal view of carapace (e) and suborbital ridge and pterygostomian region (f). a, c, e: note arrow highlighting relative size of extra-orbital angle and solid line indicating direction of ridge behind the extra-orbital angle (which is diagnostic between S. ryukyuensis sp. nov. and S. globosa). b, d, f: arrow noting raised branchial region viewed from front in S. ryukyuensis sp. nov. (e) and relative conspicuousness of sub-orbital ridge: that of S. ryukyuensis sp. nov. being most pronounced." httpUri="https://zenodo.org/record/205329/files/figure.png" pageId="2" pageNumber="45">Fig. 1</figureCitation>
b); external orbital angle obtuse, directing slightly anterior, leaving single notch along lateral border, followed by longitudinal crest extending two-thirds of carapace length (
<figureCitation id="D1D99FF6FF94ED114A636A4CFF66F82A" captionStart="FIGURE 1" captionStartId="3.[151,256,934,958]" captionTargetBox="[280,1300,202,901]" captionTargetId="figure@3.[265,1321,155,923]" captionTargetPageId="3" captionText="FIGURE 1. Scopimera globosa De Haan, 1835: dorsal view of carapace (a) and suborbital ridge and pterygostomian region (b); S. intermedia Balss, 1934: dorsal view of carapace (c) and suborbital ridge and pterygostomian region (d); S. ryukyuensis sp. nov.: dorsal view of carapace (e) and suborbital ridge and pterygostomian region (f). a, c, e: note arrow highlighting relative size of extra-orbital angle and solid line indicating direction of ridge behind the extra-orbital angle (which is diagnostic between S. ryukyuensis sp. nov. and S. globosa). b, d, f: arrow noting raised branchial region viewed from front in S. ryukyuensis sp. nov. (e) and relative conspicuousness of sub-orbital ridge: that of S. ryukyuensis sp. nov. being most pronounced." httpUri="https://zenodo.org/record/205329/files/figure.png" pageId="2" pageNumber="45">Figs. 1</figureCitation>
a, 6). Pterygostomian regions covered with rounded granules (
<figureCitation id="D1D99FF6FF94ED114C8E6A63FC15F82A" box="[908,976,1898,1924]" captionStart="FIGURE 1" captionStartId="3.[151,256,934,958]" captionTargetBox="[280,1300,202,901]" captionTargetId="figure@3.[265,1321,155,923]" captionTargetPageId="3" captionText="FIGURE 1. Scopimera globosa De Haan, 1835: dorsal view of carapace (a) and suborbital ridge and pterygostomian region (b); S. intermedia Balss, 1934: dorsal view of carapace (c) and suborbital ridge and pterygostomian region (d); S. ryukyuensis sp. nov.: dorsal view of carapace (e) and suborbital ridge and pterygostomian region (f). a, c, e: note arrow highlighting relative size of extra-orbital angle and solid line indicating direction of ridge behind the extra-orbital angle (which is diagnostic between S. ryukyuensis sp. nov. and S. globosa). b, d, f: arrow noting raised branchial region viewed from front in S. ryukyuensis sp. nov. (e) and relative conspicuousness of sub-orbital ridge: that of S. ryukyuensis sp. nov. being most pronounced." httpUri="https://zenodo.org/record/205329/files/figure.png" pageId="2" pageNumber="45">Fig. 1</figureCitation>
b). External maxillipeds convex, outer surface covered with rounded tubercles, merus slightly smaller than ischium, suture between them oblique, dactylus of palp just reaching suture. Ventral surface devoid of setae except short dense tufts of light-colored soft setae between bases of first and second ambulatory legs.
</paragraph>
<caption id="1D9DD3FBFF95ED104F956EAFFE5CFBD3" httpUri="https://zenodo.org/record/205329/files/figure.png" pageId="3" pageNumber="46" targetBox="[280,1300,202,901]" targetPageId="3">
<paragraph id="495D8373FF95ED104F956EAFFE5CFBD3" blockId="3.[151,1436,934,1149]" pageId="3" pageNumber="46">
<emphasis id="7B965F61FF95ED104F956EAFFED1FC10" bold="true" box="[151,276,934,958]" pageId="3" pageNumber="46">FIGURE 1</emphasis>
.
<taxonomicName id="8EE2F8F0FF95ED104E216EAFFD64FC13" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[291,673,934,957]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF95ED104E216EAFFE30FC13" box="[291,501,934,957]" italics="true" pageId="3" pageNumber="46">Scopimera globosa</emphasis>
De Haan, 1835
</taxonomicName>
: dorsal view of carapace (a) and suborbital ridge and pterygostomian region (b);
<taxonomicName id="8EE2F8F0FF95ED104E0D6ECFFDE4FC73" authority="Balss, 1934" authorityName="Balss" authorityYear="1934" box="[271,545,966,989]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="7B965F61FF95ED104E0D6ECFFE5BFC73" box="[271,414,966,989]" italics="true" pageId="3" pageNumber="46">S. intermedia</emphasis>
Balss, 1934
</taxonomicName>
: dorsal view of carapace (c) and suborbital ridge and pterygostomian region (d);
<taxonomicName id="8EE2F8F0FF95ED104A8A6ECFFED0FC53" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="7B965F61FF95ED104A8A6ECFFED0FC53" italics="true" pageId="3" pageNumber="46">S. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60A5E21AFF95ED104E1E6EEFFEB7FC50" box="[284,370,998,1022]" pageId="3" pageNumber="46" rank="species">
<emphasis id="7B965F61FF95ED104E1E6EEFFEB7FC50" bold="true" box="[284,370,998,1022]" pageId="3" pageNumber="46">sp. nov.</emphasis>
</taxonomicNameLabel>
: dorsal view of carapace (e) and suborbital ridge and pterygostomian region (f). a, c, e: note arrow highlighting relative size of extra-orbital angle and solid line indicating direction of ridge behind the extra-orbital angle (which is diagnostic between
<taxonomicName id="8EE2F8F0FF95ED104ED0692FFDAFFB93" box="[466,618,1062,1085]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="7B965F61FF95ED104ED0692FFDAFFB93" box="[466,618,1062,1085]" italics="true" pageId="3" pageNumber="46">S. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60A5E21AFF95ED104D72692FFD00FB90" box="[624,709,1062,1086]" pageId="3" pageNumber="46" rank="species">
<emphasis id="7B965F61FF95ED104D72692FFD00FB90" bold="true" box="[624,709,1062,1086]" pageId="3" pageNumber="46">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="8EE2F8F0FF95ED104DFA692FFCA3FB93" box="[760,870,1062,1085]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF95ED104DFA692FFCA3FB93" box="[760,870,1062,1085]" italics="true" pageId="3" pageNumber="46">S. globosa</emphasis>
</taxonomicName>
). b, d, f: arrow noting raised branchial region viewed from front in
<taxonomicName id="8EE2F8F0FF95ED104E25694FFE7AFBF3" box="[295,447,1094,1117]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="7B965F61FF95ED104E25694FFE7AFBF3" box="[295,447,1094,1117]" italics="true" pageId="3" pageNumber="46">S. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60A5E21AFF95ED104EC4694FFDDEFBF0" box="[454,539,1094,1118]" pageId="3" pageNumber="46" rank="species">
<emphasis id="7B965F61FF95ED104EC4694FFDDEFBF0" bold="true" box="[454,539,1094,1118]" pageId="3" pageNumber="46">sp. nov.</emphasis>
</taxonomicNameLabel>
(e) and relative conspicuousness of sub-orbital ridge: that of
<taxonomicName id="8EE2F8F0FF95ED104BAA694FFA84FBF3" box="[1192,1345,1094,1117]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="7B965F61FF95ED104BAA694FFA84FBF3" box="[1192,1345,1094,1117]" italics="true" pageId="3" pageNumber="46">S. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60A5E21AFF95ED104A45694FFA59FBF0" box="[1351,1436,1094,1118]" pageId="3" pageNumber="46" rank="species">
<emphasis id="7B965F61FF95ED104A45694FFA59FBF0" bold="true" box="[1351,1436,1094,1118]" pageId="3" pageNumber="46">sp. nov.</emphasis>
</taxonomicNameLabel>
being most pronounced.
</paragraph>
</caption>
<paragraph id="495D8373FF95ED104FC669AFFBE6FA6B" blockId="3.[151,1437,1190,2037]" pageId="3" pageNumber="46">
Entire cheliped covered with fine granules, total length about twice of carapace length in adult males, merus with single longitudinal ovate tympana on inner surface; carpus longitudinally ovate, slightly shorter than merus in adult males; palm as long as merus and fingers; tips of both fingers tapered to sharp points, inner margins of both weakly serrated, movable finger with faint triangular tooth on inner margin (
<figureCitation id="D1D99FF6FF95ED104A34681FFABAFA9E" box="[1334,1407,1302,1328]" captionStart="FIGURE 2" captionStartId="4.[151,256,1383,1407]" captionTargetBox="[280,1309,721,1345]" captionTargetId="figure@4.[265,1321,677,1362]" captionTargetPageId="4" captionText="FIGURE 2. Scopimera globosa De Haan, 1835: chela (a) and male abdomen (b); S. intermedia Balss, 1934: chela (c) and male abdomen (d); S. ryukyuensis sp. nov.: chela (e) and male abdomen (f). a, c, e: arrow pointing tooth at inner margin of movable finger: that of S. intermedia Balss, 1934, being most rudimentary. b, d, f: large arrows showing dimensions of the sixth abdominal somite and small arrow indicating the convexity of distal border of the fourth somite." httpUri="https://zenodo.org/record/205330/files/figure.png" pageId="3" pageNumber="46">Fig. 2</figureCitation>
a). Ambulatory legs relatively elongated, slender, first and second legs longest, subequal to each other; fourth leg shortest; merus laterally compressed, tympana entire, occupying most of segment; dactylus tapered to sharp tip, slightly longer than propodus; fourth dactylus curved slightly upwards. Posterior border of merus, carpus, propodus of ambulatory legs lined sparsely with long stiff dark-colored setae.
</paragraph>
<paragraph id="495D8373FF95ED104FC668D8FEDAF92E" blockId="3.[151,1437,1190,2037]" pageId="3" pageNumber="46">
Male abdomen elongated, telson distally rounded, sixth somite as long as broad, lateral borders of fifth conspicuously concave on proximal half, proximal margin much narrower; fourth somite broad, distal margin very convex (
<figureCitation id="D1D99FF6FF95ED104E3B6B12FE44F99B" box="[313,385,1563,1589]" captionStart="FIGURE 2" captionStartId="4.[151,256,1383,1407]" captionTargetBox="[280,1309,721,1345]" captionTargetId="figure@4.[265,1321,677,1362]" captionTargetPageId="4" captionText="FIGURE 2. Scopimera globosa De Haan, 1835: chela (a) and male abdomen (b); S. intermedia Balss, 1934: chela (c) and male abdomen (d); S. ryukyuensis sp. nov.: chela (e) and male abdomen (f). a, c, e: arrow pointing tooth at inner margin of movable finger: that of S. intermedia Balss, 1934, being most rudimentary. b, d, f: large arrows showing dimensions of the sixth abdominal somite and small arrow indicating the convexity of distal border of the fourth somite." httpUri="https://zenodo.org/record/205330/files/figure.png" pageId="3" pageNumber="46">Fig. 2</figureCitation>
b). G1 slender, curved dorsally, distal end rounded with inward-dorsal radiating brush of longer setae of approximately same length, and a row of short setae around the opening on the inner surface (
<figureCitation id="D1D99FF6FF95ED104F9D6B6FFF21F92E" box="[159,228,1638,1664]" captionStart="FIGURE 3" captionStartId="6.[151,255,1641,1665]" captionTargetBox="[156,1430,197,1610]" captionTargetId="figure@6.[151,1436,194,1619]" captionTargetPageId="6" captionText="FIGURE 3. Scopimera globosa De Haan, 1835: G 1 entire view (a), tip (b) and magnification of setae (c); S. intermedia Balss, 1934: G 1 entire view (d), tip (e) and magnification of setae (f); S. ryukyuensis sp. nov.: G 1 entire view (g), tip (h) and tip observed under light microscope (i). Scale bars in µm." httpUri="https://zenodo.org/record/205331/files/figure.png" pageId="3" pageNumber="46">Fig. 3</figureCitation>
ac).
</paragraph>
<paragraph id="495D8373FF95ED104FC66B82FD9BF95E" blockId="3.[151,1437,1190,2037]" pageId="3" pageNumber="46">
<emphasis id="7B965F61FF95ED104FC66B82FE93F90B" bold="true" box="[196,342,1675,1701]" pageId="3" pageNumber="46">Coloration.</emphasis>
Grayish carapace, merus of external maxillipeds dark or with random blotches, chelipeds gray with pale palm and fingers, ambulatory legs banded dark and light gray (
<figureCitation id="D1D99FF6FF95ED104B4A6BB8FB54F965" box="[1096,1169,1713,1739]" captionStart="FIGURE 4" captionStartId="8.[151,255,1858,1882]" captionTargetBox="[183,1394,200,1816]" captionTargetId="figure@8.[178,1403,194,1825]" captionTargetPageId="8" captionText="FIGURE 4. Photographs in life (a e). Scopimera globosa De Haan, 1835: male frontal view (a) and dorsal view (b); S. intermedia Balss, 1934: male frontal view (c) and dorsal view (d); and S. ryukyuensis sp. nov.: male frontal view (e) and close-up of external maxillipeds from preserved specimens, showing ring-like marking on merus (f)." httpUri="https://zenodo.org/record/205332/files/figure.png" pageId="3" pageNumber="46">Fig. 4</figureCitation>
a, b) (also see
<bibRefCitation id="2D73FE82FF95ED104A436BB8FF13F95E" author="Miyake, S." bookContentInfo="277 pp." journalOrPublisher="Hoikusha, Osaka" pageId="3" pageNumber="46" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in Color. II Brachyura (Crabs). Hoikusha, Osaka, 277 pp. (in Japanese)" title="Japanese Crustacean Decapods and Stomatopods in Color. II Brachyura (Crabs)" type="book" year="1983">Miyake 1983</bibRefCitation>
: Pl. 56(8);
<bibRefCitation id="2D73FE82FF95ED104E556BDFFDD3F95E" author="Minemizu, R." bookContentInfo="344 pp." box="[343,534,1750,1776]" journalOrPublisher="Tokyo" pageId="3" pageNumber="46" refString="Minemizu, R. (2000) Marine Decapod and Stomatopod Crustaceans Mainly from Japan. Bunnichi-Sogo Shuppan, Tokyo, 344 pp. (in Japanese)" title="Marine Decapod and Stomatopod Crustaceans Mainly from Japan. Bunnichi-Sogo Shuppan" type="book" year="2000">Minemizu 2000</bibRefCitation>
: 309).
</paragraph>
</subSubSection>
<subSubSection id="01F8D0F8FF95ED104FC66BF2FA96F8BB" box="[196,1363,1787,1813]" pageId="3" pageNumber="46" type="distribution">
<paragraph id="495D8373FF95ED104FC66BF2FA96F8BB" blockId="3.[151,1437,1190,2037]" box="[196,1363,1787,1813]" pageId="3" pageNumber="46">
<emphasis id="7B965F61FF95ED104FC66BF2FE98F8BB" bold="true" box="[196,349,1787,1813]" pageId="3" pageNumber="46">Distribution</emphasis>
.
<collectingCountry id="31F5C3E3FF95ED104E6E6BF2FE6AF8BB" box="[364,431,1787,1813]" name="Japan" pageId="3" pageNumber="46">Japan</collectingCountry>
(
<typeStatus id="96593DD1FF95ED104EBC6BF2FE37F8BB" box="[446,498,1787,1813]" pageId="3" pageNumber="46">type</typeStatus>
locality),
<collectingCountry id="31F5C3E3FF95ED104D6B6BF2FD74F8BB" box="[617,689,1787,1813]" name="South Korea" pageId="3" pageNumber="46">Korea</collectingCountry>
(
<bibRefCitation id="2D73FE82FF95ED104DC26BF2FC9BF8BB" author="Kamita, T." bookContentInfo="289 pp." box="[704,862,1787,1813]" journalOrPublisher="The Fisheries Society of Chosen, Keijo, Korea" pageId="3" pageNumber="46" refString="Kamita, T. (1941) Studies of the Decapod Crustaceans of Chosen. Pt. 1. Crabs. The Fisheries Society of Chosen, Keijo, Korea, 289 pp. (in Japanese)" title="Studies of the Decapod Crustaceans of Chosen. Pt. 1. Crabs" type="book" year="1941">Kamita 1941</bibRefCitation>
;
<bibRefCitation id="2D73FE82FF95ED104C6B6BF2FC24F8BB" author="Kim, H. S." box="[873,993,1787,1813]" journalOrPublisher="Seoul, pp. 589 - 682. (in Korean)" pageId="3" pageNumber="46" refString="Kim, H. S. (1973) Anomura-Brachyura. In: Illustrated Encyclopedia of Fauna &amp; Flora of Korea. Vol. 14. Samhwa, Seoul, pp. 589 - 682. (in Korean)" title="Anomura-Brachyura" type="book" volumeTitle="Illustrated Encyclopedia of Fauna &amp; Flora of Korea. Vol. 14. Samhwa" year="1973">Kim 1973</bibRefCitation>
) and North
<collectingCountry id="31F5C3E3FF95ED104B6C6BF2FB71F8BB" box="[1134,1204,1787,1813]" name="China" pageId="3" pageNumber="46">China</collectingCountry>
(
<bibRefCitation id="2D73FE82FF95ED104BC16BF2FA83F8BB" author="Shen, C. - J." box="[1219,1350,1787,1813]" journalOrPublisher="Zoologia Sinica, Series A" pageId="3" pageNumber="46" pagination="1 - 300" part="9" refString="Shen, C. - J. (1932) The brachyuran Crustacea of North China. Zoologia Sinica, Series A, 9, 1 - 300." title="The brachyuran Crustacea of North China" type="journal article" year="1932">Shen 1932</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="01F8D0F8FF95ED174FC66A29FCA7FD39" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46" type="discussion">
<paragraph id="495D8373FF95ED174FC66A29FCA7FD39" blockId="3.[151,1437,1190,2037]" lastBlockId="4.[151,1437,152,663]" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46">
<emphasis id="7B965F61FF95ED104FC66A29FEF3F894" bold="true" box="[196,310,1824,1850]" pageId="3" pageNumber="46">Remarks</emphasis>
.
<taxonomicName id="8EE2F8F0FF95ED104E476A28FDEDF894" box="[325,552,1825,1850]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF95ED104E476A28FDEDF894" box="[325,552,1825,1850]" italics="true" pageId="3" pageNumber="46">Scopimera globosa</emphasis>
</taxonomicName>
is a common species on muddy and sandy shores in Honshu,
<collectingCountry id="31F5C3E3FF95ED104A046A28FA8CF895" box="[1286,1353,1825,1851]" name="Japan" pageId="3" pageNumber="46">Japan</collectingCountry>
(
<bibRefCitation id="2D73FE82FF95ED104A5B6A28FF11F8CE" author="Sakai, T." journalOrPublisher="Tokyo, pp. 365 - 741 + pls" pageId="3" pageNumber="46" pagination="42 - 111" refString="Sakai, T. (1939) Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo, Tokyo, pp. 365 - 741 + pls. 42 - 111." title="Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo" type="book chapter" year="1939">Sakai 1939</bibRefCitation>
,
<bibRefCitation id="2D73FE82FF95ED104FE26A4FFEDAF8CE" author="Sakai, T." bookContentInfo="773 pp" box="[224,287,1862,1888]" journalOrPublisher="Kodansha, Tokyo" pageId="3" pageNumber="46" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, 773 pp + 251 pls." title="Crabs of Japan and the Adjacent Seas" type="book" year="1976">1976</bibRefCitation>
). Sketch of G1 of
<taxonomicName id="8EE2F8F0FF95ED104EF16A4FFCE0F8CE" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[499,805,1862,1888]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF95ED104EF16A4FFDA8F8F1" box="[499,621,1862,1887]" italics="true" pageId="3" pageNumber="46">S. globosa</emphasis>
De Haan, 1835
</taxonomicName>
<typeStatus id="96593DD1FF95ED104C296A4FFC53F8CE" box="[811,918,1862,1888]" pageId="3" pageNumber="46" type="lectotype">lectotype</typeStatus>
matches the present material collected from Honshu,
<collectingCountry id="31F5C3E3FF95ED104E006A62FE8CF82B" box="[258,329,1899,1925]" name="Japan" pageId="3" pageNumber="46">Japan</collectingCountry>
. However, of the over 300
<taxonomicName id="8EE2F8F0FF95ED104D896A65FCCCF82B" box="[651,777,1900,1925]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="7B965F61FF95ED104D896A65FCCCF82B" box="[651,777,1900,1925]" italics="true" pageId="3" pageNumber="46">Scopimera</emphasis>
</taxonomicName>
specimens collected and observed from more than four localities from
<collectingCountry id="31F5C3E3FF95ED104E526A98FE1BF805" box="[336,478,1937,1963]" name="Hong Kong" pageId="3" pageNumber="46">Hong Kong</collectingCountry>
(in South
<collectingCountry id="31F5C3E3FF95ED104D626A98FD6FF805" box="[608,682,1937,1963]" name="China" pageId="3" pageNumber="46">China</collectingCountry>
) and more than six localities from
<collectingCountry id="31F5C3E3FF95ED104B556A98FB76F805" box="[1111,1203,1937,1963]" name="Taiwan" pageId="3" pageNumber="46">Taiwan</collectingCountry>
, no
<taxonomicName id="8EE2F8F0FF95ED104BE46A98FAA3F804" box="[1254,1382,1937,1962]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF95ED104BE46A98FAA3F804" box="[1254,1382,1937,1962]" italics="true" pageId="3" pageNumber="46">S. globosa</emphasis>
</taxonomicName>
was found.
<bibRefCitation id="2D73FE82FF95ED104FEB6ABFFEBDF87E" author="Shen, C. - J." box="[233,376,1974,2000]" journalOrPublisher="Hong Kong Fisheries Research Station Journal" pageId="3" pageNumber="46" pagination="211 - 242" part="1" refString="Shen, C. - J. (1940) The brachyuran fauna of Hong Kong. Hong Kong Fisheries Research Station Journal, 1, 211 - 242." title="The brachyuran fauna of Hong Kong" type="journal article" year="1940">Shen (1940)</bibRefCitation>
reported
<taxonomicName id="8EE2F8F0FF95ED104EE56ABFFDA4F861" box="[487,609,1974,1999]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF95ED104EE56ABFFDA4F861" box="[487,609,1974,1999]" italics="true" pageId="3" pageNumber="46">S. globosa</emphasis>
</taxonomicName>
from Tsuen Wan, Wong Chuk Hang, and Tai Po in
<collectingCountry id="31F5C3E3FF95ED104BB76ABFFA84F87E" box="[1205,1345,1974,2000]" name="Hong Kong" pageId="3" pageNumber="46">Hong Kong</collectingCountry>
, but the first two sites are today already heavily urbanized and almost devoid of any natural coasts.
<bibRefCitation id="2D73FE82FF95ED174BC46AD2FEE3FF1C" author="Morton, B. &amp; Morton, J." bookContentInfo="350 pp." journalOrPublisher="Hong Kong University Press, Hong Kong" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46" refString="Morton, B. &amp; Morton, J. (1983) The Sea Shore Ecology of Hong Kong. Hong Kong University Press, Hong Kong, 350 pp." title="The Sea Shore Ecology of Hong Kong" type="book" year="1983">Morton &amp; Morton (1983: 185)</bibRefCitation>
stated “
<taxonomicName id="8EE2F8F0FF92ED174E896D91FDB1FF1F" box="[395,628,152,177]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF92ED174E896D91FDB1FF1F" box="[395,628,152,177]" italics="true" pageId="4" pageNumber="47">Scopimera globosa</emphasis>
</taxonomicName>
is as common at Tai Tam as
<taxonomicName id="8EE2F8F0FF92ED174CD86D91FBB9FF1F" box="[986,1148,152,177]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="7B965F61FF92ED174CD86D91FBB9FF1F" box="[986,1148,152,177]" italics="true" pageId="4" pageNumber="47">S. intermedia</emphasis>
</taxonomicName>
is at Shiu Hau…”. The second author visited both Tai Po and Tai Tam, and specimens from Tai Po are
<taxonomicName id="8EE2F8F0FF92ED174B7D6DB4FAE2FF78" box="[1151,1319,189,214]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="7B965F61FF92ED174B7D6DB4FAE2FF78" box="[1151,1319,189,214]" italics="true" pageId="4" pageNumber="47">S. intermedia</emphasis>
</taxonomicName>
while no
<taxonomicName id="8EE2F8F0FF92ED174F956DEBFED2FF55" box="[151,279,226,251]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="7B965F61FF92ED174F956DEBFED2FF55" box="[151,279,226,251]" italics="true" pageId="4" pageNumber="47">Scopimera</emphasis>
</taxonomicName>
sp. was found in Tai Tam, where a reservoir has been constructed. It is thus unclear whether
<taxonomicName id="8EE2F8F0FF92ED174A876DEBFF3DFE8F" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF92ED174A876DEBFF3DFE8F" italics="true" pageId="4" pageNumber="47">S. globosa</emphasis>
</taxonomicName>
really exists in
<collectingCountry id="31F5C3E3FF92ED174EBC6C01FD94FE8C" box="[446,593,264,290]" name="Hong Kong" pageId="4" pageNumber="47">Hong Kong</collectingCountry>
. As for Taiwanese records of
<taxonomicName id="8EE2F8F0FF92ED174CC06C01FB84FE8F" box="[962,1089,264,289]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF92ED174CC06C01FB84FE8F" box="[962,1089,264,289]" italics="true" pageId="4" pageNumber="47">S. globosa</emphasis>
</taxonomicName>
, the first was reported from Danshuei (= Tamsui),
<collectingRegion id="8B264D91FF92ED174EA06C24FE29FEE9" box="[418,492,301,327]" country="Taiwan" name="Taipei" pageId="4" pageNumber="47">Taipei</collectingRegion>
County, by
<bibRefCitation id="2D73FE82FF92ED174D7C6C24FCFBFEE9" author="Takahasi, S." box="[638,830,301,327]" journalOrPublisher="Transactions of the Natural History Society of Formosa" pageId="4" pageNumber="47" pagination="1 - 14" part="24" refString="Takahasi, S. (1934) An ecological study of the littoral animals near the mouth of the Tamsui. Transactions of the Natural History Society of Formosa, 24, 1 - 14. (in Japanese)" title="An ecological study of the littoral animals near the mouth of the Tamsui" type="journal article" year="1934">Takahasi (1934)</bibRefCitation>
and have been recorded on numerous localities on the main island and
<collectingRegion id="8B264D91FF92ED174E946C5BFE3DFEC2" box="[406,504,338,364]" country="Taiwan" name="Penghu" pageId="4" pageNumber="47">Penghu</collectingRegion>
. In a review of ocypodoid crabs, however, a typical
<taxonomicName id="8EE2F8F0FF92ED174B8F6C5BFAF0FEC5" box="[1165,1333,338,363]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="7B965F61FF92ED174B8F6C5BFAF0FEC5" box="[1165,1333,338,363]" italics="true" pageId="4" pageNumber="47">S. intermedia</emphasis>
</taxonomicName>
G1 was illustrated under the name
<taxonomicName id="8EE2F8F0FF92ED174EE66C71FDA0FE3F" box="[484,613,376,401]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF92ED174EE66C71FDA0FE3F" box="[484,613,376,401]" italics="true" pageId="4" pageNumber="47">S. globosa</emphasis>
</taxonomicName>
(see
<bibRefCitation id="2D73FE82FF92ED174DAA6C71FC49FE3C" author="Huang, J. - F. &amp; Yu, H. - P. &amp; Takeda, M." box="[680,908,376,402]" journalOrPublisher="Bulletin of the Institute of Zoology, Academia Sinica" pageId="4" pageNumber="47" pagination="141 - 161" part="31" refString="Huang, J. - F., Yu, H. - P. &amp; Takeda, M. (1992) A review of the ocypodid and mictyrid crabs (Crustacea: Decapoda: Brachyura) in Taiwan. Bulletin of the Institute of Zoology, Academia Sinica, 31, 141 - 161." title="A review of the ocypodid and mictyrid crabs (Crustacea: Decapoda: Brachyura) in Taiwan" type="journal article" year="1992">
Huang
<emphasis id="7B965F61FF92ED174C006C71FC84FE3F" box="[770,833,376,401]" italics="true" pageId="4" pageNumber="47">et al.</emphasis>
1992
</bibRefCitation>
: fig. 16). Checking available specimens in National
<collectingCountry id="31F5C3E3FF92ED174E016C94FE9CFE19" box="[259,345,413,439]" name="Taiwan" pageId="4" pageNumber="47">Taiwan</collectingCountry>
Museum and National Museum of Natural Science which has been collected and named as
<taxonomicName id="8EE2F8F0FF92ED174A846C94FF30FE75" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF92ED174A846C94FF30FE75" italics="true" pageId="4" pageNumber="47">S. globosa</emphasis>
</taxonomicName>
between 1980s and today, we could find no specimens referable to
<taxonomicName id="8EE2F8F0FF92ED174B1D6CCBFB5FFE75" box="[1055,1178,450,475]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF92ED174B1D6CCBFB5FFE75" box="[1055,1178,450,475]" italics="true" pageId="4" pageNumber="47">S. globosa</emphasis>
</taxonomicName>
. All this material has proven to be
<taxonomicName id="8EE2F8F0FF92ED174E306CE1FE0AFDAF" box="[306,463,488,513]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="7B965F61FF92ED174E306CE1FE0AFDAF" box="[306,463,488,513]" italics="true" pageId="4" pageNumber="47">S. intermedia</emphasis>
</taxonomicName>
instead. Thus the presence of
<taxonomicName id="8EE2F8F0FF92ED174C376CE1FC75FDAF" box="[821,944,488,513]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF92ED174C376CE1FC75FDAF" box="[821,944,488,513]" italics="true" pageId="4" pageNumber="47">S. globosa</emphasis>
</taxonomicName>
in
<collectingCountry id="31F5C3E3FF92ED174CD46CE1FBE9FDAC" box="[982,1068,488,514]" name="Taiwan" pageId="4" pageNumber="47">Taiwan</collectingCountry>
and
<collectingCountry id="31F5C3E3FF92ED174B646CE1FB35FDAC" box="[1126,1264,488,514]" name="Hong Kong" pageId="4" pageNumber="47">Hong Kong</collectingCountry>
is not verified in this study and its southern limit along the Chinese coast remains unclear. However, our extensive collections in
<collectingCountry id="31F5C3E3FF92ED174E3F6F3BFE56FDE2" box="[317,403,562,588]" name="Taiwan" pageId="4" pageNumber="47">Taiwan</collectingCountry>
and
<collectingCountry id="31F5C3E3FF92ED174ECC6F3BFD9DFDE2" box="[462,600,562,588]" name="Hong Kong" pageId="4" pageNumber="47">Hong Kong</collectingCountry>
fail to yield any
<taxonomicName id="8EE2F8F0FF92ED174C266F3BFC65FDE5" box="[804,928,562,587]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF92ED174C266F3BFC65FDE5" box="[804,928,562,587]" italics="true" pageId="4" pageNumber="47">S. globosa</emphasis>
</taxonomicName>
, suggesting this species is probably absent in these two regions. There was one record of “
<taxonomicName id="8EE2F8F0FF92ED174DCF6F51FC8DFDDF" box="[717,840,600,625]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="7B965F61FF92ED174DCF6F51FC8DFDDF" box="[717,840,600,625]" italics="true" pageId="4" pageNumber="47">S. globosa</emphasis>
</taxonomicName>
” from
<collectingCountry id="31F5C3E3FF92ED174C9D6F51FBD2FDDC" box="[927,1047,600,626]" name="Singapore" pageId="4" pageNumber="47">Singapore</collectingCountry>
(
<bibRefCitation id="2D73FE82FF92ED174B246F51FB21FDDC" author="Tan, C. G. S." box="[1062,1252,600,626]" journalOrPublisher="Hydrobiologia" pageId="4" pageNumber="47" pagination="75 - 84" part="285" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1994) An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia, 285, 75 - 84." title="An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore" type="journal article" year="1994">Tan &amp; Ng 1994</bibRefCitation>
) which may be misidentified with
<taxonomicName id="8EE2F8F0FF92ED174E766F74FDD4FD38" box="[372,529,637,662]" class="Malacostraca" family="Dotillidae" genus="Scopimera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="7B965F61FF92ED174E766F74FDD4FD38" box="[372,529,637,662]" italics="true" pageId="4" pageNumber="47">S. intermedia</emphasis>
</taxonomicName>
or other unrecorded species.
</paragraph>
</subSubSection>
</treatment>
</document>