975 lines
189 KiB
XML
975 lines
189 KiB
XML
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<mods:title id="4C905BAF0362B2B5040048A498B13F2F">Redescription of the Central American genus Moribaetis Waltz & McCafferty 1985 (Ephemeroptera, Baetidae)</mods:title>
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<taxonomicName id="05B74D02FF60AC74FF4887D6FD9493CF" ID-CoL="5V72" ID-ENA="603901" authority="Waltz & McCafferty 1985" authorityName="Waltz & McCafferty" authorityYear="1985" box="[151,613,1203,1230]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
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<emphasis id="F0C3EA93FF60AC74FF4887D6FD9493CF" bold="true" box="[151,613,1203,1230]" pageId="1" pageNumber="232">
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<emphasis id="F0C3EA93FF60AC74FF4887D6FEE893CC" bold="true" box="[151,281,1203,1229]" italics="true" pageId="1" pageNumber="232">Moribaetis</emphasis>
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<bibRefCitation id="A6264B70FF60AC74FEFF87D1FD9493CF" author="Waltz, R. D. & McCafferty, W. P." box="[288,613,1204,1230]" pageId="1" pageNumber="232" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985</bibRefCitation>
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<subSubSection id="8AAD650AFF60AC74FF4887B3FED693EE" box="[151,295,1238,1263]" pageId="1" pageNumber="232" type="description">
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<paragraph id="C2083681FF60AC74FF4887B3FED693EE" blockId="1.[151,613,1203,1263]" box="[151,295,1238,1263]" pageId="1" pageNumber="232">
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(
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<figureCitation id="5A8C2A04FF60AC74FF4087B3FEEF93EE" box="[159,286,1238,1263]" captionStart-0="FIGURES 1–15" captionStart-1="FIGURES 16–21" captionStart-10="FIGURES 71–76" captionStart-11="FIGURES 77–84" captionStart-12="FIGURES 85–88" captionStart-13="FIGURES 89–95" captionStart-14="FIGURES 96–101" captionStart-15="FIGURES 102–105" captionStart-2="FIGURES 22–24" captionStart-3="FIGURES 25–33" captionStart-4="FIGURES 34–43" captionStart-5="FIGURES 44–47" captionStart-6="FIGURES 48–52" captionStart-7="FIGURES 53–55" captionStart-8="FIGURES 56–61" captionStart-9="FIGURES 62–70" captionStartId-0="7.[151,264,1892,1915]" captionStartId-1="8.[151,264,1850,1873]" captionStartId-10="19.[151,264,1752,1775]" captionStartId-11="20.[151,264,1888,1911]" captionStartId-12="21.[151,264,1737,1760]" captionStartId-13="22.[151,264,1940,1963]" captionStartId-14="23.[151,264,1536,1559]" captionStartId-15="24.[151,264,1973,1996]" captionStartId-2="9.[151,264,1680,1703]" captionStartId-3="10.[151,264,1857,1880]" captionStartId-4="11.[151,264,1837,1860]" captionStartId-5="12.[151,264,1365,1388]" captionStartId-6="13.[151,264,1889,1912]" captionStartId-7="14.[151,264,1761,1784]" captionStartId-8="15.[151,264,1780,1803]" captionStartId-9="18.[151,264,1878,1901]" captionTargetBox-0="[177,1402,193,1870]" captionTargetBox-1="[151,1436,213,1829]" captionTargetBox-10="[150,1436,193,1728]" captionTargetBox-11="[183,1389,193,1851]" captionTargetBox-12="[152,1435,195,1675]" captionTargetBox-13="[151,1436,409,1919]" captionTargetBox-14="[151,1437,193,1515]" captionTargetBox-15="[151,1436,574,1953]" captionTargetBox-2="[159,1392,210,1656]" captionTargetBox-3="[151,1436,193,1836]" captionTargetBox-4="[150,1436,193,1817]" captionTargetBox-5="[153,1434,197,1339]" captionTargetBox-6="[150,1436,193,1868]" captionTargetBox-7="[150,1436,193,1740]" captionTargetBox-8="[150,1435,301,1759]" captionTargetBox-9="[197,1385,217,1850]" captionTargetId-0="figure@7.[177,1402,193,1870]" captionTargetId-1="figure@8.[151,1436,193,1829]" captionTargetId-10="figure@19.[150,1436,193,1731]" captionTargetId-11="figure@20.[183,1389,193,1851]" captionTargetId-12="figure@21.[152,1435,193,1717]" captionTargetId-13="figure@22.[151,1436,373,1919]" captionTargetId-14="figure@23.[151,1437,193,1515]" captionTargetId-15="figure@24.[151,1436,574,1953]" captionTargetId-2="figure@9.[151,1436,193,1660]" captionTargetId-3="figure@10.[151,1436,193,1836]" captionTargetId-4="figure@11.[150,1436,193,1817]" captionTargetId-5="figure@12.[151,1436,193,1345]" captionTargetId-6="figure@13.[150,1436,193,1868]" captionTargetId-7="figure@14.[150,1436,193,1740]" captionTargetId-8="figure@15.[150,1435,301,1759]" captionTargetId-9="figure@18.[197,1385,217,1850]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-10="19" captionTargetPageId-11="20" captionTargetPageId-12="21" captionTargetPageId-13="22" captionTargetPageId-14="23" captionTargetPageId-15="24" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionTargetPageId-6="13" captionTargetPageId-7="14" captionTargetPageId-8="15" captionTargetPageId-9="18" captionText-0="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." captionText-1="FIGURES 16–21. Mandibles of Moribaetis maculipennis. 16–17, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 18–21, left and right mandibles of last instar larva just after moult and their fragments enlarged. Abbreviations: kd, kinetodontium; prs, prostheca." captionText-10="FIGURES 71–76. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 71–72, labrum, focus on dorsal side; 73, focus on anterior margin; 74, focus on ventral side; 75, head with labrum; 76, labium." captionText-11="FIGURES 77–84. Larvae of Moribaetis latipennis sp. n. 77–78, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 79–82, left and right mandibles of last instar larva just after moult and their fragments enlarged; 83, apex of maxilla; 84, apices of glossae and paraglossae before moult to next (last) instar, with new paraglossae visible inside (arrows show three rows of apical setae on paraglossae). Abbreviations: ds1, ds2, ds3, dentisetae; kd, kinetodontium; prs, prostheca." captionText-12="FIGURES 85–88. Larva of Moribaetis latipennis sp. n. 85, bases of fore legs; 86, the same, with black lines showing colorless tracheal gill and its colorless tracheae, poorly visible on this photo; 87–88, claws of the same two legs." captionText-13="FIGURES 89–95. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 89–91, fore, middle and hind legs; 92–94, apex of femur of fore, middle and hind legs; 95, inner margin of femur of fore leg." captionText-14="FIGURES 96–101. Moribaetis latipennis sp. n. (holotype). 96–97, subimago ready to moult to imago; 98, fragment of subimaginal exuviae of wing with fragment of vein; 99, imaginal hind wing extracted from subimago; 100, imaginal fore wing extracted from subimago; 101, subimaginal genitals." captionText-15="FIGURES 102–105. Moribaetis salvini (lectotype, photo by Ben Price). 102, lateral view; 103, fragment of cercus; 104, dorsal view and tube with genitals; 105, labels." captionText-2="FIGURES 22–24. Larva of Moribaetis maculipennis. 22, labium (at left ventral view, at right dorsal view); 23, claw (anterior view); 24, its cross section. Abbreviation: d-m, seta of dorso-median row." captionText-3="FIGURES 25–33. Legs of Moribaetis maculipennis. 25–27, larval fore, middle and hind legs; 28, fragment of fore tibia of female subimago; 29, the same, another specimen; 30, fragment of middle tibia of female subimago; 31–32, fragments of fore and middle leg of female subimago; 33, fragment of femur of larval fore leg (25–27 and 29–32, the same individual). Abbreviation: pt-ti, patella-tibial suture." captionText-4="FIGURES 34–43. Moribaetis maculipennis. 34–35, male subimago just after moult; 36, abdominal sternum IX of last instar male larva with gonostyli visible inside; 37, gonostyli of male subimago; 38, exuviae of subimaginal gonostyli; 39, genitals of male imago; 40, caudalii of last instar larva with crumpled sibimaginal cerci visible inside; 41–42, proximal part of imaginal cercus and its subimaginal exuviae (arrows show correspondence of their joints); 43, proximal part of imaginal cercus of another specimen." captionText-5="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." captionText-6="FIGURES 48–52. Moribaetis maculipennis. 48–50, male imago; 51, subimaginal exuviae of left postsubalar sclerite and lateropostnotal crest; 52, subimaginal exuviae of left half of mesonotum." captionText-7="FIGURES 53–55. Moribaetis maculipennis. 53–54, female imago; 55, hind wing." captionText-8="FIGURES 56–61. Eggs of Moribaetis maculipennis. 56, view in transmitted light; 57–59, eggs covered with coat; 60–61, egg without coat." captionText-9="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri-0="https://zenodo.org/record/2609839/files/figure.png" httpUri-1="https://zenodo.org/record/2609841/files/figure.png" httpUri-10="https://zenodo.org/record/2609859/files/figure.png" httpUri-11="https://zenodo.org/record/2609861/files/figure.png" httpUri-12="https://zenodo.org/record/2609863/files/figure.png" httpUri-13="https://zenodo.org/record/2609865/files/figure.png" httpUri-14="https://zenodo.org/record/2609867/files/figure.png" httpUri-15="https://zenodo.org/record/2609869/files/figure.png" httpUri-2="https://zenodo.org/record/2609843/files/figure.png" httpUri-3="https://zenodo.org/record/2609845/files/figure.png" httpUri-4="https://zenodo.org/record/2609847/files/figure.png" httpUri-5="https://zenodo.org/record/2609849/files/figure.png" httpUri-6="https://zenodo.org/record/2609851/files/figure.png" httpUri-7="https://zenodo.org/record/2609853/files/figure.png" httpUri-8="https://zenodo.org/record/2609855/files/figure.png" httpUri-9="https://zenodo.org/record/2609857/files/figure.png" pageId="1" pageNumber="232">Figs 1–105</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="8AAD650AFF60AC74FF48867DFCEE924D" pageId="1" pageNumber="232" type="reference_group">
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<paragraph id="C2083681FF60AC74FF48867DFCBF922F" blockId="1.[151,846,1304,1387]" box="[151,846,1304,1326]" pageId="1" pageNumber="232">
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<treatmentCitationGroup id="E2A711AFFF60AC74FF48867DFCBF922F" box="[151,846,1304,1326]" pageId="1" pageNumber="232">
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Subgenus
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<taxonomicName id="05B74D02FF60AC74FF20867DFD40922F" authority="Waltz & McCafferty 1985: 243" authorityName="Waltz & McCafferty" authorityPageNumber="243" authorityYear="1985" box="[255,689,1304,1326]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="subGenus" subGenus="Moribaetis">
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<emphasis id="F0C3EA93FF60AC74FF20867DFE9F922F" box="[255,366,1304,1326]" italics="true" pageId="1" pageNumber="232">Moribaetis</emphasis>
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<treatmentCitation id="43161090FF60AC74FEAA867DFD40922F" author="Waltz, R. D. & McCafferty, W. P." box="[373,689,1304,1326]" page="243" pageId="1" pageNumber="232" year="1985">
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<bibRefCitation id="A6264B70FF60AC74FEAA867DFD73922F" author="Waltz, R. D. & McCafferty, W. P." box="[373,642,1304,1326]" pageId="1" pageNumber="232" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985</bibRefCitation>
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: 243
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</treatmentCitation>
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(imago, larva).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="C2083681FF60AC74FF488653FCEE924D" blockId="1.[151,846,1304,1387]" box="[151,799,1334,1356]" pageId="1" pageNumber="232">
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<treatmentCitationGroup id="E2A711AFFF60AC74FF488653FCEE924D" box="[151,799,1334,1356]" pageId="1" pageNumber="232">
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Genus
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<taxonomicName id="05B74D02FF60AC74FF018653FD3F924D" ID-CoL="5V72" authority=": Lugo-Ortiz & McCafferty 1996: 368" authorityName=": Lugo-Ortiz & McCafferty" authorityPageNumber="368" authorityYear="1996" box="[222,718,1334,1356]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
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<emphasis id="F0C3EA93FF60AC74FF018653FEBC924D" box="[222,333,1334,1356]" italics="true" pageId="1" pageNumber="232">Moribaetis</emphasis>
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:
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<treatmentCitation id="43161090FF60AC74FE858653FD3F924D" author="Lugo-Ortiz, C. R. & McCafferty, W. P." box="[346,718,1334,1356]" page="368" pageId="1" pageNumber="232" year="1996">
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<bibRefCitation id="A6264B70FF60AC74FE858653FD6F924D" author="Lugo-Ortiz, C. R. & McCafferty, W. P." box="[346,670,1334,1356]" pageId="1" pageNumber="232" pagination="367 - 380" refId="ref11489" refString="Lugo-Ortiz, C. R. & McCafferty, W. P. (1996) Phylogeny and classification of the Baetodes complex (Ephemeroptera: Baetidae), with description of a new genus. Journal of the North American Benthological Society, 15 (3), 367 - 380. https: // doi. org / 10.2307 / 1467283" type="journal article" year="1996">Lugo-Ortiz & McCafferty 1996</bibRefCitation>
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: 368
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</treatmentCitation>
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</taxonomicName>
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(larva).
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|
||
<paragraph id="C2083681FF60AC74FF488633FD7B926A" blockId="1.[151,846,1304,1387]" box="[151,650,1365,1387]" pageId="1" pageNumber="232">
|
||
<typeStatus id="1D0C8823FF60AC74FF488633FF38926A" box="[151,201,1366,1387]" pageId="1" pageNumber="232">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="05B74D02FF60AC74FEF98633FD76926A" authority="Flowers 1979" authorityName="Flowers" authorityYear="1979" box="[294,647,1365,1387]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF60AC74FEF98633FE02926A" box="[294,499,1365,1387]" italics="true" pageId="1" pageNumber="232">Baetis maculipennis</emphasis>
|
||
<bibRefCitation id="A6264B70FF60AC74FE268630FD76926A" author="Flowers, R. W." box="[505,647,1365,1387]" pageId="1" pageNumber="232" pagination="187 - 191" refId="ref10974" refString="Flowers, R. W. (1979) A new species of Baetis from Panama (Ephemeroptera: Baetidae). Pan-Pacific Entomologist, 55 (3), 187 - 191." type="journal article" year="1979">Flowers 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF60AC74FF4886F2FB2B9019" blockId="1.[151,1436,1431,2033]" pageId="1" pageNumber="232">
|
||
<emphasis id="F0C3EA93FF60AC74FF4886F2FE7492B1" bold="true" box="[151,389,1431,1456]" pageId="1" pageNumber="232">Systematic position.</emphasis>
|
||
In the rank-free classification
|
||
<taxonomicName id="05B74D02FF60AC74FD0586FDFCA492B1" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[730,853,1432,1456]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF60AC74FD0586FDFCA492B1" box="[730,853,1432,1456]" italics="true" pageId="1" pageNumber="232">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
belongs to the following subsequently subordinated taxa: Tetramerotarsata
|
||
<bibRefCitation id="A6264B70FF60AC74FE4886D9FDEB92D5" author="Kluge, N. J." box="[407,538,1468,1493]" pageId="1" pageNumber="232" pagination="527 - 535" refId="ref11223" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. & Sartori, M. (Eds.), Ephemeroptera & Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely & Lacht SA, Fribourg, pp. 527 - 535." type="book chapter" year="1997">Kluge 1997</bibRefCitation>
|
||
> Liberevenata
|
||
<bibRefCitation id="A6264B70FF60AC74FD0E86D9FCA492D5" author="Kluge, N. J." box="[721,853,1468,1493]" pageId="1" pageNumber="232" pagination="527 - 535" refId="ref11223" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. & Sartori, M. (Eds.), Ephemeroptera & Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely & Lacht SA, Fribourg, pp. 527 - 535." type="book chapter" year="1997">Kluge 1997</bibRefCitation>
|
||
(i.e.
|
||
<taxonomicName id="05B74D02FF60AC74FC5186D9FC0192D4" authorityName="Leach" authorityYear="1815" box="[910,1008,1468,1493]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
s.l.)> Turbanoculata
|
||
<bibRefCitation id="A6264B70FF60AC74FB3986D9FA9892D5" author="Kluge, N. J." box="[1254,1385,1468,1493]" pageId="1" pageNumber="232" pagination="527 - 535" refId="ref11223" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. & Sartori, M. (Eds.), Ephemeroptera & Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely & Lacht SA, Fribourg, pp. 527 - 535." type="book chapter" year="1997">Kluge 1997</bibRefCitation>
|
||
(i.e.
|
||
<taxonomicName id="05B74D02FF60AC74FF4886BAFF0892F9" authorityName="Leach" authorityYear="1815" box="[151,249,1503,1528]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
s.str.)> Anteropatellata
|
||
<bibRefCitation id="A6264B70FF60AC74FDC786BAFD6F92F9" author="Kluge, N. J." box="[536,670,1503,1528]" pageId="1" pageNumber="232" pagination="527 - 535" refId="ref11223" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. & Sartori, M. (Eds.), Ephemeroptera & Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely & Lacht SA, Fribourg, pp. 527 - 535." type="book chapter" year="1997">Kluge 1997</bibRefCitation>
|
||
> Baetovectata
|
||
<bibRefCitation id="A6264B70FF60AC74FC8386BAFB8992F9" author="Kluge, N. J. & Novikova, E. A." box="[860,1144,1503,1528]" pageId="1" pageNumber="232" pagination="1 - 56" refId="ref11380" refString="Kluge, N. J. & Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge & Novikova 2011</bibRefCitation>
|
||
> Baetungulata
|
||
<bibRefCitation id="A6264B70FF60AC74FAE786BAFEB9911D" author="Kluge, N. J. & Novikova, E. A." pageId="1" pageNumber="232" pagination="1 - 56" refId="ref11380" refString="Kluge, N. J. & Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge & Novikova 2011</bibRefCitation>
|
||
. In male genitals, gonovectes are bent in the manner characteristic for Baetovectata (
|
||
<figureCitation id="5A8C2A04FF60AC74FADA8561FAAB911D" box="[1285,1370,1540,1565]" captionStart="FIGURES 44–47" captionStartId="12.[151,264,1365,1388]" captionTargetBox="[153,1434,197,1339]" captionTargetId="figure@12.[151,1436,193,1345]" captionTargetPageId="12" captionText="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." httpUri="https://zenodo.org/record/2609849/files/figure.png" pageId="1" pageNumber="232">Fig. 47</figureCitation>
|
||
); fore wings have two marginal intercalaries per space, that is also characteristic for Baetovectata (
|
||
<figureCitation id="5A8C2A04FF60AC74FB1D8542FAE99141" box="[1218,1304,1575,1600]" captionStart="FIGURES 53–55" captionStartId="14.[151,264,1761,1784]" captionTargetBox="[150,1436,193,1740]" captionTargetId="figure@14.[150,1436,193,1740]" captionTargetPageId="14" captionText="FIGURES 53–55. Moribaetis maculipennis. 53–54, female imago; 55, hind wing." httpUri="https://zenodo.org/record/2609853/files/figure.png" pageId="1" pageNumber="232">Fig. 53</figureCitation>
|
||
); in mature larva ready to moult to subimago, subimaginal gonostyli are folded under larval cuticle in such a manner that 2nd segment of gonostylus as a whole is directed caudally-medially (
|
||
<figureCitation id="5A8C2A04FF60AC74FCA1850AFC229189" box="[894,979,1647,1672]" captionStart="FIGURES 44–47" captionStartId="12.[151,264,1365,1388]" captionTargetBox="[153,1434,197,1339]" captionTargetId="figure@12.[151,1436,193,1345]" captionTargetPageId="12" captionText="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." httpUri="https://zenodo.org/record/2609849/files/figure.png" pageId="1" pageNumber="232">Fig. 44</figureCitation>
|
||
), but not laterally, in contrast to baetids other than Baetovectata [see character (17) below]. Claw of mature larva has one asymmetric row of denticles, characteristic for most Baetungulata (
|
||
<figureCitation id="5A8C2A04FF60AC74FDE485D2FD4E91D1" box="[571,703,1719,1744]" captionStart="FIGURES 22–24" captionStartId="9.[151,264,1680,1703]" captionTargetBox="[159,1392,210,1656]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 22–24. Larva of Moribaetis maculipennis. 22, labium (at left ventral view, at right dorsal view); 23, claw (anterior view); 24, its cross section. Abbreviation: d-m, seta of dorso-median row." httpUri="https://zenodo.org/record/2609843/files/figure.png" pageId="1" pageNumber="232">Figs 23–24</figureCitation>
|
||
); as in other Baetungulata, tergalii are unable to make respiratory movements; maxillary palp is two-segmented (
|
||
<bibRefCitation id="A6264B70FF60AC74FD6F85B9FCA091F5" author="Flowers, R. W." box="[688,849,1756,1781]" pageId="1" pageNumber="232" pagination="187 - 191" refId="ref10974" refString="Flowers, R. W. (1979) A new species of Baetis from Panama (Ephemeroptera: Baetidae). Pan-Pacific Entomologist, 55 (3), 187 - 191." type="journal article" year="1979">Flowers 1979</bibRefCitation>
|
||
: fig. 9;
|
||
<bibRefCitation id="A6264B70FF60AC74FC7385B9FB2C91F4" author="Waltz, R. D. & McCafferty, W. P." box="[940,1245,1756,1781]" pageId="1" pageNumber="232" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985</bibRefCitation>
|
||
: figs 28, 37); on fore leg of male subimago tarsomeres 2–4 are covered by blunt microlepides [see character (10)].
|
||
</paragraph>
|
||
<paragraph id="C2083681FF60AC74FF188441FDDD9084" blockId="1.[151,1436,1431,2033]" pageId="1" pageNumber="232">
|
||
All characters testify that
|
||
<taxonomicName id="05B74D02FF60AC74FE338441FD96903D" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[492,615,1828,1852]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF60AC74FE338441FD96903D" box="[492,615,1828,1852]" italics="true" pageId="1" pageNumber="232">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
belongs to Baetovectata-Baetungulata, that means that it is a member of the higher taxon Anteropatellata; therefore, the lack of the character peculiar for Anteropatellata [see character (9)], is here assumed as a secondary loss.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF60AC74FF1884EAFBA090F0" blockId="1.[151,1436,1431,2033]" pageId="1" pageNumber="232">
|
||
Being a member of Baetungulata,
|
||
<taxonomicName id="05B74D02FF60AC74FDBA84F5FD1190A9" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[613,736,1936,1960]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF60AC74FDBA84F5FD1190A9" box="[613,736,1936,1960]" italics="true" pageId="1" pageNumber="232">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
does not belong to a large taxon Baetofemorata
|
||
<bibRefCitation id="A6264B70FF60AC74FAEA84EAFEB890CD" author="Kluge, N. J. & Novikova, E. A." pageId="1" pageNumber="232" pagination="1 - 56" refId="ref11380" refString="Kluge, N. J. & Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge & Novikova 2011</bibRefCitation>
|
||
subordinated to Baetungulata: in contrast to Baetofemorata, larval femur of
|
||
<taxonomicName id="05B74D02FF60AC74FB0484D1FAA790CD" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[1243,1366,1972,1996]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF60AC74FB0484D1FAA790CD" box="[1243,1366,1972,1996]" italics="true" pageId="1" pageNumber="232">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
lacks femoral patch (a compact group of minute setae on inner side of femur near its base).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8AAD650AFF63AC77FF1883F2FB2397F9" pageId="2" pageNumber="233" type="diagnosis">
|
||
<paragraph id="C2083681FF63AC77FF1883F2FB2397F9" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
<emphasis id="F0C3EA93FF63AC77FF1883F2FD8897B1" bold="true" box="[199,633,151,176]" pageId="2" pageNumber="233">
|
||
Annotated diagnosis of
|
||
<taxonomicName id="05B74D02FF63AC77FE2A83F2FD8097B1" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[501,625,151,176]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF63AC77FE2A83F2FD8097B1" bold="true" box="[501,625,151,176]" italics="true" pageId="2" pageNumber="233">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
This diagnosis is based on two species,
|
||
<taxonomicName id="05B74D02FF63AC77FBA883FCFAC897B1" authorityName="Flowers" authorityYear="1979" box="[1143,1337,152,176]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF63AC77FBA883FCFAC897B1" box="[1143,1337,152,176]" italics="true" pageId="2" pageNumber="233">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="05B74D02FF63AC77FAA283FCFEF497D5" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF63AC77FAA283FCFEF497D5" italics="true" pageId="2" pageNumber="233">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF63AC77FED183D8FEBA97D5" box="[270,331,189,212]" pageId="2" pageNumber="233" rank="species">sp. n.</taxonomicNameLabel>
|
||
, which are known as larvae, subimagoes and imagoes; the third species,
|
||
<taxonomicName id="05B74D02FF63AC77FB4D83D8FAF597D5" authorityName="Eaton" authorityYear="1885" box="[1170,1284,188,212]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis id="F0C3EA93FF63AC77FB4D83D8FAF597D5" box="[1170,1284,188,212]" italics="true" pageId="2" pageNumber="233">M. salvini</emphasis>
|
||
</taxonomicName>
|
||
, is known as male imago only (see below), and the fourth species,
|
||
<taxonomicName id="05B74D02FF63AC77FD328384FC4797F9" authorityName="Romero & Esquivel" authorityYear="2018" box="[749,950,224,248]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="brachiostrinus">
|
||
<emphasis id="F0C3EA93FF63AC77FD328384FC4797F9" box="[749,950,224,248]" italics="true" pageId="2" pageNumber="233">M. brachiostrinus</emphasis>
|
||
</taxonomicName>
|
||
, is known as larvae only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8AAD650AFF63AC71FF188261FB6992C1" lastPageId="4" lastPageNumber="235" pageId="2" pageNumber="233" type="description">
|
||
<paragraph id="C2083681FF63AC77FF188261FD209686" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
<emphasis id="F0C3EA93FF63AC77FF188261FF16961C" bold="true" box="[199,231,260,285]" pageId="2" pageNumber="233">(1)</emphasis>
|
||
Labrum is widened in distal part, thickened in proximal part; ventrally with sclerotized lateral margins (
|
||
<figureCitation id="5A8C2A04FF63AC77FAB08261FF469641" captionStart="FIGURES 71–76" captionStartId="19.[151,264,1752,1775]" captionTargetBox="[150,1436,193,1728]" captionTargetId="figure@19.[150,1436,193,1731]" captionTargetPageId="19" captionText="FIGURES 71–76. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 71–72, labrum, focus on dorsal side; 73, focus on anterior margin; 74, focus on ventral side; 75, head with labrum; 76, labium." httpUri="https://zenodo.org/record/2609859/files/figure.png" pageId="2" pageNumber="233">Fig. 74</figureCitation>
|
||
); on its dorsal surface, number of long setae in the pair of antero-lateral rows is increased, these rows nearly meet the submedian pair of long setae, so that all these long setae form nearly integral transverse row (
|
||
<figureCitation id="5A8C2A04FF63AC77FAE58229FA7E9665" box="[1338,1423,332,357]" captionStart="FIGURES 71–76" captionStartId="19.[151,264,1752,1775]" captionTargetBox="[150,1436,193,1728]" captionTargetId="figure@19.[150,1436,193,1731]" captionTargetPageId="19" captionText="FIGURES 71–76. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 71–72, labrum, focus on dorsal side; 73, focus on anterior margin; 74, focus on ventral side; 75, head with labrum; 76, labium." httpUri="https://zenodo.org/record/2609859/files/figure.png" pageId="2" pageNumber="233">Fig. 71</figureCitation>
|
||
). Such a labral structure is found in some other taxa.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF1882F1FE22953D" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
<emphasis id="F0C3EA93FF63AC77FF1882F1FF1996AC" bold="true" box="[199,232,404,429]" pageId="2" pageNumber="233">(2)</emphasis>
|
||
Incisor of each mandible is blade-like, i.e. greatly elongated (several times longer than kinetodontium), straight, with denticles diminished or lost (
|
||
<figureCitation id="5A8C2A04FF63AC77FDA282D2FCF196D1" box="[637,768,439,464]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1850,1873]" captionTargetBox="[151,1436,213,1829]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURES 16–21. Mandibles of Moribaetis maculipennis. 16–17, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 18–21, left and right mandibles of last instar larva just after moult and their fragments enlarged. Abbreviations: kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609841/files/figure.png" pageId="2" pageNumber="233">Figs 18–21</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF63AC77FCD382DDFCA696D1" box="[780,855,440,464]" captionStart="FIGURES 77–84" captionStartId="20.[151,264,1888,1911]" captionTargetBox="[183,1389,193,1851]" captionTargetId="figure@20.[183,1389,193,1851]" captionTargetPageId="20" captionText="FIGURES 77–84. Larvae of Moribaetis latipennis sp. n. 77–78, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 79–82, left and right mandibles of last instar larva just after moult and their fragments enlarged; 83, apex of maxilla; 84, apices of glossae and paraglossae before moult to next (last) instar, with new paraglossae visible inside (arrows show three rows of apical setae on paraglossae). Abbreviations: ds1, ds2, ds3, dentisetae; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609861/files/figure.png" pageId="2" pageNumber="233">79–82</figureCitation>
|
||
). Incisors are often broken (
|
||
<figureCitation id="5A8C2A04FF63AC77FB4982D2FAE996D1" box="[1174,1304,439,464]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1850,1873]" captionTargetBox="[151,1436,213,1829]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURES 16–21. Mandibles of Moribaetis maculipennis. 16–17, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 18–21, left and right mandibles of last instar larva just after moult and their fragments enlarged. Abbreviations: kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609841/files/figure.png" pageId="2" pageNumber="233">Figs 16–17</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF63AC77FAFA82DDFA9E96D1" box="[1317,1391,440,464]" captionStart="FIGURES 77–84" captionStartId="20.[151,264,1888,1911]" captionTargetBox="[183,1389,193,1851]" captionTargetId="figure@20.[183,1389,193,1851]" captionTargetPageId="20" captionText="FIGURES 77–84. Larvae of Moribaetis latipennis sp. n. 77–78, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 79–82, left and right mandibles of last instar larva just after moult and their fragments enlarged; 83, apex of maxilla; 84, apices of glossae and paraglossae before moult to next (last) instar, with new paraglossae visible inside (arrows show three rows of apical setae on paraglossae). Abbreviations: ds1, ds2, ds3, dentisetae; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609861/files/figure.png" pageId="2" pageNumber="233">77–78</figureCitation>
|
||
), so their initial shape should be observed either on freshly-moulted specimens, or inside old mandible just before moulting. Besides
|
||
<taxonomicName id="05B74D02FF63AC77FEA68165FE059519" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[377,500,512,536]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF63AC77FEA68165FE059519" box="[377,500,512,536]" italics="true" pageId="2" pageNumber="233">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
, similar blade-like mandibular incisors independently evolved in some other distantly related baetid taxa.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF188122FD5095CD" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
<emphasis id="F0C3EA93FF63AC77FF188122FF199561" bold="true" box="[199,232,583,608]" pageId="2" pageNumber="233">(3)</emphasis>
|
||
Prostheca of right mandible is slender, straight, directed parallel to incisor, pointed or terminated by few slender processes (
|
||
<figureCitation id="5A8C2A04FF63AC77FEB18109FE399585" box="[366,456,620,645]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1850,1873]" captionTargetBox="[151,1436,213,1829]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURES 16–21. Mandibles of Moribaetis maculipennis. 16–17, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 18–21, left and right mandibles of last instar larva just after moult and their fragments enlarged. Abbreviations: kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609841/files/figure.png" pageId="2" pageNumber="233">Figs 21</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF63AC77FE098109FE079585" box="[470,502,620,644]" captionStart="FIGURES 77–84" captionStartId="20.[151,264,1888,1911]" captionTargetBox="[183,1389,193,1851]" captionTargetId="figure@20.[183,1389,193,1851]" captionTargetPageId="20" captionText="FIGURES 77–84. Larvae of Moribaetis latipennis sp. n. 77–78, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 79–82, left and right mandibles of last instar larva just after moult and their fragments enlarged; 83, apex of maxilla; 84, apices of glossae and paraglossae before moult to next (last) instar, with new paraglossae visible inside (arrows show three rows of apical setae on paraglossae). Abbreviations: ds1, ds2, ds3, dentisetae; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609861/files/figure.png" pageId="2" pageNumber="233">82</figureCitation>
|
||
); this is one of the
|
||
<typeStatus id="1D0C8823FF63AC77FD058108FCE49584" box="[730,789,621,645]" pageId="2" pageNumber="233">types</typeStatus>
|
||
of right prostheca, which are repeated among
|
||
<taxonomicName id="05B74D02FF63AC77FAEC8109FA689584" authorityName="Leach" authorityYear="1815" box="[1331,1433,620,645]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
. Prostheca of left mandible varies from the form usual for
|
||
<taxonomicName id="05B74D02FF63AC77FCF881EAFC7895A9" authorityName="Leach" authorityYear="1815" box="[807,905,655,680]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
(in
|
||
<taxonomicName id="05B74D02FF63AC77FC6781F4FBBC95A9" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[952,1101,656,680]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF63AC77FC6781F4FBBC95A9" box="[952,1101,656,680]" italics="true" pageId="2" pageNumber="233">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF63AC77FB8A81F5FB6395A6" box="[1109,1170,656,679]" pageId="2" pageNumber="233" rank="species">sp. n.</taxonomicNameLabel>
|
||
–
|
||
<figureCitation id="5A8C2A04FF63AC77FB6E81EAFAF395A9" box="[1201,1282,655,680]" captionStart="FIGURES 77–84" captionStartId="20.[151,264,1888,1911]" captionTargetBox="[183,1389,193,1851]" captionTargetId="figure@20.[183,1389,193,1851]" captionTargetPageId="20" captionText="FIGURES 77–84. Larvae of Moribaetis latipennis sp. n. 77–78, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 79–82, left and right mandibles of last instar larva just after moult and their fragments enlarged; 83, apex of maxilla; 84, apices of glossae and paraglossae before moult to next (last) instar, with new paraglossae visible inside (arrows show three rows of apical setae on paraglossae). Abbreviations: ds1, ds2, ds3, dentisetae; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609861/files/figure.png" pageId="2" pageNumber="233">Fig. 80</figureCitation>
|
||
) to similar to right prostheca (in
|
||
<taxonomicName id="05B74D02FF63AC77FEB481D0FDD495CD" authorityName="Flowers" authorityYear="1979" box="[363,549,692,716]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF63AC77FEB481D0FDD495CD" box="[363,549,692,716]" italics="true" pageId="2" pageNumber="233">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<figureCitation id="5A8C2A04FF63AC77FD9E81D1FD6595CD" box="[577,660,692,717]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1850,1873]" captionTargetBox="[151,1436,213,1829]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURES 16–21. Mandibles of Moribaetis maculipennis. 16–17, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 18–21, left and right mandibles of last instar larva just after moult and their fragments enlarged. Abbreviations: kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609841/files/figure.png" pageId="2" pageNumber="233">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF1881B2FF3C9436" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
<emphasis id="F0C3EA93FF63AC77FF1881B2FF1995F1" bold="true" box="[199,232,727,752]" pageId="2" pageNumber="233">(4)</emphasis>
|
||
Maxilla of the «
|
||
<taxonomicName id="05B74D02FF63AC77FE7F81BDFE1795EE" authorityName="Leach" authorityYear="1815" box="[416,486,728,751]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF63AC77FE7F81BDFE1795EE" box="[416,486,728,751]" italics="true" pageId="2" pageNumber="233">Baetis</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="1D0C8823FF63AC77FE3281BDFDD395F1" box="[493,546,728,752]" pageId="2" pageNumber="233">type</typeStatus>
|
||
», i.e. with 1st dentiseta canine-like, heavily sclerotized and bent toward canines (
|
||
<figureCitation id="5A8C2A04FF63AC77FF408199FF029415" box="[159,243,764,789]" captionStart="FIGURES 77–84" captionStartId="20.[151,264,1888,1911]" captionTargetBox="[183,1389,193,1851]" captionTargetId="figure@20.[183,1389,193,1851]" captionTargetPageId="20" captionText="FIGURES 77–84. Larvae of Moribaetis latipennis sp. n. 77–78, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 79–82, left and right mandibles of last instar larva just after moult and their fragments enlarged; 83, apex of maxilla; 84, apices of glossae and paraglossae before moult to next (last) instar, with new paraglossae visible inside (arrows show three rows of apical setae on paraglossae). Abbreviations: ds1, ds2, ds3, dentisetae; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609861/files/figure.png" pageId="2" pageNumber="233">Fig. 83</figureCitation>
|
||
;
|
||
<bibRefCitation id="A6264B70FF63AC77FF208199FE6C9415" author="Flowers, R. W." box="[255,413,764,789]" pageId="2" pageNumber="233" pagination="187 - 191" refId="ref10974" refString="Flowers, R. W. (1979) A new species of Baetis from Panama (Ephemeroptera: Baetidae). Pan-Pacific Entomologist, 55 (3), 187 - 191." type="journal article" year="1979">Flowers 1979</bibRefCitation>
|
||
: fig. 9,
|
||
<bibRefCitation id="A6264B70FF63AC77FE2D8199FCEC9414" author="Waltz, R. D. & McCafferty, W. P." box="[498,797,764,789]" pageId="2" pageNumber="233" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985</bibRefCitation>
|
||
: figs 28, 37). The same in many other non-related baetid taxa.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF188021FBDF94ED" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
<emphasis id="F0C3EA93FF63AC77FF188021FF19945C" bold="true" box="[199,232,836,861]" pageId="2" pageNumber="233">(5)</emphasis>
|
||
Labium has peculiar structure and differs from other taxa by shape of paraglossa and one seta instead of the dorso-median row (
|
||
<figureCitation id="5A8C2A04FF63AC77FEAD8002FE329481" box="[370,451,871,896]" captionStart="FIGURES 22–24" captionStartId="9.[151,264,1680,1703]" captionTargetBox="[159,1392,210,1656]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 22–24. Larva of Moribaetis maculipennis. 22, labium (at left ventral view, at right dorsal view); 23, claw (anterior view); 24, its cross section. Abbreviation: d-m, seta of dorso-median row." httpUri="https://zenodo.org/record/2609843/files/figure.png" pageId="2" pageNumber="233">Fig. 22</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF63AC77FE108002FE1E9481" box="[463,495,871,896]" captionStart="FIGURES 71–76" captionStartId="19.[151,264,1752,1775]" captionTargetBox="[150,1436,193,1728]" captionTargetId="figure@19.[150,1436,193,1731]" captionTargetPageId="19" captionText="FIGURES 71–76. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 71–72, labrum, focus on dorsal side; 73, focus on anterior margin; 74, focus on ventral side; 75, head with labrum; 76, labium." httpUri="https://zenodo.org/record/2609859/files/figure.png" pageId="2" pageNumber="233">76</figureCitation>
|
||
). Paraglossa is sharply narrowing toward apex; its long latero-apical setae are sparse on lateral margin and compactly concentrated on extreme apex of paraglossa, forming here 3 rows; ventro-median setal row is completely absent; dorso-median row is reduced to a single long seta (
|
||
<figureCitation id="5A8C2A04FF63AC77FBF280CAFB7094C9" box="[1069,1153,943,968]" captionStart="FIGURES 22–24" captionStartId="9.[151,264,1680,1703]" captionTargetBox="[159,1392,210,1656]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 22–24. Larva of Moribaetis maculipennis. 22, labium (at left ventral view, at right dorsal view); 23, claw (anterior view); 24, its cross section. Abbreviation: d-m, seta of dorso-median row." httpUri="https://zenodo.org/record/2609843/files/figure.png" pageId="2" pageNumber="233">Fig. 22</figureCitation>
|
||
: d-m). Glossa is long and narrow, its dorso-lateral row is reduced to a few setae on lateral margin near apex.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF188092FE029359" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
Formerly paraglossae of
|
||
<taxonomicName id="05B74D02FF63AC77FE3C809CFD519311" authorityName="Flowers" authorityYear="1979" box="[483,672,1016,1040]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF63AC77FE3C809CFD519311" box="[483,672,1016,1040]" italics="true" pageId="2" pageNumber="233">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
and the larva ascribed to
|
||
<taxonomicName id="05B74D02FF63AC77FC15809CFBCD9311" authorityName="Eaton" authorityYear="1885" box="[970,1084,1016,1040]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis id="F0C3EA93FF63AC77FC15809CFBCD9311" box="[970,1084,1016,1040]" italics="true" pageId="2" pageNumber="233">M. salvini</emphasis>
|
||
</taxonomicName>
|
||
were erroneously described as having only 2 rows of apical setae (
|
||
<bibRefCitation id="A6264B70FF63AC77FDFB8779FD329335" author="Flowers, R. W." box="[548,707,1052,1077]" pageId="2" pageNumber="233" pagination="187 - 191" refId="ref10974" refString="Flowers, R. W. (1979) A new species of Baetis from Panama (Ephemeroptera: Baetidae). Pan-Pacific Entomologist, 55 (3), 187 - 191." type="journal article" year="1979">Flowers 1979</bibRefCitation>
|
||
: 190, fig. 10;
|
||
<bibRefCitation id="A6264B70FF63AC77FC828779FB769334" author="Waltz, R. D. & McCafferty, W. P." box="[861,1159,1052,1077]" pageId="2" pageNumber="233" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985</bibRefCitation>
|
||
: 246, 247, 250); actually they have 3 rows (
|
||
<figureCitation id="5A8C2A04FF63AC77FEBB875AFE4A9359" box="[356,443,1087,1112]" captionStart="FIGURES 22–24" captionStartId="9.[151,264,1680,1703]" captionTargetBox="[159,1392,210,1656]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 22–24. Larva of Moribaetis maculipennis. 22, labium (at left ventral view, at right dorsal view); 23, claw (anterior view); 24, its cross section. Abbreviation: d-m, seta of dorso-median row." httpUri="https://zenodo.org/record/2609843/files/figure.png" pageId="2" pageNumber="233">Figs 22</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF63AC77FE188725FE169359" box="[455,487,1088,1112]" captionStart="FIGURES 77–84" captionStartId="20.[151,264,1888,1911]" captionTargetBox="[183,1389,193,1851]" captionTargetId="figure@20.[183,1389,193,1851]" captionTargetPageId="20" captionText="FIGURES 77–84. Larvae of Moribaetis latipennis sp. n. 77–78, left and right mandibles before moult to next (last) instar, with new mandibles visible inside; 79–82, left and right mandibles of last instar larva just after moult and their fragments enlarged; 83, apex of maxilla; 84, apices of glossae and paraglossae before moult to next (last) instar, with new paraglossae visible inside (arrows show three rows of apical setae on paraglossae). Abbreviations: ds1, ds2, ds3, dentisetae; kd, kinetodontium; prs, prostheca." httpUri="https://zenodo.org/record/2609861/files/figure.png" pageId="2" pageNumber="233">84</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF188700FC8893C5" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
<bibRefCitation id="A6264B70FF63AC77FF188700FDF8937D" author="Romero, D. & Esquivel, C." box="[199,521,1124,1149]" pageId="2" pageNumber="233" pagination="531 - 541" refId="ref11819" refString="Romero, D. & Esquivel, C. (2018) Moribaetis brachiostrinus (Ephemeroptera: Baetidae), a new species of mayfly from Costa Rica. Zootaxa, 4433 (3), 531 - 541. http: // dx. doi. org / 10.11646 / zootaxa. 4433.3.8" type="journal article" year="2018">Romero & Esquivel (2018)</bibRefCitation>
|
||
reported 2 rows for
|
||
<taxonomicName id="05B74D02FF63AC77FCD48700FC29937D" authorityName="Romero & Esquivel" authorityYear="2018" box="[779,984,1124,1148]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="brachiostrinus">
|
||
<emphasis id="F0C3EA93FF63AC77FCD48700FC29937D" box="[779,984,1124,1148]" italics="true" pageId="2" pageNumber="233">M. brachiostrinus</emphasis>
|
||
</taxonomicName>
|
||
. Judging by the shape of paraglossae (
|
||
<bibRefCitation id="A6264B70FF63AC77FF4187EDFDE893A1" author="Romero, D. & Esquivel, C." box="[158,537,1159,1184]" pageId="2" pageNumber="233" pagination="531 - 541" refId="ref11819" refString="Romero, D. & Esquivel, C. (2018) Moribaetis brachiostrinus (Ephemeroptera: Baetidae), a new species of mayfly from Costa Rica. Zootaxa, 4433 (3), 531 - 541. http: // dx. doi. org / 10.11646 / zootaxa. 4433.3.8" type="journal article" year="2018">Romero & Esquivel 2018: fig. 6</bibRefCitation>
|
||
), which are the same as in the two species mentioned above, we assume that paraglossae of
|
||
<taxonomicName id="05B74D02FF63AC77FEE087C8FDF693C5" authorityName="Romero & Esquivel" authorityYear="2018" box="[319,519,1196,1220]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="brachiostrinus">
|
||
<emphasis id="F0C3EA93FF63AC77FEE087C8FDF693C5" box="[319,519,1196,1220]" italics="true" pageId="2" pageNumber="233">M. brachiostrinus</emphasis>
|
||
</taxonomicName>
|
||
also have 3 rows of apical setae.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF1887AAFB0E9109" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
<emphasis id="F0C3EA93FF63AC77FF1887AAFF1993E9" bold="true" box="[199,232,1231,1256]" pageId="2" pageNumber="233">(6)</emphasis>
|
||
Larval prothorax with a pair of tracheal gills, each representing a short and simple (non-branched) outgrowth arising from the membrane connecting prosternum with coxal bases (
|
||
<bibRefCitation id="A6264B70FF63AC77FBEB8791FF479231" author="Waltz, R. D. & McCafferty, W. P." pageId="2" pageNumber="233" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985: fig. 31</bibRefCitation>
|
||
). These outgrows were termed «osmobranchia» (
|
||
<bibRefCitation id="A6264B70FF63AC77FD328672FBD19231" author="Waltz, R. D. & McCafferty, W. P." box="[749,1056,1303,1328]" pageId="2" pageNumber="233" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985</bibRefCitation>
|
||
), but actually they represent true tracheal gills, because they are supplied with special tracheae: just proximad of coxal base, the leg trachea has a thick branch which passes into the gill (
|
||
<figureCitation id="5A8C2A04FF63AC77FD89863AFD2B9279" box="[598,730,1375,1400]" captionStart="FIGURES 85–88" captionStartId="21.[151,264,1737,1760]" captionTargetBox="[152,1435,195,1675]" captionTargetId="figure@21.[152,1435,193,1717]" captionTargetPageId="21" captionText="FIGURES 85–88. Larva of Moribaetis latipennis sp. n. 85, bases of fore legs; 86, the same, with black lines showing colorless tracheal gill and its colorless tracheae, poorly visible on this photo; 87–88, claws of the same two legs." httpUri="https://zenodo.org/record/2609863/files/figure.png" pageId="2" pageNumber="233">Figs 85–86</figureCitation>
|
||
). In contrast to prothorax, mesothorax and metathorax have no gills, and their leg tracheae have no such branches. While the trachea passing into the gill is rather thick, it is unclear if these gills can play any role in respiration because they are very small in relation to the body size and in relation to tergalii, which are large, well-tracheated (
|
||
<figureCitation id="5A8C2A04FF63AC77FCCD86A9FC7892E4" box="[786,905,1484,1509]" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="2" pageNumber="233">Figs 5–11</figureCitation>
|
||
) and also serve as tracheal gills. Larvae of
|
||
<taxonomicName id="05B74D02FF63AC77FF488695FEE39109" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[151,274,1520,1544]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF63AC77FF488695FEE39109" box="[151,274,1520,1544]" italics="true" pageId="2" pageNumber="233">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
inhabit places with the fastest water current, where the role of any gills is likely minimal.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF188571FD699151" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
<emphasis id="F0C3EA93FF63AC77FF188571FF19912C" bold="true" box="[199,232,1556,1581]" pageId="2" pageNumber="233">(7)</emphasis>
|
||
Larval legs are differentiated so that fore, middle and hind legs have the following differences in proportions and setation (
|
||
<figureCitation id="5A8C2A04FF63AC77FE6B8552FDC79151" box="[436,566,1591,1616]" captionStart="FIGURES 25–33" captionStartId="10.[151,264,1857,1880]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@10.[151,1436,193,1836]" captionTargetPageId="10" captionText="FIGURES 25–33. Legs of Moribaetis maculipennis. 25–27, larval fore, middle and hind legs; 28, fragment of fore tibia of female subimago; 29, the same, another specimen; 30, fragment of middle tibia of female subimago; 31–32, fragments of fore and middle leg of female subimago; 33, fragment of femur of larval fore leg (25–27 and 29–32, the same individual). Abbreviation: pt-ti, patella-tibial suture." httpUri="https://zenodo.org/record/2609845/files/figure.png" pageId="2" pageNumber="233">Figs 25–27</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF63AC77FD9E855DFD7D9151" box="[577,652,1592,1616]" captionStart="FIGURES 89–95" captionStartId="22.[151,264,1940,1963]" captionTargetBox="[151,1436,409,1919]" captionTargetId="figure@22.[151,1436,373,1919]" captionTargetPageId="22" captionText="FIGURES 89–95. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 89–91, fore, middle and hind legs; 92–94, apex of femur of fore, middle and hind legs; 95, inner margin of femur of fore leg." httpUri="https://zenodo.org/record/2609865/files/figure.png" pageId="2" pageNumber="233">89–91</figureCitation>
|
||
):
|
||
</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF188538FAB69029" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">Fore, middle and hind femora slightly differ in length, with fore femur being the shortest and hind femur being the longest; fore and middle tibia are of equal length, while hind tibia is 1.2 times shorter; fore, middle and hind tarsi slightly differ in length, with fore tarsus being the longest and hind tarsus being the shortest. Thus, on fore leg tibia and tarsus combined are 1.3 times longer than femur, while on hind leg tibia and tarsus combined are subequal to femur or only 1.1 times longer. In course of development to subimago and to imago, legs become significantly longer, and their proportions are changed, but hind tibia and tarsus continue to be shorter than middle ones.</paragraph>
|
||
<paragraph id="C2083681FF63AC77FF188450FEBB90DC" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="233">
|
||
Fore femur is the thickest; short spine-like setae on inner side of femur (
|
||
<figureCitation id="5A8C2A04FF63AC77FBDB8451FB93904D" box="[1028,1122,1844,1869]" captionStart="FIGURES 25–33" captionStartId="10.[151,264,1857,1880]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@10.[151,1436,193,1836]" captionTargetPageId="10" captionText="FIGURES 25–33. Legs of Moribaetis maculipennis. 25–27, larval fore, middle and hind legs; 28, fragment of fore tibia of female subimago; 29, the same, another specimen; 30, fragment of middle tibia of female subimago; 31–32, fragments of fore and middle leg of female subimago; 33, fragment of femur of larval fore leg (25–27 and 29–32, the same individual). Abbreviation: pt-ti, patella-tibial suture." httpUri="https://zenodo.org/record/2609845/files/figure.png" pageId="2" pageNumber="233">Figs. 33</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF63AC77FBAF8451FB7C904C" box="[1136,1165,1844,1869]" captionStart="FIGURES 89–95" captionStartId="22.[151,264,1940,1963]" captionTargetBox="[151,1436,409,1919]" captionTargetId="figure@22.[151,1436,373,1919]" captionTargetPageId="22" captionText="FIGURES 89–95. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 89–91, fore, middle and hind legs; 92–94, apex of femur of fore, middle and hind legs; 95, inner margin of femur of fore leg." httpUri="https://zenodo.org/record/2609865/files/figure.png" pageId="2" pageNumber="233">95</figureCitation>
|
||
) occupy widest area on fore femur, narrower area on middle femur and narrowest area on hind femur. Apices of fore and middle femora with dense bunch of 4–9 spine-like setae of the same structure as spine-like setae of the longitudinal row on outer side [see (8)] (
|
||
<figureCitation id="5A8C2A04FF63AC77FEE184C5FE3590B9" box="[318,452,1952,1977]" captionStart="FIGURES 89–95" captionStartId="22.[151,264,1940,1963]" captionTargetBox="[151,1436,409,1919]" captionTargetId="figure@22.[151,1436,373,1919]" captionTargetPageId="22" captionText="FIGURES 89–95. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 89–91, fore, middle and hind legs; 92–94, apex of femur of fore, middle and hind legs; 95, inner margin of femur of fore leg." httpUri="https://zenodo.org/record/2609865/files/figure.png" pageId="2" pageNumber="233">Figs 92–93</figureCitation>
|
||
); apex of hind femur usually with two such setae (
|
||
<figureCitation id="5A8C2A04FF63AC77FBD384C5FB9190B9" box="[1036,1120,1952,1977]" captionStart="FIGURES 89–95" captionStartId="22.[151,264,1940,1963]" captionTargetBox="[151,1436,409,1919]" captionTargetId="figure@22.[151,1436,373,1919]" captionTargetPageId="22" captionText="FIGURES 89–95. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 89–91, fore, middle and hind legs; 92–94, apex of femur of fore, middle and hind legs; 95, inner margin of femur of fore leg." httpUri="https://zenodo.org/record/2609865/files/figure.png" pageId="2" pageNumber="233">Fig. 94</figureCitation>
|
||
) (as on all femora of many other
|
||
<taxonomicName id="05B74D02FF63AC77FF0884A1FECF90DC" authorityName="Leach" authorityYear="1815" box="[215,318,1988,2013]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF1883F2FC609641" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF1883F2FF1997B1" bold="true" box="[199,232,151,176]" pageId="3" pageNumber="234">(8)</emphasis>
|
||
Larval legs with stout pointed spine-like setae, fine setae and scales in angulated bases [similar to scales of abdomen—see (13)]. Each femur [different on different leg pairs – see (7)] on outer side bears a regular sparse row of stout pointed spine-like setae; between them stretches an interrupted irregular row of fine setae (
|
||
<figureCitation id="5A8C2A04FF62AC76FB2983BAFABA97F9" box="[1270,1355,223,248]" captionStart="FIGURES 89–95" captionStartId="22.[151,264,1940,1963]" captionTargetBox="[151,1436,409,1919]" captionTargetId="figure@22.[151,1436,373,1919]" captionTargetPageId="22" captionText="FIGURES 89–95. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 89–91, fore, middle and hind legs; 92–94, apex of femur of fore, middle and hind legs; 95, inner margin of femur of fore leg." httpUri="https://zenodo.org/record/2609865/files/figure.png" pageId="3" pageNumber="234">Fig. 92</figureCitation>
|
||
). Apex of each femur with smaller spine-like setae (
|
||
<figureCitation id="5A8C2A04FF62AC76FD528261FCE3961D" box="[653,786,260,285]" captionStart="FIGURES 89–95" captionStartId="22.[151,264,1940,1963]" captionTargetBox="[151,1436,409,1919]" captionTargetId="figure@22.[151,1436,373,1919]" captionTargetPageId="22" captionText="FIGURES 89–95. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 89–91, fore, middle and hind legs; 92–94, apex of femur of fore, middle and hind legs; 95, inner margin of femur of fore leg." httpUri="https://zenodo.org/record/2609865/files/figure.png" pageId="3" pageNumber="234">Figs 92–94</figureCitation>
|
||
). Each tibia with row of small pointed spine-like setae on outer side (
|
||
<figureCitation id="5A8C2A04FF62AC76FECB8242FE949641" box="[276,357,295,320]" captionStart="FIGURES 89–95" captionStartId="22.[151,264,1940,1963]" captionTargetBox="[151,1436,409,1919]" captionTargetId="figure@22.[151,1436,373,1919]" captionTargetPageId="22" captionText="FIGURES 89–95. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 89–91, fore, middle and hind legs; 92–94, apex of femur of fore, middle and hind legs; 95, inner margin of femur of fore leg." httpUri="https://zenodo.org/record/2609865/files/figure.png" pageId="3" pageNumber="234">Fig. 92</figureCitation>
|
||
) and with irregular spine-like setae on inner side.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF188229FE429561" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF188229FF199664" bold="true" box="[199,232,332,357]" pageId="3" pageNumber="234">(9)</emphasis>
|
||
Patella-tibial suture is modified as the following: on each larval leg it stretches longitudinally and gradually disappears not reaching inner (ventral) side of tibia (
|
||
<figureCitation id="5A8C2A04FF62AC76FD00820AFC909689" box="[735,865,367,392]" captionStart="FIGURES 25–33" captionStartId="10.[151,264,1857,1880]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@10.[151,1436,193,1836]" captionTargetPageId="10" captionText="FIGURES 25–33. Legs of Moribaetis maculipennis. 25–27, larval fore, middle and hind legs; 28, fragment of fore tibia of female subimago; 29, the same, another specimen; 30, fragment of middle tibia of female subimago; 31–32, fragments of fore and middle leg of female subimago; 33, fragment of femur of larval fore leg (25–27 and 29–32, the same individual). Abbreviation: pt-ti, patella-tibial suture." httpUri="https://zenodo.org/record/2609845/files/figure.png" pageId="3" pageNumber="234">Figs 25–27</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF62AC76FCB38215FC479689" box="[876,950,368,392]" captionStart="FIGURES 89–95" captionStartId="22.[151,264,1940,1963]" captionTargetBox="[151,1436,409,1919]" captionTargetId="figure@22.[151,1436,373,1919]" captionTargetPageId="22" captionText="FIGURES 89–95. Larval exuviae of Moribaetis latipennis sp. n. (holotype). 89–91, fore, middle and hind legs; 92–94, apex of femur of fore, middle and hind legs; 95, inner margin of femur of fore leg." httpUri="https://zenodo.org/record/2609865/files/figure.png" pageId="3" pageNumber="234">89–91</figureCitation>
|
||
). On middle and hind legs of subimago and imago patella-tibial suture reaches inner side of tibia (
|
||
<figureCitation id="5A8C2A04FF62AC76FD2582F1FC8896AD" box="[762,889,404,429]" captionStart="FIGURES 25–33" captionStartId="10.[151,264,1857,1880]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@10.[151,1436,193,1836]" captionTargetPageId="10" captionText="FIGURES 25–33. Legs of Moribaetis maculipennis. 25–27, larval fore, middle and hind legs; 28, fragment of fore tibia of female subimago; 29, the same, another specimen; 30, fragment of middle tibia of female subimago; 31–32, fragments of fore and middle leg of female subimago; 33, fragment of femur of larval fore leg (25–27 and 29–32, the same individual). Abbreviation: pt-ti, patella-tibial suture." httpUri="https://zenodo.org/record/2609845/files/figure.png" pageId="3" pageNumber="234">Fig. 30, 32</figureCitation>
|
||
), but on fore leg of female subimago and imago (known only for
|
||
<taxonomicName id="05B74D02FF62AC76FE8382DCFDEB96D1" authorityName="Flowers" authorityYear="1979" box="[348,538,440,464]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF62AC76FE8382DCFDEB96D1" box="[348,538,440,464]" italics="true" pageId="3" pageNumber="234">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
) patella-tibial suture stretches longitudinally and does not reach inner side of tibia (
|
||
<figureCitation id="5A8C2A04FF62AC76FF0682B9FEDC96F5" box="[217,301,476,501]" captionStart="FIGURES 25–33" captionStartId="10.[151,264,1857,1880]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@10.[151,1436,193,1836]" captionTargetPageId="10" captionText="FIGURES 25–33. Legs of Moribaetis maculipennis. 25–27, larval fore, middle and hind legs; 28, fragment of fore tibia of female subimago; 29, the same, another specimen; 30, fragment of middle tibia of female subimago; 31–32, fragments of fore and middle leg of female subimago; 33, fragment of femur of larval fore leg (25–27 and 29–32, the same individual). Abbreviation: pt-ti, patella-tibial suture." httpUri="https://zenodo.org/record/2609845/files/figure.png" pageId="3" pageNumber="234">Fig. 28</figureCitation>
|
||
), sometimes being shortened and vestigial (
|
||
<figureCitation id="5A8C2A04FF62AC76FCC382B9FC6C96F5" box="[796,925,476,501]" captionStart="FIGURES 25–33" captionStartId="10.[151,264,1857,1880]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@10.[151,1436,193,1836]" captionTargetPageId="10" captionText="FIGURES 25–33. Legs of Moribaetis maculipennis. 25–27, larval fore, middle and hind legs; 28, fragment of fore tibia of female subimago; 29, the same, another specimen; 30, fragment of middle tibia of female subimago; 31–32, fragments of fore and middle leg of female subimago; 33, fragment of femur of larval fore leg (25–27 and 29–32, the same individual). Abbreviation: pt-ti, patella-tibial suture." httpUri="https://zenodo.org/record/2609845/files/figure.png" pageId="3" pageNumber="234">Figs 29, 31</figureCitation>
|
||
) (male subimago and imago have no patellatibial suture on fore legs, as in all other
|
||
<taxonomicName id="05B74D02FF62AC76FD5D829AFD189519" authorityName="Leach" authorityYear="1815" box="[642,745,511,536]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
). In this respect
|
||
<taxonomicName id="05B74D02FF62AC76FC6C8165FBDF9519" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[947,1070,512,536]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF62AC76FC6C8165FBDF9519" box="[947,1070,512,536]" italics="true" pageId="3" pageNumber="234">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
somewhat resembles the basal plesiomorphon Protopatellata, which in all stages of both sexes have patella-tibial suture only on middle and hind legs, but not on fore legs.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF188109FB8E95A9" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF188109FF079584" bold="true" box="[199,246,620,645]" pageId="3" pageNumber="234">(10)</emphasis>
|
||
In subimagoes of both sexes, all tarsal segments on all legs are entirely covered with blunt microlepides only. The same in some other Baetungulata-non-Baetofemorata and in all Baetofemorata.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF1881D1FB4E9414" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF1881D1FF0295CC" bold="true" box="[199,243,692,717]" pageId="3" pageNumber="234">(11)</emphasis>
|
||
Larval claw bears a large seta on posterior side; this seta located equidistantly with the most distal denticle of the claw; base of this seta is inserted into a cylindrical canal (
|
||
<figureCitation id="5A8C2A04FF62AC76FCB881B2FC1895F1" box="[871,1001,727,752]" captionStart="FIGURES 22–24" captionStartId="9.[151,264,1680,1703]" captionTargetBox="[159,1392,210,1656]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 22–24. Larva of Moribaetis maculipennis. 22, labium (at left ventral view, at right dorsal view); 23, claw (anterior view); 24, its cross section. Abbreviation: d-m, seta of dorso-median row." httpUri="https://zenodo.org/record/2609843/files/figure.png" pageId="3" pageNumber="234">Figs 23–24</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF62AC76FC2B81BDFBE295F1" box="[1012,1043,728,752]" captionStart="FIGURES 85–88" captionStartId="21.[151,264,1737,1760]" captionTargetBox="[152,1435,195,1675]" captionTargetId="figure@21.[152,1435,193,1717]" captionTargetPageId="21" captionText="FIGURES 85–88. Larva of Moribaetis latipennis sp. n. 85, bases of fore legs; 86, the same, with black lines showing colorless tracheal gill and its colorless tracheae, poorly visible on this photo; 87–88, claws of the same two legs." httpUri="https://zenodo.org/record/2609863/files/figure.png" pageId="3" pageNumber="234">87</figureCitation>
|
||
). Individual claws of
|
||
<taxonomicName id="05B74D02FF62AC76FAD781BCFA6A95F1" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[1288,1435,728,752]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF62AC76FAD781BCFA6A95F1" box="[1288,1435,728,752]" italics="true" pageId="3" pageNumber="234">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF62AC76FF488198FF229415" box="[151,211,765,788]" pageId="3" pageNumber="234" rank="species">sp. n.</taxonomicNameLabel>
|
||
have two such setae (
|
||
<figureCitation id="5A8C2A04FF62AC76FE178199FDEB9415" box="[456,538,764,789]" captionStart="FIGURES 85–88" captionStartId="21.[151,264,1737,1760]" captionTargetBox="[152,1435,195,1675]" captionTargetId="figure@21.[152,1435,193,1717]" captionTargetPageId="21" captionText="FIGURES 85–88. Larva of Moribaetis latipennis sp. n. 85, bases of fore legs; 86, the same, with black lines showing colorless tracheal gill and its colorless tracheae, poorly visible on this photo; 87–88, claws of the same two legs." httpUri="https://zenodo.org/record/2609863/files/figure.png" pageId="3" pageNumber="234">Fig. 88</figureCitation>
|
||
) (sometimes three, according to
|
||
<bibRefCitation id="A6264B70FF62AC76FC568199FB429414" author="Waltz, R. D. & McCafferty, W. P." box="[905,1203,764,789]" pageId="3" pageNumber="234" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF18807AFEF194A4" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF18807AFF049439" bold="true" box="[199,245,799,824]" pageId="3" pageNumber="234">(12)</emphasis>
|
||
Hind wing is relatively wide, with short, slightly hooked costal projection and 3 longitudinal veins, among which 2nd vein is symmetrically bifurcated, with intercalary between its branches; sometimes additional intercalary(es) and cross vein(s) are present (
|
||
<figureCitation id="5A8C2A04FF62AC76FD7E8002FD069481" box="[673,759,871,896]" captionStart="FIGURES 53–55" captionStartId="14.[151,264,1761,1784]" captionTargetBox="[150,1436,193,1740]" captionTargetId="figure@14.[150,1436,193,1740]" captionTargetPageId="14" captionText="FIGURES 53–55. Moribaetis maculipennis. 53–54, female imago; 55, hind wing." httpUri="https://zenodo.org/record/2609853/files/figure.png" pageId="3" pageNumber="234">Fig. 55</figureCitation>
|
||
). Larval protopteron has corresponding form and venation (
|
||
<figureCitation id="5A8C2A04FF62AC76FF4080E9FF0294A4" box="[159,243,908,933]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="3" pageNumber="234">Fig. 67</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF1880CAFBDF9311" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF1880CAFF0794C9" bold="true" box="[199,246,943,968]" pageId="3" pageNumber="234">(13)</emphasis>
|
||
Larval abdominal terga and sterna with numerous short scales in operculate sockets (
|
||
<figureCitation id="5A8C2A04FF62AC76FB0380CAFAC294C9" box="[1244,1331,943,968]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="3" pageNumber="234">Fig. 70</figureCitation>
|
||
). Similar scales are present on thoracic pleura and on legs. Such kind of scale sockets with two operculae is found in many baetid taxa, and probably is initial for
|
||
<taxonomicName id="05B74D02FF62AC76FD9C8092FD549311" authorityName="Leach" authorityYear="1815" box="[579,677,1015,1040]" class="Insecta" family="Baetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
[
|
||
<bibRefCitation id="A6264B70FF62AC76FD6D8092FBD39311" author="Kluge, N. J." box="[690,1058,1015,1040]" pageId="3" pageNumber="234" refId="ref11293" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Springer, Dordrecht, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004: Turbanoculata (12)</bibRefCitation>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF188779FBF8939E" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF188779FF079334" bold="true" box="[199,246,1052,1077]" pageId="3" pageNumber="234">(14)</emphasis>
|
||
Larval abdominal terga with long and pointed denticles on posterior margins; in both examined species denticles are absent on tergum I and present on terga II–X; on tergum IX row of denticles is interrupted behind pair of submedian setae (
|
||
<figureCitation id="5A8C2A04FF62AC76FE5F8706FE24937A" box="[384,469,1123,1148]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="3" pageNumber="234">Fig. 70</figureCitation>
|
||
). Abdominal sterna with smaller pointed denticles, which are present either on sterna V–IX (in
|
||
<taxonomicName id="05B74D02FF62AC76FEDB87E2FE4F939F" authorityName="Flowers" authorityYear="1979" box="[260,446,1158,1182]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF62AC76FEDB87E2FE4F939F" box="[260,446,1158,1182]" italics="true" pageId="3" pageNumber="234">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
), or only on sterna VIII–IX (in
|
||
<taxonomicName id="05B74D02FF62AC76FCFD87E2FC47939F" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[802,950,1158,1182]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF62AC76FCFD87E2FC47939F" box="[802,950,1158,1182]" italics="true" pageId="3" pageNumber="234">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF62AC76FC6287E2FC0B939E" box="[957,1018,1159,1183]" pageId="3" pageNumber="234" rank="species">sp. n.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF1887CCFF30922C" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF1887CCFF0793C3" bold="true" box="[199,246,1193,1218]" pageId="3" pageNumber="234">(15)</emphasis>
|
||
Tergalii (unable to make respiratory movements, as in other Baetungulata) have the following structure. Tergalii I more or less diminished, with costal and anal ribs very short. Tergalii II–VII with costal rib welldeveloped and anal rib rather short, so that a long part of hind margin is free of rib; costal rib with serration (
|
||
<figureCitation id="5A8C2A04FF62AC76FA978795FF45922C" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="3" pageNumber="234">Figs 5– 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF188652FBC592DF" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF188652FF079251" bold="true" box="[199,246,1335,1360]" pageId="3" pageNumber="234">(16)</emphasis>
|
||
Larval paracercus is well-developed, cerci greatly elongated, longer than body (
|
||
<figureCitation id="5A8C2A04FF62AC76FB5B8652FB1B9251" box="[1156,1258,1335,1360]" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="3" pageNumber="234">Figs 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF62AC76FB288652FAE69251" box="[1271,1303,1335,1360]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="3" pageNumber="234">62</figureCitation>
|
||
); both cerci and paracercus bear very dense primary swimming setae, which are present on most their segments; bases of these setae are pressed together and regularly alternate as shifted dorsally and ventrally from the common line (
|
||
<figureCitation id="5A8C2A04FF62AC76FAE4861BFA7E9296" box="[1339,1423,1406,1431]" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="3" pageNumber="234">Fig. 14</figureCitation>
|
||
). Segments of cerci are cylindrical, with boundaries not inclined and with denticles on posterior margin not enlarged on any side of cerci or paracercus; secondary swimming setae are absent (
|
||
<figureCitation id="5A8C2A04FF62AC76FC0C86A0FBD692DF" box="[979,1063,1477,1502]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="3" pageNumber="234">Fig. 63</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF18868CFC419040" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
<emphasis id="F0C3EA93FF62AC76FF18868CFF079103" bold="true" box="[199,246,1513,1538]" pageId="3" pageNumber="234">(17)</emphasis>
|
||
Gonostyli have unusual shape and paradoxical development: in definitive condition 1st segment is longer than 2nd segment (
|
||
<figureCitation id="5A8C2A04FF62AC76FEB38569FE329124" box="[364,451,1548,1573]" captionStart="FIGURES 34–43" captionStartId="11.[151,264,1837,1860]" captionTargetBox="[150,1436,193,1817]" captionTargetId="figure@11.[150,1436,193,1817]" captionTargetPageId="11" captionText="FIGURES 34–43. Moribaetis maculipennis. 34–35, male subimago just after moult; 36, abdominal sternum IX of last instar male larva with gonostyli visible inside; 37, gonostyli of male subimago; 38, exuviae of subimaginal gonostyli; 39, genitals of male imago; 40, caudalii of last instar larva with crumpled sibimaginal cerci visible inside; 41–42, proximal part of imaginal cercus and its subimaginal exuviae (arrows show correspondence of their joints); 43, proximal part of imaginal cercus of another specimen." httpUri="https://zenodo.org/record/2609847/files/figure.png" pageId="3" pageNumber="234">Figs 39</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF62AC76FE108568FE1E9124" box="[463,495,1549,1573]" captionStart="FIGURES 44–47" captionStartId="12.[151,264,1365,1388]" captionTargetBox="[153,1434,197,1339]" captionTargetId="figure@12.[151,1436,193,1345]" captionTargetPageId="12" captionText="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." httpUri="https://zenodo.org/record/2609849/files/figure.png" pageId="3" pageNumber="234">47</figureCitation>
|
||
); in course of development, 2nd segment grows earlier, so in subimago 1st segment is much shorter than 2nd (
|
||
<figureCitation id="5A8C2A04FF62AC76FE1D854AFDEB9149" box="[450,538,1583,1608]" captionStart="FIGURES 34–43" captionStartId="11.[151,264,1837,1860]" captionTargetBox="[150,1436,193,1817]" captionTargetId="figure@11.[150,1436,193,1817]" captionTargetPageId="11" captionText="FIGURES 34–43. Moribaetis maculipennis. 34–35, male subimago just after moult; 36, abdominal sternum IX of last instar male larva with gonostyli visible inside; 37, gonostyli of male subimago; 38, exuviae of subimaginal gonostyli; 39, genitals of male imago; 40, caudalii of last instar larva with crumpled sibimaginal cerci visible inside; 41–42, proximal part of imaginal cercus and its subimaginal exuviae (arrows show correspondence of their joints); 43, proximal part of imaginal cercus of another specimen." httpUri="https://zenodo.org/record/2609847/files/figure.png" pageId="3" pageNumber="234">Figs 37</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF62AC76FDF7854AFDB99149" box="[552,584,1583,1608]" captionStart="FIGURES 44–47" captionStartId="12.[151,264,1365,1388]" captionTargetBox="[153,1434,197,1339]" captionTargetId="figure@12.[151,1436,193,1345]" captionTargetPageId="12" captionText="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." httpUri="https://zenodo.org/record/2609849/files/figure.png" pageId="3" pageNumber="234">46</figureCitation>
|
||
); in mature larva developing subimaginal gonostyli are folded under larval cuticle in a peculiar manner, which can be termed «
|
||
<taxonomicName id="05B74D02FF62AC76FD388536FC93916A" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[743,866,1619,1643]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF62AC76FD388536FC93916A" box="[743,866,1619,1643]" italics="true" pageId="3" pageNumber="234">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="1D0C8823FF62AC76FCB68531FC6F916D" box="[873,926,1620,1644]" pageId="3" pageNumber="234">type</typeStatus>
|
||
». When subimaginal wings are already fully developed and crumpled under cuticle of protopteron, developing subimaginal gonostylus has 1st segment extremely short, and 2nd segment forming a loop (3rd segment indistinguishable at this stage); apex of gonostylus locates inside apex of larval protogonostylus (
|
||
<figureCitation id="5A8C2A04FF62AC76FD7C85D8FD0D91D7" box="[675,764,1725,1750]" captionStart="FIGURES 34–43" captionStartId="11.[151,264,1837,1860]" captionTargetBox="[150,1436,193,1817]" captionTargetId="figure@11.[150,1436,193,1817]" captionTargetPageId="11" captionText="FIGURES 34–43. Moribaetis maculipennis. 34–35, male subimago just after moult; 36, abdominal sternum IX of last instar male larva with gonostyli visible inside; 37, gonostyli of male subimago; 38, exuviae of subimaginal gonostyli; 39, genitals of male imago; 40, caudalii of last instar larva with crumpled sibimaginal cerci visible inside; 41–42, proximal part of imaginal cercus and its subimaginal exuviae (arrows show correspondence of their joints); 43, proximal part of imaginal cercus of another specimen." httpUri="https://zenodo.org/record/2609847/files/figure.png" pageId="3" pageNumber="234">Figs 36</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF62AC76FCD685DBFCD891D7" box="[777,809,1726,1750]" captionStart="FIGURES 44–47" captionStartId="12.[151,264,1365,1388]" captionTargetBox="[153,1434,197,1339]" captionTargetId="figure@12.[151,1436,193,1345]" captionTargetPageId="12" captionText="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." httpUri="https://zenodo.org/record/2609849/files/figure.png" pageId="3" pageNumber="234">44</figureCitation>
|
||
). Just before moult, gonostyli become longer and meet medially by their loops of 2nd segments (
|
||
<figureCitation id="5A8C2A04FF62AC76FDAD8584FD3991FB" box="[626,712,1761,1786]" captionStart="FIGURES 44–47" captionStartId="12.[151,264,1365,1388]" captionTargetBox="[153,1434,197,1339]" captionTargetId="figure@12.[151,1436,193,1345]" captionTargetPageId="12" captionText="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." httpUri="https://zenodo.org/record/2609849/files/figure.png" pageId="3" pageNumber="234">Fig. 45</figureCitation>
|
||
). This «
|
||
<taxonomicName id="05B74D02FF62AC76FCFA8584FC5191F8" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[805,928,1761,1785]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF62AC76FCFA8584FC5191F8" box="[805,928,1761,1785]" italics="true" pageId="3" pageNumber="234">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="1D0C8823FF62AC76FC788587FC2B91FB" box="[935,986,1762,1786]" pageId="3" pageNumber="234">type</typeStatus>
|
||
» of gonostyli folding proceed from the «
|
||
<taxonomicName id="05B74D02FF62AC76FF7A8460FED9901C" authorityName="Novikova & Kluge" authorityYear="1987" box="[165,296,1797,1821]" class="Insecta" family="Baetidae" genus="Nigrobaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF62AC76FF7A8460FED9901C" box="[165,296,1797,1821]" italics="true" pageId="3" pageNumber="234">Nigrobaetis</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="1D0C8823FF62AC76FEEF8460FE92901C" box="[304,355,1797,1821]" pageId="3" pageNumber="234">type</typeStatus>
|
||
» (at which 2nd segment is directed caudally and bent with convexity directed medially): in
|
||
<taxonomicName id="05B74D02FF62AC76FF48844DFEE39041" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[151,274,1832,1856]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF62AC76FF48844DFEE39041" box="[151,274,1832,1856]" italics="true" pageId="3" pageNumber="234">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
2nd segment is longer and its bent is transformed to a loop.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC76FF18842EFD3090AA" blockId="3.[151,1437,151,2035]" pageId="3" pageNumber="234">
|
||
Other peculiar features of genitals are the following: sterno-styligeral muscle is unusually short, so that originates far from fore margin of abdominal sternum IX; a pair of protuberances is present between unistyligers; 3rd segment of gonostylus is very short (
|
||
<figureCitation id="5A8C2A04FF62AC76FDBF84F7FD4590AA" box="[608,692,1938,1963]" captionStart="FIGURES 44–47" captionStartId="12.[151,264,1365,1388]" captionTargetBox="[153,1434,197,1339]" captionTargetId="figure@12.[151,1436,193,1345]" captionTargetPageId="12" captionText="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." httpUri="https://zenodo.org/record/2609849/files/figure.png" pageId="3" pageNumber="234">Fig. 47</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF62AC71FF1884D3FC8497D5" blockId="3.[151,1437,151,2035]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="235" pageId="3" pageNumber="234">
|
||
Gonostyli have colorless cuticle and brown hypodermal coloration; their hypodermal coloration varies individually, being developed first of all in distal part of 1st segment, in distal part of 2nd segment and in small 3rd segment. This hypodermal coloration appears in last larval instar (
|
||
<figureCitation id="5A8C2A04FF65AC71FC5383F2FC1497B1" box="[908,997,151,176]" captionStart="FIGURES 34–43" captionStartId="11.[151,264,1837,1860]" captionTargetBox="[150,1436,193,1817]" captionTargetId="figure@11.[150,1436,193,1817]" captionTargetPageId="11" captionText="FIGURES 34–43. Moribaetis maculipennis. 34–35, male subimago just after moult; 36, abdominal sternum IX of last instar male larva with gonostyli visible inside; 37, gonostyli of male subimago; 38, exuviae of subimaginal gonostyli; 39, genitals of male imago; 40, caudalii of last instar larva with crumpled sibimaginal cerci visible inside; 41–42, proximal part of imaginal cercus and its subimaginal exuviae (arrows show correspondence of their joints); 43, proximal part of imaginal cercus of another specimen." httpUri="https://zenodo.org/record/2609847/files/figure.png" pageId="4" pageNumber="235">Figs 36</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF65AC71FC2D83FDFBE097B1" box="[1010,1041,152,176]" captionStart="FIGURES 44–47" captionStartId="12.[151,264,1365,1388]" captionTargetBox="[153,1434,197,1339]" captionTargetId="figure@12.[151,1436,193,1345]" captionTargetPageId="12" captionText="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." httpUri="https://zenodo.org/record/2609849/files/figure.png" pageId="4" pageNumber="235">44</figureCitation>
|
||
), that helps to reveal homology of segments at different stages of development (
|
||
<figureCitation id="5A8C2A04FF65AC71FD4D83D9FCE397D4" box="[658,786,188,213]" captionStart="FIGURES 34–43" captionStartId="11.[151,264,1837,1860]" captionTargetBox="[150,1436,193,1817]" captionTargetId="figure@11.[150,1436,193,1817]" captionTargetPageId="11" captionText="FIGURES 34–43. Moribaetis maculipennis. 34–35, male subimago just after moult; 36, abdominal sternum IX of last instar male larva with gonostyli visible inside; 37, gonostyli of male subimago; 38, exuviae of subimaginal gonostyli; 39, genitals of male imago; 40, caudalii of last instar larva with crumpled sibimaginal cerci visible inside; 41–42, proximal part of imaginal cercus and its subimaginal exuviae (arrows show correspondence of their joints); 43, proximal part of imaginal cercus of another specimen." httpUri="https://zenodo.org/record/2609847/files/figure.png" pageId="4" pageNumber="235">Figs 36–39</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF65AC71FCC183D9FC9897D5" box="[798,873,188,212]" captionStart="FIGURES 44–47" captionStartId="12.[151,264,1365,1388]" captionTargetBox="[153,1434,197,1339]" captionTargetId="figure@12.[151,1436,193,1345]" captionTargetPageId="12" captionText="FIGURES 44–47. Genital of Moribaetis maculipennis. 44, hind margin of IX sternum of mature larva and subimaginal gonostyli crumpled under larval cuticle (specimen with fully developed subimaginal wings); 45, gonostyli crumpled under larval cuticle just before moult to subimago; 46, exuviae of subimaginal gonostyli; 47, genitals of male imago (ventral view; muscles and internal sclerites shown by interrupted lines). Abbreviations: gs1, gs2, gs3, segments of gonostylus; gv, gonovectis; m.IX-X, intersegmental muscle going from base of IX sternum to base of paraproct; m.sg, sterno-styligeral muscle; pgs, larval protogonostylus; us, unistyliger." httpUri="https://zenodo.org/record/2609849/files/figure.png" pageId="4" pageNumber="235">44–47</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF65AC71FF1883BAFCA496AC" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="235">
|
||
All these features of genital structure in larval, subimaginal and imaginal stages are examined for
|
||
<taxonomicName id="05B74D02FF65AC71FAA28384FEDD961D" authorityName="Flowers" authorityYear="1979" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF65AC71FAA28384FEDD961D" italics="true" pageId="4" pageNumber="235">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
, while genitals of
|
||
<taxonomicName id="05B74D02FF65AC71FE228260FD63961D" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[509,658,260,284]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="latipennis">
|
||
<emphasis id="F0C3EA93FF65AC71FE228260FD63961D" box="[509,658,260,284]" italics="true" pageId="4" pageNumber="235">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
were examined for a single specimen of imago, which have not shed subimaginal cuticle. Genitals of this specimen (
|
||
<figureCitation id="5A8C2A04FF65AC71FD708242FCFE9641" box="[687,783,295,320]" captionStart="FIGURES 96–101" captionStartId="23.[151,264,1536,1559]" captionTargetBox="[151,1437,193,1515]" captionTargetId="figure@23.[151,1437,193,1515]" captionTargetPageId="23" captionText="FIGURES 96–101. Moribaetis latipennis sp. n. (holotype). 96–97, subimago ready to moult to imago; 98, fragment of subimaginal exuviae of wing with fragment of vein; 99, imaginal hind wing extracted from subimago; 100, imaginal fore wing extracted from subimago; 101, subimaginal genitals." httpUri="https://zenodo.org/record/2609867/files/figure.png" pageId="4" pageNumber="235">Fig. 101</figureCitation>
|
||
) have following features the same as in
|
||
<taxonomicName id="05B74D02FF65AC71FB06824CFA659641" authorityName="Flowers" authorityYear="1979" box="[1241,1428,296,320]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF65AC71FB06824CFA659641" box="[1241,1428,296,320]" italics="true" pageId="4" pageNumber="235">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
: pair of protuberances between unistyligers; subimaginal gonostyli with sharply curved 2nd segments and very short 3rd segment; inside them colored imaginal gonostyli with 1st segments strongly crumpled; the same shape of gonovectes; the same unusually short sterno-styligeral muscle.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF65AC71FF1882D2FB9496D1" blockId="4.[151,1437,151,2013]" box="[199,1125,439,464]" pageId="4" pageNumber="235">
|
||
<emphasis id="F0C3EA93FF65AC71FF1882D2FF0796D1" bold="true" box="[199,246,439,464]" pageId="4" pageNumber="235">(18)</emphasis>
|
||
Coloration is similar in both examined species and has the following features:
|
||
</paragraph>
|
||
<paragraph id="C2083681FF65AC71FF1882B9FF3295F1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="235">
|
||
Hypodermal coloration (which is present independently of moults and passes from larva to subimago and to imago) consists of peculiar contrasting brown markings on thorax (
|
||
<figureCitation id="5A8C2A04FF65AC71FC55829AFC3F9519" box="[906,974,511,536]" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="4" pageNumber="235">Fig. 3</figureCitation>
|
||
), abdomen and legs. On abdominal terga it includes paired submedian stripes, usually well developed on uromeres III–IV and VI–VIII; shape of these stripes is somewhat different in
|
||
<taxonomicName id="05B74D02FF65AC71FDCA812CFD249561" authorityName="Flowers" authorityYear="1979" box="[533,725,584,608]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF65AC71FDCA812CFD249561" box="[533,725,584,608]" italics="true" pageId="4" pageNumber="235">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="05B74D02FF65AC71FCCB812CFC5C9561" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[788,941,584,608]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF65AC71FCCB812CFC5C9561" box="[788,941,584,608]" italics="true" pageId="4" pageNumber="235">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF65AC71FC66812DFC0B955E" box="[953,1018,584,607]" pageId="4" pageNumber="235" rank="species">sp. n.</taxonomicNameLabel>
|
||
(see species characteristics below). Hypodermal coloration of legs consists of two brown or reddish spots on posterior side of femur, one at middle and another near apex (
|
||
<figureCitation id="5A8C2A04FF65AC71FEB181EAFE3495A9" box="[366,453,655,680]" captionStart="FIGURES 48–52" captionStartId="13.[151,264,1889,1912]" captionTargetBox="[150,1436,193,1868]" captionTargetId="figure@13.[150,1436,193,1868]" captionTargetPageId="13" captionText="FIGURES 48–52. Moribaetis maculipennis. 48–50, male imago; 51, subimaginal exuviae of left postsubalar sclerite and lateropostnotal crest; 52, subimaginal exuviae of left half of mesonotum." httpUri="https://zenodo.org/record/2609851/files/figure.png" pageId="4" pageNumber="235">Figs 50</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF65AC71FE0E81EAFE0195A9" box="[465,496,655,680]" captionStart="FIGURES 53–55" captionStartId="14.[151,264,1761,1784]" captionTargetBox="[150,1436,193,1740]" captionTargetId="figure@14.[150,1436,193,1740]" captionTargetPageId="14" captionText="FIGURES 53–55. Moribaetis maculipennis. 53–54, female imago; 55, hind wing." httpUri="https://zenodo.org/record/2609853/files/figure.png" pageId="4" pageNumber="235">54</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF65AC71FE2381EAFDED95A9" box="[508,540,655,680]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="4" pageNumber="235">62</figureCitation>
|
||
). In contrast to hypodermal coloration of trunk and legs, hypodermal coloration of imaginal cerci appears only at the end of last larval instar, when subimaginal cerci grow under larval cuticle (
|
||
<figureCitation id="5A8C2A04FF65AC71FAB081D1FF4695F1" captionStart="FIGURES 34–43" captionStartId="11.[151,264,1837,1860]" captionTargetBox="[150,1436,193,1817]" captionTargetId="figure@11.[150,1436,193,1817]" captionTargetPageId="11" captionText="FIGURES 34–43. Moribaetis maculipennis. 34–35, male subimago just after moult; 36, abdominal sternum IX of last instar male larva with gonostyli visible inside; 37, gonostyli of male subimago; 38, exuviae of subimaginal gonostyli; 39, genitals of male imago; 40, caudalii of last instar larva with crumpled sibimaginal cerci visible inside; 41–42, proximal part of imaginal cercus and its subimaginal exuviae (arrows show correspondence of their joints); 43, proximal part of imaginal cercus of another specimen." httpUri="https://zenodo.org/record/2609847/files/figure.png" pageId="4" pageNumber="235">Fig. 40</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF65AC71FF188199FB7794C9" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="235">
|
||
Cuticular pigmentation of larva (present only in larval stage and becoming most intensive at the end of each larval instar) consists of diffusive brown and colorless areas on head, thorax, legs and abdomen (
|
||
<figureCitation id="5A8C2A04FF65AC71FB32807AFAC99439" box="[1261,1336,799,824]" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="4" pageNumber="235">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF65AC71FA99807AFA619439" box="[1350,1424,799,824]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="4" pageNumber="235">64–66</figureCitation>
|
||
). Cuticle of larval frons has the following difference between species: in
|
||
<taxonomicName id="05B74D02FF65AC71FC708020FB98945D" authorityName="Flowers" authorityYear="1979" box="[943,1129,836,860]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF65AC71FC708020FB98945D" box="[943,1129,836,860]" italics="true" pageId="4" pageNumber="235">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
it becomes pigmented only during the last larval instar (
|
||
<figureCitation id="5A8C2A04FF65AC71FE0B8002FDD69481" box="[468,551,871,896]" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="4" pageNumber="235">Fig. 12</figureCitation>
|
||
), but remains to be colorless during previous instars (
|
||
<figureCitation id="5A8C2A04FF65AC71FB5D8002FB259481" box="[1154,1236,871,896]" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="4" pageNumber="235">Fig. 13</figureCitation>
|
||
); in
|
||
<taxonomicName id="05B74D02FF65AC71FAD7800CFA6A9481" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[1288,1435,872,896]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF65AC71FAD7800CFA6A9481" box="[1288,1435,872,896]" italics="true" pageId="4" pageNumber="235">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF65AC71FF4880E8FF2694A5" box="[151,215,909,932]" pageId="4" pageNumber="235" rank="species">sp. n.</taxonomicNameLabel>
|
||
it becomes pigmented at previous instars also (
|
||
<figureCitation id="5A8C2A04FF65AC71FCD480E9FC9294A4" box="[779,867,908,933]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="4" pageNumber="235">Fig. 69</figureCitation>
|
||
). Cuticle of larval cerci is nearly unicolor in
|
||
<taxonomicName id="05B74D02FF65AC71FAA280E8FEDC94C9" authorityName="Flowers" authorityYear="1979" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF65AC71FAA280E8FEDC94C9" italics="true" pageId="4" pageNumber="235">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="5A8C2A04FF65AC71FEE380CAFE8D94C9" box="[316,380,943,968]" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="4" pageNumber="235">Fig. 1</figureCitation>
|
||
) and has darker and lighter areas in
|
||
<taxonomicName id="05B74D02FF65AC71FCCA80D4FC5894C9" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[789,937,944,968]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF65AC71FCCA80D4FC5894C9" box="[789,937,944,968]" italics="true" pageId="4" pageNumber="235">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF65AC71FC6F80D5FC1D94C6" box="[944,1004,944,967]" pageId="4" pageNumber="235" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="5A8C2A04FF65AC71FC2480CAFB8B94C9" box="[1019,1146,943,968]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="4" pageNumber="235">Fig. 62–63</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF65AC71FF1880B1FCCB9334" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="235">
|
||
General coloration of larva is determined by combination of these cuticular and hypodermal pigments. Abdominal segments IX and X usually have both cuticular and hypodermal coloration poorly developed or absent, so tip of abdomen is often contrastingly light (
|
||
<figureCitation id="5A8C2A04FF65AC71FD428779FCF39335" box="[669,770,1052,1077]" captionStart="FIGURES 1–15" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[177,1402,193,1870]" captionTargetId="figure@7.[177,1402,193,1870]" captionTargetPageId="7" captionText="FIGURES 1–15. Larvae of Moribaetis maculipennis. 1–2, last instar male larva, dorsal and lateral view (with hypodermal and cuticular coloration combined); 3, just moulted female larva (with hypodermal coloration only); 4, part of its exuviae from abdomen (with cuticular coloration only); 5–11, tergalii of pairs I–VII; 12, head of last instar male larva; 13, the same, penultimate instar; 14, fragment of cercus with detached swimming setae, view from median side; 15, fragment of costal rib of tergalius. Abbreviations: I–X, abdominal terga." httpUri="https://zenodo.org/record/2609839/files/figure.png" pageId="4" pageNumber="235">Figs 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF65AC71FCD18779FCDF9334" box="[782,814,1052,1077]" captionStart="FIGURES 62–70" captionStartId="18.[151,264,1878,1901]" captionTargetBox="[197,1385,217,1850]" captionTargetId="figure@18.[197,1385,217,1850]" captionTargetPageId="18" captionText="FIGURES 62–70. Larvae of Moribaetis latipennis sp. n. 62, dorsal view; 63, frons of the same specimen (tissues of turbinate eyes start to grow under larval cuticle; grey arrows show boundaries between frons and turbinate eyes on larval cuticle); 64–65, exuviae of left half of pronotum, mesonotum, hind protopteron and abdominal segments I–IX; 66, hind protopteron; 67, fragment of middle part of cercus; 68, head of last instar male larva (tissues of turbinate eyes not started to grow); 69, the same, penultimate instar; 70, posterior margin of abdominal tergum IX (arrow shows midline) (64–65, 67, 70, holotype)." httpUri="https://zenodo.org/record/2609857/files/figure.png" pageId="4" pageNumber="235">62</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF65AC71FF18875AFDF493E9" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="235">
|
||
Subimaginal cuticular coloration is the following: pronotum is light brown; mesonotum is light brown with certain sutures darker brown (
|
||
<figureCitation id="5A8C2A04FF65AC71FE338701FDB3937C" box="[492,578,1124,1149]" captionStart="FIGURES 48–52" captionStartId="13.[151,264,1889,1912]" captionTargetBox="[150,1436,193,1868]" captionTargetId="figure@13.[150,1436,193,1868]" captionTargetPageId="13" captionText="FIGURES 48–52. Moribaetis maculipennis. 48–50, male imago; 51, subimaginal exuviae of left postsubalar sclerite and lateropostnotal crest; 52, subimaginal exuviae of left half of mesonotum." httpUri="https://zenodo.org/record/2609851/files/figure.png" pageId="4" pageNumber="235">Fig. 52</figureCitation>
|
||
); thoracic pleura have light and dark brown areas (
|
||
<figureCitation id="5A8C2A04FF65AC71FB578701FB2C937C" box="[1160,1245,1124,1149]" captionStart="FIGURES 48–52" captionStartId="13.[151,264,1889,1912]" captionTargetBox="[150,1436,193,1868]" captionTargetId="figure@13.[150,1436,193,1868]" captionTargetPageId="13" captionText="FIGURES 48–52. Moribaetis maculipennis. 48–50, male imago; 51, subimaginal exuviae of left postsubalar sclerite and lateropostnotal crest; 52, subimaginal exuviae of left half of mesonotum." httpUri="https://zenodo.org/record/2609851/files/figure.png" pageId="4" pageNumber="235">Fig. 51</figureCitation>
|
||
); wings are dark brown due to dark brown ring at base of each microtrichion, veins lack this coloration (
|
||
<figureCitation id="5A8C2A04FF65AC71FB1587E2FAD493A1" box="[1226,1317,1159,1184]" captionStart="FIGURES 96–101" captionStartId="23.[151,264,1536,1559]" captionTargetBox="[151,1437,193,1515]" captionTargetId="figure@23.[151,1437,193,1515]" captionTargetPageId="23" captionText="FIGURES 96–101. Moribaetis latipennis sp. n. (holotype). 96–97, subimago ready to moult to imago; 98, fragment of subimaginal exuviae of wing with fragment of vein; 99, imaginal hind wing extracted from subimago; 100, imaginal fore wing extracted from subimago; 101, subimaginal genitals." httpUri="https://zenodo.org/record/2609867/files/figure.png" pageId="4" pageNumber="235">Fig. 98</figureCitation>
|
||
); legs are unicolorous, nearly colorless, with dark brown marks at base of tibia (
|
||
<figureCitation id="5A8C2A04FF65AC71FC7287C9FBC193C5" box="[941,1072,1196,1221]" captionStart="FIGURES 25–33" captionStartId="10.[151,264,1857,1880]" captionTargetBox="[151,1436,193,1836]" captionTargetId="figure@10.[151,1436,193,1836]" captionTargetPageId="10" captionText="FIGURES 25–33. Legs of Moribaetis maculipennis. 25–27, larval fore, middle and hind legs; 28, fragment of fore tibia of female subimago; 29, the same, another specimen; 30, fragment of middle tibia of female subimago; 31–32, fragments of fore and middle leg of female subimago; 33, fragment of femur of larval fore leg (25–27 and 29–32, the same individual). Abbreviation: pt-ti, patella-tibial suture." httpUri="https://zenodo.org/record/2609845/files/figure.png" pageId="4" pageNumber="235">Figs 31–32</figureCitation>
|
||
); abdominal terga are uniformly light brown, sterna much lighter.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF65AC71FF118791FB6992C1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="235">
|
||
Imaginal wings have membrane with contrasting brown marking on colorless background; in all three known species these markings border cross veins and are present also in proximal parts of costal and subcostal fields, where cross veins are absent (
|
||
<figureCitation id="5A8C2A04FF65AC71FE358659FD9D9254" box="[490,620,1340,1365]" captionStart="FIGURES 48–52" captionStartId="13.[151,264,1889,1912]" captionTargetBox="[150,1436,193,1868]" captionTargetId="figure@13.[150,1436,193,1868]" captionTargetPageId="13" captionText="FIGURES 48–52. Moribaetis maculipennis. 48–50, male imago; 51, subimaginal exuviae of left postsubalar sclerite and lateropostnotal crest; 52, subimaginal exuviae of left half of mesonotum." httpUri="https://zenodo.org/record/2609851/files/figure.png" pageId="4" pageNumber="235">Figs 48–50</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF65AC71FDA68659FD309254" box="[633,705,1340,1365]" captionStart="FIGURES 53–55" captionStartId="14.[151,264,1761,1784]" captionTargetBox="[150,1436,193,1740]" captionTargetId="figure@14.[150,1436,193,1740]" captionTargetPageId="14" captionText="FIGURES 53–55. Moribaetis maculipennis. 53–54, female imago; 55, hind wing." httpUri="https://zenodo.org/record/2609853/files/figure.png" pageId="4" pageNumber="235">53–54</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF65AC71FD118659FD0B9255" box="[718,762,1340,1364]" captionStart="FIGURES 96–101" captionStartId="23.[151,264,1536,1559]" captionTargetBox="[151,1437,193,1515]" captionTargetId="figure@23.[151,1437,193,1515]" captionTargetPageId="23" captionText="FIGURES 96–101. Moribaetis latipennis sp. n. (holotype). 96–97, subimago ready to moult to imago; 98, fragment of subimaginal exuviae of wing with fragment of vein; 99, imaginal hind wing extracted from subimago; 100, imaginal fore wing extracted from subimago; 101, subimaginal genitals." httpUri="https://zenodo.org/record/2609867/files/figure.png" pageId="4" pageNumber="235">100</figureCitation>
|
||
,
|
||
<figureCitation id="5A8C2A04FF65AC71FCD88659FCC49255" box="[775,821,1340,1364]" captionStart="FIGURES 102–105" captionStartId="24.[151,264,1973,1996]" captionTargetBox="[151,1436,574,1953]" captionTargetId="figure@24.[151,1436,574,1953]" captionTargetPageId="24" captionText="FIGURES 102–105. Moribaetis salvini (lectotype, photo by Ben Price). 102, lateral view; 103, fragment of cercus; 104, dorsal view and tube with genitals; 105, labels." httpUri="https://zenodo.org/record/2609869/files/figure.png" pageId="4" pageNumber="235">102</figureCitation>
|
||
). At least in
|
||
<taxonomicName id="05B74D02FF65AC71FC188658FB739255" authorityName="Flowers" authorityYear="1979" box="[967,1154,1340,1364]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF65AC71FC188658FB739255" box="[967,1154,1340,1364]" italics="true" pageId="4" pageNumber="235">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="05B74D02FF65AC71FB648658FAA19255" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[1211,1360,1340,1364]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF65AC71FB648658FAA19255" box="[1211,1360,1340,1364]" italics="true" pageId="4" pageNumber="235">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF65AC71FA878658FA679255" box="[1368,1430,1341,1364]" pageId="4" pageNumber="235" rank="species">sp. n.</taxonomicNameLabel>
|
||
, this wing pigmentation appears in course of subimaginal development: just after moult from larva to subimago, wings are uniformly grey due to coloration of subimaginal cuticle (
|
||
<figureCitation id="5A8C2A04FF65AC71FC4486E1FBD1929C" box="[923,1056,1412,1437]" captionStart="FIGURES 34–43" captionStartId="11.[151,264,1837,1860]" captionTargetBox="[150,1436,193,1817]" captionTargetId="figure@11.[150,1436,193,1817]" captionTargetPageId="11" captionText="FIGURES 34–43. Moribaetis maculipennis. 34–35, male subimago just after moult; 36, abdominal sternum IX of last instar male larva with gonostyli visible inside; 37, gonostyli of male subimago; 38, exuviae of subimaginal gonostyli; 39, genitals of male imago; 40, caudalii of last instar larva with crumpled sibimaginal cerci visible inside; 41–42, proximal part of imaginal cercus and its subimaginal exuviae (arrows show correspondence of their joints); 43, proximal part of imaginal cercus of another specimen." httpUri="https://zenodo.org/record/2609847/files/figure.png" pageId="4" pageNumber="235">Figs 34–35</figureCitation>
|
||
); before moult from subimago to imago, wings get imaginal markings visible through dark subimaginal cuticle (
|
||
<figureCitation id="5A8C2A04FF65AC71FBD786C2FB7A92C1" box="[1032,1163,1447,1472]" captionStart="FIGURES 96–101" captionStartId="23.[151,264,1536,1559]" captionTargetBox="[151,1437,193,1515]" captionTargetId="figure@23.[151,1437,193,1515]" captionTargetPageId="23" captionText="FIGURES 96–101. Moribaetis latipennis sp. n. (holotype). 96–97, subimago ready to moult to imago; 98, fragment of subimaginal exuviae of wing with fragment of vein; 99, imaginal hind wing extracted from subimago; 100, imaginal fore wing extracted from subimago; 101, subimaginal genitals." httpUri="https://zenodo.org/record/2609867/files/figure.png" pageId="4" pageNumber="235">Figs 96–97</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8AAD650AFF65AC71FF1886A9FD939199" pageId="4" pageNumber="235" type="distribution">
|
||
<paragraph id="C2083681FF65AC71FF1886A9FD81912C" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="235">
|
||
<emphasis id="F0C3EA93FF65AC71FF1886A9FE9192E4" bold="true" box="[199,352,1484,1509]" pageId="4" pageNumber="235">Distribution.</emphasis>
|
||
Central America (central
|
||
<collectingCountry id="BAA07611FF65AC71FD7A86A9FD0C92E4" box="[677,765,1484,1509]" name="Mexico" pageId="4" pageNumber="235">Mexico</collectingCountry>
|
||
,
|
||
<collectingCountry id="BAA07611FF65AC71FCCE86A9FC7C92E4" box="[785,909,1484,1509]" name="Guatemala" pageId="4" pageNumber="235">Guatemala</collectingCountry>
|
||
,
|
||
<collectingCountry id="BAA07611FF65AC71FC7F86A9FBD492E4" box="[928,1061,1484,1509]" name="Costa Rica" pageId="4" pageNumber="235">Costa Rica</collectingCountry>
|
||
,
|
||
<collectingCountry id="BAA07611FF65AC71FBE686A9FB6692E5" box="[1081,1175,1484,1508]" name="Panama" pageId="4" pageNumber="235">Panama</collectingCountry>
|
||
). Species from South America and from non-tropical areas of North America, which were formerly attributed to
|
||
<taxonomicName id="05B74D02FF65AC71FB1B8695FACE9109" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[1220,1343,1520,1544]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF65AC71FB1B8695FACE9109" box="[1220,1343,1520,1544]" italics="true" pageId="4" pageNumber="235">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
, do not belong to this genus (see «Composition»).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF65AC71FF188552FD939199" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="235">
|
||
<emphasis id="F0C3EA93FF65AC71FF188552FEA49151" bold="true" box="[199,341,1591,1616]" pageId="4" pageNumber="235">Adaptation.</emphasis>
|
||
Larvae of
|
||
<taxonomicName id="05B74D02FF65AC71FE00855DFD0A9151" baseAuthorityName="Flowers" baseAuthorityYear="1979" box="[479,763,1592,1616]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF65AC71FE00855DFD0A9151" box="[479,763,1592,1616]" italics="true" pageId="4" pageNumber="235">Moribaetis maculipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="05B74D02FF65AC71FCE3855CFC249151" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[828,981,1592,1616]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF65AC71FCE3855CFC249151" box="[828,981,1592,1616]" italics="true" pageId="4" pageNumber="235">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF65AC71FC3E855CFBD39151" box="[993,1058,1593,1616]" pageId="4" pageNumber="235" rank="species">sp. n.</taxonomicNameLabel>
|
||
keep themselves at places with strongest current and stony bottom. In contrast to larvae of
|
||
<taxonomicName id="05B74D02FF65AC71FCF68539FC5D9175" authorityName="Waltz & McCafferty" authorityYear="1985" box="[809,940,1628,1652]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="subGenus" subGenus="Mayobaetis">
|
||
<emphasis id="F0C3EA93FF65AC71FCF68539FC5D9175" box="[809,940,1628,1652]" italics="true" pageId="4" pageNumber="235">Mayobaetis</emphasis>
|
||
</taxonomicName>
|
||
(which inhabit the same biotopes), larvae of
|
||
<taxonomicName id="05B74D02FF65AC71FF4885E5FEE39199" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[151,274,1664,1688]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF65AC71FF4885E5FEE39199" box="[151,274,1664,1688]" italics="true" pageId="4" pageNumber="235">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
do not crawl out of the water.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8AAD650AFF65AC70FF1885C1FC439174" lastPageId="5" lastPageNumber="236" pageId="4" pageNumber="235" type="description">
|
||
<paragraph id="C2083681FF65AC71FF1885C1FCB39071" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="235">
|
||
<emphasis id="F0C3EA93FF65AC71FF1885C1FE9691BC" bold="true" box="[199,359,1700,1725]" pageId="4" pageNumber="235">Composition.</emphasis>
|
||
<taxonomicName id="05B74D02FF65AC71FEAB85C1FE1E91BD" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[372,495,1700,1724]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF65AC71FEAB85C1FE1E91BD" box="[372,495,1700,1724]" italics="true" pageId="4" pageNumber="235">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
includes two species known as larvae and imagoes,
|
||
<taxonomicName id="05B74D02FF65AC71FBA285C1FEB491E1" authority="(Flowers 1979)" baseAuthorityName="Flowers" baseAuthorityYear="1979" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF65AC71FBA285C1FA6D91BD" box="[1149,1436,1700,1724]" italics="true" pageId="4" pageNumber="235">Moribaetis maculipennis</emphasis>
|
||
(
|
||
<bibRefCitation id="A6264B70FF65AC71FF4185A2FECD91E1" author="Flowers, R. W." box="[158,316,1735,1760]" pageId="4" pageNumber="235" pagination="187 - 191" refId="ref10974" refString="Flowers, R. W. (1979) A new species of Baetis from Panama (Ephemeroptera: Baetidae). Pan-Pacific Entomologist, 55 (3), 187 - 191." type="journal article" year="1979">Flowers 1979</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="05B74D02FF65AC71FEA185ADFD8191E1" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[382,624,1736,1760]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF65AC71FEA185ADFD8191E1" box="[382,624,1736,1760]" italics="true" pageId="4" pageNumber="235">Moribaetis latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF65AC71FDA685ACFD4691E1" box="[633,695,1737,1760]" pageId="4" pageNumber="235" rank="species">sp. n.</taxonomicNameLabel>
|
||
(see below). The third known species,
|
||
<taxonomicName id="05B74D02FF65AC71FBA585ADFF2B9004" authority="(Eaton 1885)" baseAuthorityName="Eaton" baseAuthorityYear="1885" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis id="F0C3EA93FF65AC71FBA585ADFAB891E1" box="[1146,1353,1736,1760]" italics="true" pageId="4" pageNumber="235">Moribaetis salvini</emphasis>
|
||
(Eaton 1885)
|
||
</taxonomicName>
|
||
, is known only as male imago; structure of its gonostyli, shape and venation of hind wing and coloration of fore wing testify about its belonging to
|
||
<taxonomicName id="05B74D02FF65AC71FDB28475FD199029" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[621,744,1808,1832]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF65AC71FDB28475FD199029" box="[621,744,1808,1832]" italics="true" pageId="4" pageNumber="235">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
(see below). The fourth species,
|
||
<taxonomicName id="05B74D02FF65AC71FBAC8475FE32904D" authority="Romero & Esquivel 2018" authorityName="Romero & Esquivel" authorityYear="2018" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="brachiostrinus">
|
||
<emphasis id="F0C3EA93FF65AC71FBAC8475FA6D9029" box="[1139,1436,1808,1832]" italics="true" pageId="4" pageNumber="235">Moribaetis brachiostrinus</emphasis>
|
||
<bibRefCitation id="A6264B70FF65AC71FF488450FE32904D" author="Romero, D. & Esquivel, C." box="[151,451,1844,1869]" pageId="4" pageNumber="235" pagination="531 - 541" refId="ref11819" refString="Romero, D. & Esquivel, C. (2018) Moribaetis brachiostrinus (Ephemeroptera: Baetidae), a new species of mayfly from Costa Rica. Zootaxa, 4433 (3), 531 - 541. http: // dx. doi. org / 10.11646 / zootaxa. 4433.3.8" type="journal article" year="2018">Romero & Esquivel 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
, is described as larvae; it is closely related to
|
||
<taxonomicName id="05B74D02FF65AC71FC3A8450FB8A904D" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[997,1147,1844,1868]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF65AC71FC3A8450FB8A904D" box="[997,1147,1844,1868]" italics="true" pageId="4" pageNumber="235">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF65AC71FB598450FB37904D" box="[1158,1222,1845,1868]" pageId="4" pageNumber="235" rank="species">sp. n.</taxonomicNameLabel>
|
||
Six other species, attributed by some authors to
|
||
<taxonomicName id="05B74D02FF65AC71FE3B843DFDAE9071" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[484,607,1880,1904]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF65AC71FE3B843DFDAE9071" box="[484,607,1880,1904]" italics="true" pageId="4" pageNumber="235">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
, do not belong to it.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF65AC70FF188419FC6B97D4" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,1653]" lastPageId="5" lastPageNumber="236" pageId="4" pageNumber="235">
|
||
Originally the genus
|
||
<taxonomicName id="05B74D02FF65AC71FE6E8419FDDD9095" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[433,556,1916,1940]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF65AC71FE6E8419FDDD9095" box="[433,556,1916,1940]" italics="true" pageId="4" pageNumber="235">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
included, besides the nominative subgenus, the subgenus
|
||
<taxonomicName id="05B74D02FF65AC71FB6A8419FEAD90B8" authority="Waltz & McCafferty 1985" authorityName="Waltz & McCafferty" authorityYear="1985" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="subGenus" subGenus="Mayobaetis">
|
||
<emphasis id="F0C3EA93FF65AC71FB6A8419FAC99095" box="[1205,1336,1916,1940]" italics="true" pageId="4" pageNumber="235">Mayobaetis</emphasis>
|
||
<bibRefCitation id="A6264B70FF65AC71FAE08419FEAD90B8" author="Waltz, R. D. & McCafferty, W. P." pageId="4" pageNumber="235" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
with a single species
|
||
<taxonomicName id="05B74D02FF65AC71FDB984C5FB7290B9" authority="(Mayo 1973)" baseAuthorityName="Mayo" baseAuthorityYear="1973" box="[614,1155,1952,1976]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="ellenae" subGenus="Mayobaetis">
|
||
<emphasis id="F0C3EA93FF65AC71FDB984C5FD1090B9" box="[614,737,1952,1976]" italics="true" pageId="4" pageNumber="235">Moribaetis</emphasis>
|
||
(
|
||
<emphasis id="F0C3EA93FF65AC71FD2A84C5FC8990B9" box="[757,888,1952,1976]" italics="true" pageId="4" pageNumber="235">Mayobaetis</emphasis>
|
||
)
|
||
<emphasis id="F0C3EA93FF65AC71FC5384C5FC2C90B9" box="[908,989,1952,1976]" italics="true" pageId="4" pageNumber="235">ellenae</emphasis>
|
||
(
|
||
<bibRefCitation id="A6264B70FF65AC71FC2F84C5FB8B90B9" author="Mayo, V. K." box="[1008,1146,1952,1976]" pageId="4" pageNumber="235" pagination="285 - 288" refId="ref11612" refString="Mayo, V. K. (1973) A new species of Baetis from Ecuador (Ephemeroptera: Baetidae). Pan-Pacific Entomologist, 49 (4), 285 - 288." type="journal article" year="1973">Mayo 1973</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; subsequently the taxon
|
||
<taxonomicName id="05B74D02FF65AC71FF4884A1FEEB90DD" authorityName="Waltz & McCafferty" authorityYear="1985" box="[151,282,1988,2012]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="subGenus" subGenus="Mayobaetis">
|
||
<emphasis id="F0C3EA93FF65AC71FF4884A1FEEB90DD" box="[151,282,1988,2012]" italics="true" pageId="4" pageNumber="235">Mayobaetis</emphasis>
|
||
</taxonomicName>
|
||
got a generic rank (
|
||
<bibRefCitation id="A6264B70FF65AC71FDC384A1FC6B90DC" author="Lugo-Ortiz, C. R. & McCafferty, W. P." box="[540,922,1988,2013]" pageId="4" pageNumber="235" pagination="367 - 380" refId="ref11489" refString="Lugo-Ortiz, C. R. & McCafferty, W. P. (1996) Phylogeny and classification of the Baetodes complex (Ephemeroptera: Baetidae), with description of a new genus. Journal of the North American Benthological Society, 15 (3), 367 - 380. https: // doi. org / 10.2307 / 1467283" type="journal article" year="1996">Lugo-Ortiz & McCafferty 1996</bibRefCitation>
|
||
). In contrast to
|
||
<taxonomicName id="05B74D02FF65AC71FBBA84A1FB1190DD" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[1125,1248,1988,2012]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF65AC71FBBA84A1FB1190DD" box="[1125,1248,1988,2012]" italics="true" pageId="4" pageNumber="235">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
, in
|
||
<taxonomicName id="05B74D02FF65AC71FAC684A1FA6D90DD" authorityName="Waltz & McCafferty" authorityYear="1985" box="[1305,1436,1988,2012]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="subGenus" subGenus="Mayobaetis">
|
||
<emphasis id="F0C3EA93FF65AC71FAC684A1FA6D90DD" box="[1305,1436,1988,2012]" italics="true" pageId="4" pageNumber="235">Mayobaetis</emphasis>
|
||
</taxonomicName>
|
||
paraglossa has usual shape and bears three setae of the dorso-median row (
|
||
<bibRefCitation id="A6264B70FF64AC70FC2283F2FA7F97B1" author="Waltz, R. D. & McCafferty, W. P." box="[1021,1422,151,176]" pageId="5" pageNumber="236" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985: fig. 10</bibRefCitation>
|
||
); patella-tibial suture is fully developed and crosses inner side of tibia.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF64AC70FF1883BAFB0B9641" blockId="5.[151,1437,151,1653]" pageId="5" pageNumber="236">
|
||
Three species known as winged stages only,
|
||
<taxonomicName id="05B74D02FF64AC70FD688384FC7597F9" authority="Traver 1971" authorityName="Traver" authorityYear="1971" box="[695,900,224,248]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="aneto">
|
||
<emphasis id="F0C3EA93FF64AC70FD688384FD0497F9" box="[695,757,225,248]" italics="true" pageId="5" pageNumber="236">aneto</emphasis>
|
||
<bibRefCitation id="A6264B70FF64AC70FD238385FC7597F9" author="Traver, J. R." box="[764,900,224,248]" pageId="5" pageNumber="236" pagination="58 - 63" refId="ref11873" refString="Traver, J. R. (1971) Four new species of Neotropical Baetis (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 73 (1), 58 - 63." type="journal article" year="1971">Traver 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF64AC70FC4B8385FC2B97F6" authorityName="Leach" authorityYear="1815" box="[916,986,224,247]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FC4B8385FC2B97F6" box="[916,986,224,247]" italics="true" pageId="5" pageNumber="236">Baetis</emphasis>
|
||
</taxonomicName>
|
||
],
|
||
<taxonomicName id="05B74D02FF64AC70FC2E8384FB3397F9" authority="Navas 1912" authorityName="Navas" authorityYear="1912" box="[1009,1218,224,248]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="comes">
|
||
<emphasis id="F0C3EA93FF64AC70FC2E8384FBC697F9" box="[1009,1079,225,248]" italics="true" pageId="5" pageNumber="236">comes</emphasis>
|
||
<bibRefCitation id="A6264B70FF64AC70FBE28385FB3397F9" author="Navas, L." box="[1085,1218,224,248]" pageId="5" pageNumber="236" pagination="194 - 202" refId="ref11761" refString="Navas, L. (1912) Neuropteros nuevos de America. Broteria, Serie Zoologica, 10, 194 - 202." type="journal article" year="1912">Navás 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF64AC70FB0E8385FAE697F6" authorityName="Leach" authorityYear="1815" box="[1233,1303,224,247]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FB0E8385FAE697F6" box="[1233,1303,224,247]" italics="true" pageId="5" pageNumber="236">Baetis</emphasis>
|
||
</taxonomicName>
|
||
] and
|
||
<taxonomicName id="05B74D02FF64AC70FA898385FE38961D" authority="Needham & Murphy 1924" authorityName="Needham & Murphy" authorityYear="1924" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="socius">
|
||
<emphasis id="F0C3EA93FF64AC70FA898385FA6D97F6" box="[1366,1436,224,247]" italics="true" pageId="5" pageNumber="236">socius</emphasis>
|
||
<bibRefCitation id="A6264B70FF64AC70FF488261FE38961D" author="Needham, J. G. & Murphy, H. E." box="[151,457,260,285]" pageId="5" pageNumber="236" pagination="1 - 79" refId="ref11784" refString="Needham, J. G. & Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, Entomological Series 4, 24, 1 - 79." type="journal article" year="1924">Needham & Murphy 1924</bibRefCitation>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF64AC70FE028260FDD2961D" authorityName="Leach" authorityYear="1815" box="[477,547,261,284]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FE028260FDD2961D" box="[477,547,261,284]" italics="true" pageId="5" pageNumber="236">Baetis</emphasis>
|
||
</taxonomicName>
|
||
], were placed in the genus
|
||
<taxonomicName id="05B74D02FF64AC70FCB58261FC14961D" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[874,997,260,284]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FCB58261FC14961D" box="[874,997,260,284]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="A6264B70FF64AC70FBCA8261FA69961D" author="Lugo-Ortiz, C. R. & McCafferty, W. P." box="[1045,1432,260,285]" pageId="5" pageNumber="236" pagination="257 - 262" refId="ref11552" refString="Lugo-Ortiz, C. R. & McCafferty, W. P. (1999) Revision of South American species of Baetidae (Ephemeroptera) previously placed in Baetis Leach and Pseudocloeon Klapalek. Annales de Limnologie, 35 (4), 257 - 262. https: // doi. org / 10.1051 / limn / 1999034" type="journal article" year="1999">Lugo-Ortiz & McCafferty (1999)</bibRefCitation>
|
||
. Their placement in
|
||
<taxonomicName id="05B74D02FF64AC70FEAE824DFE1D9641" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[369,492,296,320]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FEAE824DFE1D9641" box="[369,492,296,320]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
is regarded to be groundless (
|
||
<bibRefCitation id="A6264B70FF64AC70FCE48242FBF39641" author="McCafferty, W. P." box="[827,1026,295,320]" pageId="5" pageNumber="236" pagination="375 - 379" refId="ref11646" refString="McCafferty, W. P. (2000) Notations on South American Baetidae (Ephemeroptera). Entomological News, 111 (5), 375 - 379." type="journal article" year="2000">McCafferty 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A6264B70FF64AC70FBD2824DFB1F9641" author="Cruz, P. V. & Salles, F. F." box="[1037,1262,295,320]" pageId="5" pageNumber="236" pagination="398 - 400" refId="ref10842" refString="Cruz, P. V. & Salles, F. F. (2014) Further notes on South American species of Baetidae (Ephemeroptera) assigned to Moribaetis Waltz & McCafferty, 1985. Zootaxa, 3793 (3), 398 - 400. https: // doi. org / 10.11646 / zootaxa. 3793.3.9" type="journal article" year="2014">Cruz & Salles 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF64AC70FF188228FDD496AD" blockId="5.[151,1437,151,1653]" pageId="5" pageNumber="236">
|
||
In contrast to
|
||
<taxonomicName id="05B74D02FF64AC70FEB78229FE129665" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[360,483,332,356]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FEB78229FE129665" box="[360,483,332,356]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
, the species
|
||
<taxonomicName id="05B74D02FF64AC70FDA78228FD479665" authorityName="Traver" authorityYear="1971" box="[632,694,333,356]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="aneto">
|
||
<emphasis id="F0C3EA93FF64AC70FDA78228FD479665" box="[632,694,333,356]" italics="true" pageId="5" pageNumber="236">aneto</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF64AC70FD178228FCFF9665" authorityName="Leach" authorityYear="1815" box="[712,782,333,356]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FD178228FCFF9665" box="[712,782,333,356]" italics="true" pageId="5" pageNumber="236">Baetis</emphasis>
|
||
</taxonomicName>
|
||
] has unforked 2nd vein of hind wing and usual shape of gonostyli with 1st segment much shorter and thicker than 2nd, and with 3rd segment elongate (
|
||
<bibRefCitation id="A6264B70FF64AC70FB628215FABD9689" author="Traver, J. R." box="[1213,1356,368,392]" pageId="5" pageNumber="236" pagination="58 - 63" refId="ref11873" refString="Traver, J. R. (1971) Four new species of Neotropical Baetis (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 73 (1), 58 - 63." type="journal article" year="1971">Traver 1971</bibRefCitation>
|
||
: figs 1, 4;
|
||
<bibRefCitation id="A6264B70FF64AC70FF6B82F1FE6596AD" author="Cruz, P. V. & Salles, F. F." box="[180,404,404,429]" pageId="5" pageNumber="236" pagination="398 - 400" refId="ref10842" refString="Cruz, P. V. & Salles, F. F. (2014) Further notes on South American species of Baetidae (Ephemeroptera) assigned to Moribaetis Waltz & McCafferty, 1985. Zootaxa, 3793 (3), 398 - 400. https: // doi. org / 10.11646 / zootaxa. 3793.3.9" type="journal article" year="2014">Cruz & Salles 2014</bibRefCitation>
|
||
: figs 10–11).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF64AC70FF1882D2FEDB9519" blockId="5.[151,1437,151,1653]" pageId="5" pageNumber="236">
|
||
The species
|
||
<taxonomicName id="05B74D02FF64AC70FE8A82DCFE6A96D1" authorityName="Navas" authorityYear="1912" box="[341,411,441,464]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="comes">
|
||
<emphasis id="F0C3EA93FF64AC70FE8A82DCFE6A96D1" box="[341,411,441,464]" italics="true" pageId="5" pageNumber="236">comes</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF64AC70FE7282DDFE0296CE" authorityName="Leach" authorityYear="1815" box="[429,499,440,463]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FE7282DDFE0296CE" box="[429,499,440,463]" italics="true" pageId="5" pageNumber="236">Baetis</emphasis>
|
||
</taxonomicName>
|
||
] was said to be «similis
|
||
<taxonomicName id="05B74D02FF64AC70FCCC82DDFC6496CE" authority="Etn." authorityName="Eaton" authorityYear="1885" box="[787,917,440,464]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis id="F0C3EA93FF64AC70FCCC82DDFCAF96D1" box="[787,862,440,464]" italics="true" pageId="5" pageNumber="236">salvini</emphasis>
|
||
Etn.
|
||
</taxonomicName>
|
||
» and having hind wing with forked 2nd vein (
|
||
<bibRefCitation id="A6264B70FF64AC70FF4182B9FE4696F5" author="Navas, L." box="[158,439,476,501]" pageId="5" pageNumber="236" pagination="194 - 202" refId="ref11761" refString="Navas, L. (1912) Neuropteros nuevos de America. Broteria, Serie Zoologica, 10, 194 - 202." type="journal article" year="1912">Navás 1912: 194, fig. 1a</bibRefCitation>
|
||
); however, these characters are not enough to determine its systematic position (
|
||
<bibRefCitation id="A6264B70FF64AC70FA9882B9FEEF9519" author="Cruz, P. V. & Salles, F. F." pageId="5" pageNumber="236" pagination="398 - 400" refId="ref10842" refString="Cruz, P. V. & Salles, F. F. (2014) Further notes on South American species of Baetidae (Ephemeroptera) assigned to Moribaetis Waltz & McCafferty, 1985. Zootaxa, 3793 (3), 398 - 400. https: // doi. org / 10.11646 / zootaxa. 3793.3.9" type="journal article" year="2014">Cruz & Salles 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C2083681FF64AC70FF188141FED195A6" blockId="5.[151,1437,151,1653]" pageId="5" pageNumber="236">
|
||
The species
|
||
<taxonomicName id="05B74D02FF64AC70FE868140FE6E953D" authorityName="Needham & Murphy" authorityYear="1924" box="[345,415,549,572]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="socius">
|
||
<emphasis id="F0C3EA93FF64AC70FE868140FE6E953D" box="[345,415,549,572]" italics="true" pageId="5" pageNumber="236">socius</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF64AC70FE6C8140FE08953D" authorityName="Leach" authorityYear="1815" box="[435,505,549,572]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FE6C8140FE08953D" box="[435,505,549,572]" italics="true" pageId="5" pageNumber="236">Baetis</emphasis>
|
||
</taxonomicName>
|
||
] was described as female subimago; in contrast to
|
||
<taxonomicName id="05B74D02FF64AC70FB888141FB23953D" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[1111,1234,548,572]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FB888141FB23953D" box="[1111,1234,548,572]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
, its hind wing is narrow, curved, with fork of 2nd vein asymmetric and without 3rd vein (
|
||
<bibRefCitation id="A6264B70FF64AC70FC078122FF289584" author="Needham, J. G. & Murphy, H. E." pageId="5" pageNumber="236" pagination="1 - 79" refId="ref11784" refString="Needham, J. G. & Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, Entomological Series 4, 24, 1 - 79." type="journal article" year="1924">Needham & Murphy 1924: pl. XII: fig. 153A</bibRefCitation>
|
||
). Judging by this structure of hind wing and relatively large size (body length
|
||
<quantity id="054F9B64FF64AC70FBA58109FB329585" box="[1146,1219,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="236" unit="mm" value="8.0">8 mm</quantity>
|
||
), it may belong to
|
||
<taxonomicName id="05B74D02FF64AC70FF4881F5FEEB95A9" authorityName="Waltz & McCafferty" authorityYear="1985" box="[151,282,656,680]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="235" phylum="Arthropoda" rank="subGenus" subGenus="Mayobaetis">
|
||
<emphasis id="F0C3EA93FF64AC70FF4881F5FEEB95A9" box="[151,282,656,680]" italics="true" pageId="5" pageNumber="236">Mayobaetis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C2083681FF64AC70FF1881D1FD1693A1" blockId="5.[151,1437,151,1653]" pageId="5" pageNumber="236">
|
||
The species originally named
|
||
<taxonomicName id="05B74D02FF64AC70FDFA81D1FC9095CC" authority="Waltz" authorityName="Waltz" authorityYear="1985" box="[549,865,692,717]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="macaferti">
|
||
<emphasis id="F0C3EA93FF64AC70FDFA81D1FCE495CD" box="[549,789,692,716]" italics="true" pageId="5" pageNumber="236">Moribaetis macaferti</emphasis>
|
||
Waltz
|
||
</taxonomicName>
|
||
(in
|
||
<bibRefCitation id="A6264B70FF64AC70FC4A81D1FB3F95CC" author="Waltz, R. D. & McCafferty, W. P." box="[917,1230,692,717]" pageId="5" pageNumber="236" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty) 1985</bibRefCitation>
|
||
was described as larvae from
|
||
<collectingCountry id="BAA07611FF64AC70FEFE81BDFE6A95F1" box="[289,411,727,752]" name="Costa Rica" pageId="5" pageNumber="236">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="BAA07611FF64AC70FE0C81B2FDA195F1" box="[467,592,727,752]" name="Guatemala" pageId="5" pageNumber="236">Guatemala</collectingCountry>
|
||
. Male imago, ascribed to this species by
|
||
<bibRefCitation id="A6264B70FF64AC70FBC181B2FA6695F1" author="McCafferty, W. P. & Lugo-Ortiz, C. R." box="[1054,1431,727,752]" pageId="5" pageNumber="236" pagination="117 - 121" refId="ref11722" refString="McCafferty, W. P. & Lugo-Ortiz, C. R. (1998) The adult of Moribaetis macaferti (Ephemeroptera: Baetidae). Entomological News, 109 (2), 117 - 121." type="journal article" year="1998">McCafferty & Lugo-Ortiz (1998)</bibRefCitation>
|
||
, actually belongs to the new species
|
||
<taxonomicName id="05B74D02FF64AC70FD878199FCA19415" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" box="[600,848,764,788]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF64AC70FD878199FCA19415" box="[600,848,764,788]" italics="true" pageId="5" pageNumber="236">Moribaetis latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF64AC70FCBF8198FC559415" box="[864,932,765,788]" pageId="5" pageNumber="236" rank="species">sp. n.</taxonomicNameLabel>
|
||
(see below), so true imago of
|
||
<taxonomicName id="05B74D02FF64AC70FAEE8199FA6D9415" authorityName="Waltz" authorityYear="1985" box="[1329,1436,764,788]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="macaferti">
|
||
<emphasis id="F0C3EA93FF64AC70FAEE8199FA6D9415" box="[1329,1436,764,788]" italics="true" pageId="5" pageNumber="236">macaferti</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF64AC70FF7F8045FEEA9439" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[160,283,800,824]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FF7F8045FEEA9439" box="[160,283,800,824]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
] is unknown. In contrast to true
|
||
<taxonomicName id="05B74D02FF64AC70FD4B8045FCFE9439" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[660,783,800,824]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FD4B8045FCFE9439" box="[660,783,800,824]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
, larva of
|
||
<taxonomicName id="05B74D02FF64AC70FC5F8045FC1A9439" authorityName="Waltz" authorityYear="1985" box="[896,1003,800,824]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="macaferti">
|
||
<emphasis id="F0C3EA93FF64AC70FC5F8045FC1A9439" box="[896,1003,800,824]" italics="true" pageId="5" pageNumber="236">macaferti</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF64AC70FC228045FB899439" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[1021,1144,800,824]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FC228045FB899439" box="[1021,1144,800,824]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
] has labrum not widened and bearing usual «1+5–6 stout submarginal setae»; mandibular incisors are not blade-like; paraglossae are not narrowed apically and bear 3 setae in the dorso-median row; patella-tibial suture is fully developed and crosses inner side of tibia (
|
||
<bibRefCitation id="A6264B70FF64AC70FEA980E9FC5894A4" author="Waltz, R. D. & McCafferty, W. P." box="[374,937,908,933]" pageId="5" pageNumber="236" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985: 243–245, figs 15–21</bibRefCitation>
|
||
). This species was attributed to
|
||
<taxonomicName id="05B74D02FF64AC70FAFE80E9FA6D94A5" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[1313,1436,908,932]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FAFE80E9FA6D94A5" box="[1313,1436,908,932]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
based on presence of unpaired subapical seta on larval claw [see character of
|
||
<taxonomicName id="05B74D02FF64AC70FC2080D5FB8B94C9" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[1023,1146,944,968]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FC2080D5FB8B94C9" box="[1023,1146,944,968]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
(11) above] and presence of minute procoxal «osmobranchia» [see character of
|
||
<taxonomicName id="05B74D02FF64AC70FCFA80B1FC5194ED" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[805,928,980,1004]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FCFA80B1FC5194ED" box="[805,928,980,1004]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
(6)]. Combination of its larval characters (absence of frontal keel; non-widened labrum with few submarginal setae; non-blade-like mandibles; nonnarrowed paraglossae with several setae in the dorso-median row; fully-developed patella-tibial suture; unpaired subapical seta on claw; presence of hind protopteron) is in agreement with the characteristics of the taxon
|
||
<taxonomicName id="05B74D02FF64AC70FF488701FE6F937D" authority="Kluge 1992" authorityName="Kluge" authorityYear="1992" box="[151,414,1124,1149]" class="Insecta" family="Baetidae" genus="Caribaetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="genus" status="subgen. nov.">
|
||
<emphasis id="F0C3EA93FF64AC70FF488701FEFF937D" box="[151,270,1124,1148]" italics="true" pageId="5" pageNumber="236">Caribaetis</emphasis>
|
||
<bibRefCitation id="A6264B70FF64AC70FEC98701FE6F937D" author="Kluge, N. J." box="[278,414,1124,1149]" pageId="5" pageNumber="236" pagination="13 - 20" refId="ref11163" refString="Kluge, N. J. (1992) [Cuban mayflies of the family Baetidae (Ephemeroptera). 2. Subgenera Caribaetis subgen. n. and Americabaetis subgen. n. of the genus Baetis s. l.]. Zoologicheskiy Zhurnal, 71 (4), 13 - 20. [in Russian]" type="journal article" year="1992">Kluge 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
, which is recently accepted as a genus (
|
||
<bibRefCitation id="A6264B70FF64AC70FCB58701FB7D937D" author="Kluge, N. J. & Novikova, E. A." box="[874,1164,1124,1149]" pageId="5" pageNumber="236" pagination="209 - 236" refId="ref11430" refString="Kluge, N. J. & Novikova, E. A. (2014) Systematics of Indobaetis Muller-Liebenau & Morihara 1982, and related implications for some other Baetidae genera (Ephemeroptera). Zootaxa, 3835 (2), 209 - 236. https: // doi. org / 10.11646 / zootaxa. 3835.2.3" type="journal article" year="2014">Kluge & Novikova 2014</bibRefCitation>
|
||
). So this species can be presumably treated as
|
||
<taxonomicName id="05B74D02FF64AC70FE4D87EDFD8B93A1" authorityName="Kluge & Bernal Vega" authorityYear="2018" baseAuthorityName=": McCafferty & Lugo-Ortiz" baseAuthorityYear="1985" box="[402,634,1160,1184]" class="Insecta" family="Baetidae" genus="Caribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="macaferti" status="comb. nov.">
|
||
<emphasis id="F0C3EA93FF64AC70FE4D87EDFD8B93A1" box="[402,634,1160,1184]" italics="true" pageId="5" pageNumber="236">Caribaetis macaferti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F0C3EA93FF64AC70FD5E87E2FD1693A1" bold="true" box="[641,743,1159,1184]" pageId="5" pageNumber="236">
|
||
<taxonomicNameLabel id="EBF057E8FF64AC70FD5E87E2FD1693A1" box="[641,743,1159,1184]" pageId="5" pageNumber="236" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="C2083681FF64AC70FF1887C9FC439174" blockId="5.[151,1437,151,1653]" pageId="5" pageNumber="236">
|
||
The species described under the name
|
||
<taxonomicName id="05B74D02FF64AC70FDA387C9FB9993C5" authority="McCafferty 2007" authorityName="McCafferty" authorityYear="2007" box="[636,1128,1196,1221]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="mimbresaurus">
|
||
<emphasis id="F0C3EA93FF64AC70FDA387C9FC6F93C5" box="[636,926,1196,1220]" italics="true" pageId="5" pageNumber="236">Moribaetis mimbresaurus</emphasis>
|
||
<bibRefCitation id="A6264B70FF64AC70FC7987C9FB9993C5" author="McCafferty, W. P." box="[934,1128,1196,1221]" pageId="5" pageNumber="236" pagination="696 - 699" refId="ref11676" refString="McCafferty, W. P. (2007) Moribaetis mimbresaurus, new species (Ephemeroptera: Baetidae): first representative of the genus north of Mexico. Proceedings of the Entomological Society of Washington, 109 (3), 696 - 699." type="journal article" year="2007">McCafferty 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
is known only as imagoes. The only arguments to attribute it to
|
||
<taxonomicName id="05B74D02FF64AC70FD9887B5FD3393E9" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[583,706,1232,1256]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FD9887B5FD3393E9" box="[583,706,1232,1256]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
were «large size, membrane pigmentation in the forewing, the basally margined, distally oriented, sharp costal process of the hindwing, short stalk of the turbinate eye, and small terminal segment of the genital forceps» (
|
||
<bibRefCitation id="A6264B70FF64AC70FDB58672FCC29231" author="McCafferty, W. P." box="[618,819,1303,1328]" pageId="5" pageNumber="236" pagination="696 - 699" refId="ref11676" refString="McCafferty, W. P. (2007) Moribaetis mimbresaurus, new species (Ephemeroptera: Baetidae): first representative of the genus north of Mexico. Proceedings of the Entomological Society of Washington, 109 (3), 696 - 699." type="journal article" year="2007">McCafferty 2007</bibRefCitation>
|
||
). However, all these characters are found in other taxa, including
|
||
<emphasis id="F0C3EA93FF64AC70FED28659FE689255" box="[269,409,1340,1364]" italics="true" pageId="5" pageNumber="236">Rhodobaetis</emphasis>
|
||
<bibRefCitation id="A6264B70FF64AC70FE7A8659FDDF9255" author="Jacob, U." box="[421,558,1340,1365]" pageId="5" pageNumber="236" pagination="59 - 129" refId="ref11008" refString="Jacob, U. (2003) Baetis Leach 1815, sensu stricto oder sensu lato. Ein Beitrag zum Gattungskonzept auf der Grundlage von Artengruppen mit Bestimmungsschlusseln. Lauterbornia, 47, 59 - 129." type="journal article" year="2003">Jacob 2003</bibRefCitation>
|
||
. Size of some species of
|
||
<emphasis id="F0C3EA93FF64AC70FCB98659FC039255" box="[870,1010,1340,1364]" italics="true" pageId="5" pageNumber="236">Rhodobaetis</emphasis>
|
||
varies in great range, reaching one centimeter of fore wing length in individuals developed in cold water. Pigmentation of distal parts of costal and subcostal fields of fore wings is found in many taxa, including
|
||
<emphasis id="F0C3EA93FF64AC70FCB586E1FC07929D" box="[874,1014,1412,1436]" italics="true" pageId="5" pageNumber="236">Rhodobaetis</emphasis>
|
||
. Among
|
||
<emphasis id="F0C3EA93FF64AC70FBBD86E1FB1F929D" box="[1122,1262,1412,1436]" italics="true" pageId="5" pageNumber="236">Rhodobaetis</emphasis>
|
||
, costal process varies from directed anteriorly to hooked distally. Stalk of turbinate eye of
|
||
<taxonomicName id="05B74D02FF64AC70FBF186CDFB3C92C1" authorityName="McCafferty" authorityYear="2007" box="[1070,1229,1448,1472]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="mimbresaurus">
|
||
<emphasis id="F0C3EA93FF64AC70FBF186CDFB3C92C1" box="[1070,1229,1448,1472]" italics="true" pageId="5" pageNumber="236">mimbresaurus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF64AC70FB3B86CDFAAE92C1" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[1252,1375,1448,1472]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FB3B86CDFAAE92C1" box="[1252,1375,1448,1472]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
] has moderate height, as in many baetid taxa. Small terminal segment of the gonostylus is usual for
|
||
<emphasis id="F0C3EA93FF64AC70FB3786A9FA8592E5" box="[1256,1396,1484,1508]" italics="true" pageId="5" pageNumber="236">Rhodobaetis</emphasis>
|
||
. In contrast to
|
||
<taxonomicName id="05B74D02FF64AC70FECB8695FE7E9109" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[276,399,1520,1544]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF64AC70FECB8695FE7E9109" box="[276,399,1520,1544]" italics="true" pageId="5" pageNumber="236">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
, this species has fore wing without dark spots around cross veins (
|
||
<bibRefCitation id="A6264B70FF64AC70FBA2868AFA7D9109" author="McCafferty, W. P." box="[1149,1420,1519,1544]" pageId="5" pageNumber="236" pagination="696 - 699" refId="ref11676" refString="McCafferty, W. P. (2007) Moribaetis mimbresaurus, new species (Ephemeroptera: Baetidae): first representative of the genus north of Mexico. Proceedings of the Entomological Society of Washington, 109 (3), 696 - 699." type="journal article" year="2007">McCafferty 2007: fig. 3</bibRefCitation>
|
||
), hind wing without furcation on 2nd vein (
|
||
<bibRefCitation id="A6264B70FF64AC70FDB98571FC82912D" author="McCafferty, W. P." box="[614,883,1556,1581]" pageId="5" pageNumber="236" pagination="696 - 699" refId="ref11676" refString="McCafferty, W. P. (2007) Moribaetis mimbresaurus, new species (Ephemeroptera: Baetidae): first representative of the genus north of Mexico. Proceedings of the Entomological Society of Washington, 109 (3), 696 - 699." type="journal article" year="2007">McCafferty 2007: fig. 4</bibRefCitation>
|
||
) and a usual shape of gonostyli, with 1st segment much shorter and thicker than 2nd segment (
|
||
<bibRefCitation id="A6264B70FF64AC70FD5B8552FC639151" author="McCafferty, W. P." box="[644,914,1591,1616]" pageId="5" pageNumber="236" pagination="696 - 699" refId="ref11676" refString="McCafferty, W. P. (2007) Moribaetis mimbresaurus, new species (Ephemeroptera: Baetidae): first representative of the genus north of Mexico. Proceedings of the Entomological Society of Washington, 109 (3), 696 - 699." type="journal article" year="2007">McCafferty 2007: fig. 5</bibRefCitation>
|
||
). Until larva of this species is known, it should be preliminary treated as
|
||
<taxonomicName id="05B74D02FF64AC70FE6C8538FCB49175" authorityName="Kluge & Bernal Vega" authorityYear="2018" baseAuthorityName="Kluge & Bernal Vega" baseAuthorityYear="2018" box="[435,837,1628,1652]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="subGenus" status="comb. nov." subGenus="Rhodobaetis">
|
||
<emphasis id="F0C3EA93FF64AC70FE6C8538FE089175" box="[435,505,1629,1652]" italics="true" pageId="5" pageNumber="236">Baetis</emphasis>
|
||
(
|
||
<emphasis id="F0C3EA93FF64AC70FDD58539FD679175" box="[522,662,1628,1652]" italics="true" pageId="5" pageNumber="236">Rhodobaetis</emphasis>
|
||
)
|
||
<emphasis id="F0C3EA93FF64AC70FD798539FCB49175" box="[678,837,1628,1652]" italics="true" pageId="5" pageNumber="236">mimbresaurus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F0C3EA93FF64AC70FC938539FC439174" bold="true" box="[844,946,1628,1653]" pageId="5" pageNumber="236">
|
||
<taxonomicNameLabel id="EBF057E8FF64AC70FC938539FC439174" box="[844,946,1628,1653]" pageId="5" pageNumber="236" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |