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<document id="9310E41D162CB112FC59DCFF6418103C" ID-DOI="10.5281/zenodo.5353780" ID-ISSN="2345-7600" ID-Zenodo-Dep="5353780" ID-ZooBank="8493E540-DB48-4CA6-8A73-9D232D5546E1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212606084" checkinUser="tatiana" docAuthor="Grinang, Jongkar &amp; Ng, Peter K. L." docDate="2014" docId="03AE760B7025FFB8FC3BF9DBC43D4808" docLanguage="en" docName="62rbz389-395.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Thelphusula Bott 1969" docType="treatment" docVersion="4" lastPageNumber="390" masterDocId="FF970E737025FFB9FFC5FFC8C56C4C7D" masterDocTitle="Taxonomy of the freshwater crab Coccusa cristicervix Ng &amp; Grinang, 2004 (Decapoda: Brachyura: Gecarcinucidae), with description of a new species of Thelphusula from Sarawak, Malaysia, Borneo" masterLastPageNumber="395" masterPageNumber="389" pageNumber="389" updateTime="1699187930052" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4649FA17BDFC824C9C014AE714110B37">Taxonomy of the freshwater crab Coccusa cristicervix Ng &amp; Grinang, 2004 (Decapoda: Brachyura: Gecarcinucidae), with description of a new species of Thelphusula from Sarawak, Malaysia, Borneo</mods:title>
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<mods:namePart id="9F34AF75C843F99ECE0C6052EEFFDAB7">Grinang, Jongkar</mods:namePart>
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<mods:namePart id="C8EB4037023F978714793AE060FCD1BE">Ng, Peter K. L.</mods:namePart>
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<mods:title id="B59682C983CCD2BEE58ADF2A3566A4E5">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="7F9D834BD689829C837CDA1DF587C160">2014</mods:date>
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<mods:number id="79743B55BC07446EA5BBDFCBCE9F43F3">62</mods:number>
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<mods:classification id="42FA3A9172494BC32BD3D291D2C3153B">journal article</mods:classification>
<mods:identifier id="FCDF1ABBDFD6124B579C5FD18C53095E" type="DOI">10.5281/zenodo.5353780</mods:identifier>
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<treatment id="03AE760B7025FFB8FC3BF9DBC43D4808" ID-DOI="http://doi.org/10.5281/zenodo.5451317" ID-Zenodo-Dep="5451317" LSID="urn:lsid:plazi:treatment:03AE760B7025FFB8FC3BF9DBC43D4808" httpUri="http://treatment.plazi.org/id/03AE760B7025FFB8FC3BF9DBC43D4808" lastPageId="1" lastPageNumber="390" pageId="0" pageNumber="389">
<subSubSection id="C31D94967025FFB9FC3BF9DBC06B4A57" box="[1022,1287,1555,1578]" pageId="0" pageNumber="389" type="nomenclature">
<paragraph id="8BB8C71D7025FFB9FC3BF9DBC06B4A57" blockId="0.[1022,1287,1555,1578]" box="[1022,1287,1555,1578]" pageId="0" pageNumber="389">
<heading id="D0F070717025FFB9FC3BF9DBC06B4A57" bold="true" box="[1022,1287,1555,1578]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="389" reason="2">
<taxonomicName id="4C07BC9E7025FFB9FC3BF9DBC06B4A57" ID-CoL="7Q4C6" authority="Bott, 1969" authorityName="Bott" authorityYear="1969" box="[1022,1287,1555,1578]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7025FFB9FC3BF9DBC06B4A57" bold="true" box="[1022,1287,1555,1578]" pageId="0" pageNumber="389">
<emphasis id="B9731B0F7025FFB9FC3BF9DBC1E54A57" bold="true" box="[1022,1161,1555,1578]" italics="true" pageId="0" pageNumber="389">Thelphusula</emphasis>
<bibRefCitation id="EF96BAEC7025FFB9FB57F9DBC06B4A57" author="Bott R" box="[1170,1287,1555,1578]" pageId="0" pageNumber="389" pagination="359 - 366" refId="ref3387" refString="Bott R (1969) Flusskrabben aus Asien und ihre Klassifikation (Crustacea, Decapoda). Senckenbergiana Biologica, 50 (5 - 6): 359 - 366." type="journal article" year="1969">Bott, 1969</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C31D94967025FFB8FC86F99BC43D4808" lastPageId="1" lastPageNumber="390" pageId="0" pageNumber="389" type="discussion">
<paragraph id="8BB8C71D7025FFB8FC86F99BC42B4E88" blockId="0.[835,1474,1619,2090]" lastBlockId="1.[113,752,190,757]" lastPageId="1" lastPageNumber="390" pageId="0" pageNumber="389">
<emphasis id="B9731B0F7025FFB9FC86F99BC6D84A17" bold="true" box="[835,948,1619,1642]" pageId="0" pageNumber="389">Remarks.</emphasis>
<taxonomicName id="4C07BC9E7025FFB9FC7BF99BC1A64A17" authority="Bott, 1969" authorityName="Bott" authorityYear="1969" box="[958,1226,1619,1642]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7025FFB9FC7BF99BC1294A17" box="[958,1093,1619,1642]" italics="true" pageId="0" pageNumber="389">Thelphusula</emphasis>
<bibRefCitation id="EF96BAEC7025FFB9FB95F99BC1A64A17" author="Bott R" box="[1104,1226,1619,1642]" pageId="0" pageNumber="389" pagination="359 - 366" refId="ref3387" refString="Bott R (1969) Flusskrabben aus Asien und ihre Klassifikation (Crustacea, Decapoda). Senckenbergiana Biologica, 50 (5 - 6): 359 - 366." type="journal article" year="1969">Bott, 1969</bibRefCitation>
</taxonomicName>
, was established with
<taxonomicName id="4C07BC9E7025FFB9FC86F9BBC01D4AF7" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" box="[835,1393,1651,1674]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="buergeri" subGenus="Geothelphusa">
<emphasis id="B9731B0F7025FFB9FC86F9BBC6C94AF7" box="[835,933,1651,1674]" italics="true" pageId="0" pageNumber="389">Potamon</emphasis>
(
<emphasis id="B9731B0F7025FFB9FC7CF9BBC13C4AF7" box="[953,1104,1651,1674]" italics="true" pageId="0" pageNumber="389">Geothelphusa</emphasis>
)
<emphasis id="B9731B0F7025FFB9FBA1F9BBC1AD4AF7" box="[1124,1217,1651,1674]" italics="true" pageId="0" pageNumber="389">buergeri</emphasis>
De
<bibRefCitation id="EF96BAEC7025FFB9FB30F9BBC01D4AF7" author="Man JG De" box="[1269,1393,1651,1674]" pageId="0" pageNumber="389" pagination="53 - 144" refId="ref3657" refString="Man JG De (1899) Zoological results of the Dutch scientific expedition to central Borneo. The Crustacea. Part II, Brachyura. Notes from the Leyden Museum, 21: 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
</taxonomicName>
, as the
<typeStatus id="54BC79BF7025FFB9FC86F95BC61D4AD7" box="[835,881,1683,1706]" pageId="0" pageNumber="389">type</typeStatus>
species.
<bibRefCitation id="EF96BAEC7025FFB9FC15F95BC1274AD7" author="Bott R" box="[976,1099,1683,1706]" pageId="0" pageNumber="389" pagination="359 - 366" refId="ref3387" refString="Bott R (1969) Flusskrabben aus Asien und ihre Klassifikation (Crustacea, Decapoda). Senckenbergiana Biologica, 50 (5 - 6): 359 - 366." type="journal article" year="1969">Bott (1969)</bibRefCitation>
recognised five other species in the genus as he defined it at the time:
<taxonomicName id="4C07BC9E7025FFB9FB75F97BC1EA4A97" authority="De Man, 1902" authorityName="De Man" authorityYear="1902" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="baramensis" subGenus="Potamonautes">
<emphasis id="B9731B0F7025FFB9FB75F97BC07C4AB7" box="[1200,1296,1715,1738]" italics="true" pageId="0" pageNumber="389">Potamon</emphasis>
(
<emphasis id="B9731B0F7025FFB9FAE4F97BC0D64AB7" box="[1313,1466,1715,1738]" italics="true" pageId="0" pageNumber="389">Potamonautes</emphasis>
)
<emphasis id="B9731B0F7025FFB9FC86F91BC6A54A97" box="[835,969,1747,1770]" italics="true" pageId="0" pageNumber="389">baramensis</emphasis>
De
<bibRefCitation id="EF96BAEC7025FFB9FBC6F91BC1EA4A97" author="Man JG De" box="[1027,1158,1747,1770]" pageId="0" pageNumber="389" pagination="467 - 929" refId="ref3700" refString="Man JG De (1902) Die von Herrn Professor Kukenthal im indischen Archipel gesammelten Dekapoden und Stomatopoden. In: Kukenthal W (ed.), Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo, in Auftrage der Senckenberg. Naturforschenden Gesellschaft ausgefuhrt von Dr. Willy Kukenthal. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 25 (3): 467 - 929, pls. 19 - 27." type="journal article" year="1902">Man, 1902</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C07BC9E7025FFB9FB5DF91BC1CF4B77" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="hendersonianum" subGenus="Geothelphusa">
<emphasis id="B9731B0F7025FFB9FB5DF91BC06C4A97" box="[1176,1280,1747,1770]" italics="true" pageId="0" pageNumber="389">Potamon</emphasis>
(
<emphasis id="B9731B0F7025FFB9FAD3F91BC0D44A97" box="[1302,1464,1747,1770]" italics="true" pageId="0" pageNumber="389">Geothelphusa</emphasis>
)
<emphasis id="B9731B0F7025FFB9FC86F93BC6994B77" box="[835,1013,1779,1802]" italics="true" pageId="0" pageNumber="389">hendersonianum</emphasis>
De
<bibRefCitation id="EF96BAEC7025FFB9FBECF93BC1CF4B77" author="Man JG De" box="[1065,1187,1779,1802]" pageId="0" pageNumber="389" pagination="53 - 144" refId="ref3657" refString="Man JG De (1899) Zoological results of the Dutch scientific expedition to central Borneo. The Crustacea. Part II, Brachyura. Notes from the Leyden Museum, 21: 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C07BC9E7025FFB9FB74F93BC1EB4B57" authority="Borradaile, 1900" authorityName="Borradaile" authorityYear="1900" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="kadamaianum" subGenus="Geothelphusa">
<emphasis id="B9731B0F7025FFB9FB74F93BC07E4B77" box="[1201,1298,1779,1802]" italics="true" pageId="0" pageNumber="389">Potamon</emphasis>
(
<emphasis id="B9731B0F7025FFB9FAE1F93BC0D54B77" box="[1316,1465,1779,1802]" italics="true" pageId="0" pageNumber="389">Geothelphusa</emphasis>
)
<emphasis id="B9731B0F7025FFB9FC86F8DBC6BA4B57" box="[835,982,1811,1834]" italics="true" pageId="0" pageNumber="389">kadamaianum</emphasis>
<bibRefCitation id="EF96BAEC7025FFB9FC1FF8DBC1EB4B57" author="Borradaile LA" box="[986,1159,1811,1834]" pageId="0" pageNumber="389" pagination="93 - 95" refId="ref3352" refString="Borradaile LA (1900) On a small collection of decapod crustaceans from freshwaters in North Borneo. Proceedings of the Zoological Society of London, 1900 (1): 93 - 95." type="journal article" year="1900">Borradaile, 1900</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C07BC9E7025FFB9FB4AF8DBC1724B37" authority="Chace, 1938" authorityName="Chace" authorityYear="1938" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="luidana" subGenus="Liothelphusa">
<emphasis id="B9731B0F7025FFB9FB4AF8DBC04B4B57" box="[1167,1319,1811,1834]" italics="true" pageId="0" pageNumber="389">Parathelphusa</emphasis>
(
<emphasis id="B9731B0F7025FFB9FAF6F8DBC0D54B57" box="[1331,1465,1811,1834]" italics="true" pageId="0" pageNumber="389">Liothelphusa</emphasis>
)
<emphasis id="B9731B0F7025FFB9FC86F8FBC6F84B37" box="[835,916,1843,1866]" italics="true" pageId="0" pageNumber="389">luidana</emphasis>
<bibRefCitation id="EF96BAEC7025FFB9FC5CF8FBC1724B37" author="Chace FA Jr" box="[921,1054,1843,1866]" pageId="0" pageNumber="389" pagination="9 - 22" refId="ref3467" refString="Chace FA Jr (1938) Freshwater water decapod crustacea from Mount Kinabalu, British North Borneo. Proceedings of the New England Zoological Club, 17: 9 - 22, figs. 1, 2, pls. 1 - 4." type="book chapter" year="1938">Chace, 1938</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C07BC9E7025FFB9FB91F8FBC6D44B17" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="melanippe" subGenus="Potamon">
<emphasis id="B9731B0F7025FFB9FB91F8FBC1D84B37" box="[1108,1204,1843,1866]" italics="true" pageId="0" pageNumber="389">Potamon</emphasis>
(
<emphasis id="B9731B0F7025FFB9FB07F8FBC04E4B37" box="[1218,1314,1843,1866]" italics="true" pageId="0" pageNumber="389">Potamon</emphasis>
)
<emphasis id="B9731B0F7025FFB9FAEAF8FBC0F14B37" box="[1327,1437,1843,1866]" italics="true" pageId="0" pageNumber="389">melanippe</emphasis>
De
<bibRefCitation id="EF96BAEC7025FFB9FC86F89BC6D44B17" author="Man JG De" box="[835,952,1875,1898]" pageId="0" pageNumber="389" pagination="53 - 144" refId="ref3657" refString="Man JG De (1899) Zoological results of the Dutch scientific expedition to central Borneo. The Crustacea. Part II, Brachyura. Notes from the Leyden Museum, 21: 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
</taxonomicName>
(see also
<bibRefCitation id="EF96BAEC7025FFB9FBE3F89BC1A94B17" author="Bott R" box="[1062,1221,1875,1898]" pageId="0" pageNumber="389" pagination="1 - 338" refId="ref3418" refString="Bott R (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt, 526: 1 - 338, pls. 1 - 58." type="journal article" year="1970">Bott, 1970: 58</bibRefCitation>
).
<taxonomicName id="4C07BC9E7025FFB9FB1CF89BC1EA4BF7" authority="Lanchester, 1900" authorityName="Lanchester" authorityYear="1900" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="bidiensis" subGenus="Thelphusa">
<emphasis id="B9731B0F7025FFB9FB1CF89BC0554B17" box="[1241,1337,1875,1898]" italics="true" pageId="0" pageNumber="389">Potamon</emphasis>
(
<emphasis id="B9731B0F7025FFB9FA8FF89BC0D54B17" box="[1354,1465,1875,1898]" italics="true" pageId="0" pageNumber="389">Thelphusa</emphasis>
)
<emphasis id="B9731B0F7025FFB9FC86F8BBC6C74BF7" box="[835,939,1907,1930]" italics="true" pageId="0" pageNumber="389">bidiensis</emphasis>
<bibRefCitation id="EF96BAEC7025FFB9FC7DF8BBC1EA4BF7" author="Lanchester WF" box="[952,1158,1907,1930]" pageId="0" pageNumber="389" pagination="249 - 265" refId="ref3615" refString="Lanchester WF (1900) On some malacostracous crustaceans from Malaysia in the collection of the Sarawak Museum. Annals and Magazine of Natural History, (7) 6 (33): 249 - 265, pl. 12." type="journal article" year="1900">Lanchester, 1900</bibRefCitation>
</taxonomicName>
, was regarded as a junior subjective synonym of
<taxonomicName id="4C07BC9E7025FFB9FB98F85BC0034BD7" box="[1117,1391,1939,1962]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="melanippe">
<emphasis id="B9731B0F7025FFB9FB98F85BC0034BD7" box="[1117,1391,1939,1962]" italics="true" pageId="0" pageNumber="389">Thelphusula melanippe</emphasis>
</taxonomicName>
, while
<taxonomicName id="4C07BC9E7025FFB9FC86F87BC12C4BB7" authorityName="Borradaile" authorityYear="1900" box="[835,1088,1971,1994]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="species" species="kadamaianum">
<emphasis id="B9731B0F7025FFB9FC86F87BC12C4BB7" box="[835,1088,1971,1994]" italics="true" pageId="0" pageNumber="389">Potamon kadamaianum</emphasis>
</taxonomicName>
was treated as its subspecies by
<bibRefCitation id="EF96BAEC7025FFB9FA51F87BC6E24B97" author="Bott R" pageId="0" pageNumber="389" pagination="1 - 338" refId="ref3418" refString="Bott R (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt, 526: 1 - 338, pls. 1 - 58." type="journal article" year="1970">Bott (1970)</bibRefCitation>
.
<bibRefCitation id="EF96BAEC7025FFB9FC5DF81BC1504B97" author="Ng PKL" box="[920,1084,2003,2026]" pageId="0" pageNumber="389" pagination="63 - 72" refId="ref3818" refString="Ng PKL (1989 a) The identity of the cavernicolous freshwater crab Potamon (Thelphusa) bidiense Lanchester, 1900 (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from Sarawak, Borneo, with description of a new genus. Raffles Bulletin of Zoology, 37 (1 &amp; 2): 63 - 72." type="journal article" year="1989">Ng (1989a: 64)</bibRefCitation>
questioned
<bibRefCitation id="EF96BAEC7025FFB9FB78F81BC0274B97" author="Bott R" box="[1213,1355,2003,2026]" pageId="0" pageNumber="389" pagination="1 - 338" refId="ref3418" refString="Bott R (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt, 526: 1 - 338, pls. 1 - 58." type="journal article" year="1970">Bott’s (1970)</bibRefCitation>
concept of
<taxonomicName id="4C07BC9E7025FFB9FC86F83BC6AB4477" authorityName="Bott" authorityYear="1969" box="[835,967,2035,2058]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7025FFB9FC86F83BC6AB4477" box="[835,967,2035,2058]" italics="true" pageId="0" pageNumber="389">Thelphusula</emphasis>
</taxonomicName>
and argued that it should be separated into three distinct morphological groups. He disputed
<bibRefCitation id="EF96BAEC7025FFB9FAEAF7DBC0AE4457" author="Bott R" box="[1327,1474,2067,2090]" pageId="0" pageNumber="389" pagination="1 - 338" refId="ref3418" refString="Bott R (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt, 526: 1 - 338, pls. 1 - 58." type="journal article" year="1970">Bott’s (1970)</bibRefCitation>
synonymisation of
<taxonomicName id="4C07BC9E7024FFB8FE8DFF76C7794CA8" authorityName="Lanchester" authorityYear="1900" box="[328,533,190,213]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="bidiensis">
<emphasis id="B9731B0F7024FFB8FE8DFF76C7794CA8" box="[328,533,190,213]" italics="true" pageId="1" pageNumber="390">Potamon bidiensis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C07BC9E7024FFB8FD9CFF76C7854CA8" box="[601,745,190,213]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="melanippe">
<emphasis id="B9731B0F7024FFB8FD9CFF76C7854CA8" box="[601,745,190,213]" italics="true" pageId="1" pageNumber="390">T. melanippe</emphasis>
</taxonomicName>
; recognised it as a distinct species and referred it to a monotypic genus,
<taxonomicName id="4C07BC9E7024FFB8FE87FF36C7024D68" authority="Ng, 1989 a" authorityName="Ng" authorityYear="1989" box="[322,622,254,277]" class="Malacostraca" family="Gecarcinucidae" genus="Stygothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FE87FF36C4864D68" box="[322,490,254,277]" italics="true" pageId="1" pageNumber="390">Stygothelphusa</emphasis>
<bibRefCitation id="EF96BAEC7024FFB8FE30FF36C7024D68" author="Ng PKL" box="[501,622,254,277]" pageId="1" pageNumber="390" pagination="63 - 72" refId="ref3818" refString="Ng PKL (1989 a) The identity of the cavernicolous freshwater crab Potamon (Thelphusa) bidiense Lanchester, 1900 (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from Sarawak, Borneo, with description of a new genus. Raffles Bulletin of Zoology, 37 (1 &amp; 2): 63 - 72." type="journal article" year="1989">Ng, 1989a</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EF96BAEC7024FFB8FDBEFF36C79C4D68" author="Ng PKL" box="[635,752,254,277]" pageId="1" pageNumber="390" pagination="1 - 12" refId="ref3924" refString="Ng PKL (1991) Bornean freshwater crabs of the genus Arachnothelphusa gen. nov. (Crustacea: Decapoda: Brachyura: Gecarcinucidae). Zoologische Mededelingen, 65 (1): 1 - 12." type="journal article" year="1991">Ng (1991)</bibRefCitation>
subsequently established a new genus,
<taxonomicName id="4C07BC9E7024FFB8FDC0FED6C7AA4D48" authorityName="P. K. L. Ng" authorityYear="1991" box="[517,710,286,309]" class="Malacostraca" family="Gecarcinucidae" genus="Arachnothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FDC0FED6C7AA4D48" box="[517,710,286,309]" italics="true" pageId="1" pageNumber="390">Arachnothelphusa</emphasis>
</taxonomicName>
, for
<taxonomicName id="4C07BC9E7024FFB8FFB4FEF6C7374D28" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" box="[113,603,318,341]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="melanippe" subGenus="Potamon">
<emphasis id="B9731B0F7024FFB8FFB4FEF6C5BD4D28" box="[113,209,318,341]" italics="true" pageId="1" pageNumber="390">Potamon</emphasis>
(
<emphasis id="B9731B0F7024FFB8FF25FEF6C42C4D28" box="[224,320,318,341]" italics="true" pageId="1" pageNumber="390">Potamon</emphasis>
)
<emphasis id="B9731B0F7024FFB8FE8BFEF6C4D14D28" box="[334,445,318,341]" italics="true" pageId="1" pageNumber="390">melanippe</emphasis>
De
<bibRefCitation id="EF96BAEC7024FFB8FE23FEF6C7374D28" author="Man JG De" box="[486,603,318,341]" pageId="1" pageNumber="390" pagination="53 - 144" refId="ref3657" refString="Man JG De (1899) Zoological results of the Dutch scientific expedition to central Borneo. The Crustacea. Part II, Brachyura. Notes from the Leyden Museum, 21: 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C07BC9E7024FFB8FD55FEF6C7084D08" authority="Borradaile, 1900" authorityName="Borradaile" authorityYear="1900" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="kadamaianum" subGenus="Geothelphusa">
<emphasis id="B9731B0F7024FFB8FD55FEF6C79C4D28" box="[656,752,318,341]" italics="true" pageId="1" pageNumber="390">Potamon</emphasis>
(
<emphasis id="B9731B0F7024FFB8FFBFFE96C4674D08" box="[122,267,350,373]" italics="true" pageId="1" pageNumber="390">Geothelphusa</emphasis>
)
<emphasis id="B9731B0F7024FFB8FEDCFE96C4C24D08" box="[281,430,350,373]" italics="true" pageId="1" pageNumber="390">kadamaianum</emphasis>
<bibRefCitation id="EF96BAEC7024FFB8FE76FE96C7084D08" author="Borradaile LA" box="[435,612,350,373]" pageId="1" pageNumber="390" pagination="93 - 95" refId="ref3352" refString="Borradaile LA (1900) On a small collection of decapod crustaceans from freshwaters in North Borneo. Proceedings of the Zoological Society of London, 1900 (1): 93 - 95." type="journal article" year="1900">Borradaile, 1900</bibRefCitation>
</taxonomicName>
, treating both as distinct species.
<bibRefCitation id="EF96BAEC7024FFB8FE8CFEB6C4D24DE8" author="Ng PKL" box="[329,446,382,405]" pageId="1" pageNumber="390" pagination="181 - 205" refId="ref3962" refString="Ng PKL (1995) The freshwater crabs and prawns (Crustacea: Decapoda) of Bako National Park, Sarawak, Malaysia, with descriptions of one new genus and three new species. Raffles Bulletin of Zoology, 43 (1): 181 - 205." type="journal article" year="1995">Ng (1995)</bibRefCitation>
, in describing a new genus from western
<collectingCountry id="F310878D7024FFB8FEC5FE56C4314DC8" box="[256,349,414,437]" name="Malaysia" pageId="1" pageNumber="390">Sarawak</collectingCountry>
, showed that
<taxonomicName id="4C07BC9E7024FFB8FE2DFE56C4AF4DA8" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="hendersonianum" subGenus="Geothelphusa">
<emphasis id="B9731B0F7024FFB8FE2DFE56C72B4DC8" box="[488,583,414,437]" italics="true" pageId="1" pageNumber="390">Potamon</emphasis>
(
<emphasis id="B9731B0F7024FFB8FD90FE56C78B4DC8" box="[597,743,414,437]" italics="true" pageId="1" pageNumber="390">Geothelphusa</emphasis>
)
<emphasis id="B9731B0F7024FFB8FFB4FE76C44F4DA8" box="[113,291,446,469]" italics="true" pageId="1" pageNumber="390">hendersonianum</emphasis>
De
<bibRefCitation id="EF96BAEC7024FFB8FE8BFE76C4AF4DA8" author="Man JG De" box="[334,451,446,469]" pageId="1" pageNumber="390" pagination="53 - 144" refId="ref3657" refString="Man JG De (1899) Zoological results of the Dutch scientific expedition to central Borneo. The Crustacea. Part II, Brachyura. Notes from the Leyden Museum, 21: 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
</taxonomicName>
, had been incorrectly placed in
<taxonomicName id="4C07BC9E7024FFB8FF4EFE16C4624D88" authorityName="Bott" authorityYear="1969" box="[139,270,478,501]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FF4EFE16C4624D88" box="[139,270,478,501]" italics="true" pageId="1" pageNumber="390">Thelphusula</emphasis>
</taxonomicName>
as it has differently structured G1 and G2, and should be transferred to
<taxonomicName id="4C07BC9E7024FFB8FEBFFE36C72A4E68" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[378,582,510,533]" class="Malacostraca" family="Gecarcinucidae" genus="Bakousa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FEBFFE36C4BA4E68" box="[378,470,510,533]" italics="true" pageId="1" pageNumber="390">Bakousa</emphasis>
<bibRefCitation id="EF96BAEC7024FFB8FE1BFE36C72A4E68" author="Ng PKL" box="[478,582,510,533]" pageId="1" pageNumber="390" pagination="181 - 205" refId="ref3962" refString="Ng PKL (1995) The freshwater crabs and prawns (Crustacea: Decapoda) of Bako National Park, Sarawak, Malaysia, with descriptions of one new genus and three new species. Raffles Bulletin of Zoology, 43 (1): 181 - 205." type="journal article" year="1995">Ng, 1995</bibRefCitation>
</taxonomicName>
, instead.
<bibRefCitation id="EF96BAEC7024FFB8FD69FE36C58D4E48" author="Tan SH &amp; Ng PKL" pageId="1" pageNumber="390" pagination="808 - 822" refId="ref4350" refString="Tan SH &amp; Ng PKL (1998) Revision of the genus Thelphusula (Decapoda: Brachyura: Gecarcinucidae) with description of a new genus and three new species. Journal of Crustacean Biology, 18 (4): 808 - 822." type="journal article" year="1998">Tan &amp; Ng (1998)</bibRefCitation>
revised
<taxonomicName id="4C07BC9E7024FFB8FEFBFDD6C4AE4E48" authorityName="Bott" authorityYear="1969" box="[318,450,542,565]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FEFBFDD6C4AE4E48" box="[318,450,542,565]" italics="true" pageId="1" pageNumber="390">Thelphusula</emphasis>
</taxonomicName>
, restricted the diagnosis for nine species and established a new genus,
<taxonomicName id="4C07BC9E7024FFB8FDFDFDF6C7F84E28" authorityName="Tan &amp; Ng" authorityYear="1998" box="[568,660,574,597]" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FDFDFDF6C7F84E28" box="[568,660,574,597]" italics="true" pageId="1" pageNumber="390">Coccusa</emphasis>
</taxonomicName>
, for two species from Borneo in which the carapace is ovate and much smoother, and most significantly, the G1 terminal segment is very long, slender and as long as the subterminal segment (versus distinctly shorter than the subterminal segment in
<taxonomicName id="4C07BC9E7024FFB8FFB4FD16C5994E88" authorityName="Bott" authorityYear="1969" box="[113,245,734,757]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B9731B0F7024FFB8FFB4FD16C5994E88" box="[113,245,734,757]" italics="true" pageId="1" pageNumber="390">Thelphusula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A240A6747024FFB8FF3BFD16C4574E88" box="[254,315,734,757]" pageId="1" pageNumber="390" sensu="stricto">s. str.</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8BB8C71D7024FFB8FFB4FCD6C43D4808" blockId="1.[113,752,798,1141]" pageId="1" pageNumber="390">
<bibRefCitation id="EF96BAEC7024FFB8FFB4FCD6C4344F48" author="Ng PKL &amp; Grinang J" box="[113,344,798,821]" pageId="1" pageNumber="390" pagination="299 - 325" refId="ref4094" refString="Ng PKL &amp; Grinang J (2004) Decapod crustaceans with descriptions of three new species. In: Yong HS Ng FSP &amp; Yen EEL (eds.) Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, 59, No. 80 (New Series), Special Issue No. 6: 299 - 325." type="book chapter" year="2004">Ng &amp; Grinang (2004)</bibRefCitation>
recognised a third species of
<taxonomicName id="4C07BC9E7024FFB8FD51FCD6C79C4F48" authorityName="Tan &amp; Ng" authorityYear="1998" box="[660,752,798,821]" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FD51FCD6C79C4F48" box="[660,752,798,821]" italics="true" pageId="1" pageNumber="390">Coccusa</emphasis>
</taxonomicName>
from western
<collectingCountry id="F310878D7024FFB8FED7FCF6C4194F28" box="[274,373,830,853]" name="Malaysia" pageId="1" pageNumber="390">Sarawak</collectingCountry>
. Although only
<specimenCount id="9D010C947024FFB8FDF5FCF6C7F64F28" box="[560,666,830,853]" pageId="1" pageNumber="390" type="adult">one adult</specimenCount>
female specimen was available, they noted that the general carapace shape and features agreed better with
<taxonomicName id="4C07BC9E7024FFB8FDC4FCB6C7314FE8" authorityName="Tan &amp; Ng" authorityYear="1998" box="[513,605,894,917]" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FDC4FCB6C7314FE8" box="[513,605,894,917]" italics="true" pageId="1" pageNumber="390">Coccusa</emphasis>
</taxonomicName>
and therefore referred it to this genus, even though it was the most granulated of the three known taxa. The collection of more specimens of the Sarawakian species, including males, shows that
<taxonomicName id="4C07BC9E7024FFB8FF65FC36C5904868" authorityName="Tan &amp; Ng" authorityYear="1998" box="[160,252,1022,1045]" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FF65FC36C5904868" box="[160,252,1022,1045]" italics="true" pageId="1" pageNumber="390">Coccusa</emphasis>
</taxonomicName>
should be restricted for species with smoother carapaces. On the basis of the gonopod structures,
<taxonomicName id="4C07BC9E7024FFB8FD51FBD6C48B4828" authority="Ng &amp; Grinang, 2004" authorityName="Ng &amp; Grinang" authorityYear="2004" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="cristicervix">
<emphasis id="B9731B0F7024FFB8FD51FBD6C5814828" italics="true" pageId="1" pageNumber="390">Coccusa cristicervix</emphasis>
<bibRefCitation id="EF96BAEC7024FFB8FF3DFBF6C48B4828" author="Ng PKL &amp; Grinang J" box="[248,487,1086,1109]" pageId="1" pageNumber="390" pagination="299 - 325" refId="ref4094" refString="Ng PKL &amp; Grinang J (2004) Decapod crustaceans with descriptions of three new species. In: Yong HS Ng FSP &amp; Yen EEL (eds.) Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, 59, No. 80 (New Series), Special Issue No. 6: 299 - 325." type="book chapter" year="2004">Ng &amp; Grinang, 2004</bibRefCitation>
</taxonomicName>
, must be transferred to
<taxonomicName id="4C07BC9E7024FFB8FFB4FB96C5994808" authorityName="Bott" authorityYear="1969" box="[113,245,1118,1141]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis id="B9731B0F7024FFB8FFB4FB96C5994808" box="[113,245,1118,1141]" italics="true" pageId="1" pageNumber="390">Thelphusula</emphasis>
</taxonomicName>
instead.
</paragraph>
</subSubSection>
</treatment>
</document>
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