298 lines
40 KiB
XML
298 lines
40 KiB
XML
<document id="ED2CDB4FD306BB4BDC49C59C4F0CA7EF" ID="10.11646/zootaxa.4179.2.6" ID-DOI="10.11646/zootaxa.4179.2.6" ID-GBIF-Dataset="06e87c9b-0077-437a-b313-4aa8cd55cce3" ID-ISSN="1175-5326" ID-PMC="PMC5321562" ID-PubMed="27811673" ID-Zenodo-Dep="163902" ID-ZooBank="7BEE4FFC-36E8-473D-8D4E-53D151A8CD2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477899714536" checkinUser="plazi" docAuthor="Hyžný, Matúš & Gross, Martin" docDate="2016" docId="9605E23F9271FFC896D8F9A5FC13FAC3" docLanguage="en" docName="zootaxa.4179.2.6.pdf" docOrigin="Zootaxa 4179 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Iphiculus eliasi Hyžný & Gross, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="268" masterDocId="6A3C9A479272FFCD964FFFADFFDAFFFC" masterDocTitle="A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus" masterLastPageNumber="270" masterPageNumber="263" pageNumber="266" updateTime="1698391355973" updateUser="plazi">
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<mods:titleInfo id="2D39392373FC6E982DA47E733871C784">
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<mods:title id="3CFF32D6EEDD31D8C33369A124E9A1DD">A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus</mods:title>
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<mods:namePart id="566C4C54417AC5F9BA110EAEB3B03E9C">Hyžný, Matúš</mods:namePart>
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<mods:name id="BC7BF3F0A6FC703391C60D34906385C3" type="personal">
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<mods:roleTerm id="F9CA3E259E72473A759102C4C838F2F7">Author</mods:roleTerm>
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<mods:namePart id="05C72298D820D6FF858EF091A1675A06">Gross, Martin</mods:namePart>
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<mods:typeOfResource id="9620C746E8101E74C978901920C4CA4E">text</mods:typeOfResource>
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<mods:title id="A54BCC40DEB087577F422E47CDB002C7">Zootaxa</mods:title>
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<mods:part id="89278EE68A8DE528AD881BA629A2A5F1">
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<mods:date id="4D5CEC660D50A62F7776B2DEE05AF39C">2016</mods:date>
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<mods:detail id="E42444BCE40203EE149AAC6F4290FAF5" type="volume">
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<mods:number id="2DB6CB29F2220912D8531DE99DD96561">4179</mods:number>
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<mods:classification id="A46EE336A32246BAFBD545E52A51B90C">journal article</mods:classification>
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<mods:identifier id="F22553268D49720DE005B2F58C0A0DCB" type="DOI">10.11646/zootaxa.4179.2.6</mods:identifier>
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<treatment id="9605E23F9271FFC896D8F9A5FC13FAC3" ID-DOI="http://doi.org/10.5281/zenodo.6083811" ID-GBIF-Taxon="124560777" ID-Zenodo-Dep="6083811" LSID="urn:lsid:plazi:treatment:9605E23F9271FFC896D8F9A5FC13FAC3" httpUri="http://treatment.plazi.org/id/9605E23F9271FFC896D8F9A5FC13FAC3" lastPageId="5" lastPageNumber="268" pageId="3" pageNumber="266">
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<subSubSection id="56B600A29271FFCE96D8F9A5FE79F9B8" pageId="3" pageNumber="266" type="nomenclature">
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<paragraph id="1E1353299271FFCE96D8F9A5FE6CF9DF" blockId="3.[151,438,1544,1604]" box="[151,438,1544,1571]" pageId="3" pageNumber="266">
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<heading id="455BE4459271FFCE96D8F9A5FE6CF9DF" bold="true" box="[151,438,1544,1571]" fontSize="11" level="1" pageId="3" pageNumber="266" reason="1">
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<emphasis id="2CD88F3B9271FFCE96D8F9A5FE6CF9DF" bold="true" box="[151,438,1544,1571]" pageId="3" pageNumber="266">
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<taxonomicName id="D9AC28AA9271FFCE96D8F9A5FE88F9DE" ID-CoL="7bd60306-bd5d-4dbb-b5e1-a692dd82011c" box="[151,338,1544,1570]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="266" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
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<emphasis id="2CD88F3B9271FFCE96D8F9A5FE88F9DE" bold="true" box="[151,338,1544,1570]" italics="true" pageId="3" pageNumber="266">Iphiculus eliasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="37EB32409271FFCE9716F9A4FE6CF9DF" box="[345,438,1545,1571]" pageId="3" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="1E1353299271FFCE96D8F986FE79F9B8" blockId="3.[151,438,1544,1604]" box="[151,419,1579,1604]" pageId="3" pageNumber="266">
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(
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<figureCitation id="86974FAC9271FFCE96D0F986FF3FF9B8" box="[159,229,1579,1604]" captionStart="FIGURE 2" captionStartId="3.[151,249,1386,1408]" captionTargetBox="[151,1435,447,1361]" captionTargetId="figure@3.[151,1435,447,1361]" captionTargetPageId="3" captionText="FIGURE 2. Fossil and extant Iphiculus. A, Iphiculus eliasi sp. nov., holotype UMJGP 75.612 in dorsal view (dry and uncoated); B – D, Iphiculus convexus Ihle, 1918, ZRC 2009.0462 (male specimen from Vanuatu) in dorsal (B), posterior (C), and posterodorsal view (D). Photos in B – D by P. K. L. Ng." httpUri="https://zenodo.org/record/163904/files/figure.png" pageId="3" pageNumber="266">Figs 2</figureCitation>
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A, 3A–E, 4A–C)
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</paragraph>
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</subSubSection>
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<subSubSection id="56B600A29271FFCE96D8F9DEFAC9F928" pageId="3" pageNumber="266" type="diagnosis">
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<paragraph id="1E1353299271FFCE96D8F9DEFAC9F928" blockId="3.[151,1437,1651,1928]" pageId="3" pageNumber="266">
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<emphasis id="2CD88F3B9271FFCE96D8F9DEFED0F970" bold="true" box="[151,266,1651,1676]" pageId="3" pageNumber="266">Diagnosis</emphasis>
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. Carapace transversely subovate in outline, widest at level of posteriormost anterolateral spine; lateral margins bearing 6 short triangular spines; dorsal carapace surface covered evenly with large rounded tubercles: hepatic region with tubercle, gastric region with 3 pairs of tubercles, branchial region with 3 tubercles.
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</paragraph>
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</subSubSection>
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<subSubSection id="56B600A29271FFCE9688F972FBD3F904" box="[199,1033,1759,1784]" pageId="3" pageNumber="266" type="etymology">
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<paragraph id="1E1353299271FFCE9688F972FBD3F904" blockId="3.[151,1437,1651,1928]" box="[199,1033,1759,1784]" pageId="3" pageNumber="266">
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<emphasis id="2CD88F3B9271FFCE9688F972FE94F904" bold="true" box="[199,334,1759,1784]" pageId="3" pageNumber="266">Etymology.</emphasis>
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The species name is dedicated to Eliáš, son of the first author.
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</paragraph>
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</subSubSection>
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<subSubSection id="56B600A29271FFCE9688F8AEFE5AF874" pageId="3" pageNumber="266" type="materials_examined">
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<paragraph id="1E1353299271FFCE9688F8AEFE5AF874" blockId="3.[151,1437,1651,1928]" pageId="3" pageNumber="266">
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<emphasis id="2CD88F3B9271FFCE9688F8AEFE6BF8E0" bold="true" box="[199,433,1795,1820]" pageId="3" pageNumber="266">Material examined.</emphasis>
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Holotype: near-complete carapace UMJGP Inv.No. 75.612, max. length,
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<quantity id="D954FECC9271FFCE92BEF8AEFA83F8E7" box="[1265,1369,1795,1820]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="3" pageNumber="266" unit="mm" value="15.7">15.7 mm</quantity>
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; max. width, 19.0 mm (
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<figureCitation id="86974FAC9271FFCE9711F88AFE7CF8BC" box="[350,422,1831,1856]" captionStart="FIGURE 3" captionStartId="4.[151,249,1250,1272]" captionTargetBox="[151,1434,183,1228]" captionTargetId="figure@4.[151,1435,183,1228]" captionTargetPageId="4" captionText="FIGURE 3. Type material of Iphiculus eliasi sp. nov. A – D, holotype UMJGP 75.612 in dorsal (A), posterodorsal (B), frontal (C), and posterior view (D); E, paratype UMJGP 75.613 in dorsal view. Specimens were coated with ammonium chloride prior the photography. All specimens to scale." httpUri="https://zenodo.org/record/257320/files/figure.png" pageId="3" pageNumber="266">Figs 3</figureCitation>
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A–D). Paratype: near-complete carapace UMJGP Inv.No. 75.613, max. length, 13.0 mm (preserved portion); max. width,
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<quantity id="D954FECC9271FFCE9456F8E1FD5EF89F" box="[537,644,1868,1892]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="266" unit="mm" value="17.2">17.2 mm</quantity>
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;
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<figureCitation id="86974FAC9271FFCE94DCF8E6FD03F898" box="[659,729,1867,1892]" captionStart="FIGURE 3" captionStartId="4.[151,249,1250,1272]" captionTargetBox="[151,1434,183,1228]" captionTargetId="figure@4.[151,1435,183,1228]" captionTargetPageId="4" captionText="FIGURE 3. Type material of Iphiculus eliasi sp. nov. A – D, holotype UMJGP 75.612 in dorsal (A), posterodorsal (B), frontal (C), and posterior view (D); E, paratype UMJGP 75.613 in dorsal view. Specimens were coated with ammonium chloride prior the photography. All specimens to scale." httpUri="https://zenodo.org/record/257320/files/figure.png" pageId="3" pageNumber="266">Fig. 3</figureCitation>
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E). Additional specimen: carapace fragment UMJGP Inv.No. 211339;
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<figureCitation id="86974FAC9271FFCE96B6F8C2FEE1F874" box="[249,315,1903,1928]" captionStart="FIGURE 4" captionStartId="5.[151,249,645,667]" captionTargetBox="[151,1436,337,623]" captionTargetId="figure@5.[150,1436,337,623]" captionTargetPageId="5" captionText="FIGURE 4. Additional material of Iphiculus eliasi sp. nov. A – C, UMJGP 21.1339 with partially degraded cuticular surfaces (B – C). Specimen in A was coated with ammonium chloride prior the photography." httpUri="https://zenodo.org/record/257321/files/figure.png" pageId="3" pageNumber="266">Fig. 4</figureCitation>
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A–C).
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</paragraph>
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</subSubSection>
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<caption id="4AD303A19276FFC996D8FB4FFDEBFAC9" httpUri="https://zenodo.org/record/257320/files/figure.png" pageId="4" pageNumber="267" targetBox="[151,1434,183,1228]" targetPageId="4">
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<paragraph id="1E1353299276FFC996D8FB4FFDEBFAC9" blockId="4.[151,1436,1250,1333]" pageId="4" pageNumber="267">
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<emphasis id="2CD88F3B9276FFC996D8FB4FFEC9FB0B" bold="true" box="[151,275,1250,1272]" pageId="4" pageNumber="267">FIGURE 3.</emphasis>
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Type material of
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<taxonomicName id="D9AC28AA9276FFC99786FB4FFDBFFB0B" box="[457,613,1250,1271]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
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<emphasis id="2CD88F3B9276FFC99786FB4FFDBFFB0B" box="[457,613,1250,1271]" italics="true" pageId="4" pageNumber="267">Iphiculus eliasi</emphasis>
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</taxonomicName>
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<emphasis id="2CD88F3B9276FFC99423FB4FFD64FB0B" bold="true" box="[620,702,1250,1271]" pageId="4" pageNumber="267">
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<taxonomicNameLabel id="37EB32409276FFC99423FB4FFD64FB0B" box="[620,702,1250,1271]" pageId="4" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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A–D, holotype UMJGP 75.612 in dorsal (A), posterodorsal (B), frontal (C), and posterior view (D); E, paratype UMJGP 75.613 in dorsal view. Specimens were coated with ammonium chloride prior the photography. All specimens to scale.
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</paragraph>
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</caption>
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<subSubSection id="56B600A29276FFC99688FA2DFB4CF8B5" pageId="4" pageNumber="267" type="description">
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<paragraph id="1E1353299276FFC99688FA2DFEDDF8D9" blockId="4.[151,1437,1408,2009]" pageId="4" pageNumber="267">
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<emphasis id="2CD88F3B9276FFC99688FA2DFE83FA65" bold="true" box="[199,345,1408,1433]" pageId="4" pageNumber="267">Description.</emphasis>
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Small carapace; transversely subovate in outline; slightly wider than long, widest at midlength, at level of posteriormost anterolateral spine; dorsal surface moderately convex in both directions. Front not projected, slightly raised, narrow, not well preserved, presumably bilobed. Orbits small, concave, anteriorly directed. Lateral margins bearing 6 short triangular spines, anterolateral margin with 4 spines, posterolateral margin with 2 spines; corners between posterolateral, posterior margins pointed; posterior margin straight, narrow. Dorsal surface of carapace evenly covered with many large, densely packed granules, nearly identical in size (if cuticular surface preserved) or with round concave pustules (if cuticular surface not preserved). Carapace surface covered evenly with large rounded tubercles: hepatic region with 1 tubercle, gastric region with 3 pairs of tubercles, branchial region with 3 tubercles. Carapace grooves absent in anterior carapace portion, well developed in posterior carapace portion. Gastric region large, indistinctly demarcated with grooves. Cardiac region ovate in outline, strongly arched. Branchial regions broad. Intestinal region narrow without large spines. Sternum, pleon, pereiopods unknown.
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</paragraph>
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<paragraph id="1E1353299276FFC99688F89DFB4CF8B5" blockId="4.[151,1437,1408,2009]" box="[199,1174,1840,1865]" pageId="4" pageNumber="267">
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<materialsCitation id="AEC459749276FFC99688F89DFB4CF8B5" ID-GBIF-Occurrence="1324332092" box="[199,1174,1840,1865]" country="Austria" location="Wetzelsdorf" pageId="4" pageNumber="267" specimenCount="1" typeStatus="holotype">
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<emphasis id="2CD88F3B9276FFC99688F89DFE83F8B5" bold="true" box="[199,345,1840,1865]" pageId="4" pageNumber="267">Occurrence.</emphasis>
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The species is known only from its
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<typeStatus id="C117ED8B9276FFC994A0F89CFCC5F8B5" box="[751,799,1841,1865]" pageId="4" pageNumber="267">type</typeStatus>
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locality at
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<location id="1B7305F29276FFC995D3F89DFBF9F8B5" LSID="urn:lsid:plazi:treatment:9605E23F9271FFC896D8F9A5FC13FAC3:1B7305F29276FFC995D3F89DFBF9F8B5" box="[924,1059,1840,1865]" country="Austria" name="Wetzelsdorf" pageId="4" pageNumber="267">Wetzelsdorf</location>
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(
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<collectingCountry id="66BB13B99276FFC9927DF89DFB51F8B5" box="[1074,1163,1840,1865]" name="Austria" pageId="4" pageNumber="267">Austria</collectingCountry>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="56B600A29276FFC89688F8F9FC40FEE1" lastPageId="5" lastPageNumber="268" pageId="4" pageNumber="267" type="discussion">
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<paragraph id="1E1353299276FFC89688F8F9FC40FEE1" blockId="4.[151,1437,1408,2009]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="268" pageId="4" pageNumber="267">
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<emphasis id="2CD88F3B9276FFC99688F8F9FEEEF891" bold="true" box="[199,308,1876,1901]" pageId="4" pageNumber="267">Remarks</emphasis>
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.
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<taxonomicName id="D9AC28AA9276FFC9970DF8F9FE37F891" box="[322,493,1876,1901]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
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<emphasis id="2CD88F3B9276FFC9970DF8F9FE37F891" box="[322,493,1876,1901]" italics="true" pageId="4" pageNumber="267">Iphiculus eliasi</emphasis>
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</taxonomicName>
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<emphasis id="2CD88F3B9276FFC997BBF8F9FD94F891" bold="true" box="[500,590,1876,1901]" pageId="4" pageNumber="267">
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<taxonomicNameLabel id="37EB32409276FFC997BBF8F9FD94F891" box="[500,590,1876,1901]" pageId="4" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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is closest to the extant
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<taxonomicName id="D9AC28AA9276FFC9951DF8FBFC0BF891" box="[850,977,1878,1901]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="convexus">
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<emphasis id="2CD88F3B9276FFC9951DF8FBFC0BF891" box="[850,977,1878,1901]" italics="true" pageId="4" pageNumber="267">I. convexus</emphasis>
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</taxonomicName>
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in terms of the morphology of the dorsal carapace. The latter species, however, has a relatively wider carapace closer to the anterior half and possesses more large round tubercles on the dorsal carapace, especially on the hepatic and gastric regions (
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<figureCitation id="86974FAC9276FFC992C3F831FB14F849" box="[1164,1230,1948,1973]" captionStart="FIGURE 2" captionStartId="3.[151,249,1386,1408]" captionTargetBox="[151,1435,447,1361]" captionTargetId="figure@3.[151,1435,447,1361]" captionTargetPageId="3" captionText="FIGURE 2. Fossil and extant Iphiculus. A, Iphiculus eliasi sp. nov., holotype UMJGP 75.612 in dorsal view (dry and uncoated); B – D, Iphiculus convexus Ihle, 1918, ZRC 2009.0462 (male specimen from Vanuatu) in dorsal (B), posterior (C), and posterodorsal view (D). Photos in B – D by P. K. L. Ng." httpUri="https://zenodo.org/record/163904/files/figure.png" pageId="4" pageNumber="267">Fig. 2</figureCitation>
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A
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<emphasis id="2CD88F3B9276FFC992A6F833FAEBF849" box="[1257,1329,1950,1973]" italics="true" pageId="4" pageNumber="267">versus</emphasis>
|
||
<figureCitation id="86974FAC9276FFC99377F831FAA1F849" box="[1336,1403,1948,1973]" captionStart="FIGURE 2" captionStartId="3.[151,249,1386,1408]" captionTargetBox="[151,1435,447,1361]" captionTargetId="figure@3.[151,1435,447,1361]" captionTargetPageId="3" captionText="FIGURE 2. Fossil and extant Iphiculus. A, Iphiculus eliasi sp. nov., holotype UMJGP 75.612 in dorsal view (dry and uncoated); B – D, Iphiculus convexus Ihle, 1918, ZRC 2009.0462 (male specimen from Vanuatu) in dorsal (B), posterior (C), and posterodorsal view (D). Photos in B – D by P. K. L. Ng." httpUri="https://zenodo.org/record/163904/files/figure.png" pageId="4" pageNumber="267">Fig. 2</figureCitation>
|
||
B). Another extant species,
|
||
<taxonomicName id="D9AC28AA9276FFC997EDF86FFDEEF825" box="[418,564,1986,2009]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="spongiosus">
|
||
<emphasis id="2CD88F3B9276FFC997EDF86FFDEEF825" box="[418,564,1986,2009]" italics="true" pageId="4" pageNumber="267">I. spongiosus</emphasis>
|
||
</taxonomicName>
|
||
, differs from
|
||
<taxonomicName id="D9AC28AA9276FFC99481F86FFCFAF825" box="[718,800,1984,2009]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
|
||
<emphasis id="2CD88F3B9276FFC99481F86FFCFAF825" box="[718,800,1984,2009]" italics="true" pageId="4" pageNumber="267">I. eliasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2CD88F3B9276FFC99568F86DFC5AF825" bold="true" box="[807,896,1984,2009]" pageId="4" pageNumber="267">
|
||
<taxonomicNameLabel id="37EB32409276FFC99568F86DFC5AF825" box="[807,896,1984,2009]" pageId="4" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the presence of large anterolateral spines and the possession of dorsal carapace tubercles which are more projected outwards (Ng
|
||
<emphasis id="2CD88F3B9277FFC89209FF34FBA5FF4C" box="[1094,1151,151,176]" italics="true" pageId="5" pageNumber="268">et al.</emphasis>
|
||
2008: fig. 79). Similarly, all fossil species of
|
||
<taxonomicName id="D9AC28AA9277FFC8973DFF11FE00FF29" box="[370,474,188,213]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2CD88F3B9277FFC8973DFF11FE00FF29" box="[370,474,188,213]" italics="true" pageId="5" pageNumber="268">Iphiculus</emphasis>
|
||
</taxonomicName>
|
||
so far known, i.e.
|
||
<taxonomicName id="D9AC28AA9277FFC894E7FF10FB9CFF28" authority="Morris & Collins, 1991" authorityName="Morris & Collins" authorityYear="1991" box="[680,1094,188,213]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="2CD88F3B9277FFC894E7FF10FCEDFF29" box="[680,823,188,213]" italics="true" pageId="5" pageNumber="268">I. granulatus</emphasis>
|
||
<bibRefCitation id="7A3D2ED89277FFC89571FF11FB9CFF28" author="Morris" box="[830,1094,188,213]" pageId="5" pageNumber="268" refString="Morris, S. F. & Collins, J. S. H. (1991) Neogene crabs from Brunei, Sabah and Sarawak. Bulletin of the British Museum of Natural History (Geology), 47, 1 - 33." type="journal article" year="1991">Morris & Collins, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D9AC28AA9277FFC8921EFF10FF0BFF04" authority="Morris & Collins, 1991" authorityName="Morris & Collins" authorityYear="1991" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="miriensis">
|
||
<emphasis id="2CD88F3B9277FFC8921EFF10FB80FF28" box="[1105,1114,189,212]" italics="true" pageId="5" pageNumber="268">I</emphasis>
|
||
.
|
||
<emphasis id="2CD88F3B9277FFC89227FF10FB14FF28" box="[1128,1230,189,212]" italics="true" pageId="5" pageNumber="268">miriensis</emphasis>
|
||
<bibRefCitation id="7A3D2ED89277FFC8929AFF11FF0BFF04" author="Morris" pageId="5" pageNumber="268" refString="Morris, S. F. & Collins, J. S. H. (1991) Neogene crabs from Brunei, Sabah and Sarawak. Bulletin of the British Museum of Natural History (Geology), 47, 1 - 33." type="journal article" year="1991">Morris & Collins, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="D9AC28AA9277FFC89744FF4CFD6EFF04" authority="Morris & Collins, 1991" authorityName="Morris & Collins" authorityYear="1991" box="[267,692,223,248]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="sexspinosus">
|
||
<emphasis id="2CD88F3B9277FFC89744FF4CFE7CFF04" box="[267,422,225,248]" italics="true" pageId="5" pageNumber="268">I. sexspinosus</emphasis>
|
||
<bibRefCitation id="7A3D2ED89277FFC897E3FF72FD6EFF04" author="Morris" box="[428,692,223,248]" pageId="5" pageNumber="268" refString="Morris, S. F. & Collins, J. S. H. (1991) Neogene crabs from Brunei, Sabah and Sarawak. Bulletin of the British Museum of Natural History (Geology), 47, 1 - 33." type="journal article" year="1991">Morris & Collins, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
, have well-developed anterolateral spines (
|
||
<bibRefCitation id="7A3D2ED89277FFC892DBFF72FE2DFEE1" author="Morris" pageId="5" pageNumber="268" refString="Morris, S. F. & Collins, J. S. H. (1991) Neogene crabs from Brunei, Sabah and Sarawak. Bulletin of the British Museum of Natural History (Geology), 47, 1 - 33." type="journal article" year="1991">Morris & Collins 1991: figs 15, 14 and 17, respectively</bibRefCitation>
|
||
) which are absent in
|
||
<taxonomicName id="D9AC28AA9277FFC894A9FEA8FCE3FEE1" box="[742,825,260,285]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
|
||
<emphasis id="2CD88F3B9277FFC894A9FEA8FCE3FEE1" box="[742,825,260,285]" italics="true" pageId="5" pageNumber="268">I. eliasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2CD88F3B9277FFC8950EFEA9FC40FEE1" bold="true" box="[833,922,260,285]" pageId="5" pageNumber="268">
|
||
<taxonomicNameLabel id="37EB32409277FFC8950EFEA9FC40FEE1" box="[833,922,260,285]" pageId="5" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="4AD303A19277FFC896D8FD28FC04FD45" httpUri="https://zenodo.org/record/257321/files/figure.png" pageId="5" pageNumber="268" targetBox="[151,1436,337,623]" targetPageId="5">
|
||
<paragraph id="1E1353299277FFC896D8FD28FC04FD45" blockId="5.[151,1436,644,697]" pageId="5" pageNumber="268">
|
||
<emphasis id="2CD88F3B9277FFC896D8FD28FEC9FD66" bold="true" box="[151,275,645,667]" pageId="5" pageNumber="268">FIGURE 4.</emphasis>
|
||
Additional material of
|
||
<taxonomicName id="D9AC28AA9277FFC8944AFD28FD78FD66" box="[517,674,645,666]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
|
||
<emphasis id="2CD88F3B9277FFC8944AFD28FD78FD66" box="[517,674,645,666]" italics="true" pageId="5" pageNumber="268">Iphiculus eliasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2CD88F3B9277FFC894E6FD28FD21FD66" bold="true" box="[681,763,645,666]" pageId="5" pageNumber="268">
|
||
<taxonomicNameLabel id="37EB32409277FFC894E6FD28FD21FD66" box="[681,763,645,666]" pageId="5" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A–C, UMJGP 21.1339 with partially degraded cuticular surfaces (B–C). Specimen in A was coated with ammonium chloride prior the photography.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="56B600A29277FFC89688FD4BFE21FB77" pageId="5" pageNumber="268" type="reference_group">
|
||
<paragraph id="1E1353299277FFC89688FD4BFCEBFC73" blockId="5.[151,1437,742,1343]" pageId="5" pageNumber="268">
|
||
<taxonomicName id="D9AC28AA9277FFC89688FD4BFEAEFD03" ID-CoL="7bd60306-bd5d-4dbb-b5e1-a692dd82011c" box="[199,372,742,767]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
|
||
<emphasis id="2CD88F3B9277FFC89688FD4BFEAEFD03" box="[199,372,742,767]" italics="true" pageId="5" pageNumber="268">Iphiculus eliasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2CD88F3B9277FFC89731FD4BFE03FD03" bold="true" box="[382,473,742,767]" pageId="5" pageNumber="268">
|
||
<taxonomicNameLabel id="37EB32409277FFC89731FD4BFE03FD03" box="[382,473,742,767]" pageId="5" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also resembles some
|
||
<taxonomicName id="D9AC28AA9277FFC8949AFD4BFC9AFD03" ID-CoL="9F4M9" box="[725,832,742,767]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2CD88F3B9277FFC8949AFD4BFC9AFD03" box="[725,832,742,767]" italics="true" pageId="5" pageNumber="268">Typilobus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="D9AC28AA9277FFC895E3FD4BFA82FD03" ID-CoL="b754450a-0811-43ff-ba92-5cd55d44619f" authority="Karasawa, 1998" authorityName="Karasawa" authorityYear="1998" box="[940,1368,742,767]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="kishimotoi">
|
||
<emphasis id="2CD88F3B9277FFC895E3FD4BFB4CFD03" box="[940,1174,742,767]" italics="true" pageId="5" pageNumber="268">Typilobus kishimotoi</emphasis>
|
||
<bibRefCitation id="7A3D2ED89277FFC892D0FD4AFA82FD03" author="Karasawa" box="[1183,1368,743,767]" pageId="5" pageNumber="268" refString="Karasawa, H. (1998) Typilobus kishimotoi, a new leucosiid crab (Crustacea: Decapoda: Brachyura) from the Miocene Katsuta Group, Japan. Proceedings of the Biological Society of Washington, 111, 97 - 101." type="journal article" year="1998">Karasawa, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Miocene of
|
||
<collectingCountry id="66BB13B99277FFC89701FCA6FE54FCDE" box="[334,398,779,802]" name="Japan" pageId="5" pageNumber="268">Japan</collectingCountry>
|
||
has similarly shaped carapace outline and possesses large rounded tubercles on the dorsal surface; but the tubercles are fewer than in
|
||
<taxonomicName id="D9AC28AA9277FFC894C9FC82FD01FCBB" ID-CoL="7bd60306-bd5d-4dbb-b5e1-a692dd82011c" box="[646,731,814,839]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
|
||
<emphasis id="2CD88F3B9277FFC894C9FC82FD01FCBB" box="[646,731,814,839]" italics="true" pageId="5" pageNumber="268">I. eliasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2CD88F3B9277FFC894ABFC83FCE4FCBB" bold="true" box="[740,830,814,839]" pageId="5" pageNumber="268">
|
||
<taxonomicNameLabel id="37EB32409277FFC894ABFC83FCE4FCBB" box="[740,830,814,839]" pageId="5" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="86974FAC9277FFC89500FC83FC4BFCBB" box="[847,913,814,839]" captionStart="FIGURE 3" captionStartId="4.[151,249,1250,1272]" captionTargetBox="[151,1434,183,1228]" captionTargetId="figure@4.[151,1435,183,1228]" captionTargetPageId="4" captionText="FIGURE 3. Type material of Iphiculus eliasi sp. nov. A – D, holotype UMJGP 75.612 in dorsal (A), posterodorsal (B), frontal (C), and posterior view (D); E, paratype UMJGP 75.613 in dorsal view. Specimens were coated with ammonium chloride prior the photography. All specimens to scale." httpUri="https://zenodo.org/record/257320/files/figure.png" pageId="5" pageNumber="268">Fig. 3</figureCitation>
|
||
A vs.
|
||
<bibRefCitation id="7A3D2ED89277FFC89597FC82FB51FCBB" author="Karasawa" box="[984,1163,815,839]" pageId="5" pageNumber="268" refString="Karasawa, H. (1998) Typilobus kishimotoi, a new leucosiid crab (Crustacea: Decapoda: Brachyura) from the Miocene Katsuta Group, Japan. Proceedings of the Biological Society of Washington, 111, 97 - 101." type="journal article" year="1998">Karasawa 1998</bibRefCitation>
|
||
: fig. 2). Additionally,
|
||
<taxonomicName id="D9AC28AA9277FFC893C8FC82FED1FC97" ID-CoL="b754450a-0811-43ff-ba92-5cd55d44619f" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="kishimotoi">
|
||
<emphasis id="2CD88F3B9277FFC893C8FC82FED1FC97" italics="true" pageId="5" pageNumber="268">T. kishimotoi</emphasis>
|
||
</taxonomicName>
|
||
has two longitudinal ridges behind the median sulcus of the frontal region (
|
||
<bibRefCitation id="7A3D2ED89277FFC89213FCFFFAA6FC97" author="Karasawa" box="[1116,1404,850,875]" pageId="5" pageNumber="268" refString="Karasawa, H. (1998) Typilobus kishimotoi, a new leucosiid crab (Crustacea: Decapoda: Brachyura) from the Miocene Katsuta Group, Japan. Proceedings of the Biological Society of Washington, 111, 97 - 101." type="journal article" year="1998" yearSuffix="b">Karasawa 1998: fig. 2.2b</bibRefCitation>
|
||
), a feature that is absent in the new species (
|
||
<figureCitation id="86974FAC9277FFC8942EFCDBFD74FC73" box="[609,686,886,911]" captionStart="FIGURE 2" captionStartId="3.[151,249,1386,1408]" captionTargetBox="[151,1435,447,1361]" captionTargetId="figure@3.[151,1435,447,1361]" captionTargetPageId="3" captionText="FIGURE 2. Fossil and extant Iphiculus. A, Iphiculus eliasi sp. nov., holotype UMJGP 75.612 in dorsal view (dry and uncoated); B – D, Iphiculus convexus Ihle, 1918, ZRC 2009.0462 (male specimen from Vanuatu) in dorsal (B), posterior (C), and posterodorsal view (D). Photos in B – D by P. K. L. Ng." httpUri="https://zenodo.org/record/163904/files/figure.png" pageId="5" pageNumber="268">Figs. 2</figureCitation>
|
||
A, 3A, 3C).
|
||
</paragraph>
|
||
<paragraph id="1E1353299277FFC89688FC37FE21FB77" blockId="5.[151,1437,742,1343]" pageId="5" pageNumber="268">
|
||
One specimen of
|
||
<taxonomicName id="D9AC28AA9277FFC897D1FC37FD95FC4F" ID-CoL="7bd60306-bd5d-4dbb-b5e1-a692dd82011c" box="[414,591,922,947]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
|
||
<emphasis id="2CD88F3B9277FFC897D1FC37FD95FC4F" box="[414,591,922,947]" italics="true" pageId="5" pageNumber="268">Iphiculus eliasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2CD88F3B9277FFC89412FC37FD67FC4F" bold="true" box="[605,701,922,947]" pageId="5" pageNumber="268">
|
||
<taxonomicNameLabel id="37EB32409277FFC89412FC37FD67FC4F" box="[605,701,922,947]" pageId="5" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(UMJGP 21.1339) exhibits partial cuticle degradation on the preserved portion of the carapace (
|
||
<figureCitation id="86974FAC9277FFC89452FC13FDBBFC2B" box="[541,609,958,983]" captionStart="FIGURE 4" captionStartId="5.[151,249,645,667]" captionTargetBox="[151,1436,337,623]" captionTargetId="figure@5.[150,1436,337,623]" captionTargetPageId="5" captionText="FIGURE 4. Additional material of Iphiculus eliasi sp. nov. A – C, UMJGP 21.1339 with partially degraded cuticular surfaces (B – C). Specimen in A was coated with ammonium chloride prior the photography." httpUri="https://zenodo.org/record/257321/files/figure.png" pageId="5" pageNumber="268">Fig. 4</figureCitation>
|
||
A–C). It suggests rather complex internal structure of the tuberculation of leucosioids; tubercles are expressed differently in various cuticular layers. It was already noted that taphonomic aspects of the cuticle preservation in fossil crabs may have major impact on taxonomical evaluation of the characters such as carapace ornamentation (e.g.
|
||
<bibRefCitation id="7A3D2ED89277FFC8948FFB87FC32FBBE" author="Feldmann" box="[704,1000,1066,1091]" pageId="5" pageNumber="268" refString="Feldmann, R. M. & Portell, R. W. (2007) First report of Costacopluma Collins and Morris, 1975 (Decapoda: Brachyura: Retroplumidae) from the Eocene of Alabama, U. S. A. Journal of Crustacean Biology, 27, 90 - 96. http: // dx. doi. org / 10.1651 / S- 2712.1" type="journal article" year="2007">Feldmann & Portell 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7A3D2ED89277FFC895BAFB87FAC8FBBF" author="Klompmaker" box="[1013,1298,1066,1091]" pageId="5" pageNumber="268" refString="Klompmaker, A. A., Hyzny, M. & Jakobsen, S. L. (2015) Taphonomy of decapod crustacean cuticle and its effect on the appearance as exemplified by new and known taxa from the Cretaceous - Danian crab Caloxanthus. Cretaceous Research, 55, 141 - 151. http: // dx. doi. org / 10.1016 / j. cretres. 2014.11.011" type="journal article" year="2015">
|
||
Klompmaker
|
||
<emphasis id="2CD88F3B9277FFC892DDFB86FB16FBBF" box="[1170,1228,1066,1091]" italics="true" pageId="5" pageNumber="268">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). Therefore, only large rounded tubercles present also on the specimen without preserved cuticle (
|
||
<figureCitation id="86974FAC9277FFC8921DFBE3FB48FB9B" box="[1106,1170,1102,1127]" captionStart="FIGURE 3" captionStartId="4.[151,249,1250,1272]" captionTargetBox="[151,1434,183,1228]" captionTargetId="figure@4.[151,1435,183,1228]" captionTargetPageId="4" captionText="FIGURE 3. Type material of Iphiculus eliasi sp. nov. A – D, holotype UMJGP 75.612 in dorsal (A), posterodorsal (B), frontal (C), and posterior view (D); E, paratype UMJGP 75.613 in dorsal view. Specimens were coated with ammonium chloride prior the photography. All specimens to scale." httpUri="https://zenodo.org/record/257320/files/figure.png" pageId="5" pageNumber="268">Fig. 3</figureCitation>
|
||
A–D) are considered of taxonomical importance herein.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="56B600A29277FFC89688FB3BFC13FAC3" pageId="5" pageNumber="268" type="description">
|
||
<paragraph id="1E1353299277FFC89688FB3BFC13FAC3" blockId="5.[151,1437,742,1343]" pageId="5" pageNumber="268">
|
||
<materialsCitation id="AEC459749277FFC89688FB3BFC13FAC3" ID-GBIF-Occurrence="1324332091" country="Spain" location="Catalonia" pageId="5" pageNumber="268" specimenCount="1" stateProvince="Catalonia">
|
||
<bibRefCitation id="7A3D2ED89277FFC89688FB3BFE9EFB53" author="Via" box="[199,324,1174,1199]" pageId="5" pageNumber="268" refString="Via, L. (1941) Los cancrejos fosiles de Cataluna. Boletin del Instituto Geologico de Espana, 55, 55 - 128." type="journal article" year="1941">Vía (1941)</bibRefCitation>
|
||
reported an “
|
||
<taxonomicName id="D9AC28AA9277FFC897ABFB3BFDEAFB53" box="[484,560,1174,1199]" class="Malacostraca" family="Leucosiidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="subFamily" subFamily="Iliinae">Iliinae</taxonomicName>
|
||
,
|
||
<taxonomicName id="D9AC28AA9277FFC89471FB3BFD76FB53" box="[574,684,1174,1199]" class="Malacostraca" family="Leucosiidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="subFamily" subFamily="Ebaliinae">Ebaliinae</taxonomicName>
|
||
?” from the Middle Miocene of
|
||
<collectingRegion id="DC689DCB9277FFC8926FFB3BFB54FB53" box="[1056,1166,1174,1199]" country="Spain" name="Catalunya" pageId="5" pageNumber="268">Catalonia</collectingRegion>
|
||
,
|
||
<collectingCountry id="66BB13B99277FFC892D3FB3BFB05FB53" box="[1180,1247,1174,1199]" name="Spain" pageId="5" pageNumber="268">Spain</collectingCountry>
|
||
. His figure (
|
||
<bibRefCitation id="7A3D2ED89277FFC8933DFB3BFE5CFB2F" author="Via" pageId="5" pageNumber="268" refString="Via, L. (1941) Los cancrejos fosiles de Cataluna. Boletin del Instituto Geologico de Espana, 55, 55 - 128." type="journal article" year="1941">Vía 1941: pl. 10, fig. 75</bibRefCitation>
|
||
) clearly represents a leucosioid crab with striking similarities to
|
||
<taxonomicName id="D9AC28AA9277FFC892CCFB17FAE8FB2F" box="[1155,1330,1210,1235]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">
|
||
<emphasis id="2CD88F3B9277FFC892CCFB17FAE8FB2F" box="[1155,1330,1210,1235]" italics="true" pageId="5" pageNumber="268">Iphiculus eliasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2CD88F3B9277FFC89371FB17FA46FB2F" bold="true" box="[1342,1436,1210,1235]" pageId="5" pageNumber="268">
|
||
<taxonomicNameLabel id="37EB32409277FFC89371FB17FA46FB2F" box="[1342,1436,1210,1235]" pageId="5" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<bibRefCitation id="7A3D2ED89277FFC896D8FB73FE00FB0B" author="Muller" box="[151,474,1246,1271]" pageId="5" pageNumber="268" refString="Muller, P. (1993) Neogene decapod crustaceans from Catalonia. Scripta Musei Geologici Seminarii Barcinonensis, 225, 1 - 39." type="journal article" year="1993">Müller (1993: figs 5M–N)</bibRefCitation>
|
||
refigured the specimen in greater detail (although still insufficient for closer comparison) and assigned it questionably to the leucosiid genus
|
||
<taxonomicName id="D9AC28AA9277FFC89532FAAFFB7DFAE7" authority="Stimpson, 1857" authorityName="Stimpson" authorityYear="1857" box="[893,1191,1282,1307]" class="Malacostraca" family="Leucosiidae" genus="Randallia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2CD88F3B9277FFC89532FAAFFC31FAE7" box="[893,1003,1282,1307]" italics="true" pageId="5" pageNumber="268">Randallia</emphasis>
|
||
<bibRefCitation id="7A3D2ED89277FFC895BAFAAFFB7DFAE7" author="Stimpson" box="[1013,1191,1282,1307]" pageId="5" pageNumber="268" refString="Stimpson, W. (1857) Notices of new species of Crustacea of western North America; being an abstract from a paper to be published in the journal of the Society. Proceedings of the Boston Society of Natural History, 6, 84 - 89. http: // dx. doi. org / 10.5962 / bhl. part. 15468" type="journal article" year="1857">Stimpson, 1857</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="2CD88F3B9277FFC892FEFAAEFAFEFAE7" box="[1201,1316,1282,1307]" italics="true" pageId="5" pageNumber="268">sensu lato</emphasis>
|
||
(see
|
||
<bibRefCitation id="7A3D2ED89277FFC8932BFAAFFF0EFAC3" author="Galil" pageId="5" pageNumber="268" refString="Galil, B. S. (2003) Four new genera of leucosiid crabs (Crustacea: Brachyura: Leucosiidae) for three new species and nine species previously in the genus Randallia Stimpson, 1857, with a redescription of the type species, R. ornata (Randall, 1939). Proceedings of the Biological Society of Washington, 116, 395 - 422." type="journal article" year="2003">Galil 2003</bibRefCitation>
|
||
). Re-examination of the specimen is needed to resolve its affinities.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |