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<document id="7059B56327C696E9C50604219152619F" ID-DOI="10.5281/zenodo.5352924" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352924" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630209239027" checkinUser="felipe" docAuthor="Edmonds, W. D. &amp; Zidek, J." docDate="2010" docId="3A7F7B2C7F6A886BFF34FF72224AEF52" docLanguage="en" docName="0129.pdf" docOrigin="Insecta Mundi 2010 (129)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Coprophanaeus (Megaphanaeus) bonariensis" docType="treatment" docVersion="4" lastPageNumber="19" masterDocId="C64603547F7C887DFFF4FFB02050E849" masterDocTitle="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)" masterLastPageNumber="111" masterPageNumber="1" pageNumber="19" updateTime="1699186523840" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="F0372938EDE650317F8262D2A5580E5D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="02285DF74954D1202378BBF76A5EBD0F">
<mods:title id="4D2FF39A36C8059E93EADDB1B1D89B72">A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="F64D2E6BC5B1C6B7119E1C5176DD78EB" type="personal">
<mods:role id="8184DFDFDB34CE0465B06AF92364995B">
<mods:roleTerm id="8C8E5AD774EDD5A6A702611DA5F6E2FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A40C1E5F3BB3104AECFE194AA2C188F6">Edmonds, W. D.</mods:namePart>
</mods:name>
<mods:name id="1119567835E2531373334DB4BE38265C" type="personal">
<mods:role id="828C0E8810050149CAB4576EDA9D180B">
<mods:roleTerm id="E256F573C1D9082BB747BD492537C57A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7CB9DF844D01FF583FC2CFC8FB017343">Zidek, J.</mods:namePart>
<mods:affiliation id="DC58C69FF1E8D0964A38679800162C47">Coprophanaeus bonariensis (Gory)</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6351E5667EA61FF6D5C23C8640364887">text</mods:typeOfResource>
<mods:relatedItem id="B6DEAE4F4AF40B9A850B72DBD20C1B14" type="host">
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<mods:title id="22DE0B23E57BD0DAD3D8AC01FDD6EB69">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="6CE10F3115882BADEB82F968112EC7FF">
<mods:date id="B4B9EDD7060319C0C8987E2ECF21EFBA">2010</mods:date>
<mods:detail id="BB526569F007A2970E2C515F654F6301" type="pubDate">
<mods:number id="E7EC2D22FB455082CCD20278F0E96F06">2010-07-02</mods:number>
</mods:detail>
<mods:detail id="77D2FF135B455E02122E332064FAC120" type="volume">
<mods:number id="C9675E8C1706F5D5CF14C0EF75C9FD52">2010</mods:number>
</mods:detail>
<mods:detail id="9744CA39FF38553C3A89AF93C835DDA1" type="issue">
<mods:number id="156CFD4FD6AB4E398949FBE2D741320F">129</mods:number>
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<mods:extent id="67B71F9AD7AFDDB76AA640AF5D33736D" unit="page">
<mods:start id="D60B483C907A47F73CDC9E6E90EEFD5E">1</mods:start>
<mods:end id="7809836D87D373B15439A70633726E27">111</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CF2C9F9E3A960AB301AC274D3CB81D3F">journal article</mods:classification>
<mods:identifier id="EB9E62ADC9D73D023E8F881BF23B13AA" type="DOI">10.5281/zenodo.5352924</mods:identifier>
<mods:identifier id="452D6C2D20A9E1B104DF28ED54F4739C" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="6729F5E36FDE170C641C86791F9FC6EA" type="Zenodo-Dep">5352924</mods:identifier>
</mods:mods>
<treatment id="3A7F7B2C7F6A886BFF34FF72224AEF52" ID-DOI="http://doi.org/10.5281/zenodo.5450868" ID-Zenodo-Dep="5450868" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52" httpUri="http://treatment.plazi.org/id/3A7F7B2C7F6A886BFF34FF72224AEF52" lastPageNumber="19" pageId="22" pageNumber="19">
<subSubSection id="FACC99B17F6A886BFF34FF722396E892" box="[192,966,194,219]" pageId="22" pageNumber="19" type="nomenclature">
<paragraph id="B269CA3A7F6A886BFF34FF722396E892" blockId="22.[192,966,194,251]" box="[192,966,194,219]" pageId="22" pageNumber="19">
<heading id="E9217D567F6A886BFF34FF722396E892" bold="true" box="[192,966,194,219]" fontSize="10" level="2" pageId="22" pageNumber="19" reason="3">
<taxonomicName id="75D6B1B97F6A886BFF34FF722396E892" ID-CoL="Y6CB" authority="(Gory, 1844)" baseAuthorityName="Gory" baseAuthorityYear="1844" box="[192,966,194,219]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="species" species="bonariensis" subGenus="Megaphanaeus">
<emphasis id="80A216287F6A886BFF34FF722396E892" bold="true" box="[192,966,194,219]" pageId="22" pageNumber="19">
<emphasis id="80A216287F6A886BFF34FF7221C7E892" bold="true" box="[192,407,194,219]" italics="true" pageId="22" pageNumber="19">Coprophanaeus</emphasis>
(
<emphasis id="80A216287F6A886BFE51FF722223E892" bold="true" box="[421,627,194,219]" italics="true" pageId="22" pageNumber="19">Megaphanaeus</emphasis>
)
<emphasis id="80A216287F6A886BFD76FF722371E892" bold="true" box="[642,801,194,219]" italics="true" pageId="22" pageNumber="19">bonariensis</emphasis>
(
<bibRefCitation id="D647B7CB7F6A886BFCD9FF7223EEE892" author="Gory, H. L." box="[813,958,194,219]" pageId="22" pageNumber="19" pagination="79 - 80" refId="ref57874" refString="Gory, H. L. 1844. (Description of Phanaeus bonariensis). p. 79 - 80. In: F. E. Guerin-Meneville. Iconographie du Regne Animal de G. Cuvier, representation d'apres nature de l'une des especes les plus remarquables, et souvent non encore figurees, de chaque genre d'animaux. Insectes. Part 7. J. B. Bailliere; Paris. 576 p." type="book chapter" year="1844">Gory, 1844</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F6A886BFF34FF5321C2E8B2" box="[192,402,227,251]" pageId="22" pageNumber="19" type="description">
<paragraph id="B269CA3A7F6A886BFF34FF5321C2E8B2" blockId="22.[192,966,194,251]" box="[192,402,227,251]" pageId="22" pageNumber="19">
<figureCitation id="2AEDD6BF7F6A886BFF34FF532146E8B2" box="[192,278,227,251]" captionStart="Figure 13-27" captionStartId="15.[192,276,1691,1713]" captionTargetBox="[280,1357,196,1662]" captionTargetId="figure-18@15.[280,1357,195,1662]" captionTargetPageId="15" captionText="Figure 13-27. Characters of the Coprophanaeus subgenus Megaphanaeus. 13) Elytral surface of C. lancifer. 14) Same, C. ensifer. 15) Same, C. bonariensis. 16) Same, C. bellicosus. 17) Mesosternum and anterior portion of metasternum of C. lancifer (arrow indicates smooth area of metasternum). 18) Metasternum of C.lancifer. 19) Same, C. ensifer. 20) Dorsal view of posteromedian pronotal prominence of female C. ensifer (broad form). 21) Same (narrow form). 22) Partial lateral view of pronotum male C. ensifer (arrow indicates lateral spine). 23) Same, C. lancifer (arrow indicates vertical ridge). 24) Dorsal view of posteromedian pronotal prominence of male C. lancifer. 25) Same, female (broad form). 26) Same (narrow form). 27) Anterior view of cephalic process of female C. bellicosus." figureDoi="http://doi.org/10.5281/zenodo.5352934" httpUri="https://zenodo.org/record/5352934/files/figure.png" pageId="22" pageNumber="19">Fig. 15</figureCitation>
,
<figureCitation id="2AEDD6BF7F6A886BFED6FF532112E8B2" box="[290,322,227,251]" captionStart="Figure 28" captionStartId="16.[192,276,1892,1914]" captionTargetBox="[258,1375,196,1848]" captionTargetId="figure-17@16.[258,1378,193,1848]" captionTargetPageId="16" captionText="Figure 28. Approximate geographic distribution of Coprophanaeus (Megaphanaeus) species." figureDoi="http://doi.org/10.5281/zenodo.5352936" httpUri="https://zenodo.org/record/5352936/files/figure.png" pageId="22" pageNumber="19">28</figureCitation>
,
<figureCitation id="2AEDD6BF7F6A886BFEB9FF5321C2E8B2" box="[333,402,227,251]" captionStart="Figure 40-44" captionStartId="23.[192,276,1619,1641]" captionTargetBox="[198,1430,204,1588]" captionTargetId="figure-58@23.[190,1443,191,1601]" captionTargetPageId="23" captionText="Figure 40-44. Coprophanaeus bonariensis. 40-41) Female habitus. 42-43) Male habitus. 44) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352942" httpUri="https://zenodo.org/record/5352942/files/figure.png" pageId="22" pageNumber="19">40-44</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F6A886BFF34FE9323A0E9F2" pageId="22" pageNumber="19" type="reference_group">
<paragraph id="B269CA3A7F6A886BFF34FE9322D3E972" blockId="22.[192,1282,291,443]" box="[192,643,291,315]" pageId="22" pageNumber="19">
<treatmentCitationGroup id="92C6ED147F6A886BFF34FE9322D3E972" box="[192,643,291,315]" pageId="22" pageNumber="19">
<treatmentCitation id="3377EC2B7F6A886BFF34FE9322D3E972" author="Gory, H. L." box="[192,643,291,315]" page="79" pageId="22" pageNumber="19" year="1844">
<taxonomicName id="75D6B1B97F6A886BFF34FE9322D3E972" ID-CoL="4FSLQ" authority="Gory, 1844: 79" authorityName="Gory" authorityPageNumber="79" authorityYear="1844" box="[192,643,291,315]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="80A216287F6A886BFF34FE93219DE972" box="[192,461,291,315]" italics="true" pageId="22" pageNumber="19">Phanaeus bonariensis</emphasis>
<bibRefCitation id="D647B7CB7F6A886BFE26FE9322D3E972" author="Gory, H. L." box="[466,643,291,315]" pageId="22" pageNumber="19" pagination="79 - 80" refId="ref57874" refString="Gory, H. L. 1844. (Description of Phanaeus bonariensis). p. 79 - 80. In: F. E. Guerin-Meneville. Iconographie du Regne Animal de G. Cuvier, representation d'apres nature de l'une des especes les plus remarquables, et souvent non encore figurees, de chaque genre d'animaux. Insectes. Part 7. J. B. Bailliere; Paris. 576 p." type="book chapter" year="1844">Gory, 1844: 79</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="B269CA3A7F6A886BFF34FEF3245EE912" blockId="22.[192,1282,291,443]" box="[192,1038,323,347]" pageId="22" pageNumber="19">
<treatmentCitationGroup id="92C6ED147F6A886BFF34FEF3245EE912" box="[192,1038,323,347]" pageId="22" pageNumber="19">
<taxonomicName id="75D6B1B97F6A886BFF34FEF3220DE912" ID-CoL="Y6CB" authority="(Gory)" baseAuthorityName="Gory" baseAuthorityYear="1844" box="[192,605,323,347]" class="Insecta" family="Scarabaeidae" genus="Megaphanaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="80A216287F6A886BFF34FEF3225AE912" box="[192,522,323,347]" italics="true" pageId="22" pageNumber="19">Megaphanaeus bonariensis</emphasis>
(Gory)
</taxonomicName>
(recomb. by
<treatmentCitation id="3377EC2B7F6A886BFD07FEF32456E912" author="Blackwelder, R. E." box="[755,1030,323,347]" page="209" pageId="22" pageNumber="19" year="1944">
<bibRefCitation id="D647B7CB7F6A886BFD07FEF32456E912" author="Blackwelder, R. E." box="[755,1030,323,347]" pageId="22" pageNumber="19" pagination="189 - 341" refId="ref56650" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2. Bulletin of the United States National Museum 185: 189 - 341." type="journal article" year="1944">Blackwelder 1944: 209</bibRefCitation>
</treatmentCitation>
)
</treatmentCitationGroup>
</paragraph>
<paragraph id="B269CA3A7F6A886BFF34FED32426E932" blockId="22.[192,1282,291,443]" box="[192,1142,355,379]" pageId="22" pageNumber="19">
<treatmentCitationGroup id="92C6ED147F6A886BFF34FED32426E932" box="[192,1142,355,379]" pageId="22" pageNumber="19">
<treatmentCitation id="3377EC2B7F6A886BFF34FED322C1E932" author="Martinez, A." box="[192,657,355,379]" page="187" pageId="22" pageNumber="19" year="1944">
<taxonomicName id="75D6B1B97F6A886BFF34FED322C1E932" authority="Martinez, 1944: 187" authorityName="Martinez" authorityPageNumber="187" authorityYear="1944" box="[192,657,355,379]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="80A216287F6A886BFF34FED321C7E932" box="[192,407,355,379]" italics="true" pageId="22" pageNumber="19">Phanaeus vicinus</emphasis>
<bibRefCitation id="D647B7CB7F6A886BFE69FED322C1E932" author="Martinez, A." box="[413,657,355,379]" pageId="22" pageNumber="19" pagination="184 - 189" refId="ref58947" refString="Martinez, A. 1944. Insectos nuevos o poco conocidos. Revista de la Sociedad Entomologica Argentina 12: 184 - 189, plate 17." type="journal article" year="1944">Martínez, 1944: 187</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(syn. by
<treatmentCitation id="3377EC2B7F6A886BFD0EFED3243EE932" author="Martinez, A. &amp; F. S. Pereira" box="[762,1134,355,379]" page="61" pageId="22" pageNumber="19" year="1967">
<bibRefCitation id="D647B7CB7F6A886BFD0EFED3243EE932" author="Martinez, A. &amp; F. S. Pereira" box="[762,1134,355,379]" pageId="22" pageNumber="19" pagination="53 - 69" refId="ref59045" refString="Martinez, A., and F. S. Pereira. 1967. Notas escarabeidologicas III. (Col. Scarabaeidae - Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez and Pereira 1967: 61</bibRefCitation>
</treatmentCitation>
)
</treatmentCitationGroup>
</paragraph>
<paragraph id="B269CA3A7F6A886BFF34FE3323A0E9F2" blockId="22.[192,1282,291,443]" pageId="22" pageNumber="19">
<treatmentCitationGroup id="92C6ED147F6A886BFF34FE3323A0E9F2" pageId="22" pageNumber="19">
<treatmentCitation id="3377EC2B7F6A886BFF34FE33234CE9D2" author="Martinez, A." box="[192,796,387,411]" page="187" pageId="22" pageNumber="19" year="1944">
<taxonomicName id="75D6B1B97F6A886BFF34FE33234CE9D2" authority="Martinez, 1944: 187" authorityName="Martinez" authorityPageNumber="187" authorityYear="1944" box="[192,796,387,411]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="vicinus" subSpecies="argentinus">
<emphasis id="80A216287F6A886BFF34FE332272E9D2" box="[192,546,387,411]" italics="true" pageId="22" pageNumber="19">Phanaeus vicinus argentinus</emphasis>
<bibRefCitation id="D647B7CB7F6A886BFDDCFE33234CE9D2" author="Martinez, A." box="[552,796,387,411]" pageId="22" pageNumber="19" pagination="184 - 189" refId="ref58947" refString="Martinez, A. 1944. Insectos nuevos o poco conocidos. Revista de la Sociedad Entomologica Argentina 12: 184 - 189, plate 17." type="journal article" year="1944">Martínez, 1944: 187</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(syn. by
<treatmentCitation id="3377EC2B7F6A886BFC72FE3324A9E9D2" author="Martinez, A. &amp; F. S. Pereira" box="[902,1273,387,411]" page="61" pageId="22" pageNumber="19" year="1967">
<bibRefCitation id="D647B7CB7F6A886BFC72FE3324A9E9D2" author="Martinez, A. &amp; F. S. Pereira" box="[902,1273,387,411]" pageId="22" pageNumber="19" pagination="53 - 69" refId="ref59045" refString="Martinez, A., and F. S. Pereira. 1967. Notas escarabeidologicas III. (Col. Scarabaeidae - Scarabaeinae). Revista de la Sociedad Entomologica Argentina 29: 53 - 69." type="journal article" year="1967">Martínez and Pereira 1967: 61</bibRefCitation>
</treatmentCitation>
)
<taxonomicName id="75D6B1B97F6A886BFF34FE132234E9F2" ID-CoL="Y6CB" authority="(Gory)" baseAuthorityName="Gory" baseAuthorityYear="1844" box="[192,612,419,443]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="80A216287F6A886BFF34FE132241E9F2" box="[192,529,419,443]" italics="true" pageId="22" pageNumber="19">Coprophanaeus bonariensis</emphasis>
(Gory)
</taxonomicName>
(recomb. by
<treatmentCitation id="3377EC2B7F6A886BFD0EFE1323B8E9F2" author="Edmonds, W. D." box="[762,1000,419,443]" page="841" pageId="22" pageNumber="19" year="1972">
<bibRefCitation id="D647B7CB7F6A886BFD0EFE1323B8E9F2" author="Edmonds, W. D." box="[762,1000,419,443]" pageId="22" pageNumber="19" pagination="731 - 874" refId="ref56889" refString="Edmonds, W. D. 1972. Comparative skeletal morphology, systematic and evolution of the phanaeine dung beetles (Coleoptera: Scarabaeidae). The University of Kansas Science Bulletin 49: 731 - 874." type="journal article" year="1972">Edmonds 1972: 841</bibRefCitation>
</treatmentCitation>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F6A886BFF34FE5323C7EA72" pageId="22" pageNumber="19" type="materials_examined">
<paragraph id="B269CA3A7F6A886BFF34FE5323C7EA72" blockId="22.[192,1440,483,571]" pageId="22" pageNumber="19">
<emphasis id="80A216287F6A886BFF34FE53215DE9B2" bold="true" box="[192,269,483,507]" pageId="22" pageNumber="19">Type.</emphasis>
<taxonomicName id="75D6B1B97F6A886BFEE0FE542194E9B2" baseAuthorityName="Gory" baseAuthorityYear="1844" box="[276,452,483,507]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="80A216287F6A886BFEE0FE542194E9B2" box="[276,452,483,507]" italics="true" pageId="22" pageNumber="19">P. bonariensis</emphasis>
</taxonomicName>
– unknown to us;
<emphasis id="80A216287F6A886BFD5EFE54230CE9B2" box="[682,860,484,507]" italics="true" pageId="22" pageNumber="19">
<taxonomicName id="75D6B1B97F6A886BFD5EFE542374E9B2" authorityName="Martinez" authorityYear="1944" box="[682,804,484,507]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="species" species="vicinus">P. vicinus</taxonomicName>
s. st
</emphasis>
<taxonomicName id="75D6B1B97F6A886BFCA8FE5423B1E9B2" box="[860,993,483,507]" pageId="22" pageNumber="19" rank="subSpecies" subSpecies="holotype">
r.
<typeStatus id="6D6D74987F6A886BFC83FE5323B1E9B2" box="[887,993,483,507]" pageId="22" pageNumber="19" type="holotype">holotype</typeStatus>
</taxonomicName>
male, Museo Argentino de Ciencias Naturales “Bernadino Rivadavia,”
<collectingRegion id="701204D87F6A886BFD96FDB42356EA52" box="[610,774,515,539]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="22" pageNumber="19">Buenos Aires</collectingRegion>
;
<taxonomicName id="75D6B1B97F6A886BFCFBFDB4245EEA52" authorityName="Martinez" authorityYear="1944" box="[783,1038,516,539]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="vicinus" subSpecies="argentinus">
<emphasis id="80A216287F6A886BFCFBFDB4245EEA52" box="[783,1038,516,539]" italics="true" pageId="22" pageNumber="19">P. vicinus argentinus</emphasis>
</taxonomicName>
,
<typeStatus id="6D6D74987F6A886BFBEEFDB324D2EA52" box="[1050,1154,515,539]" pageId="22" pageNumber="19" type="holotype">holotype</typeStatus>
male, Museo Argentino de Ciencias Naturales “Bernadino Rivadavia,”
<collectingRegion id="701204D87F6A886BFD1BFD9423C2EA72" box="[751,914,547,571]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="22" pageNumber="19">Buenos Aires</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F6A886BFF34FDD3226DEB92" pageId="22" pageNumber="19" type="diagnosis">
<paragraph id="B269CA3A7F6A886BFF34FDD320A8EB72" blockId="22.[192,1440,611,987]" pageId="22" pageNumber="19">
<emphasis id="80A216287F6A886BFF34FDD3211FEA32" bold="true" box="[192,335,611,635]" pageId="22" pageNumber="19">Diagnosis.</emphasis>
<emphasis id="80A216287F6A886BFEA0FDD321E4EA32" box="[340,436,611,635]" italics="true" pageId="22" pageNumber="19">General</emphasis>
– Pronotum strongly granulate posteromedially. Posterolateral pronotal ridge (
<figureCitation id="2AEDD6BF7F6A886BFA84FDD320B1EAD2" captionStart="Figure 40-44" captionStartId="23.[192,276,1619,1641]" captionTargetBox="[198,1430,204,1588]" captionTargetId="figure-58@23.[190,1443,191,1601]" captionTargetPageId="23" captionText="Figure 40-44. Coprophanaeus bonariensis. 40-41) Female habitus. 42-43) Male habitus. 44) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352942" httpUri="https://zenodo.org/record/5352942/files/figure.png" pageId="22" pageNumber="19">Fig. 40</figureCitation>
, cf.
<figureCitation id="2AEDD6BF7F6A886BFEE6FD332139EAD2" box="[274,361,643,667]" captionStart="Figure 35-39" captionStartId="20.[192,277,1517,1539]" captionTargetBox="[241,1410,217,1492]" captionTargetId="figure-142@20.[235,1413,210,1500]" captionTargetPageId="20" captionText="Figure 35-39. Coprophanaeus lancifer. 35-36) Female habitus. 37-38) Male habitus (arrow in 37 indicates posterolateral pronotal ridge). 39) Aedeagus (lateral view on left; dorsal view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352940" httpUri="https://zenodo.org/record/5352940/files/figure.png" pageId="22" pageNumber="19">Fig. 37</figureCitation>
) short, not extending to lateral pronotal fossa. Metasternal shield completely coarsely punctured (as in
<figureCitation id="2AEDD6BF7F6A886BFE7AFD1321B4EAF2" box="[398,484,675,699]" captionStart="Figure 13-27" captionStartId="15.[192,276,1691,1713]" captionTargetBox="[280,1357,196,1662]" captionTargetId="figure-18@15.[280,1357,195,1662]" captionTargetPageId="15" captionText="Figure 13-27. Characters of the Coprophanaeus subgenus Megaphanaeus. 13) Elytral surface of C. lancifer. 14) Same, C. ensifer. 15) Same, C. bonariensis. 16) Same, C. bellicosus. 17) Mesosternum and anterior portion of metasternum of C. lancifer (arrow indicates smooth area of metasternum). 18) Metasternum of C.lancifer. 19) Same, C. ensifer. 20) Dorsal view of posteromedian pronotal prominence of female C. ensifer (broad form). 21) Same (narrow form). 22) Partial lateral view of pronotum male C. ensifer (arrow indicates lateral spine). 23) Same, C. lancifer (arrow indicates vertical ridge). 24) Dorsal view of posteromedian pronotal prominence of male C. lancifer. 25) Same, female (broad form). 26) Same (narrow form). 27) Anterior view of cephalic process of female C. bellicosus." figureDoi="http://doi.org/10.5281/zenodo.5352934" httpUri="https://zenodo.org/record/5352934/files/figure.png" pageId="22" pageNumber="19">Fig. 19</figureCitation>
); anterior portion of metasternum finely granulate laterally, smooth medially. Carinulate margins of striae 2-4 gently undulate such that width of stria at narrowings is at least threefourths of that at widest points (
<figureCitation id="2AEDD6BF7F6A886BFDA5FD5322F7EAB2" box="[593,679,739,763]" captionStart="Figure 13-27" captionStartId="15.[192,276,1691,1713]" captionTargetBox="[280,1357,196,1662]" captionTargetId="figure-18@15.[280,1357,195,1662]" captionTargetPageId="15" captionText="Figure 13-27. Characters of the Coprophanaeus subgenus Megaphanaeus. 13) Elytral surface of C. lancifer. 14) Same, C. ensifer. 15) Same, C. bonariensis. 16) Same, C. bellicosus. 17) Mesosternum and anterior portion of metasternum of C. lancifer (arrow indicates smooth area of metasternum). 18) Metasternum of C.lancifer. 19) Same, C. ensifer. 20) Dorsal view of posteromedian pronotal prominence of female C. ensifer (broad form). 21) Same (narrow form). 22) Partial lateral view of pronotum male C. ensifer (arrow indicates lateral spine). 23) Same, C. lancifer (arrow indicates vertical ridge). 24) Dorsal view of posteromedian pronotal prominence of male C. lancifer. 25) Same, female (broad form). 26) Same (narrow form). 27) Anterior view of cephalic process of female C. bellicosus." figureDoi="http://doi.org/10.5281/zenodo.5352934" httpUri="https://zenodo.org/record/5352934/files/figure.png" pageId="22" pageNumber="19">Fig. 15</figureCitation>
); carinulae simple, rarely tuberculate. Interstriae completely covered by large, round granules. Dorsal color usually shade of green or yellowish green. Length
<quantity id="752E67DF7F6A886BFAAFFCB320A4EB72" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.5" metricValueMin="2.1" pageId="22" pageNumber="19" unit="mm" value="28.0" valueMax="35.0" valueMin="21.0">21-35 mm</quantity>
.
</paragraph>
<paragraph id="B269CA3A7F6A886BFF04FCF32283EBD2" blockId="22.[192,1440,611,987]" pageId="22" pageNumber="19">
<emphasis id="80A216287F6A886BFF04FCF3217EEB12" box="[240,302,835,859]" italics="true" pageId="22" pageNumber="19">Male</emphasis>
(
<figureCitation id="2AEDD6BF7F6A886BFEC9FCF321EAEB12" box="[317,442,835,859]" captionStart="Figure 40-44" captionStartId="23.[192,276,1619,1641]" captionTargetBox="[198,1430,204,1588]" captionTargetId="figure-58@23.[190,1443,191,1601]" captionTargetPageId="23" captionText="Figure 40-44. Coprophanaeus bonariensis. 40-41) Female habitus. 42-43) Male habitus. 44) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352942" httpUri="https://zenodo.org/record/5352942/files/figure.png" pageId="22" pageNumber="19">Fig. 42-44</figureCitation>
) – Pronotal concavity with dorsally inclined spiniform process on each side below prominence (
<figureCitation id="2AEDD6BF7F6A886BFEAFFCD321FFEB32" box="[347,431,867,891]" captionStart="Figure 40-44" captionStartId="23.[192,276,1619,1641]" captionTargetBox="[198,1430,204,1588]" captionTargetId="figure-58@23.[190,1443,191,1601]" captionTargetPageId="23" captionText="Figure 40-44. Coprophanaeus bonariensis. 40-41) Female habitus. 42-43) Male habitus. 44) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352942" httpUri="https://zenodo.org/record/5352942/files/figure.png" pageId="22" pageNumber="19">Fig. 43</figureCitation>
). Dorsal prominence narrow, scoop-shaped, width of saddle less than length (
<figureCitation id="2AEDD6BF7F6A886BFACBFCD325C4EB32" box="[1343,1428,867,891]" captionStart="Figure 40-44" captionStartId="23.[192,276,1619,1641]" captionTargetBox="[198,1430,204,1588]" captionTargetId="figure-58@23.[190,1443,191,1601]" captionTargetPageId="23" captionText="Figure 40-44. Coprophanaeus bonariensis. 40-41) Female habitus. 42-43) Male habitus. 44) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352942" httpUri="https://zenodo.org/record/5352942/files/figure.png" pageId="22" pageNumber="19">Fig. 42</figureCitation>
). Apical process of parameres acute in profile.
</paragraph>
<paragraph id="B269CA3A7F6A886BFF04FC132490EBF2" blockId="22.[192,1440,611,987]" box="[240,1216,931,955]" pageId="22" pageNumber="19">
<emphasis id="80A216287F6A886BFF04FC13211AEBF2" box="[240,330,931,955]" italics="true" pageId="22" pageNumber="19">Female</emphasis>
(
<figureCitation id="2AEDD6BF7F6A886BFEACFC132183EBF2" box="[344,467,931,955]" captionStart="Figure 40-44" captionStartId="23.[192,276,1619,1641]" captionTargetBox="[198,1430,204,1588]" captionTargetId="figure-58@23.[190,1443,191,1601]" captionTargetPageId="23" captionText="Figure 40-44. Coprophanaeus bonariensis. 40-41) Female habitus. 42-43) Male habitus. 44) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352942" httpUri="https://zenodo.org/record/5352942/files/figure.png" pageId="22" pageNumber="19">Fig. 40-41</figureCitation>
) – Pronotal concavity lacking lateral spines. Protarsi lacking.
</paragraph>
<paragraph id="B269CA3A7F6A886BFF04FC74226DEB92" blockId="22.[192,1440,611,987]" box="[240,573,963,987]" pageId="22" pageNumber="19">
<emphasis id="80A216287F6A886BFF04FC7421A0EB92" box="[240,496,963,987]" italics="true" pageId="22" pageNumber="19">Specimens examined</emphasis>
– 222.
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F6A886BFF34FBB3213CEE12" pageId="22" pageNumber="19" type="distribution">
<paragraph id="B269CA3A7F6A886BFF34FBB3240CEC52" blockId="22.[192,1116,1027,1051]" box="[192,1116,1027,1051]" pageId="22" pageNumber="19">
<emphasis id="80A216287F6A886BFF34FBB32124EC52" bold="true" box="[192,372,1027,1051]" pageId="22" pageNumber="19">Distribution.</emphasis>
Cerrado and
<collectingRegion id="701204D87F6A886BFDE3FBB32233EC52" box="[535,611,1027,1051]" country="Argentina" name="Chaco" pageId="22" pageNumber="19">Chaco</collectingRegion>
provinces of Chacoan subregion (
<figureCitation id="2AEDD6BF7F6A886BFC0CFBB3241FEC52" box="[1016,1103,1027,1051]" captionStart="Figure 28" captionStartId="16.[192,276,1892,1914]" captionTargetBox="[258,1375,196,1848]" captionTargetId="figure-17@16.[258,1378,193,1848]" captionTargetPageId="16" captionText="Figure 28. Approximate geographic distribution of Coprophanaeus (Megaphanaeus) species." figureDoi="http://doi.org/10.5281/zenodo.5352936" httpUri="https://zenodo.org/record/5352936/files/figure.png" pageId="22" pageNumber="19">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="B269CA3A7F6A886BFF34FBF3213CEE12" blockId="22.[192,1440,1091,1627]" pageId="22" pageNumber="19">
<emphasis id="80A216287F6A886BFF34FBF32184EC12" bold="true" box="[192,468,1091,1115]" pageId="22" pageNumber="19">Collection Records.</emphasis>
<materialsCitation id="02BEC0677F6A886BFE2BFBF3246FEC12" box="[479,1087,1091,1115]" country="Argentina" location="Catamarca" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Catamarca">
<emphasis id="80A216287F6A886BFE2BFBF322C4EC12" bold="true" box="[479,660,1091,1115]" pageId="22" pageNumber="19">
<collectingCountry id="CAC18AAA7F6A886BFE2BFBF322C4EC12" box="[479,660,1091,1115]" name="Argentina" pageId="22" pageNumber="19">ARGENTINA</collectingCountry>
</emphasis>
:
<emphasis id="80A216287F6A886BFD5CFBF32311EC12" bold="true" box="[680,833,1091,1115]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFD5CFBF32311EC12" box="[680,833,1091,1115]" country="Argentina" name="Catamarca" pageId="22" pageNumber="19">Catamarca</collectingRegion>
</emphasis>
– Catamarca (Feb)
</materialsCitation>
.
<emphasis id="80A216287F6A886BFBBAFBF324F5EC12" bold="true" box="[1102,1189,1091,1115]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFBBAFBF324F5EC12" box="[1102,1189,1091,1115]" country="Argentina" name="Chaco" pageId="22" pageNumber="19">Chaco</collectingRegion>
</emphasis>
– Miraflores (Apr).
<emphasis id="80A216287F6A886BFF34FBD22167EC32" bold="true" box="[192,311,1122,1147]" pageId="22" pageNumber="19">Córdoba</emphasis>
– San Javier, Las Rosas (Jan); Quilino (Dec).
<emphasis id="80A216287F6A886BFCAAFBD323A3EC32" bold="true" box="[862,1011,1123,1147]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFCAAFBD323A3EC32" box="[862,1011,1123,1147]" country="Argentina" name="Corrientes" pageId="22" pageNumber="19">Corrientes</collectingRegion>
</emphasis>
– Bella Vista (Feb); Ituzaingo (Jan, Nov); Ituzaingo [Alto Paraná] (Sep).
<emphasis id="80A216287F6A886BFD71FB3322AFECD2" bold="true" box="[645,767,1155,1179]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFD71FB3322AFECD2" box="[645,767,1155,1179]" country="Argentina" name="Formosa" pageId="22" pageNumber="19">Formosa</collectingRegion>
</emphasis>
– Gran Guardia (Jan).
<materialsCitation id="02BEC0677F6A886BFBD4FB332529ECD2" box="[1056,1401,1155,1179]" country="Argentina" location="6 km W Yuto" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Jujuy">
<emphasis id="80A216287F6A886BFBD4FB332420ECD2" bold="true" box="[1056,1136,1155,1179]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFBD4FB332420ECD2" box="[1056,1136,1155,1179]" country="Argentina" name="Jujuy" pageId="22" pageNumber="19">Jujuy</collectingRegion>
</emphasis>
–
<location id="B7099CE17F6A886BFB79FB33257EECD2" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFB79FB33257EECD2" box="[1165,1326,1155,1179]" country="Argentina" name="6 km W Yuto" pageId="22" pageNumber="19" stateProvince="Jujuy">
<quantity id="752E67DF7F6A886BFB79FB33249BECD2" box="[1165,1227,1155,1179]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="19" unit="km" value="6.0">6 km</quantity>
W Yuto
</location>
(Feb)
</materialsCitation>
; El Quemado (Apr).
<emphasis id="80A216287F6A886BFE73FB132251ECF2" bold="true" box="[391,513,1187,1211]" pageId="22" pageNumber="19">Misiones</emphasis>
– Ayolas (Oct).
<materialsCitation id="02BEC0677F6A886BFD48FB132579ECF2" box="[700,1321,1187,1211]" country="Argentina" location="Termas de Rio Hondo" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Santiago del Estero">
<emphasis id="80A216287F6A886BFD48FB132395ECF2" bold="true" box="[700,965,1187,1211]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFD48FB132395ECF2" box="[700,965,1187,1211]" country="Argentina" name="Santiago del Estero" pageId="22" pageNumber="19">Santiago del Estero</collectingRegion>
</emphasis>
–
<location id="B7099CE17F6A886BFC2BFB1424B2ECF2" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFC2BFB1424B2ECF2" box="[991,1250,1187,1211]" country="Argentina" name="Termas de Rio Hondo" pageId="22" pageNumber="19" stateProvince="Santiago del Estero">Termas de Río Hondo</location>
(Dec)
</materialsCitation>
;
<materialsCitation id="02BEC0677F6A886BFAC7FB1321D9EC92" country="Argentina" location="Santiago del Estero" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Santiago del Estero">
<location id="B7099CE17F6A886BFAC7FB13216EEC92" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFAC7FB13216EEC92" country="Argentina" name="Santiago del Estero" pageId="22" pageNumber="19" stateProvince="Santiago del Estero">Santiago del Estero</location>
(Feb)
</materialsCitation>
;
<materialsCitation id="02BEC0677F6A886BFE61FB73224CEC92" box="[405,540,1219,1243]" country="Argentina" location="Rio Salado" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Santiago del Estero">
<location id="B7099CE17F6A886BFE61FB73224CEC92" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFE61FB73224CEC92" box="[405,540,1219,1243]" country="Argentina" name="Rio Salado" pageId="22" pageNumber="19" stateProvince="Santiago del Estero">Río Salado</location>
</materialsCitation>
; Campo Gallo (Mar).
<emphasis id="80A216287F6A886BFCD1FB7323F9EC92" bold="true" box="[805,937,1219,1243]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFCD1FB7323F9EC92" box="[805,937,1219,1243]" country="Argentina" name="Tucuman" pageId="22" pageNumber="19">Tucuman</collectingRegion>
</emphasis>
– Trancas (Jan); Benjamin Araoz (Mar); Gobernador Garmendia (Jan).
<materialsCitation id="02BEC0677F6A886BFDC3FB5323A0ECB2" box="[567,1008,1251,1275]" country="Bolivia" location="Beni" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Beni">
<emphasis id="80A216287F6A886BFDC3FB5322E6ECB2" bold="true" box="[567,694,1251,1275]" pageId="22" pageNumber="19">
<collectingCountry id="CAC18AAA7F6A886BFDC3FB5322E6ECB2" box="[567,694,1251,1275]" name="Bolivia" pageId="22" pageNumber="19">BOLIVIA</collectingCountry>
</emphasis>
:
<emphasis id="80A216287F6A886BFD37FB532353ECB2" bold="true" box="[707,771,1251,1275]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFD37FB532353ECB2" box="[707,771,1251,1275]" country="Bolivia" name="Beni" pageId="22" pageNumber="19">Beni</collectingRegion>
</emphasis>
– Magdalena (Feb)
</materialsCitation>
.
<materialsCitation id="02BEC0677F6A886BFC0EFB532173ED52" collectorName="San Matias" country="Bolivia" latitude="-16.566668" location="Santa Cruz" longLatPrecision="1282" longitude="-59.15" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Santa Cruz">
<emphasis id="80A216287F6A886BFC0EFB5324C3ECB2" bold="true" box="[1018,1171,1251,1275]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFC0EFB5324C3ECB2" box="[1018,1171,1251,1275]" country="Bolivia" name="Santa Cruz" pageId="22" pageNumber="19">Santa Cruz</collectingRegion>
</emphasis>
–
<collectorName id="1F23AFEC7F6A886BFB59FB53256CECB2" box="[1197,1340,1251,1275]" pageId="22" pageNumber="19">San Matias</collectorName>
,
<geoCoordinate id="D7E2ACFD7F6A886BFAB2FB5325CFECB2" box="[1350,1439,1249,1275]" degrees="16" direction="south" minutes="34" orientation="latitude" pageId="22" pageNumber="19" precision="925" value="-16.566668">
16
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34’S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F6A886BFF34FAB32173ED52" box="[192,291,1281,1307]" degrees="59" direction="west" minutes="09" orientation="longitude" pageId="22" pageNumber="19" precision="925" value="-59.15">
59
<superScript id="45A367727F6A886BFF29FAB120B5ED46" attach="none" box="[221,229,1281,1295]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
09’W
</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="02BEC0677F6A886BFED9FAB32214ED52" box="[301,580,1283,1307]" collectorName="de la Sierra" country="Bolivia" location="Santa Cruz" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Santa Cruz">
<collectingRegion id="701204D87F6A886BFED9FAB321E4ED52" box="[301,436,1283,1307]" inferredBySuffix="true" pageId="22" pageNumber="19">Santa Cruz</collectingRegion>
<collectorName id="1F23AFEC7F6A886BFE4DFAB32214ED52" box="[441,580,1283,1307]" pageId="22" pageNumber="19">de la Sierra</collectorName>
</materialsCitation>
;
<materialsCitation id="02BEC0677F6A886BFDB9FAB322B9ED52" box="[589,745,1283,1307]" country="Bolivia" elevation="400" location="Ichila" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Santa Cruz">
<location id="B7099CE17F6A886BFDB9FAB322C4ED52" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFDB9FAB322C4ED52" box="[589,660,1283,1307]" country="Bolivia" name="Ichila" pageId="22" pageNumber="19" stateProvince="Santa Cruz">Ichila</location>
,
<quantity id="752E67DF7F6A886BFD6AFAB322B9ED52" box="[670,745,1283,1307]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="19" unit="m" value="400.0">
<elevation id="39FB2D097F6A886BFD6AFAB322B9ED52" box="[670,745,1283,1307]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="19" unit="m" value="400.0">400 m</elevation>
</quantity>
</materialsCitation>
; Charagua,
<geoCoordinate id="D7E2ACFD7F6A886BFC82FAB323A5ED52" box="[886,1013,1281,1307]" degrees="20" direction="south" minutes="14" orientation="latitude" pageId="22" pageNumber="19" precision="15" seconds="11" value="-20.23639">
20
<superScript id="45A367727F6A886BFC67FAB123CBED46" attach="none" box="[915,923,1281,1295]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
14’11”S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F6A886BFC0DFAB324D4ED52" box="[1017,1156,1281,1307]" degrees="63" direction="west" minutes="13" orientation="longitude" pageId="22" pageNumber="19" precision="15" seconds="52" value="-63.231113">
63
<superScript id="45A367727F6A886BFBE2FAB1244DED46" attach="none" box="[1046,1053,1281,1295]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
13’52”W
</geoCoordinate>
,
<quantity id="752E67DF7F6A886BFB79FAB32486ED52" box="[1165,1238,1283,1307]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="22" pageNumber="19" unit="m" value="960.0">960 m</quantity>
(Apr); Urubigua,
<geoCoordinate id="D7E2ACFD7F6A886BFF34FA932110ED72" box="[192,320,1313,1339]" degrees="20" direction="south" minutes="14" orientation="latitude" pageId="22" pageNumber="19" precision="15" seconds="11" value="-20.23639">
20
<superScript id="45A367727F6A886BFF29FA9120B5ED66" attach="none" box="[221,229,1313,1327]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
14’11”S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F6A886BFEB2FA932182ED72" box="[326,466,1313,1339]" degrees="63" direction="west" minutes="13" orientation="longitude" pageId="22" pageNumber="19" precision="15" seconds="52" value="-63.231113">
63
<superScript id="45A367727F6A886BFE97FA91213BED66" attach="none" box="[355,363,1313,1327]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
13’52”W
</geoCoordinate>
; Buena Vista (Jan); La Peña (Nov); Cabezas (Jan);
<materialsCitation id="02BEC0677F6A886BFBC5FA932137ED12" country="Bolivia" latitude="-18.333334" location="Santiago de Chiquitos" longLatPrecision="1275" longitude="-59.6" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Santa Cruz">
<location id="B7099CE17F6A886BFBC5FA93256DED72" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFBC5FA93256DED72" box="[1073,1341,1315,1339]" country="Bolivia" latitude="-18.333334" longLatPrecision="1275" longitude="-59.6" name="Santiago de Chiquitos" pageId="22" pageNumber="19" stateProvince="Santa Cruz">Santiago de Chiquitos</location>
,
<geoCoordinate id="D7E2ACFD7F6A886BFAB3FA9325F0ED72" box="[1351,1440,1313,1339]" degrees="18" direction="south" minutes="20" orientation="latitude" pageId="22" pageNumber="19" precision="925" value="-18.333334">
18
<superScript id="45A367727F6A886BFA91FA91253DED66" attach="none" box="[1381,1389,1313,1327]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
20’S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F6A886BFF34FAF32171ED12" box="[192,289,1345,1371]" degrees="59" direction="west" minutes="36" orientation="longitude" pageId="22" pageNumber="19" precision="925" value="-59.6">
59
<superScript id="45A367727F6A886BFF29FAF120B5ED06" attach="none" box="[221,229,1345,1359]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
36’W
</geoCoordinate>
(Feb)
</materialsCitation>
; Santa Elena El Pantanal,
<geoCoordinate id="D7E2ACFD7F6A886BFD51FAF32374ED12" box="[677,804,1345,1371]" degrees="17" direction="south" minutes="57" orientation="latitude" pageId="22" pageNumber="19" precision="15" seconds="53" value="-17.964724">
17
<superScript id="45A367727F6A886BFD36FAF1229AED06" attach="none" box="[706,714,1345,1359]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
57’53”S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F6A886BFCDDFAF323E1ED12" box="[809,945,1345,1371]" degrees="59" direction="west" minutes="01" orientation="longitude" pageId="22" pageNumber="19" precision="15" seconds="33" value="-59.025833">
59
<superScript id="45A367727F6A886BFCB2FAF1231EED06" attach="none" box="[838,846,1345,1359]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
01’33”W
</geoCoordinate>
(Apr);
<materialsCitation id="02BEC0677F6A886BFBF5FAF322C1ED32" country="Bolivia" elevation="622" latitude="-18.33505" location="1.6 km ESE Santiago de Chiquitos" longLatPrecision="1" longitude="-59.58345" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Santa Cruz">
<location id="B7099CE17F6A886BFBF5FAF325CCED12" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFBF5FAF325CCED12" box="[1025,1436,1347,1371]" country="Bolivia" latitude="-18.33505" longLatPrecision="1" longitude="-59.58345" name="1.6 km ESE Santiago de Chiquitos" pageId="22" pageNumber="19" stateProvince="Santa Cruz">
<quantity id="752E67DF7F6A886BFBF5FAF32401ED12" box="[1025,1105,1347,1371]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="19" unit="km" value="1.6">1.6 km</quantity>
ESE Santiago de Chiquitos
</location>
,
<geoCoordinate id="D7E2ACFD7F6A886BFF34FAD3211FED32" box="[192,335,1377,1403]" degrees="18" direction="south" minutes="20.103" orientation="latitude" pageId="22" pageNumber="19" precision="1" value="-18.33505">
18
<superScript id="45A367727F6A886BFF2AFAD120B6ED26" attach="none" box="[222,230,1377,1391]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
20.103’S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F6A886BFEA0FAD321BFED32" box="[340,495,1377,1403]" degrees="59" direction="west" minutes="35.007" orientation="longitude" pageId="22" pageNumber="19" precision="1" value="-59.58345">
59
<superScript id="45A367727F6A886BFE86FAD1212AED26" attach="none" box="[370,378,1377,1391]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
35.007’W
</geoCoordinate>
,
<quantity id="752E67DF7F6A886BFE0EFAD32215ED32" box="[506,581,1379,1403]" metricMagnitude="2" metricUnit="m" metricValue="6.22" pageId="22" pageNumber="19" unit="m" value="622.0">
<elevation id="39FB2D097F6A886BFE0EFAD32215ED32" box="[506,581,1379,1403]" metricMagnitude="2" metricUnit="m" metricValue="6.22" pageId="22" pageNumber="19" unit="m" value="622.0">622 m</elevation>
</quantity>
(Nov)
</materialsCitation>
;
<materialsCitation id="02BEC0677F6A886BFD68FAD321A7EDD2" country="Bolivia" elevation="215" latitude="-18.243166" location="Tucavaca River" longLatPrecision="1" longitude="-59.46512" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Santa Cruz">
<locationDeviation id="A3DBD3187F6A886BFD68FAD3247DED32" box="[668,1069,1379,1403]" location="Santiago de Chiquitos" pageId="22" pageNumber="19">
<quantity id="752E67DF7F6A886BFD68FAD322B8ED32" box="[668,744,1379,1403]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="19" unit="km" value="20.0">20 km</quantity>
NE Santiago de Chiquitos
</locationDeviation>
(
<location id="B7099CE17F6A886BFBCFFAD324AEED32" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFBCFFAD324AEED32" box="[1083,1278,1379,1403]" country="Bolivia" latitude="-18.243166" longLatPrecision="1" longitude="-59.46512" name="Tucavaca River" pageId="22" pageNumber="19" stateProvince="Santa Cruz">Tucavaca River</location>
),
<geoCoordinate id="D7E2ACFD7F6A886BFAE6FAD325F0ED32" box="[1298,1440,1377,1403]" degrees="18" direction="south" minutes="14.590" orientation="latitude" pageId="22" pageNumber="19" precision="1" value="-18.243166">
18
<superScript id="45A367727F6A886BFADBFAD12567ED26" attach="none" box="[1327,1335,1377,1391]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
14.590’S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F6A886BFF34FA33210AEDD2" box="[192,346,1409,1435]" degrees="59" direction="west" minutes="27.907" orientation="longitude" pageId="22" pageNumber="19" precision="1" value="-59.46512">
59
<superScript id="45A367727F6A886BFF2AFA3120B6EDC6" attach="right" box="[222,230,1409,1423]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
27.907’W
</geoCoordinate>
,
<quantity id="752E67DF7F6A886BFE90FA3321FEEDD2" box="[356,430,1411,1435]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="22" pageNumber="19" unit="m" value="215.0">
<elevation id="39FB2D097F6A886BFE90FA3321FEEDD2" box="[356,430,1411,1435]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="22" pageNumber="19" unit="m" value="215.0">215 m</elevation>
</quantity>
(Nov)
</materialsCitation>
.
<materialsCitation id="02BEC0677F6A886BFDFEFA3325CCEDD2" box="[522,1436,1409,1435]" country="Bolivia" elevation="490" latitude="-21.838612" location="between Caiza and Creveaux" longLatPrecision="21" longitude="-63.41611" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Tarija">
<emphasis id="80A216287F6A886BFDFEFA33220FEDD2" bold="true" box="[522,607,1411,1435]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFDFEFA33220FEDD2" box="[522,607,1411,1435]" country="Bolivia" name="Tarija" pageId="22" pageNumber="19">Tarija</collectingRegion>
</emphasis>
–
<location id="B7099CE17F6A886BFD8DFA33238FEDD2" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFD8DFA33238FEDD2" box="[633,991,1411,1435]" country="Bolivia" latitude="-21.838612" longLatPrecision="21" longitude="-63.41611" name="between Caiza and Creveaux" pageId="22" pageNumber="19" stateProvince="Tarija">between Caiza and Creveaux</location>
,
<geoCoordinate id="D7E2ACFD7F6A886BFC1DFA33243BEDD2" box="[1001,1131,1409,1435]" degrees="21" direction="south" minutes="50" orientation="latitude" pageId="22" pageNumber="19" precision="15" seconds="19" value="-21.838612">
21
<superScript id="45A367727F6A886BFBF3FA31245FEDC6" attach="none" box="[1031,1039,1409,1423]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
50’19”S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F6A886BFB85FA3324AFEDD2" box="[1137,1279,1409,1435]" degrees="63" direction="west" minutes="24" orientation="longitude" pageId="22" pageNumber="19" precision="15" seconds="58" value="-63.41611">
63
<superScript id="45A367727F6A886BFB7AFA3124C6EDC6" attach="none" box="[1166,1174,1409,1423]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
24’58”W
</geoCoordinate>
,
<quantity id="752E67DF7F6A886BFAFDFA332503EDD2" box="[1289,1363,1411,1435]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="22" pageNumber="19" unit="m" value="490.0">
<elevation id="39FB2D097F6A886BFAFDFA332503EDD2" box="[1289,1363,1411,1435]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="22" pageNumber="19" unit="m" value="490.0">490 m</elevation>
</quantity>
(Jan)
</materialsCitation>
.
<materialsCitation id="02BEC0677F6A886BFF34FA1323B2EDF2" box="[192,994,1443,1467]" country="Brazil" location="5 km S Cuiaba" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Mato Grosso">
<emphasis id="80A216287F6A886BFF34FA132160EDF2" bold="true" box="[192,304,1443,1467]" pageId="22" pageNumber="19">
<collectingCountry id="CAC18AAA7F6A886BFF34FA132160EDF2" box="[192,304,1443,1467]" name="Brazil" pageId="22" pageNumber="19">BRAZIL</collectingCountry>
</emphasis>
:
<emphasis id="80A216287F6A886BFECBFA1321BEEDF2" bold="true" box="[319,494,1443,1467]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFECBFA1321BEEDF2" box="[319,494,1443,1467]" country="Brazil" name="Mato Grosso" pageId="22" pageNumber="19">Mato Grosso</collectingRegion>
</emphasis>
–
<location id="B7099CE17F6A886BFDFFFA132291EDF2" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFDFFFA132291EDF2" box="[523,705,1443,1467]" country="Brazil" name="5 km S Cuiaba" pageId="22" pageNumber="19" stateProvince="Mato Grosso">
<quantity id="752E67DF7F6A886BFDFFFA132219EDF2" box="[523,585,1443,1467]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="19" unit="km" value="5.0">5 km</quantity>
S Cuiabá
</location>
[Várzea Grande] (Dec)
</materialsCitation>
; Rosario Oeste (Feb).
<materialsCitation id="02BEC0677F6A886BFB05FA1323A4ED92" country="Paraguay" location="Estacion La Patria" pageId="22" pageNumber="19" specimenCount="1" stateProvince="Boqueron">
<emphasis id="80A216287F6A886BFB05FA1325C8EDF2" bold="true" box="[1265,1432,1443,1467]" pageId="22" pageNumber="19">
<collectingCountry id="CAC18AAA7F6A886BFB05FA1325C8EDF2" box="[1265,1432,1443,1467]" name="Paraguay" pageId="22" pageNumber="19">PARAGUAY</collectingCountry>
</emphasis>
:
<emphasis id="80A216287F6A886BFF34FA722119ED92" bold="true" box="[192,329,1474,1499]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFF34FA722119ED92" box="[192,329,1474,1499]" country="Paraguay" name="Boqueron" pageId="22" pageNumber="19">Boquerón</collectingRegion>
</emphasis>
–
<location id="B7099CE17F6A886BFE90FA73221BED92" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F6A886BFF34FF72224AEF52:B7099CE17F6A886BFE90FA73221BED92" box="[356,587,1475,1499]" country="Paraguay" name="Estacion La Patria" pageId="22" pageNumber="19" stateProvince="Boqueron">Estación La Patria</location>
[trans-Chaco highway] (Mar, Dec)
</materialsCitation>
; Nueva Asunción,
<geoCoordinate id="D7E2ACFD7F6A886BFB22FA73257FED92" box="[1238,1327,1473,1499]" degrees="20" direction="south" minutes="48" orientation="latitude" pageId="22" pageNumber="19" precision="925" value="-20.8">
20
<superScript id="45A367727F6A886BFB07FA7124ABED86" attach="none" box="[1267,1275,1473,1487]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
48’S
</geoCoordinate>
<geoCoordinate id="D7E2ACFD7F6A886BFAC1FA7325CBED92" box="[1333,1435,1473,1499]" degrees="61" direction="west" minutes="55" orientation="longitude" pageId="22" pageNumber="19" precision="925" value="-61.916668">
61
<superScript id="45A367727F6A886BFAA7FA71250BED86" attach="none" box="[1363,1371,1473,1487]" fontSize="6" pageId="22" pageNumber="19">o</superScript>
55’W
</geoCoordinate>
; Mariscal Estigarribia (Feb, May).
<emphasis id="80A216287F6A886BFDABFA532299EDB2" bold="true" box="[607,713,1507,1531]" pageId="22" pageNumber="19">Central</emphasis>
– Aregua (Dec); Altos; Colonia Nueva Italia (Feb); Asunción (Nov); Puerto Villeta (Jan).
<emphasis id="80A216287F6A886BFDF8F9B222FEEE52" bold="true" box="[524,686,1538,1563]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFDF8F9B222FEEE52" box="[524,686,1538,1563]" country="Paraguay" name="Concepcion" pageId="22" pageNumber="19">Concepción</collectingRegion>
</emphasis>
– Horqueta (Nov).
<emphasis id="80A216287F6A886BFC64F9B3244EEE52" bold="true" box="[912,1054,1539,1563]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFC64F9B3244EEE52" box="[912,1054,1539,1563]" country="Paraguay" name="Cordillera" pageId="22" pageNumber="19">Cordillera</collectingRegion>
</emphasis>
– Altos (Apr); Compañia Narajo (Nov).
<emphasis id="80A216287F6A886BFEFBF99321D8EE72" bold="true" box="[271,392,1571,1595]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFEFBF99321D8EE72" box="[271,392,1571,1595]" country="Paraguay" name="Misiones" pageId="22" pageNumber="19">Misiones</collectingRegion>
</emphasis>
– Ayolas (Oct);
<emphasis id="80A216287F6A886BFDB6F993229EEE72" bold="true" box="[578,718,1571,1595]" pageId="22" pageNumber="19">
<collectingRegion id="701204D87F6A886BFDB6F993229EEE72" box="[578,718,1571,1595]" country="Paraguay" name="Paraguari" pageId="22" pageNumber="19">Paraguari</collectingRegion>
</emphasis>
– Sapucay (Jan- Apr, Oct-Dec).
<emphasis id="80A216287F6A886BFBBEF9932579EE72" bold="true" box="[1098,1321,1571,1595]" pageId="22" pageNumber="19">Presidente Hays</emphasis>
– Laguna Capitán (Jan).
</paragraph>
</subSubSection>
<subSubSection id="FACC99B17F6A886BFF34F933224AEF52" pageId="22" pageNumber="19" type="discussion">
<paragraph id="B269CA3A7F6A886BFF34F9332430EEF2" blockId="22.[192,1440,1667,1819]" pageId="22" pageNumber="19">
<emphasis id="80A216287F6A886BFF34F933210CEED2" bold="true" box="[192,348,1667,1691]" pageId="22" pageNumber="19">Comments.</emphasis>
This species largely replaces
<taxonomicName id="75D6B1B97F6A886BFD30F9342367EED2" baseAuthorityName="Germar" baseAuthorityYear="1821" box="[708,823,1667,1691]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="species" species="ensifer">
<emphasis id="80A216287F6A886BFD30F9342367EED2" box="[708,823,1667,1691]" italics="true" pageId="22" pageNumber="19">C. ensifer</emphasis>
</taxonomicName>
in the chiquitano, cerrado and chacoan habitats of eastern
<collectingCountry id="CAC18AAA7F6A886BFED5F9132128EEF2" box="[289,376,1699,1723]" name="Bolivia" pageId="22" pageNumber="19">Bolivia</collectingCountry>
and surrounding portions of
<collectingCountry id="CAC18AAA7F6A886BFD20F9132303EEF2" box="[724,851,1699,1723]" name="Argentina" pageId="22" pageNumber="19">Argentina</collectingCountry>
,
<collectingCountry id="CAC18AAA7F6A886BFCAAF9142385EEF5" box="[862,981,1700,1724]" name="Paraguay" pageId="22" pageNumber="19">Paraguay</collectingCountry>
and
<collectingCountry id="CAC18AAA7F6A886BFBF9F913240CEEF2" box="[1037,1116,1699,1723]" name="Brazil" pageId="22" pageNumber="19">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="B269CA3A7F6A886BFF04F974224AEF52" blockId="22.[192,1440,1667,1819]" pageId="22" pageNumber="19">
We here correct the record concerning female protarsi in
<taxonomicName id="75D6B1B97F6A886BFC60F974246FEE92" baseAuthorityName="Gory" baseAuthorityYear="1844" box="[916,1087,1731,1755]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="19" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="80A216287F6A886BFC60F974246FEE92" box="[916,1087,1731,1755]" italics="true" pageId="22" pageNumber="19">C. bonariensis</emphasis>
</taxonomicName>
; contrary to
<bibRefCitation id="D647B7CB7F6A886BFB21F97325CCEE92" author="Edmonds, W. D." box="[1237,1436,1731,1755]" pageId="22" pageNumber="19" pagination="731 - 874" refId="ref56889" refString="Edmonds, W. D. 1972. Comparative skeletal morphology, systematic and evolution of the phanaeine dung beetles (Coleoptera: Scarabaeidae). The University of Kansas Science Bulletin 49: 731 - 874." type="journal article" year="1972">Edmonds (1972)</bibRefCitation>
, they are absent in this species. Consequently, medium-sized females and smaller males are virtually indistinguishable externally.
</paragraph>
</subSubSection>
</treatment>
</document>
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