233 lines
29 KiB
XML
233 lines
29 KiB
XML
<document id="D002EFBEC30B605D474E623CAC55FCBB" ID-DOI="10.5281/zenodo.5352924" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352924" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630209239027" checkinUser="felipe" docAuthor="Edmonds, W. D. & Zidek, J." docDate="2010" docId="3A7F7B2C7F5E8859FF34F93222D4ECF2" docLanguage="en" docName="0129.pdf" docOrigin="Insecta Mundi 2010 (129)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Coprophanaeus (Metallophanaeus) punctatus" docType="treatment" docVersion="3" lastPageNumber="33" masterDocId="C64603547F7C887DFFF4FFB02050E849" masterDocTitle="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)" masterLastPageNumber="111" masterPageNumber="1" pageNumber="31" updateTime="1699186523840" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="94F7ADBDCA5F5804F0163B3FB6219BE5">A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)</mods:title>
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<mods:name id="CC7BD224F3A1F44B28342FE38AA2F55B" type="personal">
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<mods:namePart id="E04AC03329277818CE3E569935B84E2E">Edmonds, W. D.</mods:namePart>
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<mods:name id="D7A7EFFB16A07A7639E96F66A5E0F2D9" type="personal">
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<mods:namePart id="CC8C79A8A43441BB2CCAE65183DED7DE">Zidek, J.</mods:namePart>
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<mods:affiliation id="622A060D38332BE95B97CEC9E9F207D6">Coprophanaeus bonariensis (Gory)</mods:affiliation>
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<mods:title id="796FBDD8EBEED7DEAF33EA344070D6AE">Insecta Mundi</mods:title>
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<mods:part id="C5E0417E0F62273F2574A3594719058B">
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<mods:date id="EBD3551442F6AFC90B8C944EE08A3603">2010</mods:date>
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<mods:number id="E322FC8055DEB902B29F5804E05438D7">2010-07-02</mods:number>
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<mods:detail id="C7AEBF2F972D91B791F94707C1DCE3D8" type="volume">
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<mods:number id="EDCF7D916BF5333E4834C8604AB8B4A4">2010</mods:number>
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<mods:number id="7DA692C0808E13B1EDBC25DDBAA33F43">129</mods:number>
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<mods:start id="3198635D68DCD3FCCCE69F8316FCC2C6">1</mods:start>
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<mods:classification id="6594CDDC80141D41B58B46BE84180BC5">journal article</mods:classification>
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<mods:identifier id="68ABD2F260E3FDD101A7942C2E9E7117" type="DOI">10.5281/zenodo.5352924</mods:identifier>
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<mods:identifier id="B57F29ADCEB9A05A4BECEEFBCF9AB2FF" type="ISSN">1942-1354</mods:identifier>
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<treatment id="3A7F7B2C7F5E8859FF34F93222D4ECF2" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F5E8859FF34F93222D4ECF2" httpUri="http://treatment.plazi.org/id/3A7F7B2C7F5E8859FF34F93222D4ECF2" lastPageId="36" lastPageNumber="33" pageId="34" pageNumber="31">
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<subSubSection id="FACC99B17F5E885FFF34F932245FEED2" box="[192,1039,1666,1691]" pageId="34" pageNumber="31" type="nomenclature">
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<paragraph id="B269CA3A7F5E885FFF34F932245FEED2" blockId="34.[192,1039,1666,1723]" box="[192,1039,1666,1691]" pageId="34" pageNumber="31">
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<taxonomicName id="75D6B1B97F5E885FFF34F932245FEED2" ID-CoL="Y6CJ" authority="(Olsoufieff, 1924)" baseAuthorityName="Olsoufieff" baseAuthorityYear="1924" box="[192,1039,1666,1691]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="species" species="punctatus" subGenus="Metallophanaeus">
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<emphasis id="80A216287F5E885FFF34F932245FEED2" bold="true" box="[192,1039,1666,1691]" pageId="34" pageNumber="31">
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<emphasis id="80A216287F5E885FFF34F93221C6EED2" bold="true" box="[192,406,1666,1691]" italics="true" pageId="34" pageNumber="31">Coprophanaeus</emphasis>
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(
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<emphasis id="80A216287F5E885FFE50F93222DFEED2" bold="true" box="[420,655,1666,1691]" italics="true" pageId="34" pageNumber="31">Metallophanaeus</emphasis>
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)
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<emphasis id="80A216287F5E885FFD6AF9342377EED2" bold="true" box="[670,807,1668,1691]" italics="true" pageId="34" pageNumber="31">punctatus</emphasis>
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(
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<bibRefCitation id="D647B7CB7F5E885FFCC0F9322455EED2" author="Olsoufieff, G." box="[820,1029,1666,1691]" pageId="34" pageNumber="31" pagination="4 - 201" refId="ref59457" refString="Olsoufieff, G. 1924. Les Phanaeides (Coleoptera - Lamellicornia), Famille Scarabaeidae - Tr. Coprini. Insecta 13: 4 - 201." type="journal article" year="1924">Olsoufieff, 1924</bibRefCitation>
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)
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="FACC99B17F5E885FFF34F91321C2EEF2" box="[192,402,1699,1723]" pageId="34" pageNumber="31" type="description">
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<paragraph id="B269CA3A7F5E885FFF34F91321C2EEF2" blockId="34.[192,1039,1666,1723]" box="[192,402,1699,1723]" pageId="34" pageNumber="31">
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<figureCitation id="2AEDD6BF7F5E885FFF34F9132147EEF2" box="[192,279,1699,1723]" captionStart="Figure 50-61" captionStartId="29.[192,276,1484,1506]" captionTargetBox="[191,1440,190,1454]" captionTargetId="figure-130@29.[191,1440,190,1454]" captionTargetPageId="29" captionText="Figure 50-61. Characters of the Coprophanaeus subgenus Metallophanaeus. 50) Ventral view of left protibia of C. saphirinus. 51) Same, C. pertyi. 52) Dorsal view of posterior portion of pronotum of C. thalassinus. 53) Same, C. pessoai. 54) Dorsal view of left elytron of C. punctatus. 55) Same, C. pessoai. 56) Posterior view of pygidium of C. horus. 57) Lateral view of metasternal angle of C. pertyi. 58) Same, C. pessoai. 59) Same, C. saphirinus. 60) Dorsal view of right side of head of C. horus. 61) Same, C. pessoai." figureDoi="http://doi.org/10.5281/zenodo.5352946" httpUri="https://zenodo.org/record/5352946/files/figure.png" pageId="34" pageNumber="31">Fig. 54</figureCitation>
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,
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<figureCitation id="2AEDD6BF7F5E885FFED6F9132113EEF2" box="[290,323,1699,1723]" captionStart="Figure 62" captionStartId="30.[191,275,1894,1916]" captionTargetBox="[247,1385,192,1865]" captionTargetId="figure-18@30.[247,1386,191,1865]" captionTargetPageId="30" captionText="Figure 62. Approximate geographic distribution of Coprophanaeus (Metallophanaeus) species." figureDoi="http://doi.org/10.5281/zenodo.5352948" httpUri="https://zenodo.org/record/5352948/files/figure.png" pageId="34" pageNumber="31">62</figureCitation>
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,
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<figureCitation id="2AEDD6BF7F5E885FFEB9F91321C2EEF2" box="[333,402,1699,1723]" captionStart="Figure 75-79" captionStartId="35.[192,276,1557,1579]" captionTargetBox="[202,1425,203,1532]" captionTargetId="figure-118@35.[192,1444,191,1539]" captionTargetPageId="35" captionText="Figure 75-79. Coprophanaeus punctatus. 75-76) Female habitus. 77-78) Male habitus. 79) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352954" httpUri="https://zenodo.org/record/5352954/files/figure.png" pageId="34" pageNumber="31">75-79</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="FACC99B17F5E885FFF34F95323A0EF52" pageId="34" pageNumber="31" type="reference_group">
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<paragraph id="B269CA3A7F5E885FFF34F95322F6EEB2" blockId="34.[192,1008,1763,1819]" box="[192,678,1763,1787]" pageId="34" pageNumber="31">
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<treatmentCitationGroup id="92C6ED147F5E885FFF34F95322F6EEB2" box="[192,678,1763,1787]" pageId="34" pageNumber="31">
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<treatmentCitation id="3377EC2B7F5E885FFF34F95322F6EEB2" author="Olsoufieff, G." box="[192,678,1763,1787]" page="71" pageId="34" pageNumber="31" year="1924">
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<taxonomicName id="75D6B1B97F5E885FFF34F95322F6EEB2" ID-CoL="4FSJ5" authority="Olsoufieff, 1924: 71" authorityName="Olsoufieff" authorityPageNumber="71" authorityYear="1924" box="[192,678,1763,1787]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="species" species="punctatus">
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<emphasis id="80A216287F5E885FFF34F95321E7EEB2" box="[192,439,1763,1787]" italics="true" pageId="34" pageNumber="31">Phanaeus punctatus</emphasis>
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<bibRefCitation id="D647B7CB7F5E885FFE49F95322F6EEB2" author="Olsoufieff, G." box="[445,678,1763,1787]" pageId="34" pageNumber="31" pagination="4 - 201" refId="ref59457" refString="Olsoufieff, G. 1924. Les Phanaeides (Coleoptera - Lamellicornia), Famille Scarabaeidae - Tr. Coprini. Insecta 13: 4 - 201." type="journal article" year="1924">Olsoufieff, 1924: 71</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="B269CA3A7F5E885FFF34F8B323A0EF52" blockId="34.[192,1008,1763,1819]" box="[192,1008,1795,1819]" pageId="34" pageNumber="31">
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<treatmentCitationGroup id="92C6ED147F5E885FFF34F8B323A0EF52" box="[192,1008,1795,1819]" pageId="34" pageNumber="31">
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<taxonomicName id="75D6B1B97F5E885FFF34F8B322D8EF52" ID-CoL="Y6CJ" authority="(Olsoufieff)" baseAuthorityName="Olsoufieff" baseAuthorityYear="1924" box="[192,648,1795,1819]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="species" species="punctatus">
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<emphasis id="80A216287F5E885FFF34F8B321ACEF52" box="[192,508,1795,1819]" italics="true" pageId="34" pageNumber="31">Coprophanaeus punctatus</emphasis>
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(Olsoufieff)
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</taxonomicName>
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(recomb.
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<treatmentCitation id="3377EC2B7F5E885FFD08F8B323B8EF52" author="Edmonds, W. D." box="[764,1000,1795,1819]" page="843" pageId="34" pageNumber="31" year="1972">
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||
<bibRefCitation id="D647B7CB7F5E885FFD08F8B323B8EF52" author="Edmonds, W. D." box="[764,1000,1795,1819]" pageId="34" pageNumber="31" pagination="731 - 874" refId="ref56889" refString="Edmonds, W. D. 1972. Comparative skeletal morphology, systematic and evolution of the phanaeine dung beetles (Coleoptera: Scarabaeidae). The University of Kansas Science Bulletin 49: 731 - 874." type="journal article" year="1972">Edmonds 1972: 843</bibRefCitation>
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</treatmentCitation>
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||
)
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="FACC99B17F5E885FFF34F8F32152EF32" pageId="34" pageNumber="31" type="materials_examined">
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||
<paragraph id="B269CA3A7F5E885FFF34F8F32152EF32" blockId="34.[192,1441,1859,1915]" pageId="34" pageNumber="31">
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||
<emphasis id="80A216287F5E885FFF34F8F3215BEF12" bold="true" box="[192,267,1859,1883]" pageId="34" pageNumber="31">Type.</emphasis>
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||
<typeStatus id="6D6D74987F5E885FFEE4F8F421D6EF15" box="[272,390,1860,1884]" pageId="34" pageNumber="31" type="lectotype">Lectotype</typeStatus>
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female (des. by Arnaud, 1982: 116), Muséum National d’Histoire Naturelle, Paris (examined).
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</paragraph>
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||
</subSubSection>
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||
<caption id="E6A99AB27F5F885EFF34F9A5226FEE00" ID-DOI="http://doi.org/10.5281/zenodo.5352954" ID-Zenodo-Dep="5352954" httpUri="https://zenodo.org/record/5352954/files/figure.png" pageId="35" pageNumber="32" startId="35.[192,276,1557,1579]" targetBox="[202,1425,203,1532]" targetPageId="35">
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<paragraph id="B269CA3A7F5F885EFF34F9A5226FEE00" blockId="35.[192,1440,1557,1609]" pageId="35" pageNumber="32">
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||
<emphasis id="80A216287F5F885EFF34F9A52134EE62" bold="true" box="[192,356,1557,1579]" pageId="35" pageNumber="32">Figure 75-79.</emphasis>
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<taxonomicName id="75D6B1B97F5F885EFE99F9A622C4EE65" baseAuthorityName="Olsoufieff" baseAuthorityYear="1924" box="[365,660,1558,1580]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="32" phylum="Arthropoda" rank="species" species="punctatus">
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<emphasis id="80A216287F5F885EFE99F9A622C4EE65" box="[365,660,1558,1580]" italics="true" pageId="35" pageNumber="32">Coprophanaeus punctatus</emphasis>
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</taxonomicName>
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||
.
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<emphasis id="80A216287F5F885EFD50F9A522BDEE62" bold="true" box="[676,749,1557,1579]" pageId="35" pageNumber="32">75-76)</emphasis>
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Female habitus.
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<emphasis id="80A216287F5F885EFC43F9A52450EE62" bold="true" box="[951,1024,1557,1579]" pageId="35" pageNumber="32">77-78)</emphasis>
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Male habitus.
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<emphasis id="80A216287F5F885EFB5BF9A52484EE62" bold="true" box="[1199,1236,1557,1579]" pageId="35" pageNumber="32">79)</emphasis>
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Aedeagus (dorsal view on left; lateral view on right).
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</paragraph>
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</caption>
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<subSubSection id="FACC99B17F5F8859FF34F9C82273E9B2" lastPageId="36" lastPageNumber="33" pageId="35" pageNumber="32" type="diagnosis">
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||
<paragraph id="B269CA3A7F5F8859FF34F9C823D2E8B2" blockId="35.[192,1441,1656,1872]" lastBlockId="36.[192,1440,195,507]" lastPageId="36" lastPageNumber="33" pageId="35" pageNumber="32">
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||
<emphasis id="80A216287F5F885EFF34F9C8211DEED9" bold="true" box="[192,333,1656,1680]" pageId="35" pageNumber="32">Diagnosis.</emphasis>
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<emphasis id="80A216287F5F885EFEA6F9C821E1EED9" box="[338,433,1656,1680]" italics="true" pageId="35" pageNumber="32">General</emphasis>
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||
– Paraocular areas (genae) completely punctured, usually coarsely and densely so, at most only weakly convex, not carinate lateral to eyes. Pronotum smooth posteromedially, otherwise weakly granulorugose; posterior angle normal, basal fossae small, incorporated into inner ends of sulcus paralleling posterior margin. Anteromedian angle of metasternum (seen in profile, as in
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<figureCitation id="2AEDD6BF7F5F885EFB6BF96824A4EEB9" box="[1183,1268,1752,1776]" captionStart="Figure 50-61" captionStartId="29.[192,276,1484,1506]" captionTargetBox="[191,1440,190,1454]" captionTargetId="figure-130@29.[191,1440,190,1454]" captionTargetPageId="29" captionText="Figure 50-61. Characters of the Coprophanaeus subgenus Metallophanaeus. 50) Ventral view of left protibia of C. saphirinus. 51) Same, C. pertyi. 52) Dorsal view of posterior portion of pronotum of C. thalassinus. 53) Same, C. pessoai. 54) Dorsal view of left elytron of C. punctatus. 55) Same, C. pessoai. 56) Posterior view of pygidium of C. horus. 57) Lateral view of metasternal angle of C. pertyi. 58) Same, C. pessoai. 59) Same, C. saphirinus. 60) Dorsal view of right side of head of C. horus. 61) Same, C. pessoai." figureDoi="http://doi.org/10.5281/zenodo.5352946" httpUri="https://zenodo.org/record/5352946/files/figure.png" pageId="35" pageNumber="32">Fig. 59</figureCitation>
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||
) rounded, not salient; angle capped with elongate thickening (bead). Elytral interstriae (
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<figureCitation id="2AEDD6BF7F5F885EFBC8F94824DFEF59" box="[1084,1167,1784,1808]" captionStart="Figure 50-61" captionStartId="29.[192,276,1484,1506]" captionTargetBox="[191,1440,190,1454]" captionTargetId="figure-130@29.[191,1440,190,1454]" captionTargetPageId="29" captionText="Figure 50-61. Characters of the Coprophanaeus subgenus Metallophanaeus. 50) Ventral view of left protibia of C. saphirinus. 51) Same, C. pertyi. 52) Dorsal view of posterior portion of pronotum of C. thalassinus. 53) Same, C. pessoai. 54) Dorsal view of left elytron of C. punctatus. 55) Same, C. pessoai. 56) Posterior view of pygidium of C. horus. 57) Lateral view of metasternal angle of C. pertyi. 58) Same, C. pessoai. 59) Same, C. saphirinus. 60) Dorsal view of right side of head of C. horus. 61) Same, C. pessoai." figureDoi="http://doi.org/10.5281/zenodo.5352946" httpUri="https://zenodo.org/record/5352946/files/figure.png" pageId="35" pageNumber="32">Fig. 54</figureCitation>
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||
) weakly convex; striae (x5) weakly impressed, carinulate, appearing weakly punctate (x5; see Comments below); bases of striae 1-4 impressed, not distinctly fossate. Ventral surface of protibia lateral to longitudinal carina paralleling inner margin entirely punctatorugose. Pygidium lacking basal groove. Length
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||
<quantity id="752E67DF7F588859FB8FFF7324A8E892" box="[1147,1272,195,219]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="36" pageNumber="33" unit="mm" value="15.0" valueMax="17.0" valueMin="13.0">13-17 mm</quantity>
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||
. Elytra black, pronotum with green to yellowish-green areas (
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||
<figureCitation id="2AEDD6BF7F588859FD0DFF532326E8B2" box="[761,886,227,251]" captionStart="Figure 75-79" captionStartId="35.[192,276,1557,1579]" captionTargetBox="[202,1425,203,1532]" captionTargetId="figure-118@35.[192,1444,191,1539]" captionTargetPageId="35" captionText="Figure 75-79. Coprophanaeus punctatus. 75-76) Female habitus. 77-78) Male habitus. 79) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352954" httpUri="https://zenodo.org/record/5352954/files/figure.png" pageId="36" pageNumber="33">Fig. 75-78</figureCitation>
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).
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</paragraph>
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||
<paragraph id="B269CA3A7F588859FF04FEB32302E932" blockId="36.[192,1440,195,507]" pageId="36" pageNumber="33">
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<emphasis id="80A216287F588859FF04FEB3217CE952" box="[240,300,259,283]" italics="true" pageId="36" pageNumber="33">Male</emphasis>
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(
|
||
<figureCitation id="2AEDD6BF7F588859FECEFEB321E0E952" box="[314,432,259,283]" captionStart="Figure 75-79" captionStartId="35.[192,276,1557,1579]" captionTargetBox="[202,1425,203,1532]" captionTargetId="figure-118@35.[192,1444,191,1539]" captionTargetPageId="35" captionText="Figure 75-79. Coprophanaeus punctatus. 75-76) Female habitus. 77-78) Male habitus. 79) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352954" httpUri="https://zenodo.org/record/5352954/files/figure.png" pageId="36" pageNumber="33">Fig. 77-79</figureCitation>
|
||
) – Head with evenly tapering long horn whose base is not abruptly swollen. Pronotum with pair of closely set, apically convergent processes near posterior margin, separated by strong concavity; disk anterior to processes lacking concavities. Pronotal disk punctate posteromedially, otherwise granulorugose. Apical processes of parameres rounded.
|
||
</paragraph>
|
||
<paragraph id="B269CA3A7F588859FF04FE332383E992" blockId="36.[192,1440,195,507]" pageId="36" pageNumber="33">
|
||
<emphasis id="80A216287F588859FF04FE332119E9D2" box="[240,329,387,411]" italics="true" pageId="36" pageNumber="33">Female</emphasis>
|
||
(
|
||
<figureCitation id="2AEDD6BF7F588859FEACFE332182E9D2" box="[344,466,387,411]" captionStart="Figure 75-79" captionStartId="35.[192,276,1557,1579]" captionTargetBox="[202,1425,203,1532]" captionTargetId="figure-118@35.[192,1444,191,1539]" captionTargetPageId="35" captionText="Figure 75-79. Coprophanaeus punctatus. 75-76) Female habitus. 77-78) Male habitus. 79) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352954" httpUri="https://zenodo.org/record/5352954/files/figure.png" pageId="36" pageNumber="33">Fig. 75-76</figureCitation>
|
||
) – Pronotum with small, weakly trituberculate crest adjacent to anterior margin followed by weak concavity bounded posteriorly by pair of weak tumosities near middle of disk. Pronotum finely granulorugose on disk and sides, punctate posteromedially.
|
||
</paragraph>
|
||
<paragraph id="B269CA3A7F588859FF04FE542273E9B2" blockId="36.[192,1440,195,507]" box="[240,547,483,507]" pageId="36" pageNumber="33">
|
||
<emphasis id="80A216287F588859FF04FE542254E9B2" box="[240,516,483,507]" italics="true" pageId="36" pageNumber="33">Specimens examined –</emphasis>
|
||
6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FACC99B17F588859FF34FD9320AAEA92" pageId="36" pageNumber="33" type="distribution">
|
||
<paragraph id="B269CA3A7F588859FF34FD9320BCEA12" blockId="36.[192,1440,547,603]" pageId="36" pageNumber="33">
|
||
<emphasis id="80A216287F588859FF34FD932125EA72" bold="true" box="[192,373,547,571]" pageId="36" pageNumber="33">Distribution.</emphasis>
|
||
Known only from isolated localities in northern Brazilian Atlantic Forest province (
|
||
<figureCitation id="2AEDD6BF7F588859FA84FD9320B0EA12" captionStart="Figure 62" captionStartId="30.[191,275,1894,1916]" captionTargetBox="[247,1385,192,1865]" captionTargetId="figure-18@30.[247,1386,191,1865]" captionTargetPageId="30" captionText="Figure 62. Approximate geographic distribution of Coprophanaeus (Metallophanaeus) species." figureDoi="http://doi.org/10.5281/zenodo.5352948" httpUri="https://zenodo.org/record/5352948/files/figure.png" pageId="36" pageNumber="33">Fig. 62</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B269CA3A7F588859FF34FD3320AAEA92" blockId="36.[192,1441,641,731]" pageId="36" pageNumber="33">
|
||
<emphasis id="80A216287F588859FF34FD332265EAD2" bold="true" box="[192,565,643,667]" pageId="36" pageNumber="33">
|
||
<collectingRegion id="701204D87F588859FF34FD332194EAD2" box="[192,452,643,667]" pageId="36" pageNumber="33">Collection Records.</collectingRegion>
|
||
<collectingCountry id="CAC18AAA7F588859FE3CFD332265EAD2" box="[456,565,643,667]" name="Brazil" pageId="36" pageNumber="33">BRAZIL</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="80A216287F588859FDB6FD332354EAD2" bold="true" box="[578,772,643,667]" pageId="36" pageNumber="33">
|
||
<collectingRegion id="701204D87F588859FDB6FD332354EAD2" box="[578,772,643,667]" pageId="36" pageNumber="33">Espiritu Santo</collectingRegion>
|
||
</emphasis>
|
||
– Fazenda Lagoa do Macuco [Mun. Linhares],
|
||
<geoCoordinate id="D7E2ACFD7F588859FAD7FD3325F0EAD2" box="[1315,1440,641,667]" degrees="19" direction="south" minutes="03" orientation="latitude" pageId="36" pageNumber="33" precision="15" seconds="50" value="-19.063889">
|
||
19
|
||
<superScript id="45A367727F588859FACBFD312517EAC6" attach="none" box="[1343,1351,641,655]" fontSize="6" pageId="36" pageNumber="33">o</superScript>
|
||
03’50”S
|
||
</geoCoordinate>
|
||
<geoCoordinate id="D7E2ACFD7F588859FF34FD13211FEAF2" box="[192,335,673,699]" degrees="39" direction="west" minutes="58" orientation="longitude" pageId="36" pageNumber="33" precision="15" seconds="43" value="-39.97861">
|
||
39
|
||
<superScript id="45A367727F588859FF2AFD1120B6EAE6" attach="none" box="[222,230,673,687]" fontSize="6" pageId="36" pageNumber="33">o</superScript>
|
||
58’43”W
|
||
</geoCoordinate>
|
||
,
|
||
<quantity id="752E67DF7F588859FEADFD1321C5EAF2" box="[345,405,675,699]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="33" unit="m" value="10.0">10 m</quantity>
|
||
(Jan).
|
||
<emphasis id="80A216287F588859FE1CFD1322CAEAF2" bold="true" box="[488,666,675,699]" pageId="36" pageNumber="33">
|
||
<collectingRegion id="701204D87F588859FE1CFD1322CAEAF2" box="[488,666,675,699]" country="Brazil" name="Pernambuco" pageId="36" pageNumber="33">Pernambuco</collectingRegion>
|
||
</emphasis>
|
||
– Igarassú [Reserva Ecologica Charles Darwin] (Mar, May-Jul, Oct).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FACC99B17F588859FF34FCB322D4ECF2" pageId="36" pageNumber="33" type="discussion">
|
||
<paragraph id="B269CA3A7F588859FF34FCB3246AEBD2" blockId="36.[192,1441,771,1211]" pageId="36" pageNumber="33">
|
||
<emphasis id="80A216287F588859FF34FCB32108EB52" bold="true" box="[192,344,771,795]" pageId="36" pageNumber="33">Comments.</emphasis>
|
||
<bibRefCitation id="D647B7CB7F588859FEAAFCB32276EB52" author="Olsoufieff, G." box="[350,550,771,795]" pageId="36" pageNumber="33" pagination="4 - 201" refId="ref59457" refString="Olsoufieff, G. 1924. Les Phanaeides (Coleoptera - Lamellicornia), Famille Scarabaeidae - Tr. Coprini. Insecta 13: 4 - 201." type="journal article" year="1924">Olsoufieff (1924)</bibRefCitation>
|
||
had only
|
||
<specimenCount id="A4D001B37F588859FD63FCB3236BEB52" box="[663,827,771,795]" pageId="36" pageNumber="33" type="female">three females</specimenCount>
|
||
at hand when he described this species. But he was rightly suspicious of placing it in
|
||
<emphasis id="80A216287F588859FDAAFC93233AEB72" box="[606,874,803,827]" italics="true" pageId="36" pageNumber="33">
|
||
<taxonomicName id="75D6B1B97F588859FDAAFC93234FEB72" authorityName="Olsoufieff" authorityYear="1924" box="[606,799,803,827]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="genus" sensu="stricto">Coprophanaeus</taxonomicName>
|
||
<taxonomicNameLabel id="9B91AB537F588859FCD3FC94233AEB72" box="[807,874,804,827]" pageId="36" pageNumber="33" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, noting “L’absence des tarses antérieurs et la forme de l’epistome les font placer ici [
|
||
<emphasis id="80A216287F588859FD73FCF323DCEB12" box="[647,908,835,859]" italics="true" pageId="36" pageNumber="33">
|
||
<taxonomicName id="75D6B1B97F588859FD73FCF32315EB12" authorityName="Olsoufieff" authorityYear="1924" box="[647,837,835,859]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="genus" sensu="stricto">Coprophanaeus</taxonomicName>
|
||
<taxonomicNameLabel id="9B91AB537F588859FCBFFCF423DCEB12" box="[843,908,836,859]" pageId="36" pageNumber="33" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</emphasis>
|
||
], ou les ranger au groupe du
|
||
<taxonomicName id="75D6B1B97F588859FB12FCF325C9EB12" baseAuthorityName="Sturm" baseAuthorityYear="1826" box="[1254,1433,835,859]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="saphirinus">
|
||
<emphasis id="80A216287F588859FB12FCF325C9EB12" box="[1254,1433,835,859]" italics="true" pageId="36" pageNumber="33">Ph. saphirinus</emphasis>
|
||
</taxonomicName>
|
||
, dont elles ont le caractère general. Ce n’est qu’en trouvant les [
|
||
<specimenCount id="A4D001B37F588859FC69FCD4245BEB32" box="[925,1035,867,891]" pageId="36" pageNumber="33" type="male">two male</specimenCount>
|
||
symbols] que l’on pourra résoudre cette question.”
|
||
<bibRefCitation id="D647B7CB7F588859FE75FC33226DEBD2" author="Arnaud, P." box="[385,573,899,923]" pageId="36" pageNumber="33" refId="ref56534" refString="Arnaud, P. 2002 c. Phanaeini. Les coleopteres du monde, vol. 28. Hillside Books; Canterbury. 151 p." type="book" year="2002">Arnaud (2002c)</bibRefCitation>
|
||
confirmed its placement in this subgenus.
|
||
</paragraph>
|
||
<paragraph id="B269CA3A7F588859FF04FC1322AAEC52" blockId="36.[192,1441,771,1211]" pageId="36" pageNumber="33">Olsoufieff’s name refers to apparent puncturing of the elytral striae. But strial puncturing in this species (x10) is in fact vague and virtually effaced. The illusion of obvious puncturing is produced by serial widening of the striae to produce a beaded (“moniliform”) appearance. Each wide spot of the stria encloses a flat area with a micropuncture in the middle.</paragraph>
|
||
<paragraph id="B269CA3A7F588859FF04FB9322D4ECF2" blockId="36.[192,1441,771,1211]" pageId="36" pageNumber="33">
|
||
<taxonomicName id="75D6B1B97F588859FF04FB93227AEC72" baseAuthorityName="Olsoufieff" baseAuthorityYear="1924" box="[240,554,1059,1083]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="80A216287F588859FF04FB93227AEC72" box="[240,554,1059,1083]" italics="true" pageId="36" pageNumber="33">Coprophanaeus punctatus</emphasis>
|
||
</taxonomicName>
|
||
is a scarce species of which we have seen, in addition to the
|
||
<typeStatus id="6D6D74987F588859FB18FB942571EC75" box="[1260,1313,1060,1084]" pageId="36" pageNumber="33">type</typeStatus>
|
||
series and a single female from Espiritu Santo, only one recently collected series from the Reserva Ecologica Charles Darwin in
|
||
<collectingRegion id="701204D87F588859FEB4FBD3218DEC32" box="[320,477,1123,1147]" country="Brazil" name="Pernambuco" pageId="36" pageNumber="33">Pernambuco</collectingRegion>
|
||
. The only strongly developed male in the series (see
|
||
<figureCitation id="2AEDD6BF7F588859FBBEFBD32493EC32" box="[1098,1219,1123,1147]" captionStart="Figure 75-79" captionStartId="35.[192,276,1557,1579]" captionTargetBox="[202,1425,203,1532]" captionTargetId="figure-118@35.[192,1444,191,1539]" captionTargetPageId="35" captionText="Figure 75-79. Coprophanaeus punctatus. 75-76) Female habitus. 77-78) Male habitus. 79) Aedeagus (dorsal view on left; lateral view on right)." figureDoi="http://doi.org/10.5281/zenodo.5352954" httpUri="https://zenodo.org/record/5352954/files/figure.png" pageId="36" pageNumber="33">Fig. 77-78</figureCitation>
|
||
) resembles that of
|
||
<taxonomicName id="75D6B1B97F588859FF34FB342137ECD2" baseAuthorityName="Sturm" baseAuthorityYear="1826" box="[192,359,1155,1179]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="saphirinus">
|
||
<emphasis id="80A216287F588859FF34FB342137ECD2" box="[192,359,1155,1179]" italics="true" pageId="36" pageNumber="33">C. saphirinus</emphasis>
|
||
</taxonomicName>
|
||
with the difference that the tooth-like posteromedian pronotal projections point upward rather than being inclined anteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |