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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.89864" ID-Pensoft-Pub="1860-1324-2-219" ID-Pensoft-UUID="9EE9E317E8195E3A94756A0DBBB1C35B" ID-ZooBank="BFE2BE4759E9450A807079B702A38625" ModsDocID="1860-1324-69-2-219" checkinTime="1671200981815" checkinUser="pensoft" docAuthor="Souma, Jun" docDate="2022" docId="3A2B74D8666753F7AD7C78A31C6283F9" docLanguage="en" docName="DeutEntomolZeit 69(2): 219-281" docOrigin="Deutsche Entomologische Zeitschrift 69 (2)" docPubDate="2022-12-15" docSource="http://dx.doi.org/10.3897/dez.69.89864" docTitle="Stephanitis (Norba) ishikawai Souma 2022, sp. nov." docType="treatment" docUuid="CC9E31D4-766C-4C48-ACBB-87D3297CA576" docUuidSource="ZooBank" docVersion="1" id="9EE9E317E8195E3A94756A0DBBB1C35B" lastPageNumber="219" masterDocId="9EE9E317E8195E3A94756A0DBBB1C35B" masterDocTitle="Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan" masterLastPageNumber="281" masterPageNumber="219" pageNumber="219" updateTime="1671200981815" updateUser="pensoft">
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<mods:title>Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Souma, Jun</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2238-5015</mods:nameIdentifier>
<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, Japan &amp; Research Fellowship for Young Scientists (DC 1), Japan Society for the Promotion of Science, Tokyo, Japan</mods:affiliation>
<mods:nameIdentifier type="email">kodokusignal@gmail.com</mods:nameIdentifier>
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<mods:date>2022</mods:date>
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<mods:number>2022-12-15</mods:number>
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<mods:number>69</mods:number>
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<taxonomicName LSID="https://zoobank.org/CC9E31D4-766C-4C48-ACBB-87D3297CA576" authority="Souma, 2022" authorityName="Souma" authorityYear="2022" class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" status="sp. nov." subGenus="Norba">Stephanitis (Norba) ishikawai</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="219">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="description">
<paragraph pageId="0" pageNumber="219">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male habitus of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov. from Yonaguni Island, southern part of Ryukyu Islands; B. S. (N.) mendica from Honshu; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai from Shikoku; F. S. (S.) tomokunii sp. nov. from Miyake Island, southern part of Izu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure3" httpUri="https://binary.pensoft.net/fig/782630" pageId="0" pageNumber="219">[Japanese name: Ishikawa-gunbai] Figs 3A</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female habitus of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov. from Miyako Island, southern part of Ryukyu Islands; B. S. (N.) mendica from Honshu; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai from Kyushu; F. S. (S.) tomokunii sp. nov. from Miyake Island, southern part of Izu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure5" httpUri="https://binary.pensoft.net/fig/782632" pageId="0" pageNumber="219">, 5A</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov .. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure8" httpUri="https://binary.pensoft.net/fig/782635" pageId="0" pageNumber="219">, 8A</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica possessing lateral carina; C-E. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (E) possessing lateral carina and from Honshu (D) lacking lateral carina; F. S. (S.) takeyai; G. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure10" httpUri="https://binary.pensoft.net/fig/782637" pageId="0" pageNumber="219">, 10A</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure12" httpUri="https://binary.pensoft.net/fig/782639" pageId="0" pageNumber="219">, 12A</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica possessing lateral carina; C-E. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (E) possessing lateral carina and from Honshu (D) lacking lateral carina; F. S. (S.) takeyai; G. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure14" httpUri="https://binary.pensoft.net/fig/782641" pageId="0" pageNumber="219">, 14A</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Male hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure16" httpUri="https://binary.pensoft.net/fig/782643" pageId="0" pageNumber="219">, 16A</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Female hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure18" httpUri="https://binary.pensoft.net/fig/782645" pageId="0" pageNumber="219">, 18A</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Rostra of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure19" httpUri="https://binary.pensoft.net/fig/782646" pageId="0" pageNumber="219">, 19F</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure21" httpUri="https://binary.pensoft.net/fig/782648" pageId="0" pageNumber="219">, 21F</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure23" httpUri="https://binary.pensoft.net/fig/782650" pageId="0" pageNumber="219">, 23F</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Line drawings of the pronota of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Abbreviations: ca, calli; co, collar; ho, hood; lc, lateral carina; mc, median carina; pa, paranotum; pd, pronotal disc; pp, posterior process. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure25" httpUri="https://binary.pensoft.net/fig/782652" pageId="0" pageNumber="219">, 25F</figureCitation>
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Line drawings of the pronota of six Stephanitis species from Japan, lateral view: A. S. (Norba) hiurai; B. S. (N.) ishikawai sp. nov.; C. S. (N.) mendica lacking lateral carina; D. S. (Stephanitis) tabidula lacking lateral carina from Honshu; E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure28" httpUri="https://binary.pensoft.net/fig/782655" pageId="0" pageNumber="219">, 28B</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Line drawings of the pygophore of five Stephanitis species from Japan, dorsal view: A-C. S. (Norba) ishikawai sp. nov. from Miyako (A) and Yonaguni (B) Islands, central part of Ryukyu Islands and Taiwan (C); D. S. (N.) mendica; E, F. S. (Stephanitis) tabidula from Honshu (E) and Tsushima Islands (F); G. S. (S.) takeyai; H. S. (S.) tomokunii sp. nov. Scale bars: 0.1 mm." figureDoi="10.3897/dez.69.89864.figure30" httpUri="https://binary.pensoft.net/fig/782657" pageId="0" pageNumber="219">, 30A-C</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Line drawings of the paramere of five Stephanitis species from Japan, dorsal view: A-C. S. (Norba) ishikawai sp. nov. from Miyako (A) and Yonaguni (B) Islands, central part of Ryukyu Islands and Taiwan (C); D. S. (N.) mendica; E, F. S. (Stephanitis) tabidula from Honshu (E) and Tsushima Islands (F); G. S. (S.) takeyai; H. S. (S.) tomokunii sp. nov. Scale bars: 0.1 mm. Arrows indicate important morphological differences." figureDoi="10.3897/dez.69.89864.figure32" httpUri="https://binary.pensoft.net/fig/782659" pageId="0" pageNumber="219">, 32A-C</figureCitation>
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Living individuals of three species of Stephanitis from Japan: A-D. S. (Norba) hayashii sp. nov. from the central part of Ryukyu Islands, male (A), female (B) and fifth (C) and fourth (D) instar nymphs from Kakeroma Island; E-G. S. (N.) hiurai from Amami-Oshima Island, central part of Ryukyu Islands, male (E) and female (F) and fifth instar nymph (G); H, I. S. (N.) ishikawai sp. nov. from Yonaguni Island, southern part of Ryukyu Islands, female (H) and fifth instar nymph (I)." figureDoi="10.3897/dez.69.89864.figure41" httpUri="https://binary.pensoft.net/fig/782668" pageId="0" pageNumber="219">, 41H, I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="reference_group">
<paragraph pageId="0" pageNumber="219">
<taxonomicName authority="Souma, 2022" authorityName="Souma" authorityYear="2022" class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" status="sp. nov." subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
, 1912:
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya (1931</bibRefCitation>
: 75) (distribution: part);
<bibRefCitation author="Maa, TC" journalOrPublisher="Polish Journal of Entomology" pageId="0" pageNumber="219" refId="B39" refString="Maa, TC, 1957. Nymphal stages of certain Oriental Tingidae (Hemiptera). Quarternary Journal of Taiwan Museum 10(3,4): 117-133." title="Nymphal stages of certain Oriental Tingidae (Hemiptera). Quarternary Journal of Taiwan Museum 10 (3,4): 117 - 133." year="1957">Maa (1957</bibRefCitation>
: 127) (nymph);
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya (1963</bibRefCitation>
: 38) (distribution: part);
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="271 - 280" refId="B44" refString="Miyamoto, S, 1964a. Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera). Kontyu 32 (2): 271 - 280" title="Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 a">Miyamoto (1964a</bibRefCitation>
: 274) (distribution: part);
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="523 - 528" refId="B45" refString="Miyamoto, S, 1964b. Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera). Kontyu 32 (4): 523 - 528" title="Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 b">Miyamoto (1964b</bibRefCitation>
: 524) (distribution: part);
<bibRefCitation author="Takara, T" journalOrPublisher="University of the Ryukyus" pageId="0" pageNumber="219" pagination="99 - 121" refId="B83" refString="Takara, T, Azuma, S, 1972. Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture. University of the Ryukyus 19: 99 - 121" title="Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture." volume="19" year="1972">Takara and Azuma (1972</bibRefCitation>
: 113) (distribution);
<bibRefCitation author="Azuma, S" journalOrPublisher="The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" refId="B4" refString="Azuma, S, Kinjo, M, 1987. Check-list of the insects of Okinawa. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" title="Check-list of the insects of Okinawa." year="1987">Azuma and Kinjo (1987</bibRefCitation>
: 34) (distribution);
<bibRefCitation author="Miyamoto, S" editor="Hirashima, Y" journalOrPublisher="I. Laboratory of Entomology, Faculty of Agriculture, Kyushu University, Fukuoka, Japan" pageId="0" pageNumber="219" pagination="151 - 188" refId="B50" refString="Miyamoto, S, Yasunaga, T, 1989. Heteroptera. In: Hirashima, Y, Ed., A Check List of Japanese Insects. I. Laboratory of Entomology, Faculty of Agriculture, Kyushu University, Fukuoka, Japan: 151 - 188" title="Heteroptera." volumeTitle="A Check List of Japanese Insects." year="1989">Miyamoto and Yasunaga (1989</bibRefCitation>
: 168) (checklist: Japan);
<bibRefCitation author="Yasunaga, T" journalOrPublisher=", Tokyo, Japan" pageId="0" pageNumber="219" refId="B114" refString="Yasunaga, T, Takai, M, Yamashita, I, Kawamura, M, Kawasawa, T, 1993. A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" title="A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd." year="1993">Yasunaga et al. (1993</bibRefCitation>
: 178) (monograph: part);
<bibRefCitation author="Hayashi, M" editor="Azuma, S" journalOrPublisher=")]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" pagination="127 - 150" refId="B24" refString="Hayashi, M, 2002. Heteroptera. In: Azuma, S, Yafuso, M, Kinjo, M, Hayashi, M, Kohama, T, Sasaki, T, Kimura, M, Kawamura, F, Eds., 沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2nd Edn.)]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan: 127 - 150" title="Heteroptera." volumeTitle="沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2 nd Edn." year="2002">Hayashi (2002</bibRefCitation>
: 137) (distribution);
<bibRefCitation author="Tomokuni, M" journalOrPublisher="Memoirs of the National Science Museum" pageId="0" pageNumber="219" pagination="59 - 69" refId="B100" refString="Tomokuni, M, 2006b. A small collection of Tingidae (Insecta, Heteroptera) from Taiwan, with a checklist of the known species. Memoirs of the National Science Museum 44: 59 - 69" title="A small collection of Tingidae (Insecta, Heteroptera) from Taiwan, with a checklist of the known species." volume="44" year="2006 b">Tomokuni (2006b</bibRefCitation>
: 67) (checklist: Taiwan);
<bibRefCitation author="Zheng, S" journalOrPublisher=", Taichung, Taiwan" pageId="0" pageNumber="219" refId="B117" refString="Zheng, S, Lin, Y, 2013. 椿象圖鑑 [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd., Taichung, Taiwan" title="椿象圖鑑 [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd." year="2013">Zheng and Lin (2013</bibRefCitation>
: 312) (distribution);
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni (2012</bibRefCitation>
: 205) (monograph: part);
<bibRefCitation author="Yamada, K" editor="Hayashi, M" journalOrPublisher="Touka-shobo, Fukuoka, Japan" pageId="0" pageNumber="219" pagination="429 - 435" refId="B111" refString="Yamada, K, Ishikawa, T, 2016. Family Tingidae. In: Hayashi, M, Tomokuni, M, Yoshizawa, K, Ishikawa, T, Eds., Catalogue of the insects of Japan Volume 4 Paraneoptera. Touka-shobo, Fukuoka, Japan: 429 - 435" title="Family Tingidae." volumeTitle="Catalogue of the insects of Japan Volume 4 Paraneoptera." year="2016">Yamada and Ishikawa (2016</bibRefCitation>
: 433) (checklist: Japan). Misidentifications.
</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName authority="Souma, 2022" authorityName="Souma" authorityYear="2022" class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" status="sp. nov." subGenus="Norba">Stephanitis (Norba) exigua</taxonomicName>
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
, 1912:
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya (1963</bibRefCitation>
: 38) (distribution: part);
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="271 - 280" refId="B44" refString="Miyamoto, S, 1964a. Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera). Kontyu 32 (2): 271 - 280" title="Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 a">Miyamoto (1964a</bibRefCitation>
: 274) (distribution: part);
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="523 - 528" refId="B45" refString="Miyamoto, S, 1964b. Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera). Kontyu 32 (4): 523 - 528" title="Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 b">Miyamoto (1964b</bibRefCitation>
: 524) (distribution: part);
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kyushu University" pageId="0" pageNumber="219" pagination="99 - 110" refId="B46" refString="Miyamoto, S, 1964c. Heteroptera collected by the Second Kyushu University Expedition to the Yaeyama Group, 1963 (excluding the Lygaeidae and Reduviidae). Reports of the Committee on Foreign Scientific Research. Kyushu University 2: 99 - 110" title="Heteroptera collected by the Second Kyushu University Expedition to the Yaeyama Group, 1963 (excluding the Lygaeidae and Reduviidae). Reports of the Committee on Foreign Scientific Research." volume="2" year="1964 c">Miyamoto (1964c</bibRefCitation>
: 105) (distribution);
<bibRefCitation author="Takara, T" journalOrPublisher="University of the Ryukyus" pageId="0" pageNumber="219" pagination="99 - 121" refId="B83" refString="Takara, T, Azuma, S, 1972. Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture. University of the Ryukyus 19: 99 - 121" title="Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture." volume="19" year="1972">Takara and Azuma (1972</bibRefCitation>
: 113) (distribution);
<bibRefCitation author="Azuma, S" journalOrPublisher="The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" refId="B4" refString="Azuma, S, Kinjo, M, 1987. Check-list of the insects of Okinawa. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" title="Check-list of the insects of Okinawa." year="1987">Azuma and Kinjo (1987</bibRefCitation>
: 34) (distribution);
<bibRefCitation author="Miyamoto, S" editor="Hirashima, Y" journalOrPublisher="I. Laboratory of Entomology, Faculty of Agriculture, Kyushu University, Fukuoka, Japan" pageId="0" pageNumber="219" pagination="151 - 188" refId="B50" refString="Miyamoto, S, Yasunaga, T, 1989. Heteroptera. In: Hirashima, Y, Ed., A Check List of Japanese Insects. I. Laboratory of Entomology, Faculty of Agriculture, Kyushu University, Fukuoka, Japan: 151 - 188" title="Heteroptera." volumeTitle="A Check List of Japanese Insects." year="1989">Miyamoto and Yasunaga (1989</bibRefCitation>
: 168) (checklist: Japan);
<bibRefCitation author="Hayashi, M" editor="Azuma, S" journalOrPublisher=")]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" pagination="127 - 150" refId="B24" refString="Hayashi, M, 2002. Heteroptera. In: Azuma, S, Yafuso, M, Kinjo, M, Hayashi, M, Kohama, T, Sasaki, T, Kimura, M, Kawamura, F, Eds., 沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2nd Edn.)]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan: 127 - 150" title="Heteroptera." volumeTitle="沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2 nd Edn." year="2002">Hayashi (2002</bibRefCitation>
: 137) (distribution);
<bibRefCitation author="Tomokuni, M" journalOrPublisher="Memoirs of the National Science Museum" pageId="0" pageNumber="219" pagination="59 - 69" refId="B100" refString="Tomokuni, M, 2006b. A small collection of Tingidae (Insecta, Heteroptera) from Taiwan, with a checklist of the known species. Memoirs of the National Science Museum 44: 59 - 69" title="A small collection of Tingidae (Insecta, Heteroptera) from Taiwan, with a checklist of the known species." volume="44" year="2006 b">Tomokuni (2006b</bibRefCitation>
: 67) (checklist: Taiwan);
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni (2012</bibRefCitation>
: 204) (monograph: part);
<bibRefCitation author="Yamada, K" editor="Hayashi, M" journalOrPublisher="Touka-shobo, Fukuoka, Japan" pageId="0" pageNumber="219" pagination="429 - 435" refId="B111" refString="Yamada, K, Ishikawa, T, 2016. Family Tingidae. In: Hayashi, M, Tomokuni, M, Yoshizawa, K, Ishikawa, T, Eds., Catalogue of the insects of Japan Volume 4 Paraneoptera. Touka-shobo, Fukuoka, Japan: 429 - 435" title="Family Tingidae." volumeTitle="Catalogue of the insects of Japan Volume 4 Paraneoptera." year="2016">Yamada and Ishikawa (2016</bibRefCitation>
: 433) (checklist: Japan). Misidentifications.
</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName authority="Souma, 2022" authorityName="Souma" authorityYear="2022" class="Insecta" family="Tingidae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" status="sp. nov." subGenus="Norba">Stephanitis (Norba) hiurai</taxonomicName>
Takeya, 1963:
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="271 - 280" refId="B44" refString="Miyamoto, S, 1964a. Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera). Kontyu 32 (2): 271 - 280" title="Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 a">Miyamoto (1964a</bibRefCitation>
: 272) (distribution: part);
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="523 - 528" refId="B45" refString="Miyamoto, S, 1964b. Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera). Kontyu 32 (4): 523 - 528" title="Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 b">Miyamoto (1964b</bibRefCitation>
: 524) (distribution: part). Misidentifications.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="type series">
<paragraph pageId="0" pageNumber="219">Type series.</paragraph>
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Holotype</emphasis>
(♂, ELKU), &quot;[JAPAN]: the Ryukyus, Yaeyama Isls., Yonaguni Is., Mantabaru&quot; [=JAPAN: Ryukyu Islands (southern part): Yonaguni Island: Mantabaru (approximate coordinates:
<geoCoordinate degrees="24" direction="north" minutes="27" orientation="latitude" precision="1" seconds="27.6" value="24.457668">24°27'27.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="122" direction="east" minutes="58" orientation="longitude" precision="1" seconds="16.6" value="122.971275">122°58'16.6&quot;E</geoCoordinate>
)], 11.xi.2020, leg. J. Souma.
<emphasis bold="true" pageId="0" pageNumber="219">Paratypes</emphasis>
(141 ♂♂ 252 ♀♀), JAPAN: Ryukyu Islands (southern part): Miyako Island: Karimata, 2.ix.1958, leg. T. Hidaka (1 ♀, ELKU); Hirara, Karimata, 1.iv.1991, leg. M. Hayashi (1 ♀, TUA); as above but 21.xi.1992 (2 ♂♂ 1 ♀, ELKU); as above but 22.xi.1992 (1 ♀, TUA); Gusukube, Wipya, 2.iv.1991, leg. M. Hayashi (5 ♂♂ 9 ♀♀, TUA); Mt. Nobarudake, 2.iv.1991, leg. M. Hayashi (6 ♂♂ 12 ♀♀, TUA); as above, but 11.x.1993 (7 ♂♂ 21 ♀♀, TUA); as above, but 13.v.1995 (1 ♀, TUA); as above, but 24.vi.2008 (2 ♂♂ 1 ♀, TUA); as above but 9.xi.2018, leg. J. Souma (3 ♂♂ 2 ♀♀, TUA); Ono Mountain Forest, 18.v.1991, leg. M. Hayashi (3 ♀♀, TUA); as above but 7.xi.2018, leg. J. Souma (5 ♀♀, ELKU; 1 ♂ 2 ♀♀, TUA); Ueno-son, Mt. Nobaru-dake, 31.x.1999, leg. T. Ishikawa (3 ♂♂ 4 ♀♀, TUA); as above but leg. M. Tomokuni (2 ♂♂ 1 ♀, NSMT); Karimata,
<normalizedToken originalValue="Ôno-sanrin">Ono-sanrin</normalizedToken>
, 1.xi.1999, leg. T. Ishikawa (1 ♂, TUA); Hirara, Higashi-sokobaru, 2.xi.1999, leg. T. Ishikawa (1 ♂ 1 ♀, TUA); Kamamamine Park, 28.iv.2002, leg. M. Hayashi (1 ♀, TUA); Ueno, Mt. Nobarudake, 27.xii.2017, leg. H. Yoshitake (2 ♂♂ 3 ♀♀, TUA); Gusukube, Yoshino, 8.xi.20118, leg. J. Souma (1 ♂ 7 ♀♀, TUA). Irabu Island: Irabu, 10.xi.2018, leg. J. Souma (1 ♂, ELKU); Makiyama, 10.xi.2018, leg. J. Souma (1 ♂ 2 ♀♀, ELKU); as above but 25.vii.2022, leg. S. Shimamoto (1 ♂ 3 ♀♀, TUA); near
<normalizedToken originalValue="Makiyama-tenbôdai">Makiyama-tenbodai</normalizedToken>
, 26.ix.2020, leg. H. Yoshitake (1 ♂ 1 ♀, NIAES). Kurima Island: 31.x.2007, leg. M. Hayashi (1 ♀, TUA). Ogami Island: 9.vi.2009, leg. M. Hayashi (1 ♂, TUA); 27.xii.2017, leg. H. Yoshitake (2 ♂♂ 6 ♀♀, TUA). Ishigaki Island: 5.ix.1957, leg. T. Takara (2 ♀♀, NSMT); 20.iii.1960, leg. T. Takara (1 ♂ 1 ♀, KUM); Nanbadake, 24.xi.1960, leg. K. Yasumatsu (1 ♀, KUM); Kawara, 28.x.1963, leg. Y. Hirashima (2 ♀♀, KUM); Mt. Omoto, alt. 100-526 m, 17.xi.1963, leg. G. A. Samuelson (1 ♀, KUM); Yonehara, 15.iii.1964, leg. Taira (1 ♂ 1 ♀, NIAES); Kahara-yama, 14.iii.1964, leg. Y. Miyatake (4 ♂♂ 1 ♀, KUM); as above but leg. S. Kimoto (1 ♂, KUM); as above but 18.iii.1964 (1 ♀, KUM); Yonehara, 15.iii.1964, leg. T. Shirozu (1 ♂ 3 ♀♀, KUM); as above but leg. Y. Miyatake (1 ♂ 1 ♀, KUM); as above but 12.vi.2008, leg. M. Hayashi (1 ♀, TUA); Omoto-san, 16.iii.1964, leg. Y. Miyatake (2 ♂♂ 6 ♀♀, KUM); Takada, 10.iv.1981, leg. K. Baba (1 ♀, NIAES); as above but 10.xi.1985, leg. M. Hayashi (2 ♀♀, TUA); Omoto, 15.x.1988, leg. M. Hayashi (1 ♀, TUA); as above but 2.vii.1998, leg. M. Hayashi (1 ♀, TUA); Banna-dake, 12.v.1975, leg. Y. Notsu (1 ♂, NSMT); as above but 16.xi.1984, leg. M. Tomokuni (2 ♀♀, NSMT); Mt. Bannadake, 2.iv.1976, leg. K. Murakami (1 ♂, NSMT); as above but 19.vii.2022, leg. S. Shimamoto (3 ♂♂ 6 ♀♀, TUA); Banna Park, 17.iii.1993, leg. M. Hayashi (1 ♀, TUA); Nosoko, 1.v.1998, leg. K. Takahashi (1 ♂, NSMT); Mt. Yarabu-dake, 23.iv.1999, leg. T. Ishikawa (1 ♀, TUA); as above but 6-9.v.2016, leg. N. Tsuji (2 ♀♀, ELKU); as above but 13.xi.2018, leg. J. Souma (1 ♂, TUA); Takeda-rindo, 23.v.2000, leg. H. Mizushima (1 ♂, TUA); Mt. Omoto-dake, 18.vi.1991, leg. S. Miyakawa (4 ♀♀, NSMT); as above but 22.vi.1991 (1 ♀, NSMT); as above but 12.v.1993, leg. M. Hayashi (1 ♀, TUA); as above but 24.v.2000, leg. T. Ishikawa (1 ♀, TUA); Mt. Maesedake, 20.iv.2005, leg. M. Hayashi (3 ♂♂ 3 ♀♀, TUA); as above but 21.ii.2006 (2 ♂♂ 4 ♀♀, TUA) as above but 22.iii.2006 (1 ♂, TUA); as above but 16.xi.2018, leg. J. Souma (1 ♀, TUA); as above but 6.iii.2022, leg. T. Saeki (1 ♂, TUA); as above but 20.vii.2022, leg. S. Shimamoto (5 ♀♀, TUA); Shiramizu, 29.x.2007, leg. M. Hayashi (1 ♀, TUA); Kabira, 29.x.2007, leg. M. Hayashi (1 ♀, TUA); as above but 8.x.2013, leg. R. Ito (1 ♂ 1 ♀, ELKU); Hirae,
<normalizedToken originalValue="Takeda-rindô">Takeda-rindo</normalizedToken>
, 4.x.2013, leg. R. Ito (1 ♀, ELKU); Hirae, Omoto-dake, 7.x.2013, leg. R. Ito (1 ♂, ELKU); Nagura, 7.x.2013, leg. R. Ito (3 ♂♂ 2 ♀♀, ELKU); Yarabu, Yarabu-dake, 8.x.2013, leg. R. Ito (1 ♀, ELKU); Yarabu, 8.x.2013, leg. R. Ito (1 ♀, ELKU); Okawa, 11.xi.2018, leg. J. Souma (1 ♂, TUA); Oganzaki, 13.xi.2018, leg. J. Souma (1 ♀, ELKU); Nosoko, 14.xi.2018, leg. J. Souma (1 ♂, ELKU); Itona, 14.xi.2018, leg. J. Souma (1 ♂, TUA); Inoda, 14.xi.2018, leg. J. Souma (1 ♀, ELKU); Hirae, 17.xi.2018, leg. J. Souma (1 ♀, TUA); Sakieda, 1.iii.2020, leg. Y. Obae (2 ♂♂ 1 ♀, ELKU). Iriomote Island: 24.iv.1951, leg. T. Takara (1 ♀, NSMT);
<normalizedToken originalValue="Ôhara">Ohara</normalizedToken>
, 22.xi.1960, leg. K. Yasumatsu (1 ♂, ELKU); Ushikumori, 4.x.1963, leg. S. Miyamoto (1 ♂, KUM); as above but 9.iii.1964, leg. Y. Miyatake (2 ♂♂ 1 ♀, KUM); as above leg. T. Shirozu (1 ♂ 4 ♀♀, KUM); as above but 11.iii.1964, leg. Y. Miyatake (3 ♂♂ 2 ♀♀, KUM); as above but 24.iii.1973, leg. S. Azuma (1 ♂ 2 ♀♀, NSMT); Sonai, 6.x.1963, leg. S. Miyamoto (1 ♂, KUM); as above but 8.x.1963 (1 ♂ 1 ♀, KUM); Komi, 12.vii.1963, leg. Y. Miyatake (4 ♀♀, KUM); as above but 2.iv.1978, leg. K. Baba (1 ♀, NIAES); as above but 10.xi.1984, leg. M. Tomokuni (1 ♀, NSMT); as above but 11.xi.1984, leg. M. Tomokuni (1 ♂, NSMT); as above but 20.xi.1998, leg. T. Ishikawa (1 ♀, TUA); as above but 20-23.iii.2015, leg. R. Ito (1 ♀, ELKU); as above but 24.viii.2020, leg. Y. Hisasue (1 ♂, ELKU); Shirahama, 5.xi.1963, leg. H. Hasegawa (1 ♀, NIAES); Shirahama-Sonae, 5.xi.1963, leg. H. Hasegawa (1 ♀, KUM); Shirahama-Hoshidate, 8.iii.1964, leg. Y. Miyatake (2 ♂♂ 2 ♀♀, KUM); as above but leg. S. Kimoto (1 ♀, KUM); Upstream of Nakara-gawa Riv., 12.iii.1964, leg. Y. Miyatake (1 ♀, KUM); as above but leg. S. Kimoto (1 ♂ 1 ♀, KUM); as above but leg. S. Higashihirachi (1 ♂, NIAES); Ohara, 20.viii.1971, leg. S. Azuma (1 ♂, NSMT); Funaura, 28.iii.1973, leg. S. Azuma (1 ♂, NSMT); as above but 21.viii.1976, leg. K. Kyoda (1 ♀, NSMT); as above but 8.x.1977, leg. M. Taniguchi (1 ♀, NSMT); as above but 9.x.1977, leg. M. Kinjo (1 ♀, NSMT); as above but 9.x.1977, leg. M. Arasaki (1 ♀, NSMT); as above but 11.xi.1984, leg. M. Tomokuni (1 ♀, NSMT); as above but 31.iii.1996, leg. M. Hayashi (1 ♂, TUA); Hoshidate, 7.xi.1985, leg. M. Hayashi (1 ♂, TUA); Monpanare, 29.ix.1993, leg. M. Hayashi (1 ♀, TUA); Takana, Yutsun Riv., 22.xi.1997, leg. M. Tomokuni (1 ♀, NSMT); Urauchi, 19.vi.1998, leg. T. Ishikawa (1 ♀, TUA);
<normalizedToken originalValue="Shirahama-rindô">Shirahama-rindo</normalizedToken>
, 1.iii.2002, leg. T. Ishikawa (2 ♀, TUA); Mt. Sonaidake, 1.iii.2002, leg. T. Ishikawa (2 ♂♂ 1 ♀, TUA);
<normalizedToken originalValue="Ôhara">Ohara</normalizedToken>
, Fusatoruba, 3.iii.2002, leg. T. Ishikawa (4 ♂♂ 1 ♀, TUA); Aira-gawa Rev., 7.x.2009, leg. T. Ishikawa (1 ♂, TUA); Haiminaka, 21-23.iii.2015, leg. R. Ito (1 ♀, ELKU); Sonai, 19.xi.2018, leg. J. Souma (1 ♀, ELKU); Otomi For. Rd., 20.xi.2018, leg. J. Souma (1 ♂, TUA); Unarizaki, 20.xi.2018, leg. J. Souma (1 ♂, ELKU);
<normalizedToken originalValue="Ôtomi-path">Otomi-path</normalizedToken>
, 21.ii.2020, leg. R. Ito (3 ♂♂ 3 ♀♀, ELKU); Mt. Tedo-san, 14.x.2020, leg. Y. Hisasue (1 ♀, ELKU); Inaba Trail, 16.x.2020, leg. Y. Hisasue (1 ♀, ELKU). Yonaguni Island: Mt. Kubura-dake, 2.vii.1973, leg. K. Shigematsu (1 ♂, NSMT); as above but 30.iii.2020, leg. R. Ito (1 ♀, ELKU); Mt. Urabu-dake, 28.iii.1997, leg. T. Ishikawa (5 ♂♂ 1 ♀♀, TUA); as holotype (6 ♂♂ 13 ♀♀, ELKU); as holotype but 14.xi.2020 (2 ♂♂ 8 ♀♀, ELKU); Mt. Inbidake, 11.xi.2020, leg. J. Souma (1 ♂ 3 ♀♀, ELKU); as above but 13.xi.2020, leg. J. Souma (11 ♂♂ 16 ♀♀, ELKU); Higawa, 11.xi.2020, leg. J. Souma (4 ♀♀, ELKU); Iranda For. Rd., 25.iii.2021, leg. T. Saeki (1 ♂, ELKU); Promenade of Atlas Moth Museum, 26.iii.2021, leg. K. Saito (2 ♂♂ 1 ♀, ELKU). A single paratype collected in 1958 was recorded as &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis aperta</emphasis>
</taxonomicName>
&quot;, four paratypes collected in 1960 as &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis exigua</emphasis>
</taxonomicName>
&quot; by
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">Takeya (1963)</bibRefCitation>
. Of 12 paratypes collected in 1963, eight were recorded as &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis exigua</emphasis>
</taxonomicName>
&quot; or &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis hiurai</emphasis>
</taxonomicName>
&quot; by
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="271 - 280" refId="B44" refString="Miyamoto, S, 1964a. Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera). Kontyu 32 (2): 271 - 280" title="Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 a">Miyamoto (1964a)</bibRefCitation>
. Of 42 paratypes collected in 1964, 38 individuals were recorded as &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis exigua</emphasis>
</taxonomicName>
&quot; by
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="523 - 528" refId="B45" refString="Miyamoto, S, 1964b. Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera). Kontyu 32 (4): 523 - 528" title="Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 b">Miyamoto (1964b)</bibRefCitation>
. Four paratypes from Iriomote Island collected by Y. Miyatake in 1963 were recorded as &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis exigua</emphasis>
</taxonomicName>
&quot; by
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kyushu University" pageId="0" pageNumber="219" pagination="99 - 110" refId="B46" refString="Miyamoto, S, 1964c. Heteroptera collected by the Second Kyushu University Expedition to the Yaeyama Group, 1963 (excluding the Lygaeidae and Reduviidae). Reports of the Committee on Foreign Scientific Research. Kyushu University 2: 99 - 110" title="Heteroptera collected by the Second Kyushu University Expedition to the Yaeyama Group, 1963 (excluding the Lygaeidae and Reduviidae). Reports of the Committee on Foreign Scientific Research." volume="2" year="1964 c">Miyamoto (1964c)</bibRefCitation>
. Six paratypes from Iriomote Island collected by S. Azuma or M. Kinjo were considered to be recorded as &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis aperta</emphasis>
</taxonomicName>
&quot; or &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis exigua</emphasis>
</taxonomicName>
&quot; by
<bibRefCitation author="Azuma, S" journalOrPublisher="The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" refId="B4" refString="Azuma, S, Kinjo, M, 1987. Check-list of the insects of Okinawa. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" title="Check-list of the insects of Okinawa." year="1987">Azuma and Kinjo (1987)</bibRefCitation>
.
</paragraph>
<caption doi="10.3897/dez.69.89864.figure3" httpUri="https://binary.pensoft.net/fig/782630" pageId="0" pageNumber="219" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 3.</emphasis>
Male habitus of five
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsal view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">S. (Norba) ishikawai</taxonomicName>
sp. nov. from Yonaguni Island, southern part of Ryukyu Islands;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) mendica" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="mendica" subGenus="N.">S. (N.) mendica</taxonomicName>
from Honshu;
<emphasis bold="true" pageId="0" pageNumber="219">C, D.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) tabidula" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tabidula" subGenus="Stephanitis">S. (Stephanitis) tabidula</taxonomicName>
from Honshu (
<emphasis bold="true" pageId="0" pageNumber="219">C</emphasis>
) and Kyushu (
<emphasis bold="true" pageId="0" pageNumber="219">D</emphasis>
);
<emphasis bold="true" pageId="0" pageNumber="219">E.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) takeyai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="takeyai" subGenus="S.">S. (S.) takeyai</taxonomicName>
from Shikoku;
<emphasis bold="true" pageId="0" pageNumber="219">F.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) tomokunii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tomokunii" subGenus="S.">S. (S.) tomokunii</taxonomicName>
sp. nov. from Miyake Island, southern part of Izu Islands. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption doi="10.3897/dez.69.89864.figure4" httpUri="https://binary.pensoft.net/fig/782631" pageId="0" pageNumber="219" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 4.</emphasis>
Female habitus of five
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsal view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) ambigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ambigua" subGenus="Stephanitis">S. (Stephanitis) ambigua</taxonomicName>
from Honshu;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">S. (Norba) aperta</taxonomicName>
from Honshu;
<emphasis bold="true" pageId="0" pageNumber="219">C.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
from Okinawa Island, central part of Ryukyu Islands;
<emphasis bold="true" pageId="0" pageNumber="219">D.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov. from Aguni Island, central part of Ryukyu Islands;
<emphasis bold="true" pageId="0" pageNumber="219">E, F.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
from Okinawa (
<emphasis bold="true" pageId="0" pageNumber="219">E</emphasis>
) and Takara (
<emphasis bold="true" pageId="0" pageNumber="219">F</emphasis>
) Islands, central part of Ryukyu Islands. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption doi="10.3897/dez.69.89864.figure5" httpUri="https://binary.pensoft.net/fig/782632" pageId="0" pageNumber="219" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 5.</emphasis>
Female habitus of five
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsal view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">S. (Norba) ishikawai</taxonomicName>
sp. nov. from Miyako Island, southern part of Ryukyu Islands;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) mendica" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="mendica" subGenus="N.">S. (N.) mendica</taxonomicName>
from Honshu;
<emphasis bold="true" pageId="0" pageNumber="219">C, D.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) tabidula" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tabidula" subGenus="Stephanitis">S. (Stephanitis) tabidula</taxonomicName>
from Honshu (
<emphasis bold="true" pageId="0" pageNumber="219">C</emphasis>
) and Kyushu (
<emphasis bold="true" pageId="0" pageNumber="219">D</emphasis>
);
<emphasis bold="true" pageId="0" pageNumber="219">E.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) takeyai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="takeyai" subGenus="S.">S. (S.) takeyai</taxonomicName>
from Kyushu;
<emphasis bold="true" pageId="0" pageNumber="219">F.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) tomokunii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tomokunii" subGenus="S.">S. (S.) tomokunii</taxonomicName>
sp. nov. from Miyake Island, southern part of Izu Islands. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption doi="10.3897/dez.69.89864.figure6" httpUri="https://binary.pensoft.net/fig/782633" pageId="0" pageNumber="219" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 6.</emphasis>
Teratological forms of two
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsal view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">S. (Norba) aperta</taxonomicName>
from Izu-Oshima Island, northern part of Izu Islands;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) takeyai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="takeyai" subGenus="Stephanitis">S. (Stephanitis) takeyai</taxonomicName>
from Kyushu. Arrows indicate antennae with segmental oligomery and malformed paranotum. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption doi="10.3897/dez.69.89864.figure7" httpUri="https://binary.pensoft.net/fig/782634" pageId="0" pageNumber="219" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 7.</emphasis>
Male pronota of five
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsal view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) ambigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ambigua" subGenus="Stephanitis">S. (Stephanitis) ambigua</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">S. (Norba) aperta</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">C.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">D.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov.;
<emphasis bold="true" pageId="0" pageNumber="219">E, F.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
from Amami-Oshima (
<emphasis bold="true" pageId="0" pageNumber="219">E</emphasis>
) and Kikai (
<emphasis bold="true" pageId="0" pageNumber="219">F</emphasis>
) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm.
</paragraph>
</caption>
<caption doi="10.3897/dez.69.89864.figure8" httpUri="https://binary.pensoft.net/fig/782635" pageId="0" pageNumber="219" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 8.</emphasis>
Male pronota of five
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsal view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">S. (Norba) ishikawai</taxonomicName>
sp. nov.;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) mendica" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="mendica" subGenus="N.">S. (N.) mendica</taxonomicName>
lacking lateral carina;
<emphasis bold="true" pageId="0" pageNumber="219">C, D.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) tabidula" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tabidula" subGenus="Stephanitis">S. (Stephanitis) tabidula</taxonomicName>
from Honshu (
<emphasis bold="true" pageId="0" pageNumber="219">C</emphasis>
) and Kyushu (
<emphasis bold="true" pageId="0" pageNumber="219">D</emphasis>
);
<emphasis bold="true" pageId="0" pageNumber="219">E.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) takeyai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="takeyai" subGenus="S.">S. (S.) takeyai</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">F.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) tomokunii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tomokunii" subGenus="S.">S. (S.) tomokunii</taxonomicName>
sp. nov.. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="materials_examined">
<paragraph pageId="0" pageNumber="219">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Non-types</emphasis>
(
<specimenCount type="male">13 ♂♂</specimenCount>
<specimenCount type="female">8 ♀♀</specimenCount>
13 nymphs),
<materialsCitation collectingDate="1964-03-08" collectingDateMax="2002-03-03" collectingDateMin="1964-03-08" collectorName="Y. Miyatake" country="JAPAN" county="Ryukyu Islands" location="Shirahama-Hoshidate" municipality="Iriomote Island" specimenCount="1" stateProvince="Ishikawa" typeStatus="holotype">
<collectingCountry name="Japan">JAPAN</collectingCountry>
:
<collectingCounty>Ryukyu Islands</collectingCounty>
: (southern part):
<collectingMunicipality>Iriomote Island</collectingMunicipality>
:
<location LSID="urn:lsid:plazi:treatment:3A2B74D8666753F7AD7C78A31C6283F9:C1A2DCDEAD78DEE50CC373DE59A7DC6C" country="JAPAN" county="Ryukyu Islands" municipality="Iriomote Island" name="Shirahama-Hoshidate" stateProvince="Ishikawa">Shirahama-Hoshidate</location>
,
<collectingDate value="1964-03-08">8.iii.1964</collectingDate>
, leg.
<collectorName>Y. Miyatake</collectorName>
(1 nymph);
<normalizedToken originalValue="Ôhara">
<location LSID="urn:lsid:plazi:treatment:3A2B74D8666753F7AD7C78A31C6283F9:7A977853D133851582673C4EDDD365C5" country="JAPAN" county="Ryukyu Islands" municipality="Iriomote Island" name="Ohara" stateProvince="Ishikawa">Ohara</location>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:3A2B74D8666753F7AD7C78A31C6283F9:B99B5ECE9640532ED996466AA2F6010B" country="JAPAN" county="Ryukyu Islands" municipality="Iriomote Island" name="Fusatoruba" stateProvince="Ishikawa">Fusatoruba</location>
,
<collectingDate value="2002-03-03">3.iii.2002</collectingDate>
, leg. T.
<collectingRegion country="Japan" name="Ishikawa">Ishikawa</collectingRegion>
(4 nymphs, TUA).
<location LSID="urn:lsid:plazi:treatment:3A2B74D8666753F7AD7C78A31C6283F9:25BC0563CD6B6B53F022ADA0EA54D8CD" country="JAPAN" county="Ryukyu Islands" municipality="Iriomote Island" name="Yonaguni Island" stateProvince="Ishikawa">Yonaguni Island</location>
: as
<typeStatus>holotype</typeStatus>
(1 nymph)
</materialsCitation>
.
<materialsCitation collectingDate="1917-04-16" collectingDateMax="2021-11-22" collectingDateMin="1917-04-16" collectorName="M. Maki, R. Takahashi, C. Takeya, Y. - J. Tsai" country="TAIWAN" county="City" location="Da'an District" municipality="Taihoku" specimenCount="21" specimenCount-female="8" specimenCount-male="13" stateProvince="Taipei" typeStatus="holotype">
<collectingCountry name="Taiwan">TAIWAN</collectingCountry>
:
<collectingRegion country="Taiwan" name="Taipei">Taipei</collectingRegion>
<collectingCounty>City</collectingCounty>
:
<collectingMunicipality>Taihoku</collectingMunicipality>
[=
<collectingRegion country="Taiwan" name="Taipei">Taipei</collectingRegion>
],
<collectingDate value="1917-04-16">16.iv.1917</collectingDate>
, leg.
<collectorName>M. Maki</collectorName>
(
<specimenCount type="male">2 ♂♂</specimenCount>
, ELKU); as above but
<collectingDate value="1927-01-15">15.i.1927</collectingDate>
, leg.
<collectorName>R. Takahashi</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); as above but
<collectingDate value="1930-04-02">2.iv.1930</collectingDate>
, leg.
<collectorName>C. Takeya</collectorName>
(
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
, ELKU); as above, but
<collectingDate value="1932-02-10">10.ii.1932</collectingDate>
, leg.
<collectorName>R. Takahashi</collectorName>
(
<specimenCount type="male">4 ♂♂</specimenCount>
<specimenCount type="female">2 ♀♀</specimenCount>
, ELKU;
<specimenCount type="male">1 ♂</specimenCount>
, KUM);
<location LSID="urn:lsid:plazi:treatment:3A2B74D8666753F7AD7C78A31C6283F9:B644E79422DD24318ECEFB55735F36B5" country="TAIWAN" county="City" municipality="Taihoku" name="Da'an District" stateProvince="Taipei">
<normalizedToken originalValue="Daan">Da'an</normalizedToken>
District
</location>
,
<collectingDate value="2021-10-05">5.x.2021</collectingDate>
, leg.
<collectorName>Y.-J. Tsai</collectorName>
(
<specimenCount type="male">3 ♂♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
7 nymphs, NMNS); as above but
<collectingDate value="2021-11-22">22.xi.2021</collectingDate>
(
<specimenCount type="male">2 ♂♂</specimenCount>
<specimenCount type="female">3 ♀♀</specimenCount>
, NMNS). Twelve specimens collected from Taiwan in the early 20th century were recorded as &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis aperta</emphasis>
</taxonomicName>
&quot; or &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis exigua</emphasis>
</taxonomicName>
&quot; by
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya (1931</bibRefCitation>
,
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">1963</bibRefCitation>
). The Taiwanese specimens are most similar to
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">Stephanitis (Norba) ishikawai</taxonomicName>
sp. nov. in morphological characteristics and were provisionally identified as pertaining to the new species in the present study. All 13 nymphs recorded above are in poor condition and are thus not described here
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="diagnosis">
<paragraph pageId="0" pageNumber="219">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">Stephanitis (Norba) ishikawai</taxonomicName>
sp. nov. is recognised amongst other species of
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov .. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure8" httpUri="https://binary.pensoft.net/fig/782635" pageId="0" pageNumber="219">8A</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica possessing lateral carina; C-E. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (E) possessing lateral carina and from Honshu (D) lacking lateral carina; F. S. (S.) takeyai; G. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure10" httpUri="https://binary.pensoft.net/fig/782637" pageId="0" pageNumber="219">10A</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure12" httpUri="https://binary.pensoft.net/fig/782639" pageId="0" pageNumber="219">12A</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica possessing lateral carina; C-E. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (E) possessing lateral carina and from Honshu (D) lacking lateral carina; F. S. (S.) takeyai; G. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure14" httpUri="https://binary.pensoft.net/fig/782641" pageId="0" pageNumber="219">14A</figureCitation>
,
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Male hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure16" httpUri="https://binary.pensoft.net/fig/782643" pageId="0" pageNumber="219">16A</figureCitation>
,
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Female hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure18" httpUri="https://binary.pensoft.net/fig/782645" pageId="0" pageNumber="219">18A</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Rostra of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure19" httpUri="https://binary.pensoft.net/fig/782646" pageId="0" pageNumber="219">19F</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure21" httpUri="https://binary.pensoft.net/fig/782648" pageId="0" pageNumber="219">21F</figureCitation>
,
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure23" httpUri="https://binary.pensoft.net/fig/782650" pageId="0" pageNumber="219">23F</figureCitation>
); calli light brown; body in male 2.1 times (in female 2.0 times) as long as maximum width across hemelytra (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male habitus of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov. from Yonaguni Island, southern part of Ryukyu Islands; B. S. (N.) mendica from Honshu; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai from Shikoku; F. S. (S.) tomokunii sp. nov. from Miyake Island, southern part of Izu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure3" httpUri="https://binary.pensoft.net/fig/782630" pageId="0" pageNumber="219">3A</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female habitus of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov. from Miyako Island, southern part of Ryukyu Islands; B. S. (N.) mendica from Honshu; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai from Kyushu; F. S. (S.) tomokunii sp. nov. from Miyake Island, southern part of Izu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure5" httpUri="https://binary.pensoft.net/fig/782632" pageId="0" pageNumber="219">5A</figureCitation>
); rostrum reaching metasternum; pronotum unicarinate (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Line drawings of the pronota of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Abbreviations: ca, calli; co, collar; ho, hood; lc, lateral carina; mc, median carina; pa, paranotum; pd, pronotal disc; pp, posterior process. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure25" httpUri="https://binary.pensoft.net/fig/782652" pageId="0" pageNumber="219">25F</figureCitation>
); hood pale, shorter than median carina of pronotum, wider than vertex at widest part, incompletely covering eye, as high as median carina of pronotum at highest part, posterior margin extending to middle of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc opaque; paranotum less erect, slightly narrowed posteriorly, with 3 rows of areolae at widest part, anterolateral angle protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height longer than height of eye (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Line drawings of the pronota of six Stephanitis species from Japan, lateral view: A. S. (Norba) hiurai; B. S. (N.) ishikawai sp. nov.; C. S. (N.) mendica lacking lateral carina; D. S. (Stephanitis) tabidula lacking lateral carina from Honshu; E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure28" httpUri="https://binary.pensoft.net/fig/782655" pageId="0" pageNumber="219">28B</figureCitation>
); apices of hemelytra close to each other in rest; costal area with 3-4 rows of areolae at widest part; subcostal area in male with 2 rows (in female with 3 rows) of areolae at widest part; discoidal area with 3-4 rows of areolae at widest part; sutural area with 3-4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R+M (radiomedial) vein carinate; pygophore elevated at centre of venter, posterior margin slightly emarginate in middle part (Fig.
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Line drawings of the pygophore of five Stephanitis species from Japan, dorsal view: A-C. S. (Norba) ishikawai sp. nov. from Miyako (A) and Yonaguni (B) Islands, central part of Ryukyu Islands and Taiwan (C); D. S. (N.) mendica; E, F. S. (Stephanitis) tabidula from Honshu (E) and Tsushima Islands (F); G. S. (S.) takeyai; H. S. (S.) tomokunii sp. nov. Scale bars: 0.1 mm." figureDoi="10.3897/dez.69.89864.figure30" httpUri="https://binary.pensoft.net/fig/782657" pageId="0" pageNumber="219">30A-C</figureCitation>
); and paramere stout, weakly curved inwards at apex, with outer margin not sinuate in middle part, inner margin nearly straight in basal part (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Line drawings of the paramere of five Stephanitis species from Japan, dorsal view: A-C. S. (Norba) ishikawai sp. nov. from Miyako (A) and Yonaguni (B) Islands, central part of Ryukyu Islands and Taiwan (C); D. S. (N.) mendica; E, F. S. (Stephanitis) tabidula from Honshu (E) and Tsushima Islands (F); G. S. (S.) takeyai; H. S. (S.) tomokunii sp. nov. Scale bars: 0.1 mm. Arrows indicate important morphological differences." figureDoi="10.3897/dez.69.89864.figure32" httpUri="https://binary.pensoft.net/fig/782659" pageId="0" pageNumber="219">32A-C</figureCitation>
).
</paragraph>
<caption doi="10.3897/dez.69.89864.figure9" httpUri="https://binary.pensoft.net/fig/782636" pageId="0" pageNumber="219" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 9.</emphasis>
Female pronota of five
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsal view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) ambigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ambigua" subGenus="Stephanitis">S. (Stephanitis) ambigua</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">S. (Norba) aperta</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">C.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">D.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov.;
<emphasis bold="true" pageId="0" pageNumber="219">E, F.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
from Amami-Oshima (
<emphasis bold="true" pageId="0" pageNumber="219">E</emphasis>
) and Kikai (
<emphasis bold="true" pageId="0" pageNumber="219">F</emphasis>
) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="description">
<paragraph pageId="0" pageNumber="219">Description.</paragraph>
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Male.</emphasis>
Head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown; calli light brown; eye dark red; areolae of pronotum and hemelytron transparent; hood pale; pronotal disc opaque; pubescence on body yellowish (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male habitus of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov. from Yonaguni Island, southern part of Ryukyu Islands; B. S. (N.) mendica from Honshu; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai from Shikoku; F. S. (S.) tomokunii sp. nov. from Miyake Island, southern part of Izu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure3" httpUri="https://binary.pensoft.net/fig/782630" pageId="0" pageNumber="219">3A</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov .. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure8" httpUri="https://binary.pensoft.net/fig/782635" pageId="0" pageNumber="219">8A</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure12" httpUri="https://binary.pensoft.net/fig/782639" pageId="0" pageNumber="219">12A</figureCitation>
,
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Male hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure16" httpUri="https://binary.pensoft.net/fig/782643" pageId="0" pageNumber="219">16A</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Rostra of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure19" httpUri="https://binary.pensoft.net/fig/782646" pageId="0" pageNumber="219">19F</figureCitation>
,
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure21" httpUri="https://binary.pensoft.net/fig/782648" pageId="0" pageNumber="219">21F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="219">
Body 2.1 times as long as maximum width across hemelytra (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male habitus of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov. from Yonaguni Island, southern part of Ryukyu Islands; B. S. (N.) mendica from Honshu; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai from Shikoku; F. S. (S.) tomokunii sp. nov. from Miyake Island, southern part of Izu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure3" httpUri="https://binary.pensoft.net/fig/782630" pageId="0" pageNumber="219">3A</figureCitation>
). Head (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov .. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure8" httpUri="https://binary.pensoft.net/fig/782635" pageId="0" pageNumber="219">8A</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure12" httpUri="https://binary.pensoft.net/fig/782639" pageId="0" pageNumber="219">12A</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Rostra of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure19" httpUri="https://binary.pensoft.net/fig/782646" pageId="0" pageNumber="219">19F</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Line drawings of the pronota of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Abbreviations: ca, calli; co, collar; ho, hood; lc, lateral carina; mc, median carina; pa, paranotum; pd, pronotal disc; pp, posterior process. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure25" httpUri="https://binary.pensoft.net/fig/782652" pageId="0" pageNumber="219">25F</figureCitation>
) glabrous; a pair of frontal spines close to each other at apices, not reaching apex of clypeus; median spine as long as frontal spines, reaching bases of frontal spines; pair of occipital spines longer than median spine, reaching middle part of eyes; antenniferous tubercles obtuse, slightly curved inwards; clypeus smooth. Compound eye round in dorsal view. Antenna densely covered with pubescence; segment I cylindrical; segment II cylindrical, shortest among antennal segments; segment III longest amongst antennal segments; segment IV cylindrical, longer than segment I. Bucculae closed at anterior ends, with 3 rows of areolae throughout length. Rostrum reaching metasternum.
</paragraph>
<paragraph pageId="0" pageNumber="219">
Pronotum (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov .. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure8" httpUri="https://binary.pensoft.net/fig/782635" pageId="0" pageNumber="219">8A</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure12" httpUri="https://binary.pensoft.net/fig/782639" pageId="0" pageNumber="219">12A</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Line drawings of the pronota of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Abbreviations: ca, calli; co, collar; ho, hood; lc, lateral carina; mc, median carina; pa, paranotum; pd, pronotal disc; pp, posterior process. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure25" httpUri="https://binary.pensoft.net/fig/782652" pageId="0" pageNumber="219">25F</figureCitation>
,
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Line drawings of the pronota of six Stephanitis species from Japan, lateral view: A. S. (Norba) hiurai; B. S. (N.) ishikawai sp. nov.; C. S. (N.) mendica lacking lateral carina; D. S. (Stephanitis) tabidula lacking lateral carina from Honshu; E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure28" httpUri="https://binary.pensoft.net/fig/782655" pageId="0" pageNumber="219">28B</figureCitation>
) unicarinate, 1.3 times as long as maximum width across paranota, sparsely covered with pubescence. Pronotal disc coarsely punctate. Hood shorter than median carina of pronotum, wider than vertex at widest part, incompletely covering eye, as high as median carina of pronotum at highest part, posterior margin extending to middle of pronotal disc, 4 rows of areolae at highest part, dorsal margin arched. Median carina straight, extending to apex of posterior process, 2 rows of areolae at highest part, dorsal margin arched. Calli smooth. Paranotum less erect, slightly narrowed posteriorly, with 3 rows of areolae at widest part, anterolateral angle protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height longer than height of eye. Posterior process triangular, obtuse at apex.
</paragraph>
<caption doi="10.3897/dez.69.89864.figure10" httpUri="https://binary.pensoft.net/fig/782637" pageId="0" pageNumber="219" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 10.</emphasis>
Female pronota of five
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsal view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">S. (Norba) ishikawai</taxonomicName>
sp. nov.;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) mendica" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="mendica" subGenus="N.">S. (N.) mendica</taxonomicName>
possessing lateral carina;
<emphasis bold="true" pageId="0" pageNumber="219">C-E.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) tabidula" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tabidula" subGenus="Stephanitis">S. (Stephanitis) tabidula</taxonomicName>
from Honshu (
<emphasis bold="true" pageId="0" pageNumber="219">C</emphasis>
) and Kyushu (
<emphasis bold="true" pageId="0" pageNumber="219">E</emphasis>
) possessing lateral carina and from Honshu (
<emphasis bold="true" pageId="0" pageNumber="219">D</emphasis>
) lacking lateral carina;
<emphasis bold="true" pageId="0" pageNumber="219">F.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) takeyai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="takeyai" subGenus="S.">S. (S.) takeyai</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">G.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) tomokunii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tomokunii" subGenus="S.">S. (S.) tomokunii</taxonomicName>
sp. nov. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="219">
Hemelytron (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Male hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure16" httpUri="https://binary.pensoft.net/fig/782643" pageId="0" pageNumber="219">16A</figureCitation>
) 2.4 times as long as its maximum width, extending beyond apex of abdomen, sparsely covered with pubescence; maximum width across hemelytra 1.6 times as much as maximum width across paranota; apices close to each other in rest; costal area with 3-4 rows of areolae at widest part; subcostal area with 2 rows of areolae at widest part; discoidal area with 3-4 rows of areolae at widest part; sutural area with 3-4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; C (costal), R+M (radiomedial) and Cu (cubital) veins carinate.
</paragraph>
<paragraph pageId="0" pageNumber="219">
Thoracic pleura (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure12" httpUri="https://binary.pensoft.net/fig/782639" pageId="0" pageNumber="219">12A</figureCitation>
) smooth in anterior part, coarsely punctate in posterior part. Ostiolar peritreme oblong. Sternal laminae (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Rostra of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure19" httpUri="https://binary.pensoft.net/fig/782646" pageId="0" pageNumber="219">19F</figureCitation>
) lower than bucculae; pro- and mesosternal laminae open in both anterior and posterior ends; metasternal laminae as high as mesosternal laminae, open at anterior ends, fused each other at posterior ends. Legs (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male habitus of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov. from Yonaguni Island, southern part of Ryukyu Islands; B. S. (N.) mendica from Honshu; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai from Shikoku; F. S. (S.) tomokunii sp. nov. from Miyake Island, southern part of Izu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure3" httpUri="https://binary.pensoft.net/fig/782630" pageId="0" pageNumber="219">3A</figureCitation>
) smooth, densely covered with pubescence; femora thickest at middle.
</paragraph>
<paragraph pageId="0" pageNumber="219">
Abdomen oblong in dorsal and ventral views. Pygophore (Figs
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure21" httpUri="https://binary.pensoft.net/fig/782648" pageId="0" pageNumber="219">21F</figureCitation>
,
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Line drawings of the pygophore of five Stephanitis species from Japan, dorsal view: A-C. S. (Norba) ishikawai sp. nov. from Miyako (A) and Yonaguni (B) Islands, central part of Ryukyu Islands and Taiwan (C); D. S. (N.) mendica; E, F. S. (Stephanitis) tabidula from Honshu (E) and Tsushima Islands (F); G. S. (S.) takeyai; H. S. (S.) tomokunii sp. nov. Scale bars: 0.1 mm." figureDoi="10.3897/dez.69.89864.figure30" httpUri="https://binary.pensoft.net/fig/782657" pageId="0" pageNumber="219">30A-C</figureCitation>
) compressed dorsoventrally, semicircular in ventral view, elevated at centre of venter, with posterior margin slightly emarginate in middle part, covered with pubescence. Paramere (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Line drawings of the paramere of five Stephanitis species from Japan, dorsal view: A-C. S. (Norba) ishikawai sp. nov. from Miyako (A) and Yonaguni (B) Islands, central part of Ryukyu Islands and Taiwan (C); D. S. (N.) mendica; E, F. S. (Stephanitis) tabidula from Honshu (E) and Tsushima Islands (F); G. S. (S.) takeyai; H. S. (S.) tomokunii sp. nov. Scale bars: 0.1 mm. Arrows indicate important morphological differences." figureDoi="10.3897/dez.69.89864.figure32" httpUri="https://binary.pensoft.net/fig/782659" pageId="0" pageNumber="219">32A-C</figureCitation>
) stout, expanded in middle part, weakly curved inwards at apex, outer margin not sinuate in middle part, inner margin nearly straight in basal part, covered with pubescence in middle part of outer and inner margins.
</paragraph>
<paragraph pageId="0" pageNumber="219">Measurements (n = 20). Body length with hemelytra 2.9-3.2 mm; maximum width across hemelytra 1.4-1.6 mm; length of antennal segments I to IV 0.2 mm, 0.1 mm, 1.2 mm and 0.7-0.8 mm, respectively; pronotal length 1.2-1.3 mm; pronotal width across paranota 0.8-1.0 mm; hemelytral length 2.2-2.5 mm; maximum width of hemelytron 0.9-1.1 mm.</paragraph>
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Female.</emphasis>
General habitus very similar to that of male (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female habitus of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua from Honshu; B. S. (Norba) aperta from Honshu; C. S. (N.) exigua from Okinawa Island, central part of Ryukyu Islands; D. S. (N.) hayashii sp. nov. from Aguni Island, central part of Ryukyu Islands; E, F. S. (N.) hiurai from Okinawa (E) and Takara (F) Islands, central part of Ryukyu Islands. Scale bar: 1.0 mm." figureDoi="10.3897/dez.69.89864.figure4" httpUri="https://binary.pensoft.net/fig/782631" pageId="0" pageNumber="219">4D</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure9" httpUri="https://binary.pensoft.net/fig/782636" pageId="0" pageNumber="219">9D</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure13" httpUri="https://binary.pensoft.net/fig/782640" pageId="0" pageNumber="219">13D</figureCitation>
,
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Female hemelytra of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure17" httpUri="https://binary.pensoft.net/fig/782644" pageId="0" pageNumber="219">17D</figureCitation>
,
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female terminalia of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure23" httpUri="https://binary.pensoft.net/fig/782650" pageId="0" pageNumber="219">23D</figureCitation>
), except for the following characters: body 2.0 times as long as maximum width across hemelytra; antennal segment III shorter than in male; pronotum 1.4 times as long as maximum width across paranota; hood wider than in male; hemelytron 2.3 times as long as maximum width; maximum width across hemelytra 1.7 times as much as maximum width across paranota; subcostal area wider than in male, with 3 rows of areolae at widest part; and apical part of abdomen pentagonal in ventral view.
</paragraph>
<caption doi="10.3897/dez.69.89864.figure11" httpUri="https://binary.pensoft.net/fig/782638" pageId="0" pageNumber="219" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 11.</emphasis>
Male pronota of five
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsolateral view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) ambigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ambigua" subGenus="Stephanitis">S. (Stephanitis) ambigua</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">S. (Norba) aperta</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">C.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">D.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov.;
<emphasis bold="true" pageId="0" pageNumber="219">E, F.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
from Amami-Oshima (
<emphasis bold="true" pageId="0" pageNumber="219">E</emphasis>
) and Kikai (
<emphasis bold="true" pageId="0" pageNumber="219">F</emphasis>
) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="219">Measurements (n = 20). Body length with hemelytra 3.0-3.4 mm; maximum width across hemelytra 1.5-1.7 mm; length of antennal segments I to IV 0.2 mm, 0.1 mm, 1.0 mm and 0.7-0.8 mm, respectively; pronotal length 1.2-1.4 mm; pronotal width across paranota 0.9-1.0 mm; hemelytral length 2.3-2.5 mm; maximum width of hemelytron 1.0-1.1 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="remarks">
<paragraph pageId="0" pageNumber="219">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="219">
Amongst the Japanese species of
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">S. (Norba) ishikawai</taxonomicName>
sp. nov. is most similar to
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov. in general habitus, but the former is easily distinguished from the latter by the following characters: length of antennal segment IV 0.7-0.8 mm (0.6 mm in
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov.); paranotum less erect (more erect in
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov.) (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure7" httpUri="https://binary.pensoft.net/fig/782634" pageId="0" pageNumber="219">7D</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov .. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure8" httpUri="https://binary.pensoft.net/fig/782635" pageId="0" pageNumber="219">8A</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure9" httpUri="https://binary.pensoft.net/fig/782636" pageId="0" pageNumber="219">9D</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica possessing lateral carina; C-E. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (E) possessing lateral carina and from Honshu (D) lacking lateral carina; F. S. (S.) takeyai; G. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure10" httpUri="https://binary.pensoft.net/fig/782637" pageId="0" pageNumber="219">10A</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure11" httpUri="https://binary.pensoft.net/fig/782638" pageId="0" pageNumber="219">11D</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure12" httpUri="https://binary.pensoft.net/fig/782639" pageId="0" pageNumber="219">12A</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure13" httpUri="https://binary.pensoft.net/fig/782640" pageId="0" pageNumber="219">13D</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica possessing lateral carina; C-E. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (E) possessing lateral carina and from Honshu (D) lacking lateral carina; F. S. (S.) takeyai; G. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure14" httpUri="https://binary.pensoft.net/fig/782641" pageId="0" pageNumber="219">14A</figureCitation>
); and paramere stout (slender in
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov.), with inner margin nearly straight in basal part (slightly curved inward in
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hayashii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hayashii" subGenus="N.">S. (N.) hayashii</taxonomicName>
sp. nov.) (Figs
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Line drawings of the paramere of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D, E. S. (N.) hayashii sp. nov. from Kakeroma (D) and Aguni (E) Islands, central part of Ryukyu Islands; F. S. (N.) hiurai. Scale bars: 0.1 mm. Arrows indicate important morphological differences." figureDoi="10.3897/dez.69.89864.figure31" httpUri="https://binary.pensoft.net/fig/782658" pageId="0" pageNumber="219">31C, D</figureCitation>
,
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Line drawings of the paramere of five Stephanitis species from Japan, dorsal view: A-C. S. (Norba) ishikawai sp. nov. from Miyako (A) and Yonaguni (B) Islands, central part of Ryukyu Islands and Taiwan (C); D. S. (N.) mendica; E, F. S. (Stephanitis) tabidula from Honshu (E) and Tsushima Islands (F); G. S. (S.) takeyai; H. S. (S.) tomokunii sp. nov. Scale bars: 0.1 mm. Arrows indicate important morphological differences." figureDoi="10.3897/dez.69.89864.figure32" httpUri="https://binary.pensoft.net/fig/782659" pageId="0" pageNumber="219">32A-C</figureCitation>
). Moreover, the partial COI gene pairwise sequence distances between both species are 0.042503-0.048201 (Suppl. material 3).
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">Stephanitis (Norba) ishikawai</taxonomicName>
sp. nov. was misidentified as
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
,
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
and
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
in previous studies (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya 1931</bibRefCitation>
,
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">1963</bibRefCitation>
;
<bibRefCitation author="Maa, TC" journalOrPublisher="Polish Journal of Entomology" pageId="0" pageNumber="219" refId="B39" refString="Maa, TC, 1957. Nymphal stages of certain Oriental Tingidae (Hemiptera). Quarternary Journal of Taiwan Museum 10(3,4): 117-133." title="Nymphal stages of certain Oriental Tingidae (Hemiptera). Quarternary Journal of Taiwan Museum 10 (3,4): 117 - 133." year="1957">Maa 1957</bibRefCitation>
;
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="271 - 280" refId="B44" refString="Miyamoto, S, 1964a. Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera). Kontyu 32 (2): 271 - 280" title="Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 a">Miyamoto 1964a</bibRefCitation>
,
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="523 - 528" refId="B45" refString="Miyamoto, S, 1964b. Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera). Kontyu 32 (4): 523 - 528" title="Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 b">1964b</bibRefCitation>
;
<bibRefCitation author="Takara, T" journalOrPublisher="University of the Ryukyus" pageId="0" pageNumber="219" pagination="99 - 121" refId="B83" refString="Takara, T, Azuma, S, 1972. Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture. University of the Ryukyus 19: 99 - 121" title="Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture." volume="19" year="1972">Takara and Azuma 1972</bibRefCitation>
;
<bibRefCitation author="Yasunaga, T" journalOrPublisher=", Tokyo, Japan" pageId="0" pageNumber="219" refId="B114" refString="Yasunaga, T, Takai, M, Yamashita, I, Kawamura, M, Kawasawa, T, 1993. A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" title="A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd." year="1993">Yasunaga et al. 1993</bibRefCitation>
;
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni 2012</bibRefCitation>
;
<bibRefCitation author="Zheng, S" journalOrPublisher=", Taichung, Taiwan" pageId="0" pageNumber="219" refId="B117" refString="Zheng, S, Lin, Y, 2013. 椿象圖鑑 [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd., Taichung, Taiwan" title="椿象圖鑑 [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd." year="2013">Zheng and Lin 2013</bibRefCitation>
), as these four species are very similar to each other. However,
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="N.">S. (N.) ishikawai</taxonomicName>
sp. nov. is easily distinguished from the three species by the following characters: rostrum reaching metasternum (not reaching in
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
,
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
and
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
) (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Rostra of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure19" httpUri="https://binary.pensoft.net/fig/782646" pageId="0" pageNumber="219">19B, C, E, F</figureCitation>
); and hood slightly higher than median carina of pronotum at highest part (as high as
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
,
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
and
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
), with posterior margin extending to middle of pronotal disc (not extending in
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="N.">S. (N.) aperta</taxonomicName>
,
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
and
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) hiurai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="hiurai" subGenus="N.">S. (N.) hiurai</taxonomicName>
) (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure7" httpUri="https://binary.pensoft.net/fig/782634" pageId="0" pageNumber="219">7B, C, E, F</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov .. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure8" httpUri="https://binary.pensoft.net/fig/782635" pageId="0" pageNumber="219">8A</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure9" httpUri="https://binary.pensoft.net/fig/782636" pageId="0" pageNumber="219">9B, C, E, F</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female pronota of five Stephanitis species from Japan, dorsal view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica possessing lateral carina; C-E. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (E) possessing lateral carina and from Honshu (D) lacking lateral carina; F. S. (S.) takeyai; G. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure10" httpUri="https://binary.pensoft.net/fig/782637" pageId="0" pageNumber="219">10A</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure11" httpUri="https://binary.pensoft.net/fig/782638" pageId="0" pageNumber="219">11B, C, E, F</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica lacking lateral carina; C, D. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (D); E. S. (S.) takeyai; F. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure12" httpUri="https://binary.pensoft.net/fig/782639" pageId="0" pageNumber="219">12A</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E, F. S. (N.) hiurai from Amami-Oshima (E) and Kikai (F) Islands, central part of Ryukyu Islands. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure13" httpUri="https://binary.pensoft.net/fig/782640" pageId="0" pageNumber="219">13B, C, E, F</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Female pronota of five Stephanitis species from Japan, dorsolateral view: A. S. (Norba) ishikawai sp. nov.; B. S. (N.) mendica possessing lateral carina; C-E. S. (Stephanitis) tabidula from Honshu (C) and Kyushu (E) possessing lateral carina and from Honshu (D) lacking lateral carina; F. S. (S.) takeyai; G. S. (S.) tomokunii sp. nov. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure14" httpUri="https://binary.pensoft.net/fig/782641" pageId="0" pageNumber="219">14A</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Line drawings of the pronota of six Stephanitis species from Japan, dorsal view: A. S. (Stephanitis) ambigua; B. S. (Norba) aperta; C. S. (N.) exigua; D. S. (N.) hayashii sp. nov.; E. S. (N.) hiurai; F. S. (N.) ishikawai sp. nov. Abbreviations: ca, calli; co, collar; ho, hood; lc, lateral carina; mc, median carina; pa, paranotum; pd, pronotal disc; pp, posterior process. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.89864.figure25" httpUri="https://binary.pensoft.net/fig/782652" pageId="0" pageNumber="219">25B, C, E, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/dez.69.89864.figure12" httpUri="https://binary.pensoft.net/fig/782639" pageId="0" pageNumber="219" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="219">
<emphasis bold="true" pageId="0" pageNumber="219">Figure 12.</emphasis>
Male pronota of five
<taxonomicName authorityName="Stal" authorityYear="1873" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="219">Stephanitis</emphasis>
</taxonomicName>
species from Japan, dorsolateral view:
<emphasis bold="true" pageId="0" pageNumber="219">A.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">S. (Norba) ishikawai</taxonomicName>
sp. nov.;
<emphasis bold="true" pageId="0" pageNumber="219">B.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) mendica" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="mendica" subGenus="N.">S. (N.) mendica</taxonomicName>
lacking lateral carina;
<emphasis bold="true" pageId="0" pageNumber="219">C, D.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (Stephanitis) tabidula" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tabidula" subGenus="Stephanitis">S. (Stephanitis) tabidula</taxonomicName>
from Honshu (
<emphasis bold="true" pageId="0" pageNumber="219">C</emphasis>
) and Kyushu (
<emphasis bold="true" pageId="0" pageNumber="219">D</emphasis>
);
<emphasis bold="true" pageId="0" pageNumber="219">E.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) takeyai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="takeyai" subGenus="S.">S. (S.) takeyai</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="219">F.</emphasis>
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (S.) tomokunii" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="tomokunii" subGenus="S.">S. (S.) tomokunii</taxonomicName>
sp. nov. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="distribution">
<paragraph pageId="0" pageNumber="219">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="219">
Japan (Ryukyu Islands (southern part): Miyako Island, Irabu Island, Kurima Island, Ogami Island, Ishigaki Island, Iriomote Island, Yonaguni Island); Taiwan (northern part) (Fig.
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Collection sites of Stephanitis (Norba) ishikawai sp. nov. and S. (N.) mendica used in the present study. Red-filled areas = localities; circles = small isolated islands." figureDoi="10.3897/dez.69.89864.figure47" httpUri="https://binary.pensoft.net/fig/782674" pageId="0" pageNumber="219">47</figureCitation>
) (
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="65 - 84" refId="B86" refString="Takeya, C, 1931. Some Tingitidae of the Japanese Empire. Mushi 4: 65 - 84" title="Some Tingitidae of the Japanese Empire." volume="4" year="1931">Takeya 1931</bibRefCitation>
,
<bibRefCitation author="Takeya, C" journalOrPublisher="Mushi" pageId="0" pageNumber="219" pagination="27 - 52" refId="B92" refString="Takeya, C, 1963. Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera). Mushi 37 (4): 27 - 52" title="Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 2 (Hemiptera)." volume="37" year="1963">1963</bibRefCitation>
;
<bibRefCitation author="Maa, TC" journalOrPublisher="Polish Journal of Entomology" pageId="0" pageNumber="219" refId="B39" refString="Maa, TC, 1957. Nymphal stages of certain Oriental Tingidae (Hemiptera). Quarternary Journal of Taiwan Museum 10(3,4): 117-133." title="Nymphal stages of certain Oriental Tingidae (Hemiptera). Quarternary Journal of Taiwan Museum 10 (3,4): 117 - 133." year="1957">Maa 1957</bibRefCitation>
;
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="271 - 280" refId="B44" refString="Miyamoto, S, 1964a. Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera). Kontyu 32 (2): 271 - 280" title="Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 a">Miyamoto 1964a</bibRefCitation>
,
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="523 - 528" refId="B45" refString="Miyamoto, S, 1964b. Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera). Kontyu 32 (4): 523 - 528" title="Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 b">1964b</bibRefCitation>
,
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kyushu University" pageId="0" pageNumber="219" pagination="99 - 110" refId="B46" refString="Miyamoto, S, 1964c. Heteroptera collected by the Second Kyushu University Expedition to the Yaeyama Group, 1963 (excluding the Lygaeidae and Reduviidae). Reports of the Committee on Foreign Scientific Research. Kyushu University 2: 99 - 110" title="Heteroptera collected by the Second Kyushu University Expedition to the Yaeyama Group, 1963 (excluding the Lygaeidae and Reduviidae). Reports of the Committee on Foreign Scientific Research." volume="2" year="1964 c">1964c</bibRefCitation>
;
<bibRefCitation author="Takara, T" journalOrPublisher="University of the Ryukyus" pageId="0" pageNumber="219" pagination="99 - 121" refId="B83" refString="Takara, T, Azuma, S, 1972. Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture. University of the Ryukyus 19: 99 - 121" title="Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture." volume="19" year="1972">Takara and Azuma 1972</bibRefCitation>
;
<bibRefCitation author="Azuma, S" journalOrPublisher="The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" refId="B4" refString="Azuma, S, Kinjo, M, 1987. Check-list of the insects of Okinawa. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" title="Check-list of the insects of Okinawa." year="1987">Azuma and Kinjo 1987</bibRefCitation>
;
<bibRefCitation author="Yasunaga, T" journalOrPublisher=", Tokyo, Japan" pageId="0" pageNumber="219" refId="B114" refString="Yasunaga, T, Takai, M, Yamashita, I, Kawamura, M, Kawasawa, T, 1993. A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" title="A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd." year="1993">Yasunaga et al. 1993</bibRefCitation>
;
<bibRefCitation author="Hayashi, M" editor="Azuma, S" journalOrPublisher=")]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan" pageId="0" pageNumber="219" pagination="127 - 150" refId="B24" refString="Hayashi, M, 2002. Heteroptera. In: Azuma, S, Yafuso, M, Kinjo, M, Hayashi, M, Kohama, T, Sasaki, T, Kimura, M, Kawamura, F, Eds., 沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2nd Edn.)]. The Biological Society of Okinawa, Nishihara, Okinawa, Japan: 127 - 150" title="Heteroptera." volumeTitle="沖縄産昆虫目録 [Check List of the Insect of the Ryukyu Islands (2 nd Edn." year="2002">Hayashi 2002</bibRefCitation>
;
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni 2012</bibRefCitation>
;
<bibRefCitation author="Zheng, S" journalOrPublisher=", Taichung, Taiwan" pageId="0" pageNumber="219" refId="B117" refString="Zheng, S, Lin, Y, 2013. 椿象圖鑑 [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd., Taichung, Taiwan" title="椿象圖鑑 [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd." year="2013">Zheng and Lin 2013</bibRefCitation>
;
<bibRefCitation author="Yamada, K" editor="Hayashi, M" journalOrPublisher="Touka-shobo, Fukuoka, Japan" pageId="0" pageNumber="219" pagination="429 - 435" refId="B111" refString="Yamada, K, Ishikawa, T, 2016. Family Tingidae. In: Hayashi, M, Tomokuni, M, Yoshizawa, K, Ishikawa, T, Eds., Catalogue of the insects of Japan Volume 4 Paraneoptera. Touka-shobo, Fukuoka, Japan: 429 - 435" title="Family Tingidae." volumeTitle="Catalogue of the insects of Japan Volume 4 Paraneoptera." year="2016">Yamada and Ishikawa 2016</bibRefCitation>
; present study). Judging from the photographs, living individuals identified as &quot;
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) aperta" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="aperta" subGenus="Norba">Stephanitis (Norba) aperta</taxonomicName>
&quot; or &quot;
<taxonomicName class="Insecta" family="Lauraceae" kingdom="Animalia" lsidName="S. (N.) exigua" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="exigua" subGenus="N.">S. (N.) exigua</taxonomicName>
&quot; in previous studies (Yasunaga et al. 193;
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni 2012</bibRefCitation>
;
<bibRefCitation author="Zheng, S" journalOrPublisher=", Taichung, Taiwan" pageId="0" pageNumber="219" refId="B117" refString="Zheng, S, Lin, Y, 2013. 椿象圖鑑 [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd., Taichung, Taiwan" title="椿象圖鑑 [Encyclopedia of Heteroptera from Taiwan]. Morning Star Publishing Co., Ltd." year="2013">Zheng and Lin 2013</bibRefCitation>
) corresponded to the new species.
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">Stephanitis (Norba) ishikawai</taxonomicName>
sp. nov. inhabits the laurilignosa in a subtropical climate of the southern part of the Ryukyu Islands and northern Taiwan, which is in the Oriental Region.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="etymology">
<paragraph pageId="0" pageNumber="219">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="219">The new species is named in honour of Tadashi Ishikawa, a Japanese heteropterist who collected part of paratypes and taught the author how to describe new species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="host">
<paragraph pageId="0" pageNumber="219">Host plants.</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Cinnamomum" kingdom="Plantae" lsidName="Cinnamomum camphora" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="camphora">
<emphasis italics="true" pageId="0" pageNumber="219">Cinnamomum camphora</emphasis>
</taxonomicName>
,
<normalizedToken originalValue="“Kusunoki”">&quot;Kusunoki&quot;</normalizedToken>
(
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
) (present study);
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Litsea" kingdom="Plantae" lsidName="Litsea japonica" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="japonica">
<emphasis italics="true" pageId="0" pageNumber="219">Litsea japonica</emphasis>
</taxonomicName>
,
<normalizedToken originalValue="“Hamabiwa”">&quot;Hamabiwa&quot;</normalizedToken>
(
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
) (Fig.
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Lauraceous host plants of six species of Stephanitis from Japan: A. Lindera glauca from Honshu, damaged by S. (Stephanitis) ambigua; Machilus thunbergii (B), Neolitsea sericea (C) and Cinnamomum camphora (D) from Honshu, all damaged by S. (Norba) aperta; E. M. thunbergii from Okinawa Island, damaged by S. (N.) exigua; Litsea japonica (F) from Kakeroma Island and C. yabunikkei (G) from Tokashiki Island, both damaged by S. (N.) hayashii sp. nov.; H. M. thunbergii from Amami-Oshima Island, damaged by S. (N.) hiurai; I. Lit. japonica from Miyako Island, damaged by S. (N.) ishikawai sp. nov." figureDoi="10.3897/dez.69.89864.figure43" httpUri="https://binary.pensoft.net/fig/782670" pageId="0" pageNumber="219">43I</figureCitation>
) (present study);
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Machilus" kingdom="Plantae" lsidName="Machilus thunbergii" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="species" species="thunbergii">
<emphasis italics="true" pageId="0" pageNumber="219">Machilus thunbergii</emphasis>
</taxonomicName>
,
<normalizedToken originalValue="“Tabunoki”">&quot;Tabunoki&quot;</normalizedToken>
(
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="" order="Laurales" pageId="0" pageNumber="219" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
) (present study).
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">Stephanitis (Norba) ishikawai</taxonomicName>
sp. nov. feeds only on lauraceous trees and is oligophagous.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="219" type="biology_ecology">
<paragraph pageId="0" pageNumber="219">Biology.</paragraph>
<paragraph pageId="0" pageNumber="219">
<taxonomicName authorityName="Souma" authorityYear="2022" class="Insecta" family="Platanaceae" genus="Stephanitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephanitis (Norba) ishikawai" order="Hemiptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="ishikawai" subGenus="Norba">Stephanitis (Norba) ishikawai</taxonomicName>
sp. nov. feeds on the abaxial surface of leaves of the three host plants (present study). Adults were collected in almost all seasons (
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="271 - 280" refId="B44" refString="Miyamoto, S, 1964a. Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera). Kontyu 32 (2): 271 - 280" title="Tingidae and Nabidae of the South-West Islands, Lying Between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 a">Miyamoto 1964a</bibRefCitation>
,
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="219" pagination="523 - 528" refId="B45" refString="Miyamoto, S, 1964b. Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera). Kontyu 32 (4): 523 - 528" title="Additions to the tingid fauna of the South-West Islands, lying between Kyushu and Formosa (Hemiptera)." volume="32" year="1964 b">1964b</bibRefCitation>
,
<bibRefCitation author="Miyamoto, S" journalOrPublisher="Kyushu University" pageId="0" pageNumber="219" pagination="99 - 110" refId="B46" refString="Miyamoto, S, 1964c. Heteroptera collected by the Second Kyushu University Expedition to the Yaeyama Group, 1963 (excluding the Lygaeidae and Reduviidae). Reports of the Committee on Foreign Scientific Research. Kyushu University 2: 99 - 110" title="Heteroptera collected by the Second Kyushu University Expedition to the Yaeyama Group, 1963 (excluding the Lygaeidae and Reduviidae). Reports of the Committee on Foreign Scientific Research." volume="2" year="1964 c">1964c</bibRefCitation>
;
<bibRefCitation author="Takara, T" journalOrPublisher="University of the Ryukyus" pageId="0" pageNumber="219" pagination="99 - 121" refId="B83" refString="Takara, T, Azuma, S, 1972. Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture. University of the Ryukyus 19: 99 - 121" title="Fauna of the Iriomote Island, Ryukyus. II. List of the insects (2). Order Hemiptera (Heteroptera). The Science Bulletin of the Faculty of Agriculture." volume="19" year="1972">Takara and Azuma 1972</bibRefCitation>
;
<bibRefCitation author="Yasunaga, T" journalOrPublisher=", Tokyo, Japan" pageId="0" pageNumber="219" refId="B114" refString="Yasunaga, T, Takai, M, Yamashita, I, Kawamura, M, Kawasawa, T, 1993. A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" title="A field guide to Japanese bugs. Terrestrial heteropterans. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd." year="1993">Yasunaga et al. 1993</bibRefCitation>
;
<bibRefCitation author="Yamada, K" editor="Ishikawa, T" journalOrPublisher="Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan" pageId="0" pageNumber="219" pagination="180 - 213" refId="B112" refString="Yamada, K, Tomokuni, M, 2012. Family Tingidae Laporte, 1832. In: Ishikawa, T, Takai, M, Yasunaga, T, Eds., A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Publishing Co., Ltd., Tokyo, Japan: 180 - 213" title="Family Tingidae Laporte, 1832." volumeTitle="A field guide to Japanese bugs." year="2012">Yamada and Tomokuni 2012</bibRefCitation>
; present study); nymphs were collected in March and November (present study).
</paragraph>
</subSubSection>
</treatment>
</document>