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<document id="8428B5ACA276A63FD946D8459D857C24" ID-CLB-Dataset="22520" ID-DOI="10.11646/zootaxa.4683.2.5" ID-GBIF-Dataset="c5a38230-2c50-4d80-9aa9-876d7f687b54" ID-ISSN="1175-5326" ID-Zenodo-Dep="3772795" ID-ZooBank="49879749-226D-460B-BB85-9A4FE160C7EC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588058721948" checkinUser="plazi" docAuthor="Huang, Si-Yao, Tan, Shun-Yun, Wang, Min &amp; Fan, Xiao-Ling" docDate="2019" docId="3A0B795A8227156676BFC52A5154F8DF" docLanguage="en" docName="zootaxa.4683.2.5.pdf" docOrigin="Zootaxa 4683 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paranudina Huang &amp; Tan &amp; Wang &amp; Fan 2019, gen. nov." docType="treatment" docVersion="7" lastPageNumber="273" masterDocId="C6320122822615647628C72B526FFFDB" masterDocTitle="Establishment of a new genus for Nudina xizangensis Fang, 2000 (Lepidoptera, Erebidae, Arctiinae, Lithosiini)" masterLastPageNumber="276" masterPageNumber="271" pageNumber="272" updateTime="1698837752442" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7FEA52D7B76C41FA7FDA67330211FFE2">Establishment of a new genus for Nudina xizangensis Fang, 2000 (Lepidoptera, Erebidae, Arctiinae, Lithosiini)</mods:title>
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<mods:namePart id="DD88431157CAF89EC697D7BFCE01F5A3">Huang, Si-Yao</mods:namePart>
<mods:affiliation id="120C36CB9B703A0C0888C9FEFD887AD5">Department of Entomology, College of Agriculture, South China Agricultural University, Guangzhou 510642, Guangdong, China. Corresponding author. E-mail: fanxiaol 66 @ scau. edu. cn</mods:affiliation>
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<mods:namePart id="A9786A3AE8FCBA431B9201248814273C">Tan, Shun-Yun</mods:namePart>
<mods:affiliation id="7A0470C36CFA568EA7CC33B2A9B76583">Department of Entomology, College of Agriculture, South China Agricultural University, Guangzhou 510642, Guangdong, China. Corresponding author. E-mail: fanxiaol 66 @ scau. edu. cn</mods:affiliation>
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<mods:namePart id="AF755EAD508B3E811E9DDFA961C32DEF">Wang, Min</mods:namePart>
<mods:affiliation id="4E37CB48E692A906BAE6E54B6649459E">Department of Entomology, College of Agriculture, South China Agricultural University, Guangzhou 510642, Guangdong, China. Corresponding author. E-mail: fanxiaol 66 @ scau. edu. cn</mods:affiliation>
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<mods:namePart id="1F4CC8DF4211AB294F9F339418D7EFB6">Fan, Xiao-Ling</mods:namePart>
<mods:affiliation id="DF4D7DE1C31B5EB9C83367CD8D798DB3">Department of Entomology, College of Agriculture, South China Agricultural University, Guangzhou 510642, Guangdong, China. Corresponding author. E-mail: fanxiaol 66 @ scau. edu. cn</mods:affiliation>
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<mods:title id="18D20DF016D8E860D8EC23C725E27ED8">Zootaxa</mods:title>
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<mods:date id="046460061B6976ED15D989B5ECD23A7A">2019</mods:date>
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<mods:number id="C12A50B463803C19F433FFE7ED98B258">2019-10-08</mods:number>
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<mods:number id="471EDB77E075CCD1EAF03F5EBF0F2AE6">4683</mods:number>
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<treatment id="3A0B795A8227156676BFC52A5154F8DF" ID-DOI="http://doi.org/10.5281/zenodo.3797585" ID-GBIF-Taxon="163830137" ID-Zenodo-Dep="3797585" LSID="urn:lsid:plazi:treatment:3A0B795A8227156676BFC52A5154F8DF" httpUri="http://treatment.plazi.org/id/3A0B795A8227156676BFC52A5154F8DF" lastPageId="2" lastPageNumber="273" pageId="1" pageNumber="272">
<subSubSection id="FAB89BC78227156576BFC52A501EFDC0" box="[151,625,513,540]" pageId="1" pageNumber="272" type="nomenclature">
<paragraph id="B21DC84C8227156576BFC52A501EFDC0" blockId="1.[151,625,513,540]" box="[151,625,513,540]" pageId="1" pageNumber="272">
<heading id="E9557F208227156576BFC52A501EFDC0" bold="true" box="[151,625,513,540]" fontSize="11" level="1" pageId="1" pageNumber="272" reason="1">
<emphasis id="80D6145E8227156576BFC52A501EFDC0" bold="true" box="[151,625,513,540]" pageId="1" pageNumber="272">
<taxonomicName id="75A2B3CF8227156576BFC52A5399FDC0" ID-CoL="BF8GB" authority="Huang &amp; Wang" authorityName="Huang &amp; Tan &amp; Wang &amp; Fan" authorityYear="2019" box="[151,502,513,540]" class="Insecta" family="Erebidae" genus="Paranudina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="80D6145E8227156576BFC52A5347FDC7" bold="true" box="[151,296,513,540]" italics="true" pageId="1" pageNumber="272">Paranudina</emphasis>
Huang &amp; Wang
</taxonomicName>
,
<taxonomicNameLabel id="9BE5A92582271565742BC52A501EFDC0" box="[515,625,513,539]" pageId="1" pageNumber="272" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FAB89BC78227156576BFC56D5003FD86" box="[151,620,580,606]" pageId="1" pageNumber="272" type="materials_examined">
<paragraph id="B21DC84C8227156576BFC56D5003FD86" blockId="1.[151,620,580,606]" box="[151,620,580,606]" pageId="1" pageNumber="272">
<typeStatus id="6D1976EE8227156576BFC56D52A6FD85" box="[151,201,582,606]" pageId="1" pageNumber="272">Type</typeStatus>
species:
<taxonomicName id="75A2B3CF82271565770EC56E5007FD86" authority="Fang, 2000" authorityName="Fang" authorityYear="2000" box="[294,616,581,606]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="xizangensis">
<emphasis id="80D6145E82271565770EC56E5383FD86" box="[294,492,581,605]" italics="true" pageId="1" pageNumber="272">Nudina xizangensis</emphasis>
<bibRefCitation id="D633B5BD8227156577DAC56D5007FD86" author="Fang, C. L." box="[498,616,581,606]" pageId="1" pageNumber="272" refId="ref2664" refString="Fang, C. L. (2000) Lepidoptera. Arctiidae. Fauna Sinica (Insecta). Vol. 19. Science Press, Beijing, 590 pp. [in Chinese with English summary]" type="book" year="2000">Fang, 2000</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FAB89BC78227156576BFC5A05312FC82" pageId="1" pageNumber="272" type="diagnosis">
<paragraph id="B21DC84C8227156576BFC5A05312FC82" blockId="1.[151,1437,651,2010]" pageId="1" pageNumber="272">
<emphasis id="80D6145E8227156576BFC5A0537DFD7D" bold="true" box="[151,274,651,678]" pageId="1" pageNumber="272">Diagnosis.</emphasis>
The new genus (figs. 1–4) is characterized by the following autapomorphies: 1) antenna filiform in
<specimenCount id="A4A403C58227156573AFC5A757F3FD7D" box="[1415,1436,652,678]" pageId="1" pageNumber="272" type="male">♂</specimenCount>
and hindwing upperside unmarked. 2) In
<specimenCount id="A4A403C582271565745AC59B50E8FD11" box="[626,647,688,714]" pageId="1" pageNumber="272" type="male">♂</specimenCount>
genitalia (fig. 7) uncus short and broad with rounded apex; juxta Hshaped with long and distally broadened and spiny process; costa long with lower edge fusing with dorsal edge of sacculus and forming a spatulate process and aedeagus significantly narrowing at the middle. 3) In
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genitalia (fig. 9) ductus bursae long, broad and edged by a strongly sclerotized bar anteriorly and corpus bursae large and not fully covered by spinules.
</paragraph>
</subSubSection>
<subSubSection id="FAB89BC78227156576EFC44853CAF9AA" pageId="1" pageNumber="272" type="description">
<paragraph id="B21DC84C8227156576EFC44853FCFBA2" blockId="1.[151,1437,651,2010]" pageId="1" pageNumber="272">
<emphasis id="80D6145E8227156576EFC44853CBFCA5" bold="true" box="[199,420,867,894]" pageId="1" pageNumber="272">Description. Adult</emphasis>
. Length of forewing
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in
<specimenCount id="A4A403C5822715657500C44F5150FCA5" box="[808,831,868,894]" pageId="1" pageNumber="272" type="male">♂</specimenCount>
,
<quantity id="755A65A9822715657565C44851F1FCA5" box="[845,926,867,894]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="272" unit="mm" value="15.0">15 mm</quantity>
in
<specimenCount id="A4A403C58227156575EEC44F51B3FCA5" box="[966,988,868,894]" pageId="1" pageNumber="272" type="female">♀</specimenCount>
. Medium-sized moth. Antenna filiform in both sexes. Forewing venation similar to
<taxonomicName id="75A2B3CF8227156574A6C4A3508FFC79" authorityName="Staudinger" authorityYear="1877" box="[654,736,904,930]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E8227156574A6C4A3508FFC79" box="[654,736,904,930]" italics="true" pageId="1" pageNumber="272">Nudina</emphasis>
</taxonomicName>
. Forewing ground color yellow, with brownish pattern. In
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, a black spot present in discal cell; antemedial band extending from lower portion of discal cell and significantly expanding at dorsum; medial band shifting outwards near lower angle of discal cell, extending from upper angle of discal cell and ending at dorsum; postmedial band running from subapical zone and reaching tornus, with medial portion shifting basally, touching the medial band. In
<specimenCount id="A4A403C58227156574DAC3335168FBE9" box="[754,775,1048,1074]" pageId="1" pageNumber="272" type="female">♀</specimenCount>
most of the pattern similar to male except the medial band and postmedial band fused with each other around tornus. Hindwing ground color slightly paler than forewing, without any markings.
</paragraph>
<paragraph id="B21DC84C8227156576EFC3A853F1FA42" blockId="1.[151,1437,651,2010]" pageId="1" pageNumber="272">
<emphasis id="80D6145E8227156576EFC3A853D7FB45" bold="true" box="[199,440,1155,1182]" pageId="1" pageNumber="272">
<specimenCount id="A4A403C58227156576EFC3A852B3FB45" box="[199,220,1155,1182]" pageId="1" pageNumber="272" type="male">♂</specimenCount>
genitalia. (Fig. 7)
</emphasis>
Uncus short and stout, with a rounded tip. Tegumen broad, triangular. Juxta strongly sclerotized, broad at base, with the lobes developed into long process which becoming blade-like at distal part with several small spines presenting at the surface and edge. Saccus broad, nearly triangular, with tip rounded. Valva broad at base. Cucullus long and oval. Costa short and broad, sclerotized, its lower edge fused with the dorsal edge of sacculus at the base of a well-developed spatulate process, which is covered by numerous small spines. Sacculus long and broad, ending in a trapezoid process directing dorsally, with its dorsal edge covered by numerous spinules. Aedeagus broad and moderate long, narrowing at the middle and expanding at tip. Vesica long and broad, covered by spinules thoroughly.
</paragraph>
<paragraph id="B21DC84C8227156576EFC28853CAF9AA" blockId="1.[151,1437,651,2010]" pageId="1" pageNumber="272">
<emphasis id="80D6145E8227156576EFC28853D7FA65" bold="true" box="[199,440,1443,1470]" pageId="1" pageNumber="272">
<specimenCount id="A4A403C58227156576EFC28852B3FA65" box="[199,220,1443,1470]" pageId="1" pageNumber="272" type="female">♀</specimenCount>
genitalia. (Fig. 9)
</emphasis>
Papilla analis rectangular. Apophysis posteriores and anteriores short, sclerotized, the former about twice the length of the latter.Antevaginal plate strongly sclerotized, concave at the middle portion. Antrum broad and short, strongly sclerotized. Ductus bursae very short and broad, sclerotized and gradually expanding towards corpus bursae and edged by a strongly sclerotized bar anteriorly. Corpus bursae membranous, large and oval, with most of the surface covered by numerous long spinules, leaving a semicircle area on the extreme right side without any spines.
</paragraph>
</subSubSection>
<subSubSection id="FAB89BC78227156576EFC157536DF962" pageId="1" pageNumber="272" type="etymology">
<paragraph id="B21DC84C8227156576EFC157536DF962" blockId="1.[151,1437,651,2010]" pageId="1" pageNumber="272">
<emphasis id="80D6145E8227156576EFC1575321F94D" bold="true" box="[199,334,1660,1686]" pageId="1" pageNumber="272">Etymology.</emphasis>
The genus name is a combination of
<emphasis id="80D6145E8227156574D3C157515CF94E" box="[763,819,1660,1685]" italics="true" pageId="1" pageNumber="272">Para</emphasis>
-(means “resembling” in Latin) and
<taxonomicName id="75A2B3CF8227156572E5C1575770F94D" authorityName="Staudinger" authorityYear="1877" box="[1229,1311,1660,1686]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E8227156572E5C1575770F94D" box="[1229,1311,1660,1686]" italics="true" pageId="1" pageNumber="272">Nudina</emphasis>
</taxonomicName>
. Gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="FAB89BC78227156576EFC1E85135F8DA" pageId="1" pageNumber="272" type="distribution">
<paragraph id="B21DC84C8227156576EFC1E85135F8DA" blockId="1.[151,1437,651,2010]" pageId="1" pageNumber="272">
<emphasis id="80D6145E8227156576EFC1E8530EF905" bold="true" box="[199,353,1731,1758]" pageId="1" pageNumber="272">Distribution.</emphasis>
The only species of the genus is distributed in southern slope of Himalayas, and currently only found in
<collectingRegion id="706606AE8227156576D4C1CC5322F8DA" box="[252,333,1767,1793]" country="China" name="Xizang" pageId="1" pageNumber="272">Xizang</collectingRegion>
(
<collectingRegion id="706606AE822715657774C1CC53F5F8D9" box="[348,410,1767,1794]" country="China" name="Xizang" pageId="1" pageNumber="272">Tibet</collectingRegion>
) Autonomous Prefecture in P. R.
<collectingCountry id="CAB588DC822715657538C1CC5138F8DA" box="[784,855,1767,1793]" name="China" pageId="1" pageNumber="272">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="FAB89BC78227156676EFC0275154F8DF" lastPageId="2" lastPageNumber="273" pageId="1" pageNumber="272" type="discussion">
<paragraph id="B21DC84C8227156676EFC0275653FEC4" blockId="1.[151,1437,651,2010]" lastBlockId="2.[151,1437,153,503]" lastPageId="2" lastPageNumber="273" pageId="1" pageNumber="272">
<emphasis id="80D6145E8227156576EFC0275354F8FD" bold="true" box="[199,315,1804,1830]" pageId="1" pageNumber="272">Remarks.</emphasis>
<taxonomicName id="75A2B3CF82271565776AC02753A7F8FD" authority="Huang &amp; Tan &amp; Wang &amp; Fan, 2019" authorityName="Huang &amp; Tan &amp; Wang &amp; Fan" authorityYear="2019" box="[322,456,1804,1830]" class="Insecta" family="Erebidae" genus="Paranudina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="80D6145E82271565776AC02753A7F8FD" box="[322,456,1804,1830]" italics="true" pageId="1" pageNumber="272">Paranudina</emphasis>
</taxonomicName>
<emphasis id="80D6145E8227156577F8C0275056F8FD" bold="true" box="[464,569,1804,1830]" pageId="1" pageNumber="272">
<taxonomicNameLabel id="9BE5A9258227156577F8C0275056F8FD" box="[464,569,1804,1830]" pageId="1" pageNumber="272" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="75A2B3CF822715657543C02756EBF8FD" authority="Staudinger, 1877" authorityName="Staudinger" authorityYear="1877" box="[875,1156,1803,1830]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E822715657543C02751D2F8FD" box="[875,957,1804,1830]" italics="true" pageId="1" pageNumber="272">Nudina</emphasis>
Staudinger, 1877
</taxonomicName>
by the following combination of characters: 1) antenna filiform in
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(bipectinate in
<taxonomicName id="75A2B3CF822715657565C01B51F0F891" authorityName="Staudinger" authorityYear="1877" box="[845,927,1840,1866]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E822715657565C01B51F0F891" box="[845,927,1840,1866]" italics="true" pageId="1" pageNumber="272">Nudina</emphasis>
</taxonomicName>
); 2) hindwing unmarked in both sexes (with darkish brown shade in both sexes in
<taxonomicName id="75A2B3CF82271565741DC07F50E8F8B5" authorityName="Staudinger" authorityYear="1877" box="[565,647,1876,1902]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E82271565741DC07F50E8F8B5" box="[565,647,1876,1902]" italics="true" pageId="1" pageNumber="272">Nudina</emphasis>
</taxonomicName>
); 3) in
<specimenCount id="A4A403C58227156574FFC07F5083F8B5" box="[727,748,1876,1902]" pageId="1" pageNumber="272" type="male">♂</specimenCount>
genitalia uncus shorter and broader with rounded apex (uncus longer and narrow with apex sharp in
<taxonomicName id="75A2B3CF82271565746EC05350F7F849" authorityName="Staudinger" authorityYear="1877" box="[582,664,1912,1938]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E82271565746EC05350F7F849" box="[582,664,1912,1938]" italics="true" pageId="1" pageNumber="272">Nudina</emphasis>
</taxonomicName>
); 4) juxta with long and distally broadened and spiny process (juxta simple and flat at dorsal edge in
<taxonomicName id="75A2B3CF8227156577D6C0B7503FF86D" authorityName="Staudinger" authorityYear="1877" box="[510,592,1948,1974]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E8227156577D6C0B7503FF86D" box="[510,592,1948,1974]" italics="true" pageId="1" pageNumber="272">Nudina</emphasis>
</taxonomicName>
); 5) costa longer with lower edge fusing together with dorsal edge of sacculus and forming a spatulate process (costa shorter, well separated from sacculus, distal process of sacculus forming a sharp spear-like process and a dorsal digital process in
<taxonomicName id="75A2B3CF822415667523C7B25132FF68" authorityName="Staudinger" authorityYear="1877" box="[779,861,153,179]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E822415667523C7B25132FF68" box="[779,861,153,179]" italics="true" pageId="2" pageNumber="273">Nudina</emphasis>
</taxonomicName>
); 6) aedeagus significantly narrowing at the middle (aedeagus gradually broadening towards tip in
<taxonomicName id="75A2B3CF822415667485C7965090FF0C" authorityName="Staudinger" authorityYear="1877" box="[685,767,189,215]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E822415667485C7965090FF0C" box="[685,767,189,215]" italics="true" pageId="2" pageNumber="273">Nudina</emphasis>
</taxonomicName>
); 7) in
<specimenCount id="A4A403C582241566757FC7965103FF0C" box="[855,876,189,215]" pageId="2" pageNumber="273" type="female">♀</specimenCount>
genitalia ductus bursae much longer and broader (ductus bursae very short and nearly invisible in
<taxonomicName id="75A2B3CF82241566749BC7CA516AFF20" authorityName="Staudinger" authorityYear="1877" box="[691,773,225,251]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E82241566749BC7CA516AFF20" box="[691,773,225,251]" italics="true" pageId="2" pageNumber="273">Nudina</emphasis>
</taxonomicName>
) and 8) corpus bursae much larger and not fully covered by spinules (corpus bursae much smaller and covered with spinules totally in
<taxonomicName id="75A2B3CF8224156675F2C62E5643FEC4" authorityName="Staudinger" authorityYear="1877" box="[986,1068,261,287]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E8224156675F2C62E5643FEC4" box="[986,1068,261,287]" italics="true" pageId="2" pageNumber="273">Nudina</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="B21DC84C8224156676EFC6025778FE2C" blockId="2.[151,1437,153,503]" pageId="2" pageNumber="273">
By the wing pattern, This new genus is also somewhat similar to the genus
<taxonomicName id="75A2B3CF822415667200C602534AFEBC" authority="Dubatolov, Kishida &amp; Wu, 2014" authorityName="Dubatolov, Kishida &amp; Wu" authorityYear="2014" class="Insecta" family="Arctiidae" genus="Paraheliosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E822415667200C60256D6FE98" box="[1064,1209,297,323]" italics="true" pageId="2" pageNumber="273">Paraheliosia</emphasis>
<bibRefCitation id="D633B5BD8224156672E9C602534AFEBC" author="Dubatolov, V. V. &amp; Kishida, Y. &amp; Wu, C. S." pageId="2" pageNumber="273" pagination="373 - 380" refId="ref2597" refString="Dubatolov, V. V., Kishida, Y. &amp; Wu, C. S. (2014) Review of East Asian Heliosia (Lepidoptera: Erebidae: Arctiinae: Lithosiini) species, with description of a new genus. Zootaxa, 3802 (3), 373 - 380. https: // doi. org / 10.11646 / zootaxa. 3802.3.6" type="journal article" year="2014">Dubatolov, Kishida &amp; Wu, 2014</bibRefCitation>
</taxonomicName>
, but
<taxonomicName id="75A2B3CF822415667774C666538DFEBC" authority="Huang &amp; Tan &amp; Wang &amp; Fan, 2019" authorityName="Huang &amp; Tan &amp; Wang &amp; Fan" authorityYear="2019" box="[348,482,333,359]" class="Insecta" family="Erebidae" genus="Paranudina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="80D6145E822415667774C666538DFEBC" box="[348,482,333,359]" italics="true" pageId="2" pageNumber="273">Paranudina</emphasis>
</taxonomicName>
<emphasis id="80D6145E8224156677C2C666503CFEBC" bold="true" box="[490,595,333,359]" pageId="2" pageNumber="273">
<taxonomicNameLabel id="9BE5A9258224156677C2C666503CFEBC" box="[490,595,333,359]" pageId="2" pageNumber="273" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be immediately distinguished from
<taxonomicName id="75A2B3CF822415667232C66656C4FEBC" authorityName="Dubatolov, Kishida &amp; Wu" authorityYear="2014" box="[1050,1195,333,359]" class="Insecta" family="Arctiidae" genus="Paraheliosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E822415667232C66656C4FEBC" box="[1050,1195,333,359]" italics="true" pageId="2" pageNumber="273">Paraheliosia</emphasis>
</taxonomicName>
by
<specimenCount id="A4A403C58224156672FEC6665684FEBC" box="[1238,1259,333,359]" pageId="2" pageNumber="273" type="male">♂</specimenCount>
genitalia uncus stouter with rounded tip (uncus longer and narrow with apex sharp and hooked in
<taxonomicName id="75A2B3CF82241566727CC65A568AFE50" authorityName="Dubatolov, Kishida &amp; Wu" authorityYear="2014" box="[1108,1253,369,395]" class="Insecta" family="Arctiidae" genus="Paraheliosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E82241566727CC65A568AFE50" box="[1108,1253,369,395]" italics="true" pageId="2" pageNumber="273">Paraheliosia</emphasis>
</taxonomicName>
), lobes of juxta thicker and much longer (lobes of juxta much shorter and narrower, forming tooth-like process in
<taxonomicName id="75A2B3CF8224156672D3C6BE57E3FE74" authorityName="Dubatolov, Kishida &amp; Wu" authorityYear="2014" box="[1275,1420,405,431]" class="Insecta" family="Arctiidae" genus="Paraheliosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E8224156672D3C6BE57E3FE74" box="[1275,1420,405,431]" italics="true" pageId="2" pageNumber="273">Paraheliosia</emphasis>
</taxonomicName>
), sacculus ending with a broad and trapezoid process (sacculus ending with long and slender process in
<taxonomicName id="75A2B3CF82241566732AC69257FCFE08" authorityName="Dubatolov, Kishida &amp; Wu" authorityYear="2014" box="[1282,1427,441,467]" class="Insecta" family="Arctiidae" genus="Paraheliosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E82241566732AC69257FCFE08" box="[1282,1427,441,467]" italics="true" pageId="2" pageNumber="273">Paraheliosia</emphasis>
</taxonomicName>
) and aedeagus without any spines laterally (aedeagus with strong spines at lateral wall in
<taxonomicName id="75A2B3CF82241566725EC6F65768FE2C" authorityName="Dubatolov, Kishida &amp; Wu" authorityYear="2014" box="[1142,1287,477,503]" class="Insecta" family="Arctiidae" genus="Paraheliosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E82241566725EC6F65768FE2C" box="[1142,1287,477,503]" italics="true" pageId="2" pageNumber="273">Paraheliosia</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="B21DC84C8224156676BFC1565154F8DF" blockId="2.[151,1436,1661,1796]" pageId="2" pageNumber="273">
<emphasis id="80D6145E8224156676BFC1565321F94C" bold="true" box="[151,334,1661,1688]" pageId="2" pageNumber="273">FIGURES 1–6.</emphasis>
Adults of
<taxonomicName id="75A2B3CF8224156677E3C156503EF94C" authority="Huang &amp; Tan &amp; Wang &amp; Fan, 2019" authorityName="Huang &amp; Tan &amp; Wang &amp; Fan" authorityYear="2019" box="[459,593,1661,1687]" class="Insecta" family="Erebidae" genus="Paranudina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="80D6145E8224156677E3C156503EF94C" box="[459,593,1661,1687]" italics="true" pageId="2" pageNumber="273">Paranudina</emphasis>
</taxonomicName>
<emphasis id="80D6145E822415667472C15550A9F943" bold="true" box="[602,710,1662,1688]" pageId="2" pageNumber="273">
<taxonomicNameLabel id="9BE5A925822415667472C15550A9F943" box="[602,710,1662,1688]" pageId="2" pageNumber="273" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="75A2B3CF822415667528C156513DF94C" authorityName="Staudinger" authorityYear="1877" box="[768,850,1661,1687]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80D6145E822415667528C156513DF94C" box="[768,850,1661,1687]" italics="true" pageId="2" pageNumber="273">Nudina</emphasis>
</taxonomicName>
: 1–4:
<taxonomicName id="75A2B3CF8224156675B7C1565724F943" authority="(Fang, 2000)" authorityName="Huang &amp; Tan &amp; Wang &amp; Fan" authorityYear="2019" baseAuthorityName="Fang" baseAuthorityYear="2000" box="[927,1355,1661,1688]" class="Insecta" family="Erebidae" genus="Paranudina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="species" species="xizangensis" status="comb. nov.">
<emphasis id="80D6145E8224156675B7C15656C0F94C" box="[927,1199,1661,1687]" italics="true" pageId="2" pageNumber="273">Paranudina xizangensis</emphasis>
(
<bibRefCitation id="D633B5BD8224156672E8C155572CF943" author="Fang, C. L." box="[1216,1347,1662,1688]" pageId="2" pageNumber="273" refId="ref2664" refString="Fang, C. L. (2000) Lepidoptera. Arctiidae. Fauna Sinica (Insecta). Vol. 19. Science Press, Beijing, 590 pp. [in Chinese with English summary]" type="book" year="2000">Fang, 2000</bibRefCitation>
)
</taxonomicName>
<emphasis id="80D6145E82241566737CC15552A7F967" bold="true" pageId="2" pageNumber="273">
<taxonomicNameLabel id="9BE5A92582241566737CC15552A7F967" pageId="2" pageNumber="273" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
; 1:
<specimenCount id="A4A403C58224156676DDC1895363F967" box="[245,268,1698,1724]" pageId="2" pageNumber="273" type="male">♂</specimenCount>
,
<typeStatus id="6D1976EE822415667730C18A5315F960" box="[280,378,1697,1723]" pageId="2" pageNumber="273" type="holotype">holotype</typeStatus>
of
<taxonomicName id="75A2B3CF822415667788C18A516EF967" authority="Fang, 2000" authorityName="Fang" authorityYear="2000" box="[416,769,1697,1724]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="species" species="xizangensis">
<emphasis id="80D6145E822415667788C18A5014F960" box="[416,635,1697,1723]" italics="true" pageId="2" pageNumber="273">Nudina xizangensis</emphasis>
<bibRefCitation id="D633B5BD8224156674ABC189516EF967" author="Fang, C. L." box="[643,769,1698,1724]" pageId="2" pageNumber="273" refId="ref2664" refString="Fang, C. L. (2000) Lepidoptera. Arctiidae. Fauna Sinica (Insecta). Vol. 19. Science Press, Beijing, 590 pp. [in Chinese with English summary]" type="book" year="2000">Fang, 2000</bibRefCitation>
</taxonomicName>
(IZCAS), 2:
<specimenCount id="A4A403C58224156675B0C18951C2F967" box="[920,941,1698,1724]" pageId="2" pageNumber="273" type="female">♀</specimenCount>
,
<typeStatus id="6D1976EE822415667593C18A567AF960" box="[955,1045,1697,1723]" pageId="2" pageNumber="273" type="allotype">allotype</typeStatus>
of
<taxonomicName id="75A2B3CF822415667213C18A57F3F967" authority="Fang, 2000" authorityName="Fang" authorityYear="2000" box="[1083,1436,1697,1724]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="species" species="xizangensis">
<emphasis id="80D6145E822415667213C18A5779F960" box="[1083,1302,1697,1723]" italics="true" pageId="2" pageNumber="273">Nudina xizangensis</emphasis>
<bibRefCitation id="D633B5BD822415667336C18957F3F967" author="Fang, C. L." box="[1310,1436,1698,1724]" pageId="2" pageNumber="273" refId="ref2664" refString="Fang, C. L. (2000) Lepidoptera. Arctiidae. Fauna Sinica (Insecta). Vol. 19. Science Press, Beijing, 590 pp. [in Chinese with English summary]" type="book" year="2000">Fang, 2000</bibRefCitation>
</taxonomicName>
(IZCAS), 3:
<specimenCount id="A4A403C582241566770CC1ED5354F93B" box="[292,315,1734,1760]" pageId="2" pageNumber="273" type="male">♂</specimenCount>
, Motuo,
<collectingRegion id="706606AE8224156677B6C1EE5380F904" box="[414,495,1733,1759]" country="China" name="Xizang" pageId="2" pageNumber="273">Xizang</collectingRegion>
(SCAU), 4:
<specimenCount id="A4A403C5822415667454C1ED50FEF93B" box="[636,657,1734,1760]" pageId="2" pageNumber="273" type="female">♀</specimenCount>
,
<emphasis id="80D6145E8224156674B7C1EE50BDF904" box="[671,722,1733,1759]" italics="true" pageId="2" pageNumber="273">ditto</emphasis>
(SCAU). 5–6:
<taxonomicName id="75A2B3CF822415667553C1EE568EF93B" authority="(Butler, 1881)" baseAuthorityName="Butler" baseAuthorityYear="1881" box="[891,1249,1733,1760]" class="Insecta" family="Arctiidae" genus="Nudina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="species" species="artaxidia">
<emphasis id="80D6145E822415667553C1EE5654F904" box="[891,1083,1733,1759]" italics="true" pageId="2" pageNumber="273">Nudina artaxidia</emphasis>
(
<bibRefCitation id="D633B5BD822415667260C1EE56B7F93B" author="Butler, A. G." box="[1096,1240,1733,1760]" pageId="2" pageNumber="273" pagination="401 - 426" refId="ref2474" refString="Butler, A. G. (1881) Descriptions of new Genera and Species of Heterocerous Lepidoptera from Japan. Transactions of the Entomological Society of London, 1881 (1 - 4), 1 - 23, 171 - 200, 401 - 426 &amp; 579 - 600. https: // doi. org / 10.1111 / j. 1365 - 2311.1873. tb 01519. x" type="journal article" year="1881">Butler, 1881</bibRefCitation>
)
</taxonomicName>
; 5:
<specimenCount id="A4A403C5822415667320C1ED5770F93B" box="[1288,1311,1734,1760]" pageId="2" pageNumber="273" type="male">♂</specimenCount>
, Xingshan,
<collectingRegion id="706606AE8224156676BFC1C252B4F8D8" box="[151,219,1769,1795]" country="China" name="Hubei" pageId="2" pageNumber="273">Hubei</collectingRegion>
(SCAU), 6:
<specimenCount id="A4A403C5822415667743C1C153EFF8DF" box="[363,384,1770,1796]" pageId="2" pageNumber="273" type="female">♀</specimenCount>
, Qujing,
<collectingRegion id="706606AE8224156677C0C1C15051F8DF" box="[488,574,1770,1796]" country="China" name="Yunnan" pageId="2" pageNumber="273">Yunnan</collectingRegion>
(SCAU). Scale=
<quantity id="755A65A982241566752AC1C25157F8DF" box="[770,824,1769,1796]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="273" unit="cm" value="1.0">1 cm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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