treatments-xml/data/BA/D0/E8/BAD0E8B5EBE2537383E489325B6503B3.xml
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<document ID-DOI="http://dx.doi.org/10.5091/plecevo.108472" ID-Pensoft-Pub="2032-3921-1-20" ID-Pensoft-UUID="9494A62B95C65323B2170E11C1379AD3" ModsDocID="2032-3921-157-1-20" checkinTime="1705544489675" checkinUser="pensoft" docAuthor="Flores-Argueelles, Alejandra, Hernandez-Cardenas, Rodrigo Alejandro, Rosales-Martinez, Santiago, Espejo-Serna, Adolfo, Lopez-Ferrari, Ana Rosa &amp; Siekkinen, Andrew" docDate="2024" docId="BAD0E8B5EBE2537383E489325B6503B3" docLanguage="en" docName="PlantEcolEvol 157(1): 20-31" docOrigin="Plant Ecology and Evolution 157 (1)" docPubDate="2024-01-17" docSource="http://dx.doi.org/10.5091/plecevo.108472" docTitle="Hechtia reticulata L. B. Sm. (Smith 1937" docType="treatment" docVersion="2" id="9494A62B95C65323B2170E11C1379AD3" lastPageNumber="20" masterDocId="9494A62B95C65323B2170E11C1379AD3" masterDocTitle="Two new species of Hechtia (Bromeliaceae; Hechtioideae) from Jalisco, Mexico" masterLastPageNumber="31" masterPageNumber="20" pageNumber="20" updateTime="1705544790684" updateUser="ExternalLinkService">
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<mods:title id="07CF2B2571054CB93E2BED122A0196B9">Two new species of Hechtia (Bromeliaceae; Hechtioideae) from Jalisco, Mexico</mods:title>
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<mods:namePart id="1B592197BB47D470FA1CD162D93EB794">Flores-Argueelles, Alejandra</mods:namePart>
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<mods:affiliation id="EA13A2DCF604AAE6F79941E6C5E7E18E">Departamento de Botanica y Zoologia, Centro Universitario de Ciencias Biologicas y Agropecuarias, Universidad de Guadalajara, Zapopan, Jalisco, Mexico</mods:affiliation>
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<mods:namePart id="30C91C41E4489C1B913BB1DF302132D2">Hernandez-Cardenas, Rodrigo Alejandro</mods:namePart>
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<mods:affiliation id="DD3B9564D18B997DA8E0F4F268E29968">Herbario Metropolitano, Departamento de Biologia, Division de Ciencias Biologicas y de Salud, Universidad Autonoma Metropolitana-Iztapalapa, Iztapalapa, Ciudad de Mexico, Mexico</mods:affiliation>
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<mods:namePart id="E7FA256944ED2111378B19C6BEE22EC8">Rosales-Martinez, Santiago</mods:namePart>
<mods:affiliation id="5507973E3474FA04B46F13BEB1245917">Vallarta Norte, Justo Sierra 2807, Guadalajara, Jalisco, Mexico</mods:affiliation>
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<mods:namePart id="115047F9D5B439F5E075DAEAB1DCDF56">Espejo-Serna, Adolfo</mods:namePart>
<mods:affiliation id="0DE8483E429A71CB96EE81A11A55AF2C">Herbario Metropolitano, Departamento de Biologia, Division de Ciencias Biologicas y de Salud, Universidad Autonoma Metropolitana-Iztapalapa, Iztapalapa, Ciudad de Mexico, Mexico</mods:affiliation>
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<mods:namePart id="BD294ED79350E55C2B59FBE559DAB360">Lopez-Ferrari, Ana Rosa</mods:namePart>
<mods:affiliation id="B12EBE5CC4370CFF2BED54DFDFF1A58D">Herbario Metropolitano, Departamento de Biologia, Division de Ciencias Biologicas y de Salud, Universidad Autonoma Metropolitana-Iztapalapa, Iztapalapa, Ciudad de Mexico, Mexico</mods:affiliation>
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<mods:namePart id="49F6C5E80028F0A1267D4109318E20B9">Siekkinen, Andrew</mods:namePart>
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<mods:date id="5EEF42E300F535A0684783D7D49075D9">2024</mods:date>
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<treatment id="BAD0E8B5EBE2537383E489325B6503B3" LSID="urn:lsid:plazi:treatment:BAD0E8B5EBE2537383E489325B6503B3" httpUri="http://treatment.plazi.org/id/BAD0E8B5EBE2537383E489325B6503B3" lastPageNumber="20" pageId="0" pageNumber="20">
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<paragraph id="24305001E14CE99AAC7BD71B119EAE28" pageId="0" pageNumber="20">
<taxonomicName id="172AE16A7328105ABB351937DA3E159B" LSID="BAD0E8B5-EBE2-5373-83E4-89325B6503B3" authority="L. B. Sm. (Smith 1937)" authorityName="L. B. Sm. (Smith" authorityYear="1937" class="Liliopsida" family="Bromeliaceae" genus="Hechtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hechtia reticulata" order="Poales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="reticulata">Hechtia reticulata L.B.Sm. (Smith 1937)</taxonomicName>
</paragraph>
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<subSubSection id="48FE9EE8FDBD9EC9EF941344CE918B14" pageId="0" pageNumber="20" type="description">
<paragraph id="AE2A063BC8D2961FACB3937BEA5CA2E3" pageId="0" pageNumber="20">
<figureCitation id="B523A28FD4989081C0334F86C8ABC6A7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type specimens of Hechtia reticulata. A. E. Palmer 1352 (holotype: GH). B. E. Palmer 1352 (isotype: US). C - D. E. Palmer 1352 (isotypes: F)." figureDoi="10.5091/plecevo.108472.figure1" httpUri="https://binary.pensoft.net/fig/964343" pageId="0" pageNumber="20">Figs 1</figureCitation>
<figureCitation id="3B6788FA15C1B852B616E2297A3ECF72" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of Hechtia chamelensis, H. costalegrensis, H. ibugana, H. iltisii, H. jaliscana, H. laevis, H. pacifica, H. pedicellata, H. reticulata, H. santanae, and H. subalata. Biogeographic provinces according to Morrone et al. (2017)." figureDoi="10.5091/plecevo.108472.figure3" httpUri="https://binary.pensoft.net/fig/964345" pageId="0" pageNumber="20">, 3</figureCitation>
<figureCitation id="1055CCDD8AE6DD1C2A7223ED8173B24C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hechtia reticulata. A - C. Plants at the type locality. D. Detail of the rosettes. E. Detail of the staminate spike. F. Staminate flower. F 1. Floral bract. F 2. Sepals. F 3. Petals. F 4. Stamens. G. Detail of the pistillate spike. H. Pistillate flower. H 1. Floral bract. H 2. Sepals. H 3. Petals. H 4. Pistil. Photographs A-C by Alejandra Flores-Argueelles; D-H by Rodrigo Hernandez-Cardenas." figureDoi="10.5091/plecevo.108472.figure5" httpUri="https://binary.pensoft.net/fig/964347" pageId="0" pageNumber="20">, 5</figureCitation>
</paragraph>
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<subSubSection id="BE955CF0AE51191978584CF5F7DE98CE" pageId="0" pageNumber="20" type="materials_examined">
<paragraph id="C729900510F04C96F089AAD0C957ABBE" pageId="0" pageNumber="20">
<typeStatus id="3D0824AD54BDEAEAD4FFF768C000F82D">Type</typeStatus>
.
</paragraph>
<paragraph id="53D9D29FDD789F068D8326A88A263F71" pageId="0" pageNumber="20">
<materialsCitation id="0237694E1914904E3DA8415121DB8451" collectingDate="1891-03-02" collectingDateMax="1891-03-18" collectingDateMin="1891-03-02" collectorName="E. Palmer" country="Mexico" location="Manzanillo" specimenCount="1" stateProvince="Colima" typeStatus="holotype">
<collectingCountry id="03BC697C3FDB5C339BC0DFA6B3F18A2A" name="Mexico">Mexico</collectingCountry>
-
<emphasis id="808BE0A2802A739136C13DC37B389807" bold="true" pageId="0" pageNumber="20">
<collectingRegion id="20BF4DE63E561C3F11C7EA88D4CC87C9" country="Mexico" name="Colima">Colima</collectingRegion>
</emphasis>
<location id="8ACF7209E1972E2A91DF4C29D0C89B5B" LSID="urn:lsid:plazi:treatment:BAD0E8B5EBE2537383E489325B6503B3:8ACF7209E1972E2A91DF4C29D0C89B5B" country="Mexico" name="Manzanillo" stateProvince="Colima">Manzanillo</location>
[along the ocean just above high tide and extending back to the base of the mountains,
<collectingDate id="C7055624401C4CC1E02988B57A1FEE90" value="1891-03-02" valueMax="1891-03-18" valueMin="1891-03-02">2 to 18 Mar. 1891</collectingDate>
(see
<bibRefCitation id="D68556F61B13F1949C3ADBF0CF4EC279" author="Rose, JN" journalOrPublisher="Contributions from the United State National Herbarium" pageId="0" pageNumber="20" pagination="293 - 366" publicationUrl="https://www.jstor.org/stable/23490732" refId="B16" refString="Rose, JN, 1895. Report on a collection of plants made in the States of Sonora and Colima, Mexico, by Dr. Edward Palmer, in the years 1890 and 1891. Contributions from the United State National Herbarium 1: 293 - 366, https://www.jstor.org/stable/23490732" title="Report on a collection of plants made in the States of Sonora and Colima, Mexico, by Dr. Edward Palmer, in the years 1890 and 1891." url="https://www.jstor.org/stable/23490732" volume="1" year="1895">Rose 1895</bibRefCitation>
)];
<emphasis id="32E64A36D6D524B7F58C3C72F05AD6CC" italics="true" pageId="0" pageNumber="20">
<collectorName id="A53CA41FF582E359A592AB57948B27E0">E. Palmer</collectorName>
1352
</emphasis>
;
<typeStatus id="FDCBC39546BF93D869468F1B153BB3E1" type="holotype">holotype</typeStatus>
: GH [GH00275619];
<typeStatus id="CC94AEA3C09C5318C6858F8BE0FC465D" type="isotype">isotypes</typeStatus>
: F [V0077707F, V0077708F]
</materialsCitation>
,
<collectingCountry id="FD7A1756A4CB06B2B7B0A135AF597929" name="United States of America">US</collectingCountry>
[00089102, 01095364].
</paragraph>
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<subSubSection id="77CD58C75053C37D3A711388787376F0" pageId="0" pageNumber="20" type="epitype (designated here)">
<paragraph id="15F65FF45BC4DE83BC67C7B0CE268516" pageId="0" pageNumber="20">Epitype (designated here).</paragraph>
<paragraph id="B2CE94DE160C15D563A06F30D7FADB8A" pageId="0" pageNumber="20">
Mexico -
<emphasis id="ED3E4369C56874F733A79B9A07734750" bold="true" pageId="0" pageNumber="20">Colima</emphasis>
• Manzanillo, ca 3 km sobre el camino La
<normalizedToken id="03E18962639272FE90FD79325B79D4EB" originalValue="Central-Peña">Central-Pena</normalizedToken>
Blanca, a partir de la carretera
<normalizedToken id="7ED42255280869C46CCAD4EE0C99AC9D" originalValue="Manzanillo-Cihuatlán">Manzanillo-Cihuatlan</normalizedToken>
;
<geoCoordinate id="052369342D1A04FBEC04B7E643087312" degrees="19" direction="north" minutes="07" orientation="latitude" precision="1" seconds="57.9" value="19.13275">
19°07
<normalizedToken id="ADD50C14A8BFACD0EFE84631674C28FF" originalValue="57.9”">'57.9&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="08857FE775BD08E240265B638E3D9024" degrees="104" direction="west" minutes="26" orientation="longitude" precision="1" seconds="54.6" value="-104.4485">
104°26
<normalizedToken id="55AE78C5ADED3DDD4BD5EC681EA321DE" originalValue="54.6”">'54.6&quot;</normalizedToken>
W
</geoCoordinate>
; 200 m; 19 Jul. 2022;
<emphasis id="1C215DF67D23DC49DE1BF5CDFA49CF7A" italics="true" pageId="0" pageNumber="20">
R.
<normalizedToken id="4548903476878614A9AEAC497F727857" originalValue="Hernández-Cárdenas">Hernandez-Cardenas</normalizedToken>
</emphasis>
,
<emphasis id="9D0420ECE96CCCE02A4F12B1E9975FE0" italics="true" pageId="0" pageNumber="20">
A.
<normalizedToken id="553E64386512E712EAF2204A8D6C7AA5" originalValue="Flores-Argüelles">Flores-Argueelles</normalizedToken>
&amp; S.
<normalizedToken id="2853B978E1795EA94733FC748862C5A3" originalValue="Lara-Godínez">Lara-Godinez</normalizedToken>
2618
</emphasis>
♀; UAMIZ [8 sheets: UAMIZ89782, UAMIZ89783, UAMIZ89784, UAMIZ89785, UAMIZ89786, UAMIZ89787, UAMIZ89788, UAMIZ89789].
</paragraph>
</subSubSection>
<subSubSection id="34A3AA1797DA3987DBA4B4AED335948F" pageId="0" pageNumber="20" type="description">
<paragraph id="EE1397F48A325000EFA7D9F79E942525" pageId="0" pageNumber="20">Description.</paragraph>
<paragraph id="F1C5298B5CD29E2AADA11CCA9F14701E" pageId="0" pageNumber="20">Plants saxicolous or terrestrial, in flower 170-360 cm tall, rosettes 80-100 cm tall, 60-80 cm diameter, solitary or forming clumps of 3-6 rosettes. Leaves 10-20, recurved towards the apex; sheaths purplish-grey, whitish-brown to yellowish-white when dry, widely depressed ovate to widely ovate, 7-10 cm long, 7-11 cm wide, with minute marginal sharp teeth, glabrous near the base and lepidote distally on the abaxial surface, glabrous on the adaxial surface; blades purplish-green to greyish-purple, pale brown when dry, narrowly triangular, 65-85 cm long, 3.5-5.5 cm wide at the base, long attenuate, lepidote on the abaxial surface, glabrous on the adaxial surface, margin with divaricate to ascending sharp teeth, yellowish-brown, 5-7 mm long, 4-7 mm wide, 2-3 cm apart. Inflorescences terminal, erect, once branched in pistillate plants, twice branched in staminate plants. Staminate inflorescence 190-210 cm tall; peduncle green, brown when dry, terete, 1.1-1.3 cm diameter, sparsely lepidote, internodes 5-8 cm long; peduncle bracts whitish-grey to greenish-grey, whitish-brown when dry, foliaceous, sheaths triangular, 1-2 cm long, 1.5-3 cm wide, lepidote on both surfaces, hyaline at the margin and with minute marginal sharp teeth, blades linear, 4-16 cm long, 0.6-1 cm wide, decreasing in size distally, lepidote on both surfaces, entire and spinose, the basal ones larger than the internodes, the upper ones shorter; primary bracts greenish-brown, whitish-brown when dry, triangular to narrowly triangular, 1.5-4 cm long, 0.4-1 cm wide when extended, caudate, entire, hyaline, erose to spinose at the margin, lepidote on both surfaces; primary spikes 35-40, terete, 15-25 cm long, 1.2-1.5 cm in diameter; secondary spikes 40-50, terete, 6-10 cm long, 1.2-1.5 cm in diameter; floral bracts brown in living material and dry, triangular to widely ovate, 3.8-4.2 mm long, 2.5-3 mm wide, longer than the pedicels, apiculate to acute, entire to erose at the margin, glabrous on both surfaces. Staminate flowers numerous; pedicels 0.8-1.1 mm long; sepals greenish-brown, whitish-brown when dry, ovate to widely ovate, 2.3-2.7 mm long, 2-2.3 mm wide, obtuse, entire, glabrous on both surfaces; petals white, whitish-brown when dry, oblong, 4.5-4.8 mm long, 3.1-3.5 mm wide, obtuse at the apex, entire, glabrous on both surfaces; stamens equal in length; filaments white, narrowly oblong, flattened, 4.1-4.7 mm long; anthers green, brown to pale green when dry, oblong, 1.8-2.2 mm long, versatile; pistillode conspicuous, white rose, brown when dry, glabrous. Pistillate inflorescence 250-260 cm long; peduncle green, brown when dry, terete, 1.9-2 cm in diameter, glabrous, internodes 2-6 cm long; peduncle bracts greenish-grey to brownish-grey, greyish-white when dry, foliaceous, sheaths triangular, 1-2.5 cm long, 1-2 cm wide, glabrous on both surfaces, hyaline and spinose at the margin, blades linear to narrowly oblong, lepidote on both surfaces, entire, the basal ones larger than the internodes, the upper ones shorter; primary bracts brown, triangular, 1.5-3 cm long, 0.4-0.6 cm wide when extended, caudate, entire, hyaline and spinose at the margin, glabrous on the adaxial surface, lepidote on the abaxial surface; primary spikes 30-40, terete, 8-30 cm long, 1.5-1.8 cm in diameter; floral bracts whitish-green, whitish-brown when dry, triangular, 3.7-4.2 mm long, 2.9-3.3 mm wide, longer than the pedicels, acute, entire to erose at the margin, glabrous on the adaxial surface, lepidote on the abaxial surface. Pistillate flowers numerous; pedicels 2-2.5 mm long, sulcate; sepals brownish-green in living material, whitish-brown when dry, very widely ovate, 2.3-2.7 mm long, 2.2-2.5 mm wide, acute, entire, glabrous on both surfaces; petals white, brown when dry, lanceolate, 4.2-4.8 mm long, 1.6-2 mm wide, acuminate, entire, glabrous on both surfaces; staminodes rudimentary, white, narrowly triangular, 3.2-3.6 mm long; ovary superior, green, brown when dry, oblong, 4-4.4 mm long, 2-2.4 mm diameter, glabrous; stylar branches white, brown when dry, recurved, slender, 1.7-2 mm long, stigma papillose. Capsules brownish-dark when dry, ovoid, 7.5-11 mm long, 3-6.4 mm in diameter, with prominent longitudinal ridges or creases, glabrous; seeds reddish-brown, fusiform, 3.7-6.4 mm long, caudate.</paragraph>
</subSubSection>
<subSubSection id="5D927AE4B59EF0D2E8CC7A261DA65E12" pageId="0" pageNumber="20" type="distribution">
<paragraph id="CE93654473C613A03E4CF145B471171B" pageId="0" pageNumber="20">Distribution, habitat, and ecology.</paragraph>
<paragraph id="3749988513CE37EDCF904CDF92F884D7" pageId="0" pageNumber="20">
<taxonomicName id="5BF6A02040564585D6BB30BA3DA94EB1" authorityName="L. B. Sm. (Smith" authorityYear="1937" class="Liliopsida" family="Bromeliaceae" genus="Hechtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hechtia reticulata" order="Poales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="AAC71430134DDBCAABF0CC645D57A2DF" italics="true" pageId="0" pageNumber="20">Hechtia reticulata</emphasis>
</taxonomicName>
is only known from the municipality of Manzanillo in the state of Colima and belongs to the Pacific Lowlands biogeographic province (Figs
<figureCitation id="DB3EADD3B0D0E396024FAEDF9A2D539B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of Hechtia chamelensis, H. costalegrensis, H. ibugana, H. iltisii, H. jaliscana, H. laevis, H. pacifica, H. pedicellata, H. reticulata, H. santanae, and H. subalata. Biogeographic provinces according to Morrone et al. (2017)." figureDoi="10.5091/plecevo.108472.figure3" httpUri="https://binary.pensoft.net/fig/964345" pageId="0" pageNumber="20">3</figureCitation>
,
<figureCitation id="080928ABB4A76C3CE6452DC332535899" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hechtia reticulata. A - C. Plants at the type locality. D. Detail of the rosettes. E. Detail of the staminate spike. F. Staminate flower. F 1. Floral bract. F 2. Sepals. F 3. Petals. F 4. Stamens. G. Detail of the pistillate spike. H. Pistillate flower. H 1. Floral bract. H 2. Sepals. H 3. Petals. H 4. Pistil. Photographs A-C by Alejandra Flores-Argueelles; D-H by Rodrigo Hernandez-Cardenas." figureDoi="10.5091/plecevo.108472.figure5" httpUri="https://binary.pensoft.net/fig/964347" pageId="0" pageNumber="20">5</figureCitation>
). It grows on rocky slopes in tropical deciduous forests forming colonies along with
<taxonomicName id="6C60FAF3CE8973BAD0722E0784CEE50A" class="Liliopsida" family="Agavaceae" genus="Agave" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agave colimana" order="Agavales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="colimana">
<emphasis id="035FED013F578EF4DB8D0028F7A1392F" italics="true" pageId="0" pageNumber="20">Agave colimana</emphasis>
</taxonomicName>
,
<taxonomicName id="E9F85E42993287B15CDFB4761CBC2FF9" class="Equisetopsida" family="Bixaceae" genus="Cochlospermum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cochlospermum vitifolium" order="Malvales" pageId="0" pageNumber="20" phylum="Magnoliophyta" rank="species" species="vitifolium">
<emphasis id="B40CE77D3D0F2E02E667C55EA3877C43" italics="true" pageId="0" pageNumber="20">Cochlospermum vitifolium</emphasis>
</taxonomicName>
(Willd.) Spreng.,
<taxonomicName id="865B39BB553870BC51921BE451CA4D06" class="Magnoliopsida" family="Cactaceae" genus="Pilosocereus" kingdom="Plantae" lsidName="Pilosocereus purpusii" order="Caryophyllales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="purpusii">
<emphasis id="3CED7E0574079312C2B606E322B5E9CE" italics="true" pageId="0" pageNumber="20">Pilosocereus purpusii</emphasis>
</taxonomicName>
(Britton &amp; Rose) F.M.Knuth, and other species at elevations from sea level to around 200 m.
</paragraph>
<caption id="846F0630937417985113BAB356F30B81" doi="10.5091/plecevo.108472.figure5" httpUri="https://binary.pensoft.net/fig/964347" pageId="0" pageNumber="20" start="Figure 5" startId="F5">
<paragraph id="FC61BA9B07A6393B52BEA6E6DBF6D705" pageId="0" pageNumber="20">
<emphasis id="3C762E3F1B2141303CD2AA3AB5039D70" bold="true" pageId="0" pageNumber="20">Figure 5.</emphasis>
<taxonomicName id="E5E122EDC0A35F14FB915DA6500564A7" authorityName="L. B. Sm. (Smith" authorityYear="1937" class="Liliopsida" family="Bromeliaceae" genus="Hechtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hechtia reticulata" order="Poales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="5CB874A7F71CEB1B10FEAC656C4F0E51" italics="true" pageId="0" pageNumber="20">Hechtia reticulata</emphasis>
</taxonomicName>
.
<emphasis id="D52047B1803CCABA03A5B312543E03A5" bold="true" pageId="0" pageNumber="20">A</emphasis>
-
<emphasis id="9E82A7A7D258B6A5C7AC0467A1FE0387" bold="true" pageId="0" pageNumber="20">C</emphasis>
. Plants at the type locality.
<emphasis id="52F425985BC75CC84709E8D9C9F87E15" bold="true" pageId="0" pageNumber="20">D</emphasis>
. Detail of the rosettes.
<emphasis id="1B49EB8F7442C3823DDAFA3919C75093" bold="true" pageId="0" pageNumber="20">E</emphasis>
. Detail of the staminate spike.
<emphasis id="3E077281E80200C5DD0BEABFE510DD89" bold="true" pageId="0" pageNumber="20">F</emphasis>
. Staminate flower.
<emphasis id="027448D2B116524D829FE56E40CE0618" bold="true" pageId="0" pageNumber="20">F1</emphasis>
. Floral bract.
<emphasis id="1426D138D31ECE2026A18B7E3950B540" bold="true" pageId="0" pageNumber="20">F2</emphasis>
. Sepals.
<emphasis id="57E8C89EAD1A6C6077986D073DA166C9" bold="true" pageId="0" pageNumber="20">F3</emphasis>
. Petals.
<emphasis id="B8EDDAD4350866F65BA6A8625F4CC2EA" bold="true" pageId="0" pageNumber="20">F4</emphasis>
. Stamens.
<emphasis id="9389EF2C79784EAC4DA990FB31A7BEF7" bold="true" pageId="0" pageNumber="20">G</emphasis>
. Detail of the pistillate spike.
<emphasis id="090F1F637550C0A5D80FDB938821B294" bold="true" pageId="0" pageNumber="20">H</emphasis>
. Pistillate flower.
<emphasis id="B66E7C69BA6C5A644138D3250F407842" bold="true" pageId="0" pageNumber="20">H1</emphasis>
. Floral bract.
<emphasis id="FDB8DDE3B7D565C4577F86ECC03BFF50" bold="true" pageId="0" pageNumber="20">H2</emphasis>
. Sepals.
<emphasis id="50B4FA8368AFB98E2D5DB7447B70AC72" bold="true" pageId="0" pageNumber="20">H3</emphasis>
. Petals.
<emphasis id="B114F0900931769A803D8461AA19858E" bold="true" pageId="0" pageNumber="20">H4</emphasis>
. Pistil. Photographs A-C by Alejandra
<normalizedToken id="0B5AADFA4733972D4B41EAA2941B7297" originalValue="Flores-Argüelles">Flores-Argueelles</normalizedToken>
; D-H by Rodrigo
<normalizedToken id="E14457FE8F57518DF0BDC95F5975E70D" originalValue="Hernández-Cárdenas">Hernandez-Cardenas</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E8FD373907279C9976E58711A4026C8A" pageId="0" pageNumber="20" type="phenology">
<paragraph id="D4FCAA87C3EEED065E950908BA1C9C24" pageId="0" pageNumber="20">Phenology.</paragraph>
<paragraph id="768A050FE5DC35D61045876C333A3114" pageId="0" pageNumber="20">
<taxonomicName id="DEFFE14B4B30D7F3976E0B94CA052890" authorityName="L. B. Sm. (Smith" authorityYear="1937" class="Liliopsida" family="Bromeliaceae" genus="Hechtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hechtia reticulata" order="Poales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="6CC6A3BCE77DE8D8B19887A33C52C914" italics="true" pageId="0" pageNumber="20">Hechtia reticulata</emphasis>
</taxonomicName>
blooms from July to August.
</paragraph>
</subSubSection>
<subSubSection id="95A380084D6D1AFBFDF566C62F04D040" pageId="0" pageNumber="20" type="materials_examined">
<paragraph id="53215DADDE3036BBF06F623C761BCAAD" pageId="0" pageNumber="20">Additional material examined.</paragraph>
<paragraph id="0D1CA387428D98356D8996952294E4E5" pageId="0" pageNumber="20">
<materialsCitation id="09FD6CF65D4E82675077990F39A08748" collectingDate="1925-11-28" collectingDateMax="2023-05-06" collectingDateMin="1925-11-28" collectorName="R. S. Ferris, La Central-Pena Blanca, R. Hernandez-Cardenas, A. Flores-Argueelles, S. Lara-Godinez, G. Contreras-Felix" country="MEXICO" county="Rocky" elevation="200" latitude="19.13275" location="Manzanillo" longLatPrecision="1" longitude="-104.4485" municipality="Manzanillo Bay" specimenCount="10465" specimenCount-female="7846" specimenCount-male="2619" stateProvince="Colima">
<collectingCountry id="DA931A5AA81927105EB6E9B1631CDEC4" name="Mexico">MEXICO</collectingCountry>
-
<emphasis id="04CDABCF271ADA02A26E396E39852D6A" bold="true" pageId="0" pageNumber="20">
<collectingRegion id="F39ACDF97C894FB414BE3A353A4C0040" country="Mexico" name="Colima">Colima</collectingRegion>
</emphasis>
<collectingCounty id="DFB7CDF83991853697B69A2A57CBEA1D">Rocky</collectingCounty>
cliffs southern end of
<collectingMunicipality id="FC9CA7A39A5AF7FBD1D8BE60137D1555">Manzanillo Bay</collectingMunicipality>
, vicinity of
<location id="B517E126664E14FEAD775640E1B40460" LSID="urn:lsid:plazi:treatment:BAD0E8B5EBE2537383E489325B6503B3:B517E126664E14FEAD775640E1B40460" country="MEXICO" county="Rocky" latitude="19.13275" longLatPrecision="1" longitude="-104.4485" municipality="Manzanillo Bay" name="Manzanillo" stateProvince="Colima">Manzanillo</location>
;
<collectingDate id="8254FD1466E93EF148A33AC7183BD549" value="1925-11-28">28 Nov. 1925</collectingDate>
;
<emphasis id="9EDCF3506044CC8A2176711E303A0024" italics="true" pageId="0" pageNumber="20">
<collectorName id="9C789A6BF24AC5A65CC6DB577F307585">R.S. Ferris</collectorName>
<specimenCount id="6DE848E248C457A4B7E5391EAD85B595" type="female">6111 ♀</specimenCount>
</emphasis>
; GH •
<location id="EDBC911D20356BC2383E0AF69D6DC93E" LSID="urn:lsid:plazi:treatment:BAD0E8B5EBE2537383E489325B6503B3:EDBC911D20356BC2383E0AF69D6DC93E" country="MEXICO" county="Rocky" latitude="19.13275" longLatPrecision="1" longitude="-104.4485" municipality="Manzanillo Bay" name="Manzanillo" stateProvince="Colima">Manzanillo</location>
, ca
<quantity id="E035058E18B13EF23330EE1AF2F9B5A2" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
sobre el camino
<collectorName id="0E2519C0D11B6473E0E3BE80FB920FF7">
<location id="1FB34A8361C6CE69A29A6A32147CE5FF" LSID="urn:lsid:plazi:treatment:BAD0E8B5EBE2537383E489325B6503B3:1FB34A8361C6CE69A29A6A32147CE5FF" country="MEXICO" county="Rocky" latitude="19.13275" longLatPrecision="1" longitude="-104.4485" municipality="Manzanillo Bay" name="La Central-Pena Blanca" stateProvince="Colima">
La
<normalizedToken id="1E5F71140666D192BF3684D6DBB9CBC8" originalValue="Central-Peña">Central-Pena</normalizedToken>
Blanca
</location>
</collectorName>
, a partir de la carretera
<normalizedToken id="740B0284C5DBD7F22A2D76A069668715" originalValue="Manzanillo-Cihuatlán">
<location id="5AF84DBAD9157139F26DA31515DEBAF3" LSID="urn:lsid:plazi:treatment:BAD0E8B5EBE2537383E489325B6503B3:5AF84DBAD9157139F26DA31515DEBAF3" country="MEXICO" county="Rocky" latitude="19.13275" longLatPrecision="1" longitude="-104.4485" municipality="Manzanillo Bay" name="Manzanillo-Cihuatlan" stateProvince="Colima">Manzanillo-Cihuatlan</location>
</normalizedToken>
;
<geoCoordinate id="B965459ADC9CCDE00FEE74AC69E4A831" degrees="19" direction="north" minutes="07" orientation="latitude" precision="1" seconds="57.9" value="19.13275">
19°07
<normalizedToken id="F454D150D9E4C12BA9A7564E83CF160F" originalValue="57.9”">'57.9&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="A15C2A7FB70CB1D002428DD942D0524D" degrees="104" direction="west" minutes="26" orientation="longitude" precision="1" seconds="54.6" value="-104.4485">
104°26
<normalizedToken id="2350F982F3BD933F5D294A09D5524796" originalValue="54.6”">'54.6&quot;</normalizedToken>
W
</geoCoordinate>
;
<quantity id="2E59FE66728EB2573A51139DE48F2008" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation id="7E2CBF965F1DC22A4EB60FF4BAEA4164" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
;
<collectingDate id="41D85689908B6A5B9A685508BFBDBD3D" value="2022-07-19">19 Jul. 2022</collectingDate>
;
<emphasis id="9CC696831ABE94BC7C9F6380462CB2AC" italics="true" pageId="0" pageNumber="20">
<collectorName id="F041567FF0334954980CD30C3DC7F4A5">
R.
<normalizedToken id="A229C19549941FC9634693A7D20B2B09" originalValue="Hernández-Cárdenas">Hernandez-Cardenas</normalizedToken>
</collectorName>
</emphasis>
,
<emphasis id="9D2864ABCCE74A519A40910436F31240" italics="true" pageId="0" pageNumber="20">
<collectorName id="A1BBA0EB37DD83161DD32FE98665EC6A">
A.
<normalizedToken id="DE263009CD01E30E721FAC026B3FFC08" originalValue="Flores-Argüelles">Flores-Argueelles</normalizedToken>
</collectorName>
&amp;
<collectorName id="36E34017774E46342595EFBA3A7D0A55">
S.
<normalizedToken id="74265696315AF3F5A7745B8051E9F3D0" originalValue="Lara-Godínez">Lara-Godinez</normalizedToken>
</collectorName>
<specimenCount id="88ECC57BAC9B72395C742EE1115460E8" type="male">2619 ♂</specimenCount>
</emphasis>
; UAMIZ • ca
<quantity id="1A0066F49892822FA66B0E733B002FEB" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
sobre el camino
<collectorName id="9F396F5FF3A0C5DABC851CC28934D412">
La
<normalizedToken id="C764977A538A2C16F92F8407A1242FEE" originalValue="Central-Peña">Central-Pena</normalizedToken>
Blanca
</collectorName>
, a partir de la carretera
<normalizedToken id="A07062D57527AA3A2982DDDE6D5A4D67" originalValue="Manzanillo-Cihuatlán">Manzanillo-Cihuatlan</normalizedToken>
;
<geoCoordinate id="D105043E4081A1D9034380B5BA650E3A" degrees="19" direction="north" minutes="07" orientation="latitude" precision="1" seconds="57.9" value="19.13275">
19°07
<normalizedToken id="87A80B1A64EAF50EA400DE2DA1766E82" originalValue="57.9”">'57.9&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="E0ED3CFB8249EE9D386E9E91CFA83552" degrees="104" direction="west" minutes="26" orientation="longitude" precision="1" seconds="54.6" value="-104.4485">
104°26
<normalizedToken id="F4BD58FC1ED14EDC33515B1AC77920DF" originalValue="54.6”">'54.6&quot;</normalizedToken>
W
</geoCoordinate>
;
<quantity id="AE353FCA4AF98317405164A199B7CB96" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation id="FF766AD28F18C8E5F874022DF4CBD758" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
;
<collectingDate id="00DD5F07F9E7451424D7397E24B892A7" value="2023-05-06">6 May 2023</collectingDate>
;
<emphasis id="8BF083CE9692A9DC23F0EFF12FFADCEC" italics="true" pageId="0" pageNumber="20">
<collectorName id="EAABD38A94E43B602028987D6384F0D4">
A.
<normalizedToken id="5AF3786CA67F3E82116A46BFD63C0089" originalValue="Flores-Argüelles">Flores-Argueelles</normalizedToken>
</collectorName>
&amp;
<collectorName id="4ED58B039A98202F0FDDBDF8B2A77762">
G.
<normalizedToken id="902BB181DEAFED0D6E57076BE23B0424" originalValue="Contreras-Félix">Contreras-Felix</normalizedToken>
</collectorName>
<specimenCount id="C8CC6AB5F97CB03E4D6B6F5CA1EC66C8" type="female">1735 ♀</specimenCount>
</emphasis>
; IBUG, UAMIZ
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="900AA784D0B987424D28DD63E418D227" pageId="0" pageNumber="20" type="notes">
<paragraph id="DE114FC3CDC7CFAD82AADEE4801DB495" pageId="0" pageNumber="20">Notes.</paragraph>
<paragraph id="A314628B966309ABE22022B092951E08" pageId="0" pageNumber="20">
The type specimens of
<taxonomicName id="A7AEC3EF12118D844FAAD66865476482" authorityName="L. B. Sm. (Smith" authorityYear="1937" class="Liliopsida" family="Bromeliaceae" genus="Hechtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hechtia reticulata" order="Poales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="FA6B9966E18FEE2A4EC074431469FB38" italics="true" pageId="0" pageNumber="20">Hechtia reticulata</emphasis>
</taxonomicName>
(holotype and isotypes) are composed only of fragments of mature pistillate inflorescences, collected by Edward Palmer in March 1891 so, in order to facilitate the precise application of the name, it was important to designate an epitype that included complete and representative material of the species. Specimens of
<taxonomicName id="0FB0005E2AC281A1A1EC9BCD4DC65188" authorityName="L. B. Sm. (Smith" authorityYear="1937" class="Liliopsida" family="Bromeliaceae" genus="Hechtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hechtia reticulata" order="Poales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="F5AA91B8A35FFFA139ECF537AF1FEF01" italics="true" pageId="0" pageNumber="20">Hechtia reticulata</emphasis>
</taxonomicName>
have been misidentified as
<taxonomicName id="588805BED549BDBE1A5ADAA0BBF75BEB" lsidName="H. laevis" pageId="0" pageNumber="20" rank="species" species="laevis">
<emphasis id="9DDA9FAA3FDCC1689DF46E606A9A0154" italics="true" pageId="0" pageNumber="20">H. laevis</emphasis>
</taxonomicName>
or
<taxonomicName id="396813677BAFFCB6AC62697668BB2291" lsidName="H. laxissima" pageId="0" pageNumber="20" rank="species" species="laxissima">
<emphasis id="0F789B1150957975B7F109F260BC6DE4" italics="true" pageId="0" pageNumber="20">H. laxissima</emphasis>
</taxonomicName>
. Nevertheless,
<taxonomicName id="9B7E163369A1D7C0A5DC7727A8AC8EDB" lsidName="H. reticulata" pageId="0" pageNumber="20" rank="species" species="reticulata">
<emphasis id="9059F6872EA02AFD04B05A3C6FB6BA17" italics="true" pageId="0" pageNumber="20">H. reticulata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="6E3854A06F7E611B45689E86EE4D2FF1" lsidName="H. laevis" pageId="0" pageNumber="20" rank="species" species="laevis">
<emphasis id="65A7AA48CD676DD334D7D823AFFC590F" italics="true" pageId="0" pageNumber="20">H. laevis</emphasis>
</taxonomicName>
in the length of the leaf sheath (7-10 vs 4-4.5 cm), in the size of the leaf blade (65-85
<normalizedToken id="0AF42167E8CCBA9C2C1688F49BA41568" originalValue="×">x</normalizedToken>
3.5-5.5 vs 20-60
<normalizedToken id="FA2F50AC27F6588938092005D826F88D" originalValue="×">x</normalizedToken>
2.5-3 cm), in the arrangement of the staminate inflorescence (once to twice branched vs once branched), in the length of its pedicels (0.8-1.1 vs 1.5-3 mm), and in the size of the floral bract of the pistillate inflorescence (3.7-4.2
<normalizedToken id="46EF15855B1BC9E19CF4BDB203198C60" originalValue="×">x</normalizedToken>
2.9-3.3 vs 2
<normalizedToken id="27BAA9E861DCE6B722A5BEC133101C12" originalValue="×">x</normalizedToken>
0.3-0.5 mm).
<taxonomicName id="C4842FE1766C5BB629D05F1E4D36030C" authorityName="L. B. Sm. (Smith" authorityYear="1937" class="Liliopsida" family="Bromeliaceae" genus="Hechtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hechtia reticulata" order="Poales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="D2E49409667E42E0DA09269DCB381CB2" italics="true" pageId="0" pageNumber="20">Hechtia reticulata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BB1679FF85668FDC1A6D7AE97E5028BE" lsidName="H. laxissima" pageId="0" pageNumber="20" rank="species" species="laxissima">
<emphasis id="C1ECCBD67674657F4D029B29B43AF3B8" italics="true" pageId="0" pageNumber="20">H. laxissima</emphasis>
</taxonomicName>
in the number of leaves (10-20 vs&gt; 20), in the length of the leaf blade (65-85 vs 40 cm), in the length of the sepals of the pistillate inflorescence (2.3-2.7 mm vs 3), and in the shape of its petals (lanceolate vs ovate).
<taxonomicName id="A7A1E0EDB755BCDDA887B90CE4FE073F" authorityName="L. B. Sm. (Smith" authorityYear="1937" class="Liliopsida" family="Bromeliaceae" genus="Hechtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hechtia reticulata" order="Poales" pageId="0" pageNumber="20" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="93F96DE68AA599F34CC08B493FEE74B3" italics="true" pageId="0" pageNumber="20">Hechtia reticulata</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="1125EEA8A218A7B829B58D91155459B5" lsidName="H. costalegrensis" pageId="0" pageNumber="20" rank="species" species="costalegrensis">
<emphasis id="9BA1597B600D78C8D69EDEFF1F49B617" italics="true" pageId="0" pageNumber="20">H. costalegrensis</emphasis>
</taxonomicName>
and
<taxonomicName id="F1271E4CAE9FC6C68B44261E5B49AC7D" lsidName="H. pacifica" pageId="0" pageNumber="20" rank="species" species="pacifica">
<emphasis id="BFDD64BAD3173743B77E6843BA3DF348" italics="true" pageId="0" pageNumber="20">H. pacifica</emphasis>
</taxonomicName>
, with which they were compared in this work. For more details, see Table
<tableCitation id="C80685842073ECC955E37F7F6BAD7EA2" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological differences between Hechtia costalegrensis, H. pacifica, and H. reticulata." httpUri="http://table.plazi.org/id/8070A5D181A6973B4413BFA5B0CDF13F" pageId="0" pageNumber="20" tableUuid="8070A5D181A6973B4413BFA5B0CDF13F">1</tableCitation>
. Lyman B. Smith described the fruits of
<taxonomicName id="439F8BE37DD2C7AC7DDEBF224D994DE1" lsidName="H. reticulata" pageId="0" pageNumber="20" rank="species" species="reticulata">
<emphasis id="BDF98632E1FA0A637E955BB5C2E7C679" italics="true" pageId="0" pageNumber="20">H. reticulata</emphasis>
</taxonomicName>
as &quot;capsula carpellis grosse irregulariterque reticulatis&quot;. However, in all the examined specimens, including the types, the fruits are not reticulate.
<bibRefCitation id="BDF79706976742967E7F0966217B7F57" author="McVaugh, R" journalOrPublisher="Aliso" pageId="0" pageNumber="20" refId="B10" refString="McVaugh, R, 1989. Bromeliaceae to Dioscoreaceae. Flora Novo Galiciana vol. 15. University of Michigan Press, Ann Arbor, 1-398." title="Bromeliaceae to Dioscoreaceae. Flora Novo Galiciana vol. 15. University of Michigan Press, Ann Arbor, 1 - 398." year="1989">McVaugh (1989)</bibRefCitation>
noticed the same and mentioned that &quot;I have not seen the holotype, but in other fruiting specimens (
<emphasis id="83385D0F6A2E105AD6F86BF027109116" italics="true" pageId="0" pageNumber="20">Ferris 6111</emphasis>
) the carpels are not reticulate in the ordinary sense; they are provided with prominent longitudinal ridges (nerves) that are weakly and irregularly connected by smaller lateral nerves&quot;. These ridges or creases could be due to the maturation or dehydration of the fruits and were also observed in the carpels of
<taxonomicName id="6A832604DA1810A7B151E79F4ADF034E" lsidName="H. pacifica" pageId="0" pageNumber="20" rank="species" species="pacifica">
<emphasis id="E64D4E474D33DBD43B7F8B40F1E2FD3B" italics="true" pageId="0" pageNumber="20">H. pacifica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>