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<document id="23404F384E4FC73AD477D98AFBAE1308" ID-DOI="10.5281/zenodo.210231" ID-GBIF-Dataset="c920bea6-b2d3-4dfc-8d8b-4aa1dce50dd9" ID-ISSN="1175-5326" ID-Zenodo-Dep="210231" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460479687169" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Motoh, Hiroshi" docDate="2012" docId="B16B8792FFB5483BCCA1FEEFE6D41B96" docLanguage="en" docName="zt03305p027.pdf" docOrigin="Zootaxa 3305" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pilumnus dofleini Balss 1933" docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="4D52FFEAFFB44833CC36FFAAE7481A3A" masterDocTitle="The identity of Pilumnus dofleini Balss, 1933 (Crustacea: Decapoda: Brachyura Pilumnidae), with descriptions of three new species from the Western Pacific" masterLastPageNumber="27" masterPageNumber="1" pageNumber="2" updateTime="1698307407507" updateUser="plazi">
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<mods:title id="8F384A943170E2B2830A3AB918C98D46">The identity of Pilumnus dofleini Balss, 1933 (Crustacea: Decapoda: Brachyura Pilumnidae), with descriptions of three new species from the Western Pacific</mods:title>
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<treatment id="B16B8792FFB5483BCCA1FEEFE6D41B96" ID-DOI="http://doi.org/10.5281/zenodo.6177466" ID-GBIF-Taxon="119517020" ID-Zenodo-Dep="6177466" LSID="urn:lsid:plazi:treatment:B16B8792FFB5483BCCA1FEEFE6D41B96" httpUri="http://treatment.plazi.org/id/B16B8792FFB5483BCCA1FEEFE6D41B96" lastPageId="8" lastPageNumber="9" pageId="1" pageNumber="2">
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<paragraph id="397D3684FFB54832CCA1FEEFE54A1B65" blockId="1.[151,514,325,383]" box="[151,514,325,351]" pageId="1" pageNumber="2">
<heading id="623581E8FFB54832CCA1FEEFE54A1B65" bold="true" box="[151,514,325,351]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<taxonomicName id="FEC24D07FFB54832CCA1FEEFE54A1B65" ID-CoL="4HVSW" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" box="[151,514,325,351]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB54832CCA1FEEFE54A1B65" bold="true" box="[151,514,325,351]" pageId="1" pageNumber="2">
<emphasis id="0BB6EA96FFB54832CCA1FEEFE6391B65" bold="true" box="[151,369,325,351]" italics="true" pageId="1" pageNumber="2">Pilumnus dofleini</emphasis>
<bibRefCitation id="5D534B75FFB54832CD41FEEFE54A1B65" author="Balss" box="[375,514,325,351]" pageId="1" pageNumber="2" refString="Balss, H. (1933) Beitrage zur Kentnis der Gattung Pilumnus (Crustacea Dekapoda) und verwandter Gattugen. Capita Zoologica, 4, 27 - 47, pls 1 - 7." type="journal article" year="1933">Balss, 1933</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="397D3684FFB54832CCA1FECDE6431B45" blockId="1.[151,514,325,383]" box="[151,267,359,383]" pageId="1" pageNumber="2">
(
<figureCitation id="A1F92A01FFB54832CCA9FECDE6491B45" box="[159,257,359,383]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1982,2005]" captionStartId-1="3.[151,250,1954,1977]" captionStartId-2="4.[151,250,1842,1865]" captionStartId-3="5.[151,250,1920,1943]" captionStartId-4="6.[151,250,1265,1288]" captionTargetBox-0="[151,1436,174,1959]" captionTargetBox-1="[151,1436,299,1933]" captionTargetBox-2="[244,1340,209,1801]" captionTargetBox-3="[214,1359,188,1883]" captionTargetBox-4="[151,1436,179,1244]" captionTargetId-0="figure@2.[151,1436,172,1961]" captionTargetId-1="figure@3.[151,1436,299,1934]" captionTargetId-2="figure@4.[226,1368,193,1814]" captionTargetId-3="figure@5.[186,1410,173,1906]" captionTargetId-4="figure@6.[151,1436,179,1245]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, dorsal view; B, carapace, dorsal view." captionText-1="FIGURE 2. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, ventral view; B, thoracic sternum and abdomen, ventral view." captionText-2="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." captionText-3="FIGURE 4. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, right major chela, outer view; B, carpus of right major cheliped, dorsal view; C, merus of right major cheliped, outer view; D, left minor chela, outer view; E, merus of left minor cheliped, outer view; F, left third ambulatory leg (fourth pereopod), posterior view; G, left fourth ambulatory leg (fifth pereopod), posterior view). Setae omitted. Scale bars: 2 mm." captionText-4="FIGURE 5. Pilumnus dofleini Balss, 1933, holotype, female (13.0 x 15.0 mm), ZSM A 20110199 (photography: Stefan Friedrich, ZSM). A, cephalothorax, dorsal view; B, cephalothorax, ventral view; C, right minor cheliped, ventral view; D, left major cheliped, ventral view." httpUri-0="https://zenodo.org/record/210232/files/figure.png" httpUri-1="https://zenodo.org/record/210233/files/figure.png" httpUri-2="https://zenodo.org/record/210234/files/figure.png" httpUri-3="https://zenodo.org/record/210235/files/figure.png" httpUri-4="https://zenodo.org/record/210236/files/figure.png" pageId="1" pageNumber="2">Figs 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="71D8650FFFB54832CCA1FE1AE3D01865" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="397D3684FFB54832CCA1FE1AE3A11BDE" blockId="1.[151,1436,432,608]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="19D211AAFFB54832CCA1FE1AE3A11BDE" pageId="1" pageNumber="2">
<taxonomicName id="FEC24D07FFB54832CCA1FE1AE6A71BFF" ID-CoL="4HVSW" authority="Balss, 1933: 29" authorityName="Balss" authorityPageNumber="29" authorityYear="1933" box="[151,495,432,454]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB54832CCA1FE1AE6011BFC" box="[151,329,432,454]" italics="true" pageId="1" pageNumber="2">Pilumnus dofleini</emphasis>
<treatmentCitation id="B8631095FFB54832CD66FE1AE6A71BFF" author="Balss" box="[336,495,432,454]" page="29" pageId="1" pageNumber="2" year="1933">
<bibRefCitation id="5D534B75FFB54832CD66FE1AE6821BFF" author="Balss" box="[336,458,432,454]" pageId="1" pageNumber="2" refString="Balss, H. (1933) Beitrage zur Kentnis der Gattung Pilumnus (Crustacea Dekapoda) und verwandter Gattugen. Capita Zoologica, 4, 27 - 47, pls 1 - 7." type="journal article" year="1933">Balss, 1933</bibRefCitation>
: 29
</treatmentCitation>
</taxonomicName>
, pl. 6, fig. 29. —
<treatmentCitation id="B8631095FFB54832CE97FE1AE4001BFC" author="Sakai" box="[673,840,432,454]" page="538" pageId="1" pageNumber="2" year="1939">
<bibRefCitation id="5D534B75FFB54832CE97FE1AE45D1BFF" author="Sakai" box="[673,789,432,454]" pageId="1" pageNumber="2" refString="Sakai, T. (1939) Studies on the Crabs of Japan. Brachygnatha, Brachyrhyncha, IV. Yokendo, Tokyo, pp. 365 - 741 + 1 - 23 (index), pls 42 - 111." type="book chapter" year="1939">Sakai 1939</bibRefCitation>
: 538
</treatmentCitation>
; 1965: 169, pl. 70, fig. 1; 1976: 489 (English text), pl. 175, fig. 2. —
<treatmentCitation id="B8631095FFB54832CD14FE64E50D1BDE" author="Takeda" box="[290,581,462,484]" page="157" pageId="1" pageNumber="2" year="2000">
<bibRefCitation id="5D534B75FFB54832CD14FE64E55E1BDE" author="Takeda" box="[290,534,462,484]" pageId="1" pageNumber="2" refString="Takeda, M. &amp; Manuel, M. R. (2000) Taxonomic accounts of some rare crabs new to the Philippines. National Science Museum Monographs, 18, 149 - 162." type="journal article" year="2000">Takeda &amp; Manuel 2000</bibRefCitation>
: 157
</treatmentCitation>
(part);
<treatmentCitation id="B8631095FFB54832CEA7FE65E4821BDE" author="Marumura" box="[657,970,462,484]" page="57" pageId="1" pageNumber="2" year="2003">
<bibRefCitation id="5D534B75FFB54832CEA7FE65E4EC1BDE" author="Marumura" box="[657,932,462,484]" pageId="1" pageNumber="2" refString="Marumura, M. &amp; Kosaka, A. (2003) Catalog of the Brachyuran and Anomuran Crabs Donated by the Late Mr. Seiji Nagai to the Wakayama Prefectural Museum of Natural History. 74 pp. Wakayama Prefectural Museum of Natural History, Kainan." type="book" year="2003">Marumura &amp; Kosaka 2003</bibRefCitation>
: 57
</treatmentCitation>
. Takeda
<emphasis id="0BB6EA96FFB54832C875FE7AE33E1BDF" box="[1091,1142,463,485]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
2006: 204.
</treatmentCitationGroup>
</paragraph>
<paragraph id="397D3684FFB54832CCA1FE44E4D71839" blockId="1.[151,1436,432,608]" box="[151,927,493,515]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="19D211AAFFB54832CCA1FE44E4D71839" box="[151,927,493,515]" pageId="1" pageNumber="2">
Not
<taxonomicName id="FEC24D07FFB54832CCF5FE47E63E1839" ID-CoL="4HVSW" box="[195,374,493,515]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB54832CCF5FE47E63E1839" box="[195,374,493,515]" italics="true" pageId="1" pageNumber="2">Pilumnus dofleini</emphasis>
</taxonomicName>
. —
<treatmentCitation id="B8631095FFB54832CD95FE47E5161838" author="Miyake" box="[419,606,493,515]" page="134" pageId="1" pageNumber="2" year="1983">
<bibRefCitation id="5D534B75FFB54832CD95FE47E5641838" author="Miyake" box="[419,556,493,515]" pageId="1" pageNumber="2" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in Color, Vol. II. Brachyura (Crabs). Hoikusha, Osaka, 277 pp., 64 pls." type="book" year="1983">Miyake 1983</bibRefCitation>
: 134
</treatmentCitation>
, pl. 45, fig. 5. [see Discussion].
</treatmentCitationGroup>
</paragraph>
<paragraph id="397D3684FFB54832CCA1FDA7E48F181B" blockId="1.[151,1436,432,608]" box="[151,967,523,546]" pageId="1" pageNumber="2">
Not
<taxonomicName id="FEC24D07FFB54832CCF5FDA6E63E1818" ID-CoL="4HVSW" box="[195,374,524,546]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB54832CCF5FDA6E63E1818" box="[195,374,524,546]" italics="true" pageId="1" pageNumber="2">Pilumnus dofleini</emphasis>
</taxonomicName>
.— Ng 2000: 301, figs 1, 2. [=
<taxonomicName id="FEC24D07FFB54832CE86FDA6E4C81818" ID-CoL="4HVSK" box="[688,896,524,546]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB54832CE86FDA6E4C81818" box="[688,896,524,546]" italics="true" pageId="1" pageNumber="2">Pilumnus curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB54832CFB0FDA6E4F7181B" bold="true" box="[902,959,524,545]" pageId="1" pageNumber="2">
<taxonomicNameLabel id="108557EDFFB54832CFB0FDA6E4F7181B" box="[902,959,524,545]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
]
</paragraph>
<paragraph id="397D3684FFB54832CCA1FD81E4D7187A" blockId="1.[151,1436,432,608]" box="[151,927,554,576]" pageId="1" pageNumber="2">
Not
<taxonomicName id="FEC24D07FFB54832CCF5FD80E63E187A" ID-CoL="4HVSW" box="[195,374,554,576]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB54832CCF5FD80E63E187A" box="[195,374,554,576]" italics="true" pageId="1" pageNumber="2">Pilumnus dofleini</emphasis>
</taxonomicName>
. — Ng
<emphasis id="0BB6EA96FFB54832CDFEFD81E6B3187A" box="[456,507,554,576]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
2008: fig. 112. [=
<taxonomicName id="FEC24D07FFB54832CE8EFD80E41F187A" ID-CoL="4HVRW" box="[696,855,554,576]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="bohol" status="sp. nov.">
<emphasis id="0BB6EA96FFB54832CE8EFD80E41F187A" box="[696,855,554,576]" italics="true" pageId="1" pageNumber="2">Pilumnus bohol</emphasis>
</taxonomicName>
<taxonomicNameLabel id="108557EDFFB54832CF68FD81E4DF187A" box="[862,919,555,576]" pageId="1" pageNumber="2" rank="species">
<emphasis id="0BB6EA96FFB54832CF68FD81E4DF187A" bold="true" box="[862,919,555,576]" pageId="1" pageNumber="2">n. sp.</emphasis>
</taxonomicNameLabel>
]
</paragraph>
<paragraph id="397D3684FFB54832CCA1FDE0E3D01865" blockId="1.[151,1436,432,608]" box="[151,1176,585,608]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="19D211AAFFB54832CCA1FDE0E3D01865" box="[151,1176,585,608]" pageId="1" pageNumber="2">
Not
<taxonomicName id="FEC24D07FFB54832CCF5FDE3E63E1865" ID-CoL="4HVSW" box="[195,374,585,607]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB54832CCF5FDE3E63E1865" box="[195,374,585,607]" italics="true" pageId="1" pageNumber="2">Pilumnus dofleini</emphasis>
</taxonomicName>
. —
<treatmentCitation id="B8631095FFB54832CD95FDE3E58D1865" author="Takeda" box="[419,709,585,607]" page="157" pageId="1" pageNumber="2" year="2000">
<bibRefCitation id="5D534B75FFB54832CD95FDE3E5DD1865" author="Takeda" box="[419,661,585,607]" pageId="1" pageNumber="2" refString="Takeda, M. &amp; Manuel, M. R. (2000) Taxonomic accounts of some rare crabs new to the Philippines. National Science Museum Monographs, 18, 149 - 162." type="journal article" year="2000">Takeda &amp; Manuel 2000</bibRefCitation>
: 157
</treatmentCitation>
(part), figs 3F, 5. [=
<taxonomicName id="FEC24D07FFB54832CFAEFDE3E3191865" ID-CoL="4HVRP" box="[920,1105,585,607]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="armatus" status="sp. nov.">
<emphasis id="0BB6EA96FFB54832CFAEFDE3E3191865" box="[920,1105,585,607]" italics="true" pageId="1" pageNumber="2">Pilumnus armatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="108557EDFFB54832C86EFDE0E3C71865" box="[1112,1167,585,607]" pageId="1" pageNumber="2" rank="species">
<emphasis id="0BB6EA96FFB54832C86EFDE0E3C31865" bold="true" box="[1112,1163,586,607]" pageId="1" pageNumber="2">n. sp</emphasis>
.
</taxonomicNameLabel>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="71D8650FFFB54832CCA1FD21E459190E" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="397D3684FFB54832CCA1FD21E26618FD" blockId="1.[151,1437,651,2008]" pageId="1" pageNumber="2">
<emphasis id="0BB6EA96FFB54832CCA1FD21E630189E" bold="true" box="[151,376,651,676]" pageId="1" pageNumber="2">Material examined</emphasis>
.
<typeStatus id="E6798826FFB54832CDB0FD26E6BB189E" box="[390,499,652,676]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
: female (13.0 x 15.0 mm),
<collectionCode id="5FD3AE41FFB54832CF29FD26E410189E" box="[799,856,652,676]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="2">ZSM</collectionCode>
A20110199, off Boshu (= Boso Peninsula), Sagami Sea,
<collectingCountry id="41D57614FFB54832CCFBFD1AE65818FD" box="[205,272,688,711]" name="Japan" pageId="1" pageNumber="2">Japan</collectingCountry>
,
<quantity id="FE3A9B61FFB54832CD2DFD05E62C18FD" box="[283,356,687,711]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="2" unit="m" value="180.0">180 m</quantity>
,
<date id="4D7C1044FFB54832CD46FD05E50418FD" box="[368,588,687,711]" pageId="1" pageNumber="2" value="1904-09-10">10 September 1904</date>
, coll. F. Doflein. Photographs provided by S. Friedrich examined.
</paragraph>
<paragraph id="397D3684FFB54832CCFFFD7EE459190E" blockId="1.[151,1437,651,2008]" pageId="1" pageNumber="2">
Additional material:
<specimenCount id="2FC4FD0DFFB54832CD87FD7EE6B318D6" box="[433,507,724,748]" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
(10.2 x
<quantity id="FE3A9B61FFB54832CE61FD7EE5F618D6" box="[599,702,724,748]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="2" unit="mm" value="12.5">12.5 mm</quantity>
), NSMT-CrR 2507, Kannon-zuka-dashi, Sagami Bay,
<collectingCountry id="41D57614FFB54832C913FD7FE22018D6" box="[1317,1384,725,748]" name="Japan" pageId="1" pageNumber="2">Japan</collectingCountry>
,
<quantity id="FE3A9B61FFB54832C944FD7EE79B192A" metricMagnitude="1" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="1" pageNumber="2" unit="m" value="70.0" valueMax="80.0" valueMin="60.0">60 80 m</quantity>
,
<date id="4D7C1044FFB54832CCE9FD5DE63C1935" box="[223,372,759,783]" pageId="1" pageNumber="2" value="1957-07-18">18 July 1957</date>
;
<specimenCount id="2FC4FD0DFFB54832CDB7FD5DE6841935" box="[385,460,759,783]" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
(12.5 x
<quantity id="FE3A9B61FFB54832CE1AFD5DE5DD1935" box="[556,661,759,783]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="1" pageNumber="2" unit="mm" value="15.3">15.3 mm</quantity>
), CBM-ZC 10621, off Hashidate, Ishikawa Prefecture, Sea of
<collectingCountry id="41D57614FFB54832C963FD53E2D0192A" box="[1365,1432,761,784]" name="Japan" pageId="1" pageNumber="2">Japan</collectingCountry>
,
<quantity id="FE3A9B61FFB54832CCA1FCB6E79A190E" box="[151,210,796,820]" metricMagnitude="1" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="2" unit="m" value="68.0">68 m</quantity>
, sandy mud,
<date id="4D7C1044FFB54832CD50FCB6E56D190E" box="[358,549,796,820]" pageId="1" pageNumber="2" value="2008-10-17">17 October 2008</date>
, commercial gill net.
</paragraph>
</subSubSection>
<subSubSection id="71D8650FFFB54834CCFFFC95E38A182D" lastPageId="7" lastPageNumber="8" pageId="1" pageNumber="2" type="description">
<paragraph id="397D3684FFB54832CCFFFC95E4841F16" blockId="1.[151,1437,651,2008]" pageId="1" pageNumber="2">
<emphasis id="0BB6EA96FFB54832CCFFFC95E61C1962" bold="true" box="[201,340,831,856]" pageId="1" pageNumber="2">Description</emphasis>
. Carapace (
<figureCitation id="A1F92A01FFB54832CDD2FC95E565196D" box="[484,557,831,855]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2005]" captionTargetBox="[151,1436,174,1959]" captionTargetId="figure@2.[151,1436,172,1961]" captionTargetPageId="2" captionText="FIGURE 1. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, dorsal view; B, carapace, dorsal view." httpUri="https://zenodo.org/record/210232/files/figure.png" pageId="1" pageNumber="2">Figs 1</figureCitation>
A, B, 3AD, 5A) fairly vaulted, transversely ovoid, surface covered with numerous short setae; greatest width across fourth anterolateral spines, 1.2 times of length; dorsal surface somewhat convex longitudinally, slightly convex transversely; regions well defined, surface of regions with several conical spines or spiniform tubercles; cardiac region slightly elevated, with about 10 spiniform tubercles; grooves between regions smooth, glabrous. Frontal margin not markedly produced, separated medially by wide, V-shaped notch; each frontal lobe divided in 2 parts by broad hiatus, mesial lobe with 7 small tubercles (innermost tubercle minute), lateral lobes each consisting of single conical spine slightly larger than spines on mesial lobe; inner supraorbital angle with small spine; supraorbital margin with row of small spines, with 2 shallow cleft (one at middle, one near external orbital angle). Anterolateral margin markedly spinose, with 4 main spines (including spinose external orbital angle), second to fourth anterolateral spines subequal in size, obliquely erect, each bearing several accessory spines or spinules. Posterolateral margin almost straight, converging towards posterior margin. Posterior part of carapace with submarginal row of distinct tubercles; posterior margin slightly convex. Suborbital region with covering of numerous spinules or spinulose granules; pterygostomial groove lined with granular rows dorsally and ventrally. Dorsolateral portion of buccal frame with cluster of spines.
</paragraph>
<paragraph id="397D3684FFB54832CCFFFA92E3E21F4E" blockId="1.[151,1437,651,2008]" pageId="1" pageNumber="2">
Orbits (
<figureCitation id="A1F92A01FFB54832CD28FA92E6161F6A" box="[286,350,1336,1360]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
A, D) moderately large, slightly obliquely transversal in dorsal view, transversal in anterior view; eyes filling entire orbits; suborbital margin with row of small spines, innermost spine largest.
</paragraph>
<paragraph id="397D3684FFB54832CCFFFA2AE3B01F86" blockId="1.[151,1437,651,2008]" pageId="1" pageNumber="2">
Basal segment of antennular peduncle (
<figureCitation id="A1F92A01FFB54832CEB4FA2AE58B1FA2" box="[642,707,1408,1432]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
D) with 2 obliquely transversal ridges on outer surface. Antennal peduncle just entering orbital hiatus; second segment with 1 small tubercle distally on outer surface.
</paragraph>
<paragraph id="397D3684FFB54832CCFFFA62E37B1FDA" blockId="1.[151,1437,651,2008]" box="[201,1075,1480,1504]" pageId="1" pageNumber="2">
Epistome (
<figureCitation id="A1F92A01FFB54832CD74FA62E6CB1FDA" box="[322,387,1480,1504]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
D) divided into 4 crested lobes. Endostomal ridges indistinct.
</paragraph>
<paragraph id="397D3684FFB54832CCFFFA46E58E1CAE" blockId="1.[151,1437,651,2008]" pageId="1" pageNumber="2">
Third maxillipeds (
<figureCitation id="A1F92A01FFB54832CD97FA46E6AB1C3E" box="[417,483,1516,1540]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
E) completely covering buccal cavity when closed; outer surfaces of ischium, merus exopods granular. Ischium subrectangular, longer than wide, with shallow oblique median sulcus, mesial margin weakly denticulate. Merus subrectangular, wider than long, margins granular, mesial margin somewhat produced in rounded lobe. Carpus to dactylus smooth on surfaces. Exopod moderately stout, distal margin just reaching distal margin of merus, inner subdistal spine prominent.
</paragraph>
<paragraph id="397D3684FFB54830CCFFF90AE42E1ACD" blockId="1.[151,1437,651,2008]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
Chelipeds (
<figureCitation id="A1F92A01FFB54832CD7CF90AE6C71C82" box="[330,399,1696,1720]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2005]" captionTargetBox="[151,1436,174,1959]" captionTargetId="figure@2.[151,1436,172,1961]" captionTargetPageId="2" captionText="FIGURE 1. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, dorsal view; B, carapace, dorsal view." httpUri="https://zenodo.org/record/210232/files/figure.png" pageId="1" pageNumber="2">Figs 1</figureCitation>
A, 2A, 4AE; 5C, D) slightly unequal, slightly dissimilar with right larger than left; outer surfaces with covering of short setae. Anterior margin of fused basis-ischium with row of small tubercles. Merus nearly as long as high; posterodorsal margin crested, with row of spines (2 distal spines prominent); outer (posterior) surface granular adjacent to margins, otherwise nearly smooth; lower surface bluntly carinate, with rows of acute, minute to small spines. Carpus numerous, small, occasionally curved spines on dorsal to outer (lateral) surface, inner distal angle with prominent spine. Palm with numerous spines on outer surface, arranged in irregular longitudinal rows (ventral rows extending onto fixed finger), inner dorsal margin bluntly carinate, with row of small tubercles or spinules; ventral surface spinulose, bluntly carinate; inner surface nearly smooth. Fixed finger of major cheliped not deflexed, with row of rounded teeth on cutting edge; outer surface flat medially. Fixed finger of minor cheliped slightly deflexed, with row of broadly triangular, sharply edged teeth on cutting edge. Dactylus with multiple rows of small spines or spinulose tubercles on dorsal surface extending beyond midlength, with 1 (right) or 2 (left) distinct longitudinal grooves on outer surfaces.
</paragraph>
<caption id="6DBD660CFFB64831CCA1F814E6051DC9" httpUri="https://zenodo.org/record/210232/files/figure.png" pageId="2" pageNumber="3" targetBox="[151,1436,174,1959]" targetPageId="2">
<paragraph id="397D3684FFB64831CCA1F814E6051DC9" blockId="2.[151,1436,1982,2035]" pageId="2" pageNumber="3">
<emphasis id="0BB6EA96FFB64831CCA1F814E65C1DEE" bold="true" box="[151,276,1982,2005]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
<taxonomicName id="FEC24D07FFB64831CD2AF815E51A1DEE" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" box="[284,594,1982,2005]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB64831CD2AF815E6981DEF" box="[284,464,1983,2005]" italics="true" pageId="2" pageNumber="3">Pilumnus dofleini</emphasis>
Balss, 1933
</taxonomicName>
, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, dorsal view; B, carapace, dorsal view.
</paragraph>
</caption>
<caption id="6DBD660CFFB74830CCA1F808E54E1DE2" httpUri="https://zenodo.org/record/210233/files/figure.png" pageId="3" pageNumber="4" targetBox="[151,1436,299,1933]" targetPageId="3">
<paragraph id="397D3684FFB74830CCA1F808E54E1DE2" blockId="3.[151,1436,1954,2008]" pageId="3" pageNumber="4">
<emphasis id="0BB6EA96FFB74830CCA1F808E65A1D82" bold="true" box="[151,274,1954,1977]" pageId="3" pageNumber="4">FIGURE 2.</emphasis>
<taxonomicName id="FEC24D07FFB74830CD2EF809E5001D83" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" box="[280,584,1955,1977]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB74830CD2EF809E6821D83" box="[280,458,1955,1977]" italics="true" pageId="3" pageNumber="4">Pilumnus dofleini</emphasis>
Balss, 1933
</taxonomicName>
, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, ventral view; B, thoracic sternum and abdomen, ventral view.
</paragraph>
</caption>
<caption id="6DBD660CFFB04837CCA1F898E2341DD8" httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="4" pageNumber="5" targetBox="[244,1340,209,1801]" targetPageId="4">
<paragraph id="397D3684FFB04837CCA1F898E2341DD8" blockId="4.[151,1437,1842,2018]" pageId="4" pageNumber="5">
<emphasis id="0BB6EA96FFB04837CCA1F898E65D1D72" bold="true" box="[151,277,1842,1865]" pageId="4" pageNumber="5">FIGURE 3.</emphasis>
<taxonomicName id="FEC24D07FFB04837CD2BF899E51D1D72" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" box="[285,597,1842,1865]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB04837CD2BF899E69A1D73" box="[285,466,1843,1865]" italics="true" pageId="4" pageNumber="5">Pilumnus dofleini</emphasis>
Balss, 1933
</taxonomicName>
, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for IL.
</paragraph>
</caption>
<caption id="6DBD660CFFB14836CCA1F82AE4711DC8" httpUri="https://zenodo.org/record/210235/files/figure.png" pageId="5" pageNumber="6" targetBox="[214,1359,188,1883]" targetPageId="5">
<paragraph id="397D3684FFB14836CCA1F82AE4711DC8" blockId="5.[151,1436,1920,2034]" pageId="5" pageNumber="6">
<emphasis id="0BB6EA96FFB14836CCA1F82AE65B1DAC" bold="true" box="[151,275,1920,1943]" pageId="5" pageNumber="6">FIGURE 4.</emphasis>
<taxonomicName id="FEC24D07FFB14836CD2CF82AE5051DAC" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" box="[282,589,1920,1942]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB14836CD2CF82AE6851DAC" box="[282,461,1920,1942]" italics="true" pageId="5" pageNumber="6">Pilumnus dofleini</emphasis>
Balss, 1933
</taxonomicName>
, male (12.5 x 15.3 mm), CBM-ZC 10621. A, right major chela, outer view; B, carpus of right major cheliped, dorsal view; C, merus of right major cheliped, outer view; D, left minor chela, outer view; E, merus of left minor cheliped, outer view; F, left third ambulatory leg (fourth pereopod), posterior view; G, left fourth ambulatory leg (fifth pereopod), posterior view). Setae omitted. Scale bars: 2 mm.
</paragraph>
</caption>
<caption id="6DBD660CFFB24835CCA1FB5BE6F61F7F" httpUri="https://zenodo.org/record/210236/files/figure.png" pageId="6" pageNumber="7" targetBox="[151,1436,179,1244]" targetPageId="6">
<paragraph id="397D3684FFB24835CCA1FB5BE6F61F7F" blockId="6.[151,1436,1265,1349]" pageId="6" pageNumber="7">
<emphasis id="0BB6EA96FFB24835CCA1FB5BE65F1F3D" bold="true" box="[151,279,1265,1288]" pageId="6" pageNumber="7">FIGURE 5.</emphasis>
<taxonomicName id="FEC24D07FFB24835CD15FB58E52C1F32" authority="Balss, 1933" authorityName="Balss" authorityYear="1933" box="[291,612,1266,1288]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB24835CD15FB58E6931F32" box="[291,475,1266,1288]" italics="true" pageId="6" pageNumber="7">Pilumnus dofleini</emphasis>
Balss, 1933
</taxonomicName>
, holotype, female (13.0 x 15.0 mm), ZSM A20110199 (photography: Stefan Friedrich, ZSM). A, cephalothorax, dorsal view; B, cephalothorax, ventral view; C, right minor cheliped, ventral view; D, left major cheliped, ventral view.
</paragraph>
</caption>
<paragraph id="397D3684FFB24835CCFFFAD8E7821D2C" blockId="6.[151,1436,1394,2030]" pageId="6" pageNumber="7">
Ambulatory legs (
<figureCitation id="A1F92A01FFB24835CDA5FAD8E69F1FB0" box="[403,471,1394,1418]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2005]" captionTargetBox="[151,1436,174,1959]" captionTargetId="figure@2.[151,1436,172,1961]" captionTargetPageId="2" captionText="FIGURE 1. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, dorsal view; B, carapace, dorsal view." httpUri="https://zenodo.org/record/210232/files/figure.png" pageId="6" pageNumber="7">Figs 1</figureCitation>
A, 2A, 4F, G) moderately long, relatively slender for genus; surfaces with covering of short setae, dorsal (extensor) and ventral (flexor) surfaces numerous moderately short to moderately long, stiff setae; first leg (second pereopod) longest, about 1.5 times of carapace width. Coxa with minutely granular ventrodistal margin. Basis-ischium fused segment each with sharp granules on ventral surface, that of fourth leg (fifth pereopod) with small spinulose tubercles on outer surface along distal margin. Merus with single row of slender, weakly curved spines on dorsal (extensor) margin; flexor surface flanked by distinctly delimited margins each bearing row of small acute spines or tubercles; outer (posterior) surface unarmed (first to third legs) or with some small spines (fourth leg). Carpus with row of long spines on extensor margin; outer (posterior) surface with median row of spines plus few additional spines. Propodus with 2 or 3 rows of spines on extensor surface; flexor margin unarmed (first to third legs) or with 1 or 2 small spines (fourth leg). Dactylus slender, slightly curved distally, terminating in small corneous claw, longer than respective propodus; extensor margin with 04 spinules or tiny tubercles.
</paragraph>
<paragraph id="397D3684FFB24835CCFFF888E4471DB8" blockId="6.[151,1436,1394,2030]" pageId="6" pageNumber="7">
Thoracic sternum (
<figureCitation id="A1F92A01FFB24835CDAAF888E6971D00" box="[412,479,1826,1850]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1977]" captionTargetBox="[151,1436,299,1933]" captionTargetId="figure@3.[151,1436,299,1934]" captionTargetPageId="3" captionText="FIGURE 2. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, ventral view; B, thoracic sternum and abdomen, ventral view." httpUri="https://zenodo.org/record/210233/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
B) with covering of granules; these granules becoming more conspicuous in posterior sternites. Sternites 2/3 separated by distinct, straight groove; sternites 3/4 separated by suture sloping just anterior to abdominal cavity, suture becoming obsolete medially.
</paragraph>
<paragraph id="397D3684FFB24834CCFFF824E6771ACD" blockId="6.[151,1436,1394,2030]" lastBlockId="7.[151,1437,151,2012]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Male abdomen (
<figureCitation id="A1F92A01FFB24835CD49F824E6891D9C" box="[383,449,1934,1958]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
F) moderately narrow, somites 1 and 3 widest. Somite 1 transversely trapezoidal, armed with transversal row of small tubercles. Somite 2 subtrapezoidal, narrower than flanking somites, with paired tubercles laterally. Somites 36 unarmed; somite 3 broadly trapezoidal, narrowed distally, lateral margin gently convex, unarmed; somite 4 trapezoidal, lateral margins slightly concave; somites 5 and 6 subrectangular, with straight lateral margins. Telson (
<figureCitation id="A1F92A01FFB34834CE3EFF16E5041AEE" box="[520,588,188,212]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
F) triangular with rounded tip, 1.1 times longer than wide, lateral margins nearly straight.
</paragraph>
<paragraph id="397D3684FFB34834CCFFFEAEE5D21B05" blockId="7.[151,1437,151,2012]" pageId="7" pageNumber="8">
Female abdomen (
<figureCitation id="A1F92A01FFB34834CDAFFEAEE6931B26" box="[409,475,260,284]" captionStart="FIGURE 5" captionStartId="6.[151,250,1265,1288]" captionTargetBox="[151,1436,179,1244]" captionTargetId="figure@6.[151,1436,179,1245]" captionTargetPageId="6" captionText="FIGURE 5. Pilumnus dofleini Balss, 1933, holotype, female (13.0 x 15.0 mm), ZSM A 20110199 (photography: Stefan Friedrich, ZSM). A, cephalothorax, dorsal view; B, cephalothorax, ventral view; C, right minor cheliped, ventral view; D, left major cheliped, ventral view." httpUri="https://zenodo.org/record/210236/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
B) moderately broad. Somite 1 with several tubercles on outer surface. Somite 2 with pair of tiny tubercle laterally on outer surface.
</paragraph>
<paragraph id="397D3684FFB34834CCFFFEE6E3CB1B96" blockId="7.[151,1437,151,2012]" pageId="7" pageNumber="8">
First gonopod (
<figureCitation id="A1F92A01FFB34834CD42FEE6E6FC1B5E" box="[372,436,332,356]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
GK) sinuous; distal 0.2 noticeably arcuate in lateral or mesial view, slightly directed laterally in ventral view, gradually tapering, bearing 2 rows of minute stiff setae not extending to apex. Second gonopod (
<figureCitation id="A1F92A01FFB34834CCE6FE3EE6591B96" box="[208,273,404,428]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
L) short, sigmoidal, distal part spatulate, tip slightly recurved, sharply pointed.
</paragraph>
<paragraph id="397D3684FFB34834CCFFFE1DE38A182D" blockId="7.[151,1437,151,2012]" pageId="7" pageNumber="8">
<emphasis id="0BB6EA96FFB34834CCFFFE1DE6021BEA" bold="true" box="[201,330,439,464]" pageId="7" pageNumber="8">Coloration</emphasis>
. Carapace generally brown, whitish grooves between regions, cardiac and posterolateral regions. Chelipeds light brown on dorsal surface; outer surface of palm white. Ambulatory legs whitish with brown tinge on basal parts of dactyli, propodi, carpi and distal parts of meri. Thoracic sternum generally white.
</paragraph>
</subSubSection>
<subSubSection id="71D8650FFFB34834CCFFFD89E2D41806" box="[201,1436,547,572]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="397D3684FFB34834CCFFFD89E2D41806" blockId="7.[151,1437,151,2012]" box="[201,1436,547,572]" pageId="7" pageNumber="8">
<emphasis id="0BB6EA96FFB34834CCFFFD89E6131806" bold="true" box="[201,347,547,572]" pageId="7" pageNumber="8">Distribution</emphasis>
. Presently known only from
<collectingCountry id="41D57614FFB34834CEACFD8FE5901806" box="[666,728,549,572]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
(Sagami Sea and Ishikawa Prefecture, Sea of
<collectingCountry id="41D57614FFB34834C8E7FD8EE25C1806" box="[1233,1300,548,572]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
);
<quantity id="FE3A9B61FFB34834C910FD8EE2D01806" box="[1318,1432,548,572]" metricMagnitude="2" metricUnit="m" metricValue="1.2" metricValueMax="1.8" metricValueMin="0.6" pageId="7" pageNumber="8" unit="m" value="120.0" valueMax="180.0" valueMin="60.0">
60
<specimenCount id="2FC4FD0DFFB34834C967FD8EE2D01806" box="[1361,1432,548,572]" pageId="7" pageNumber="8" type="generic">180 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="71D8650FFFB3483BCCFFFDE2E6D41B96" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
<paragraph id="397D3684FFB34834CCFFFDE2E6B718D5" blockId="7.[151,1437,151,2012]" pageId="7" pageNumber="8">
<emphasis id="0BB6EA96FFB34834CCFFFDE2E67E185A" bold="true" box="[201,310,584,608]" pageId="7" pageNumber="8">Remarks</emphasis>
. The identity of
<taxonomicName id="FEC24D07FFB34834CDCCFDE2E58B185A" box="[506,707,584,608]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CDCCFDE2E58B185A" box="[506,707,584,608]" italics="true" pageId="7" pageNumber="8">Pilumnus dofleini</emphasis>
</taxonomicName>
is here established. The present male specimens, including one specimen studied by
<bibRefCitation id="5D534B75FFB34834CDB5FDC6E54518BE" author="Sakai" box="[387,525,620,644]" pageId="7" pageNumber="8" refString="Sakai, T. (1965) The Crabs of Sagami Bay. Maruzen, Tokyo, xvi + 206 pp. (English text), 100 pls., 92 pp. (Japanese text), 32 pp. (bibliography and index)." type="book" year="1965">Sakai (1965</bibRefCitation>
;
<bibRefCitation id="5D534B75FFB34834CE2FFDC6E51D18BE" author="Sakai" box="[537,597,620,644]" pageId="7" pageNumber="8" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd., Tokyo, xxix + 773 pp. (English text), 461 pp. (Japanese text), 251 pls." type="book" year="1976">1976</bibRefCitation>
), agree well with the
<typeStatus id="E6798826FFB34834CF71FDC6E4E118BE" box="[839,937,620,644]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
female in every non-sexual diagnostic character, including the presence of a submarginal row of distinct tubercles along the posterior margin of the carapace and spines arranged in irregular rows on the outer surfaces of the chelae. These male specimens enable us to provide male diagnostic characters.
</paragraph>
<paragraph id="397D3684FFB34834CCFFFD56E41B1FDE" blockId="7.[151,1437,151,2012]" pageId="7" pageNumber="8">
The material referred to
<taxonomicName id="FEC24D07FFB34834CDEFFD56E5D7192E" box="[473,671,764,788]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CDEFFD56E5D7192E" box="[473,671,764,788]" italics="true" pageId="7" pageNumber="8">Pilumnus dofleini</emphasis>
</taxonomicName>
by Ng (2000) is here described as
<taxonomicName id="FEC24D07FFB34834C815FD57E3FB192E" box="[1059,1203,765,788]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834C815FD57E3FB192E" box="[1059,1203,765,788]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="108557EDFFB34834C88FFD56E3B0192E" box="[1209,1272,764,788]" pageId="7" pageNumber="8" rank="species">
<emphasis id="0BB6EA96FFB34834C88FFD56E3B9192E" bold="true" box="[1209,1265,764,788]" pageId="7" pageNumber="8">n. sp</emphasis>
.
</taxonomicNameLabel>
<taxonomicName id="FEC24D07FFB34834C8C8FD56E7811902" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834C8C8FD56E7811902" italics="true" pageId="7" pageNumber="8">Pilumnus dofleini</emphasis>
</taxonomicName>
is readily distinguished from this new species by the following characters: spines or tubercles on the dorsal surface and margins on the carapace are more conspicuous in
<taxonomicName id="FEC24D07FFB34834CF63FCEFE48F1966" box="[853,967,836,860]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CF63FCEFE48F1966" box="[853,967,836,860]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
than in
<taxonomicName id="FEC24D07FFB34834C813FCEFE3FF1966" box="[1061,1207,837,860]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834C813FCEFE3FF1966" box="[1061,1207,837,860]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834C888FCEEE3B61966" bold="true" box="[1214,1278,836,860]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834C888FCEEE3B61966" box="[1214,1278,836,860]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, in particular, the cardiac region is armed with about 10 distinct spines or tubercles in
<emphasis id="0BB6EA96FFB34834CFE6FCC3E37719BA" box="[976,1087,872,896]" italics="true" pageId="7" pageNumber="8">P. d o f l e i n i</emphasis>
, whereas there are only a few indistinct tubercles on that region in
<taxonomicName id="FEC24D07FFB34834CE06FC27E5F7199E" box="[560,703,909,932]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834CE06FC27E5F7199E" box="[560,703,909,932]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834CEF1FC26E44D199E" bold="true" box="[711,773,908,932]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834CEF1FC26E44D199E" box="[711,773,908,932]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A1F92A01FFB34834CF23FC26E41C199E" box="[789,852,908,932]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2005]" captionTargetBox="[151,1436,174,1959]" captionTargetId="figure@2.[151,1436,172,1961]" captionTargetPageId="2" captionText="FIGURE 1. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, dorsal view; B, carapace, dorsal view." httpUri="https://zenodo.org/record/210232/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
B versus
<figureCitation id="A1F92A01FFB34834CF89FC26E347199E" box="[959,1039,908,932]" captionStart="FIGURE 12" captionStartId="17.[151,250,1885,1908]" captionTargetBox="[232,1369,182,1849]" captionTargetId="figure@17.[204,1390,169,1875]" captionTargetPageId="17" captionText="FIGURE 12. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, carapace, dorsal view (setae omitted); B, left upper orbital and anterolateral margins, dorsal view (setae omitted); C, frontal margin, anterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer) view (setae partially omitted); F, abdomen and telson, ventral view (setae omitted). Scale bars: 2 mm for A D, F; 1 mm for E." httpUri="https://zenodo.org/record/210243/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
A); grooves defining the regions on the dorsal surface of the carapace are more distinctly marked in
<taxonomicName id="FEC24D07FFB34834CF55FC1BE49B19F2" box="[867,979,944,968]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CF55FC1BE49B19F2" box="[867,979,944,968]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
than in
<taxonomicName id="FEC24D07FFB34834C818FC1BE3F719F2" box="[1070,1215,945,968]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834C818FC1BE3F719F2" box="[1070,1215,945,968]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834C8F3FC1AE24C19F2" bold="true" box="[1221,1284,944,968]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834C8F3FC1AE24C19F2" box="[1221,1284,944,968]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A1F92A01FFB34834C922FC05E21B19FD" box="[1300,1363,943,967]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2005]" captionTargetBox="[151,1436,174,1959]" captionTargetId="figure@2.[151,1436,172,1961]" captionTargetPageId="2" captionText="FIGURE 1. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, dorsal view; B, carapace, dorsal view." httpUri="https://zenodo.org/record/210232/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
B
<emphasis id="0BB6EA96FFB34834C959FC1BE7F319D6" italics="true" pageId="7" pageNumber="8">versus</emphasis>
<figureCitation id="A1F92A01FFB34834CCF4FC7EE65A19D6" box="[194,274,980,1004]" captionStart="FIGURE 12" captionStartId="17.[151,250,1885,1908]" captionTargetBox="[232,1369,182,1849]" captionTargetId="figure@17.[204,1390,169,1875]" captionTargetPageId="17" captionText="FIGURE 12. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, carapace, dorsal view (setae omitted); B, left upper orbital and anterolateral margins, dorsal view (setae omitted); C, frontal margin, anterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer) view (setae partially omitted); F, abdomen and telson, ventral view (setae omitted). Scale bars: 2 mm for A D, F; 1 mm for E." httpUri="https://zenodo.org/record/210243/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
A); there is a submarginal row of conspicuous tubercles adjacent to the posterior margin of the carapace in
<taxonomicName id="FEC24D07FFB34834CC82FC53E66C1E2A" box="[180,292,1016,1040]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CC82FC53E66C1E2A" box="[180,292,1016,1040]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
, but such tubercles are absent in
<emphasis id="0BB6EA96FFB34834CEACFC53E46E1E2A" box="[666,806,1017,1040]" italics="true" pageId="7" pageNumber="8">P. c u r v i p e n i s</emphasis>
<emphasis id="0BB6EA96FFB34834CF07FC52E4381E2A" bold="true" box="[817,880,1016,1040]" pageId="7" pageNumber="8">n. sp.</emphasis>
(
<figureCitation id="A1F92A01FFB34834CFB6FC5DE48A1E35" box="[896,962,1015,1039]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
C versus
<figureCitation id="A1F92A01FFB34834C81DFC5DE3331E35" box="[1067,1147,1015,1039]" captionStart="FIGURE 12" captionStartId="17.[151,250,1885,1908]" captionTargetBox="[232,1369,182,1849]" captionTargetId="figure@17.[204,1390,169,1875]" captionTargetPageId="17" captionText="FIGURE 12. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, carapace, dorsal view (setae omitted); B, left upper orbital and anterolateral margins, dorsal view (setae omitted); C, frontal margin, anterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer) view (setae partially omitted); F, abdomen and telson, ventral view (setae omitted). Scale bars: 2 mm for A D, F; 1 mm for E." httpUri="https://zenodo.org/record/210243/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
A); the upper orbital margin has two small clefts in
<taxonomicName id="FEC24D07FFB34834CDF5FBB7E57B1E0E" box="[451,563,1052,1076]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CDF5FBB7E57B1E0E" box="[451,563,1052,1076]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
, whereas no distinct clefts are discernible in
<taxonomicName id="FEC24D07FFB34834C81AFBB7E3F51E0E" box="[1068,1213,1053,1076]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834C81AFBB7E3F51E0E" box="[1068,1213,1053,1076]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834C8F2FBB6E24A1E0E" bold="true" box="[1220,1282,1052,1076]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834C8F2FBB6E24A1E0E" box="[1220,1282,1052,1076]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A1F92A01FFB34834C924FBB6E21A1E0E" box="[1298,1362,1052,1076]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
A versus
<figureCitation id="A1F92A01FFB34834CCF3FBEAE6511E62" box="[197,281,1088,1112]" captionStart="FIGURE 12" captionStartId="17.[151,250,1885,1908]" captionTargetBox="[232,1369,182,1849]" captionTargetId="figure@17.[204,1390,169,1875]" captionTargetPageId="17" captionText="FIGURE 12. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, carapace, dorsal view (setae omitted); B, left upper orbital and anterolateral margins, dorsal view (setae omitted); C, frontal margin, anterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer) view (setae partially omitted); F, abdomen and telson, ventral view (setae omitted). Scale bars: 2 mm for A D, F; 1 mm for E." httpUri="https://zenodo.org/record/210243/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
B); the pterygostomial groove of the carapace is flanked by rows of distinct tubercles in
<taxonomicName id="FEC24D07FFB34834C917FBEBE2DC1E62" box="[1313,1428,1088,1112]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834C917FBEBE2DC1E62" box="[1313,1428,1088,1112]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
, whereas such rows of tubercles are absent or poorly developed in
<taxonomicName id="FEC24D07FFB34834CFBFFBCFE3531E46" box="[905,1051,1125,1148]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834CFBFFBCFE3531E46" box="[905,1051,1125,1148]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834C815FBCEE32C1E46" bold="true" box="[1059,1124,1124,1148]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834C815FBCEE32C1E46" box="[1059,1124,1124,1148]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A1F92A01FFB34834C843FBCEE3F01E46" box="[1141,1208,1124,1148]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
D versus
<figureCitation id="A1F92A01FFB34834C910FBCEE2301E46" box="[1318,1400,1124,1148]" captionStart="FIGURE 12" captionStartId="17.[151,250,1885,1908]" captionTargetBox="[232,1369,182,1849]" captionTargetId="figure@17.[204,1390,169,1875]" captionTargetPageId="17" captionText="FIGURE 12. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, carapace, dorsal view (setae omitted); B, left upper orbital and anterolateral margins, dorsal view (setae omitted); C, frontal margin, anterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer) view (setae partially omitted); F, abdomen and telson, ventral view (setae omitted). Scale bars: 2 mm for A D, F; 1 mm for E." httpUri="https://zenodo.org/record/210243/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
D); spines on the palms of the chelae are relatively less numerous and stronger in
<taxonomicName id="FEC24D07FFB34834CFC1FB23E3201E9A" box="[1015,1128,1160,1184]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CFC1FB23E3201E9A" box="[1015,1128,1160,1184]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
than in
<taxonomicName id="FEC24D07FFB34834C8F6FB23E2181E9A" box="[1216,1360,1161,1184]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834C8F6FB23E2181E9A" box="[1216,1360,1161,1184]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834C960FB22E2DD1E9A" bold="true" box="[1366,1429,1160,1184]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834C960FB22E2DD1E9A" box="[1366,1429,1160,1184]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, with those on the median area are arranged in irregular rows in
<taxonomicName id="FEC24D07FFB34834CF5EFB07E4911EFE" box="[872,985,1196,1220]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CF5EFB07E4911EFE" box="[872,985,1196,1220]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
rather than scattered in
<taxonomicName id="FEC24D07FFB34834C8DAFB07E2361EFE" box="[1260,1406,1197,1220]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834C8DAFB07E2361EFE" box="[1260,1406,1197,1220]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834C9B3FB06E7F01ED2" bold="true" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834C9B3FB06E7F01ED2" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A1F92A01FFB34834CCFEFB7AE6431ED2" box="[200,267,1232,1256]" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[214,1359,188,1883]" captionTargetId="figure@5.[186,1410,173,1906]" captionTargetPageId="5" captionText="FIGURE 4. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, right major chela, outer view; B, carpus of right major cheliped, dorsal view; C, merus of right major cheliped, outer view; D, left minor chela, outer view; E, merus of left minor cheliped, outer view; F, left third ambulatory leg (fourth pereopod), posterior view; G, left fourth ambulatory leg (fifth pereopod), posterior view). Setae omitted. Scale bars: 2 mm." httpUri="https://zenodo.org/record/210235/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
A, D versus
<figureCitation id="A1F92A01FFB34834CDA9FB7AE6A61ED2" box="[415,494,1232,1256]" captionStart="FIGURE 13" captionStartId="18.[151,250,1887,1910]" captionTargetBox="[224,1371,209,1858]" captionTargetId="figure@18.[203,1389,193,1870]" captionTargetPageId="18" captionText="FIGURE 13. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, right major chela, outer view; B, carpus of right major cheliped, dorsal view; C, left minor chela, outer view; D, left third ambulatory leg (fourth pereopod), posterior view; E, left fourth ambulatory leg (fifth pereopod), posterior view; F, left first gonopod, mesial view; G, same, lateral view; H, same, distal part, mesial view; I, same, lateral view. Setae omitted. Scale bars: 2 mm for A E; 1 mm for F, G; 0.2 mm for H, I." httpUri="https://zenodo.org/record/210244/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
A, C); the ambulatory legs are relatively more slender in
<emphasis id="0BB6EA96FFB34834C846FB7BE3961ED2" box="[1136,1246,1232,1256]" italics="true" pageId="7" pageNumber="8">P. d o f l e i n i</emphasis>
than in
<emphasis id="0BB6EA96FFB34834C90BFB7BE79A1F36" italics="true" pageId="7" pageNumber="8">P. c u r v i - penis</emphasis>
<emphasis id="0BB6EA96FFB34834CCECFB5EE6521F36" bold="true" box="[218,282,1268,1292]" pageId="7" pageNumber="8">n. sp.</emphasis>
(
<figureCitation id="A1F92A01FFB34834CD1DFB5EE6261F36" box="[299,366,1268,1292]" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[214,1359,188,1883]" captionTargetId="figure@5.[186,1410,173,1906]" captionTargetPageId="5" captionText="FIGURE 4. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, right major chela, outer view; B, carpus of right major cheliped, dorsal view; C, merus of right major cheliped, outer view; D, left minor chela, outer view; E, merus of left minor cheliped, outer view; F, left third ambulatory leg (fourth pereopod), posterior view; G, left fourth ambulatory leg (fifth pereopod), posterior view). Setae omitted. Scale bars: 2 mm." httpUri="https://zenodo.org/record/210235/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
F, G versus
<figureCitation id="A1F92A01FFB34834CDC0FB5EE50F1F36" box="[502,583,1268,1292]" captionStart="FIGURE 13" captionStartId="18.[151,250,1887,1910]" captionTargetBox="[224,1371,209,1858]" captionTargetId="figure@18.[203,1389,193,1870]" captionTargetPageId="18" captionText="FIGURE 13. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, right major chela, outer view; B, carpus of right major cheliped, dorsal view; C, left minor chela, outer view; D, left third ambulatory leg (fourth pereopod), posterior view; E, left fourth ambulatory leg (fifth pereopod), posterior view; F, left first gonopod, mesial view; G, same, lateral view; H, same, distal part, mesial view; I, same, lateral view. Setae omitted. Scale bars: 2 mm for A E; 1 mm for F, G; 0.2 mm for H, I." httpUri="https://zenodo.org/record/210244/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
D, E); the extensor margins of the dactyli of the ambulatory legs are armed with small spines or tubercles in
<taxonomicName id="FEC24D07FFB34834CE30FAB3E53E1F0A" box="[518,630,1304,1328]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CE30FAB3E53E1F0A" box="[518,630,1304,1328]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
, but unarmed in
<taxonomicName id="FEC24D07FFB34834CF02FAB3E48D1F0A" box="[820,965,1305,1328]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834CF02FAB3E48D1F0A" box="[820,965,1305,1328]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834CFFDFAB2E3421F0A" bold="true" box="[971,1034,1304,1328]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834CFFDFAB2E3421F0A" box="[971,1034,1304,1328]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A1F92A01FFB34834C82FFABDE3131F15" box="[1049,1115,1303,1327]" captionStart="FIGURE 4" captionStartId="5.[151,250,1920,1943]" captionTargetBox="[214,1359,188,1883]" captionTargetId="figure@5.[186,1410,173,1906]" captionTargetPageId="5" captionText="FIGURE 4. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, right major chela, outer view; B, carpus of right major cheliped, dorsal view; C, merus of right major cheliped, outer view; D, left minor chela, outer view; E, merus of left minor cheliped, outer view; F, left third ambulatory leg (fourth pereopod), posterior view; G, left fourth ambulatory leg (fifth pereopod), posterior view). Setae omitted. Scale bars: 2 mm." httpUri="https://zenodo.org/record/210235/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
F, G versus
<figureCitation id="A1F92A01FFB34834C8D6FABDE2671F15" box="[1248,1327,1303,1327]" captionStart="FIGURE 13" captionStartId="18.[151,250,1887,1910]" captionTargetBox="[224,1371,209,1858]" captionTargetId="figure@18.[203,1389,193,1870]" captionTargetPageId="18" captionText="FIGURE 13. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, right major chela, outer view; B, carpus of right major cheliped, dorsal view; C, left minor chela, outer view; D, left third ambulatory leg (fourth pereopod), posterior view; E, left fourth ambulatory leg (fifth pereopod), posterior view; F, left first gonopod, mesial view; G, same, lateral view; H, same, distal part, mesial view; I, same, lateral view. Setae omitted. Scale bars: 2 mm for A E; 1 mm for F, G; 0.2 mm for H, I." httpUri="https://zenodo.org/record/210244/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
D, E); the thoracic sternum is distinctly granular in
<taxonomicName id="FEC24D07FFB34834CE50FA97E59F1F6E" box="[614,727,1340,1364]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CE50FA97E59F1F6E" box="[614,727,1340,1364]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
, rather than nearly smooth in
<taxonomicName id="FEC24D07FFB34834C81DFA97E3F41F6E" box="[1067,1212,1341,1364]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834C81DFA97E3F41F6E" box="[1067,1212,1341,1364]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834C8F5FA96E24A1F6E" bold="true" box="[1219,1282,1340,1364]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834C8F5FA96E24A1F6E" box="[1219,1282,1340,1364]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A1F92A01FFB34834C924FA96E21D1F6E" box="[1298,1365,1340,1364]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1977]" captionTargetBox="[151,1436,299,1933]" captionTargetId="figure@3.[151,1436,299,1934]" captionTargetPageId="3" captionText="FIGURE 2. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, ventral view; B, thoracic sternum and abdomen, ventral view." httpUri="https://zenodo.org/record/210233/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
B versus
<figureCitation id="A1F92A01FFB34834CCF2FAF5E64C1F4D" box="[196,260,1375,1399]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2005]" captionTargetBox="[151,1436,174,1959]" captionTargetId="figure@2.[151,1436,172,1961]" captionTargetPageId="2" captionText="FIGURE 1. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, entire animal, dorsal view; B, carapace, dorsal view." httpUri="https://zenodo.org/record/210232/files/figure.png" pageId="7" pageNumber="8">
Fig.
<quantity id="FE3A9B61FFB34834CCCFFAF5E64C1F4D" box="[249,260,1375,1399]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="7" pageNumber="8" unit="g" value="1.0">1</quantity>
</figureCitation>
g of Ng 2000); the first abdominal somite is armed with several distinct tubercles in
<taxonomicName id="FEC24D07FFB34834C8EFFACBE2031F42" box="[1241,1355,1376,1400]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834C8EFFACBE2031F42" box="[1241,1355,1376,1400]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
, but is unarmed in
<taxonomicName id="FEC24D07FFB34834CD28FA2FE6F81FA6" box="[286,432,1413,1436]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834CD28FA2FE6F81FA6" box="[286,432,1413,1436]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834CD8EFA2EE6B01FA6" bold="true" box="[440,504,1412,1436]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834CD8EFA2EE6B01FA6" box="[440,504,1412,1436]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A1F92A01FFB34834CE3EFA2EE5031FA6" box="[520,587,1412,1436]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
C, F versus
<figureCitation id="A1F92A01FFB34834CEE3FA2EE46E1FA6" box="[725,806,1412,1436]" captionStart="FIGURE 12" captionStartId="17.[151,250,1885,1908]" captionTargetBox="[232,1369,182,1849]" captionTargetId="figure@17.[204,1390,169,1875]" captionTargetPageId="17" captionText="FIGURE 12. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, carapace, dorsal view (setae omitted); B, left upper orbital and anterolateral margins, dorsal view (setae omitted); C, frontal margin, anterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer) view (setae partially omitted); F, abdomen and telson, ventral view (setae omitted). Scale bars: 2 mm for A D, F; 1 mm for E." httpUri="https://zenodo.org/record/210243/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
F); the first gonopod is more strongly sinuous in
<taxonomicName id="FEC24D07FFB34834C964FA2FE7811FFA" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834C964FA2FE7811FFA" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
than in
<taxonomicName id="FEC24D07FFB34834CD13FA03E6FD1FFA" box="[293,437,1449,1472]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834CD13FA03E6FD1FFA" box="[293,437,1449,1472]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834CD8BFA02E6B41FFA" bold="true" box="[445,508,1448,1472]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834CD8BFA02E6B41FFA" box="[445,508,1448,1472]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, with its distal part arcuate in the distal
<quantity id="FE3A9B61FFB34834CFF4FA0DE34A1F85" box="[962,1026,1447,1471]" metricMagnitude="-3" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="8" unit="in" value="0.2">0.2 in</quantity>
<emphasis id="0BB6EA96FFB34834C83FFA03E33F1FFA" box="[1033,1143,1448,1472]" italics="true" pageId="7" pageNumber="8">P. d o f l e i n i</emphasis>
, but strongly curved near the apex in
<taxonomicName id="FEC24D07FFB34834CD2FFA67E6E11FDE" box="[281,425,1485,1508]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="curvipenis" status="sp. nov.">
<emphasis id="0BB6EA96FFB34834CD2FFA67E6E11FDE" box="[281,425,1485,1508]" italics="true" pageId="7" pageNumber="8">P. curvipenis</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFB34834CD86FA66E6A61FDE" bold="true" box="[432,494,1484,1508]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="108557EDFFB34834CD86FA66E6A61FDE" box="[432,494,1484,1508]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A1F92A01FFB34834CDC8FA66E5771FDE" box="[510,575,1484,1508]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
GK versus
<figureCitation id="A1F92A01FFB34834CEFAFA66E4531FDE" box="[716,795,1484,1508]" captionStart="FIGURE 13" captionStartId="18.[151,250,1887,1910]" captionTargetBox="[224,1371,209,1858]" captionTargetId="figure@18.[203,1389,193,1870]" captionTargetPageId="18" captionText="FIGURE 13. Pilumnus curvipenis n. sp., holotype, male (11.4 x 14.2 mm), NSMT-Cr 6665. A, right major chela, outer view; B, carpus of right major cheliped, dorsal view; C, left minor chela, outer view; D, left third ambulatory leg (fourth pereopod), posterior view; E, left fourth ambulatory leg (fifth pereopod), posterior view; F, left first gonopod, mesial view; G, same, lateral view; H, same, distal part, mesial view; I, same, lateral view. Setae omitted. Scale bars: 2 mm for A E; 1 mm for F, G; 0.2 mm for H, I." httpUri="https://zenodo.org/record/210244/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
FI).
</paragraph>
<paragraph id="397D3684FFB34834CCFFFA45E6D81CAD" blockId="7.[151,1437,151,2012]" pageId="7" pageNumber="8">
<bibRefCitation id="5D534B75FFB34834CCFFFA45E61A1C3D" author="Sakai" box="[201,338,1519,1543]" pageId="7" pageNumber="8" refString="Sakai, T. (1965) The Crabs of Sagami Bay. Maruzen, Tokyo, xvi + 206 pp. (English text), 100 pls., 92 pp. (Japanese text), 32 pp. (bibliography and index)." type="book" year="1965">Sakai (1965</bibRefCitation>
;
<bibRefCitation id="5D534B75FFB34834CD6BFA45E6DF1C3D" author="Sakai" box="[349,407,1519,1543]" pageId="7" pageNumber="8" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd., Tokyo, xxix + 773 pp. (English text), 461 pp. (Japanese text), 251 pls." type="book" year="1976">1976</bibRefCitation>
) reported
<taxonomicName id="FEC24D07FFB34834CE3CFA5BE5321C32" box="[522,634,1520,1544]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CE3CFA5BE5321C32" box="[522,634,1520,1544]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
on the basis of material from Sagami Bay made by the Showa Emperor of
<collectingCountry id="41D57614FFB34834CC83F9BFE7BD1C16" box="[181,245,1557,1580]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
(now housed in the collection of the Showa Memorial Institute of Tsukuba Research Center, National Science Museum, Tokyo).
<bibRefCitation id="5D534B75FFB34834CDA9F99DE5F21C75" author="Takeda" box="[415,698,1591,1615]" pageId="7" pageNumber="8" refString="Takeda, M. &amp; Manuel, M. R. (2000) Taxonomic accounts of some rare crabs new to the Philippines. National Science Museum Monographs, 18, 149 - 162." type="journal article" year="2000">Takeda &amp; Manuel (2000)</bibRefCitation>
presented a precise list of those specimens, in which the presence of ten lots was clarified. During this study, a single lot (NSMT-Cr R2507) was reexamined, of which the identification has been verified.
</paragraph>
<paragraph id="397D3684FFB34834CCFFF90EE4EF1D4A" blockId="7.[151,1437,151,2012]" pageId="7" pageNumber="8">
<bibRefCitation id="5D534B75FFB34834CCFFF90EE6391C86" author="Miyake" box="[201,369,1700,1724]" pageId="7" pageNumber="8" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in Color, Vol. II. Brachyura (Crabs). Hoikusha, Osaka, 277 pp., 64 pls." type="book" year="1983">Miyake (1983)</bibRefCitation>
reported
<taxonomicName id="FEC24D07FFB34834CDE8F90FE5181C86" box="[478,592,1700,1724]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CDE8F90FE5181C86" box="[478,592,1700,1724]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
based on a single male specimen from Tosa Bay,
<collectingCountry id="41D57614FFB34834C8B1F90EE3831C86" box="[1159,1227,1700,1724]" name="India" pageId="7" pageNumber="8">Kochi</collectingCountry>
Prefecture,
<collectingCountry id="41D57614FFB34834C963F90FE2D01C86" box="[1365,1432,1701,1724]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
. However, this specimen seems to be different from the present specimens of
<taxonomicName id="FEC24D07FFB34834CFCEF963E3211CDA" box="[1016,1129,1736,1760]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CFCEF963E3211CDA" box="[1016,1129,1736,1760]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
in the dorsally naked carapace, the weaker armature on the cheliped, the lack of long dorsal or extensor spines on the ambulatory meri to propodi, and the purplish body color. There is little doubt that the specimen used by
<bibRefCitation id="5D534B75FFB34834C870F8BAE3A51D12" author="Miyake" box="[1094,1261,1808,1832]" pageId="7" pageNumber="8" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in Color, Vol. II. Brachyura (Crabs). Hoikusha, Osaka, 277 pp., 64 pls." type="book" year="1983">Miyake (1983)</bibRefCitation>
does not represent
<taxonomicName id="FEC24D07FFB34834CCFDF89FE6741D76" box="[203,316,1844,1868]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834CCFDF89FE6741D76" box="[203,316,1844,1868]" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
. Our efforts to locate Miyakes specimen has not been successful, and the determination of the specific identity of Miyakes (1983) specimen is not possible at this time.
</paragraph>
<paragraph id="397D3684FFB3483BCCFFF8D6E6701AC2" blockId="7.[151,1437,151,2012]" lastBlockId="8.[151,1436,151,428]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<bibRefCitation id="5D534B75FFB34834CCFFF8D6E6BB1DAE" author="Takeda" box="[201,499,1916,1940]" pageId="7" pageNumber="8" refString="Takeda, M. &amp; Manuel, M. R. (2000) Taxonomic accounts of some rare crabs new to the Philippines. National Science Museum Monographs, 18, 149 - 162." type="journal article" year="2000">Takeda &amp; Manuel (2000)</bibRefCitation>
identified one female specimen from Balicasag Island, the
<collectingCountry id="41D57614FFB34834C88FF8D6E2721DAE" box="[1209,1338,1916,1940]" name="Philippines" pageId="7" pageNumber="8">Philippines</collectingCountry>
, with
<taxonomicName id="FEC24D07FFB34834C9B1F8D7E7A41D82" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFB34834C9B1F8D7E7A41D82" italics="true" pageId="7" pageNumber="8">P. dofleini</emphasis>
</taxonomicName>
. Although this specimen was not reexamined, it is assumed that it actually represents
<emphasis id="0BB6EA96FFB34834C8E0F80BE2041D82" box="[1238,1356,1953,1976]" italics="true" pageId="7" pageNumber="8">P. a r m a t u s</emphasis>
<emphasis id="0BB6EA96FFB34834C96CF80AE2D41D82" bold="true" box="[1370,1436,1952,1976]" pageId="7" pageNumber="8">n. sp.</emphasis>
based on examination of the given photograph (
<bibRefCitation id="5D534B75FFB34834CE8EF86EE3671DE6" author="Takeda" box="[696,1071,1988,2012]" pageId="7" pageNumber="8" refString="Takeda, M. &amp; Manuel, M. R. (2000) Taxonomic accounts of some rare crabs new to the Philippines. National Science Museum Monographs, 18, 149 - 162." type="journal article" year="2000">
Takeda &amp; Manuel 2000:
<figureCitation id="A1F92A01FFB34834CFEFF86EE3541DE6" box="[985,1052,1988,2012]" captionStart="FIGURE 3" captionStartId="4.[151,250,1842,1865]" captionTargetBox="[244,1340,209,1801]" captionTargetId="figure@4.[226,1368,193,1814]" captionTargetPageId="4" captionText="FIGURE 3. Pilumnus dofleini Balss, 1933, male (12.5 x 15.3 mm), CBM-ZC 10621. A, left upper orbital and anterolateral margins, dorsal view (setae omitted); B, frontal margin, anterodorsal view (setae omitted); C, posterior margin of carapace and first abdominal somite, posterodorsal view (setae omitted); D, left anterior part of carapace and cephalic appendages, anterior view (setae omitted); E, left third maxilliped, ventral (outer view; setae partially omitted); F, abdomen and telson, ventral view (setae omitted); G, left first gonopod, ventral view; H, same, mesial view; I, same, distal part, ventral view; J, same, mesial view; K, same, lateral view; L. left second gonopod. Scale bars: 2 mm for A, B, D, F; 1 mm for C, E, G, H; 0.5 mm for I L." httpUri="https://zenodo.org/record/210234/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
F
</bibRefCitation>
). The photograph clearly shows the absence of tubercles on the cardiac region of the carapace and the presence of a row of conspicuous tubercles along the posterior margin of the carapace and on the first abdominal somite, representing diagnostic features of
<taxonomicName id="FEC24D07FFBC483BC9B1FF17E7BB1AC2" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="armatus" status="sp. nov.">
<emphasis id="0BB6EA96FFBC483BC9B1FF17E7BB1AC2" italics="true" pageId="8" pageNumber="9">P. armatus</emphasis>
</taxonomicName>
<emphasis id="0BB6EA96FFBC483BCCCCFF4AE6701AC2" bold="true" box="[250,312,224,248]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="108557EDFFBC483BCCCCFF4AE6701AC2" box="[250,312,224,248]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="397D3684FFBC483BCCFFFEAEE6D41B96" blockId="8.[151,1436,151,428]" pageId="8" pageNumber="9">
The record of
<taxonomicName id="FEC24D07FFBC483BCD5DFEAFE6951B26" box="[363,477,260,284]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFBC483BCD5DFEAFE6951B26" box="[363,477,260,284]" italics="true" pageId="8" pageNumber="9">P. dofleini</emphasis>
</taxonomicName>
by
<bibRefCitation id="5D534B75FFBC483BCE3FFEAFE4021B26" author="Marumura" box="[521,842,260,284]" pageId="8" pageNumber="9" refString="Marumura, M. &amp; Kosaka, A. (2003) Catalog of the Brachyuran and Anomuran Crabs Donated by the Late Mr. Seiji Nagai to the Wakayama Prefectural Museum of Natural History. 74 pp. Wakayama Prefectural Museum of Natural History, Kainan." type="book" year="2003">Marumura &amp; Kosaka (2003)</bibRefCitation>
based on two specimens from Tosa Bay needs to be verified. Ng
<emphasis id="0BB6EA96FFBC483BCD1AFE83E62F1B7A" box="[300,359,296,320]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2008) published a color photograph of an ovigerous female specimen from the
<collectingCountry id="41D57614FFBC483BC921FE8DE2D01B05" box="[1303,1432,295,319]" name="Philippines" pageId="8" pageNumber="9">Philippines</collectingCountry>
, referred to as
<taxonomicName id="FEC24D07FFBC483BCD06FEE7E6E91B5E" box="[304,417,332,356]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFBC483BCD06FEE7E6E91B5E" box="[304,417,332,356]" italics="true" pageId="8" pageNumber="9">P. dofleini</emphasis>
</taxonomicName>
, although
<taxonomicName id="FEC24D07FFBC483BCE21FEE7E5C01B5E" box="[535,648,332,356]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dofleini">
<emphasis id="0BB6EA96FFBC483BCE21FEE7E5C01B5E" box="[535,648,332,356]" italics="true" pageId="8" pageNumber="9">P. dofleini</emphasis>
</taxonomicName>
was inadvertently omitted from their species list (Ng et al. 2008: 141 142). This specimen is referred to
<taxonomicName id="FEC24D07FFBC483BCE23FEDBE5381BB2" box="[533,624,368,392]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bohol" status="sp. nov.">
<emphasis id="0BB6EA96FFBC483BCE23FEDBE5381BB2" box="[533,624,368,392]" italics="true" pageId="8" pageNumber="9">P. bohol</emphasis>
</taxonomicName>
<taxonomicNameLabel id="108557EDFFBC483BCE40FEDAE5FD1BBD" box="[630,693,368,392]" pageId="8" pageNumber="9" rank="species">
<emphasis id="0BB6EA96FFBC483BCE40FEDAE5E61BB2" bold="true" box="[630,686,368,392]" pageId="8" pageNumber="9">n. sp</emphasis>
.
</taxonomicNameLabel>
because of the weak armature on the carapace, although it was not available for this study.
</paragraph>
</subSubSection>
</treatment>
</document>