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<document id="2E413EBB554482CEC37DEE4CCA8D6606" ID-DOI="10.5281/zenodo.5393502" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393502" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630318937109" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques &amp; Mauriès, Jean-Paul" docDate="2006" docId="03FEF24BA9776A48FC86FCDA1DB1C3CF" docLanguage="en" docName="Zoosystema.2006.28.1.75-92.pdf" docOrigin="Zoosystema 28 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Nepalella marmorata Golovatch &amp; Geoffroy &amp; Mauriès 2006, n. sp." docType="treatment" docVersion="3" lastPageNumber="80" masterDocId="FFC78A33A9756A4DFF9CFFAB1F53C162" masterDocTitle="Four new Chordeumatida (Diplopoda) from caves in China" masterLastPageNumber="92" masterPageNumber="75" pageNumber="77" updateTime="1699225294555" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="4B1903982948D592BB91AB1326EF1985" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="433C20AC3A1B4B1B0721828E34158BBE">
<mods:title id="BA863A67CF0985647ACCEC95A57808EB">Four new Chordeumatida (Diplopoda) from caves in China</mods:title>
</mods:titleInfo>
<mods:name id="83A688FA321125BAB4A7B8B6B4A7B1CE" type="personal">
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<mods:namePart id="CA71766830ABC47F1F8D9EDDE5B47E69">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="F5BFA3FF3442D20CDC8069234041101D">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
<mods:nameIdentifier id="6A268D877159D2F35C6F62F5DD9469B2" type="email">sgol@orc.ru</mods:nameIdentifier>
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<mods:namePart id="0F0FCA9E3F1A35EA4095496727474486">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="71A7D19730A722BB02784126B18E6411">Muséum national dHistoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du Sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="C0EB8095F1CD473004753CB632CB3E2C" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
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<mods:name id="539F5D45A3F4266A0B3ABAE6C2AD2534" type="personal">
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</mods:role>
<mods:namePart id="E6644B82AF412DDA3582DF0AD74F560B">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="3E4A4D1AA9E143985FFE90E61125235C">Muséum national dHistoire naturelle, Département Systématique et Évolution, USM 602 Taxonomie et Collections, case postale 53, 61 rue Buffon, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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</mods:name>
<mods:typeOfResource id="2ABB200CD101263F3C4ACC55A49EE655">text</mods:typeOfResource>
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<mods:title id="671CB3E0FD1782F05B32DEC904D77E09">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="7F6ECF9B5A1342E4F8680A8A8A72BD70">
<mods:date id="E7134A2BB3A2136819FCB6701DC82180">2006</mods:date>
<mods:detail id="6C59D19077B478D4EB43F1975FAB2497" type="volume">
<mods:number id="C136EB2152C86460B88BC5C55F9D77F6">28</mods:number>
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<mods:start id="FB90144AB3111C5A4177507D5F45782A">75</mods:start>
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<mods:identifier id="EB419DC779FBF2AF0A02CD4F313B80BD" type="DOI">10.5281/zenodo.5393502</mods:identifier>
<mods:identifier id="D5FC2EEBB1FA5FC26B5D2B69173B8CBF" type="ISSN">1638-9387</mods:identifier>
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<treatment id="03FEF24BA9776A48FC86FCDA1DB1C3CF" LSID="urn:lsid:plazi:treatment:03FEF24BA9776A48FC86FCDA1DB1C3CF" httpUri="http://treatment.plazi.org/id/03FEF24BA9776A48FC86FCDA1DB1C3CF" lastPageId="5" lastPageNumber="80" pageId="2" pageNumber="77">
<subSubSection id="C34D10D6A9776A4FFC86FCDA1B17C2EE" box="[794,1092,881,908]" pageId="2" pageNumber="77" type="nomenclature">
<paragraph id="8BE8435DA9776A4FFC86FCDA1B17C2EE" blockId="2.[794,1092,881,939]" box="[794,1092,881,908]" pageId="2" pageNumber="77">
<heading id="D0A0F431A9776A4FFC86FCDA1B17C2EE" box="[794,1092,881,908]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="77" reason="2">
<taxonomicName id="4C5738DEA9776A4FFC86FCDA1B50C2E9" ID-CoL="46WVZ" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[794,1027,881,908]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="marmorata" status="sp. nov.">
<emphasis id="B9239F4FA9776A4FFC86FCDA1B50C2E9" bold="true" box="[794,1027,881,908]" italics="true" pageId="2" pageNumber="77">Nepalella marmorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9776A4FFB95FCD91B17C2EE" box="[1033,1092,882,908]" pageId="2" pageNumber="77" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9776A4FFCA7FC3A1B70C2C9" box="[827,1059,913,939]" pageId="2" pageNumber="77" type="description">
<paragraph id="8BE8435DA9776A4FFCA7FC3A1B70C2C9" blockId="2.[794,1092,881,939]" box="[827,1059,913,939]" pageId="2" pageNumber="77">
(
<figureCitation id="136C5FD8A9776A4FFCDEFC3A1C2CC2C9" box="[834,895,913,939]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="2" pageNumber="77">Figs 1</figureCitation>
A-C; 2; 3A, B)
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9776A4FFD39FC7C1B23C55C" pageId="2" pageNumber="77" type="materials_examined">
<paragraph id="8BE8435DA9776A4FFD39FC7C1B23C55C" blockId="2.[677,1210,983,1086]" pageId="2" pageNumber="77">
<materialsCitation id="3B3F4900A9776A4FFD39FC7C1C9DC55C" collectingDate="1999-08-05" collectionCode="MNHN" collectorName="Snake Mouth Cave, No. &amp; J. &amp; B. Lips" country="China" county="Xin Long County" location="Snake Mouth Cave" pageId="2" pageNumber="77" specimenCode="DB21" specimenCount="2" specimenCount-male="2" stateProvince="Sichuan" typeStatus="holotype">
TYPE MATERIAL. —
<typeStatus id="54ECFDFFA9776A4FFCC2FC7C1CEFC28D" box="[862,956,983,1007]" pageId="2" pageNumber="77" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34003CDA9776A4FFC5FFC7C1B52C28D" box="[963,1025,983,1007]" name="China" pageId="2" pageNumber="77">China</collectingCountry>
,
<collectingRegion id="49938DBFA9776A4FFB96FC7C1BDCC28C" box="[1034,1167,983,1007]" country="China" name="Sichuan" pageId="2" pageNumber="77">Sichuan Prov.</collectingRegion>
,
<collectingCounty id="62893BD1A9776A4FFB09FC7C1C70C568" pageId="2" pageNumber="77">Xin Long County</collectingCounty>
,
<collectorName id="26A2268BA9776A4FFCB0FC5A1B5DC56B" box="[812,1038,1009,1034]" pageId="2" pageNumber="77">
<location id="8E881586A9776A4FFCB0FC5A1CB2C568" LSID="urn:lsid:plazi:treatment:03FEF24BA9776A48FC86FCDA1DB1C3CF:8E881586A9776A4FFCB0FC5A1CB2C568" box="[812,993,1009,1034]" country="China" county="Xin Long County" name="Snake Mouth Cave" pageId="2" pageNumber="77" stateProvince="Sichuan">Snake Mouth Cave</location>
, No.
</collectorName>
389,
<date id="FFE9659DA9776A4FFBDCFC5A1BE6C56B" box="[1088,1205,1009,1033]" pageId="2" pageNumber="77" value="1999-08-05">
<collectingDate id="EFAD9C75A9776A4FFBDCFC5A1BE6C56B" box="[1088,1205,1009,1033]" pageId="2" pageNumber="77" value="1999-08-05">5.VIII.1999</collectingDate>
</date>
, leg.
<collectorName id="26A2268BA9776A4FFD51FBA71C14C546" box="[717,839,1036,1060]" pageId="2" pageNumber="77">J. &amp; B. Lips</collectorName>
,
<specimenCount id="9D5188D4A9776A4FFCCEFBA41C35C541" box="[850,870,1039,1059]" pageId="2" pageNumber="77" type="male"></specimenCount>
(
<collectionCode id="ED46DB98A9776A4FFCE8FBA71C9BC546" box="[884,968,1036,1060]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="77">MNHN</collectionCode>
<specimenCode id="DBF1EB26A9776A4FFC53FBA71B5CC546" box="[975,1039,1036,1060]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="77">DB21</specimenCode>
).
<typeStatus id="54ECFDFFA9776A4FFBBCFBA71B2CC546" box="[1056,1151,1036,1060]" pageId="2" pageNumber="77" type="paratype">Paratypes</typeStatus>
: same locality,
<specimenCount id="9D5188D4A9776A4FFD6AFB8D1C4CC55C" box="[758,799,1062,1086]" pageId="2" pageNumber="77" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED46DB98A9776A4FFCB0FB8D1CD3C55C" box="[812,896,1062,1086]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="77">MNHN</collectionCode>
<specimenCode id="DBF1EB26A9776A4FFC1BFB8D1C95C55C" box="[903,966,1062,1086]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="77">DB21</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F4900A9776A4FFC44FB8D1B3FC55C" box="[984,1132,1062,1086]" collectionCode="ZMUM" country="China" pageId="2" pageNumber="77" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<specimenCount id="9D5188D4A9776A4FFC44FB8D1B52C55C" box="[984,1025,1062,1086]" pageId="2" pageNumber="77" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED46DB98A9776A4FFB93FB8D1B36C55C" box="[1039,1125,1062,1086]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="2" pageNumber="77">ZMUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9776A4FFD39FBFD1C06C5EA" pageId="2" pageNumber="77" type="etymology">
<paragraph id="8BE8435DA9776A4FFD39FBFD1C06C5EA" blockId="2.[677,1210,1110,1160]" pageId="2" pageNumber="77">ETYMOLOGY. — To emphasize the prevailing marbled, brown coloration.</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9776A4FFD39FB0A1C33C58E" pageId="2" pageNumber="77" type="diagnosis">
<paragraph id="8BE8435DA9776A4FFD39FB0A1C33C58E" blockId="2.[677,1210,1184,1261]" pageId="2" pageNumber="77">DIAGNOSIS. — Differs from congeners by peculiar male tarsal papillation, as well as certain details of leg and gonopod structure.</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9776A48FD39FAA61C0CC073" lastPageId="5" lastPageNumber="80" pageId="2" pageNumber="77" type="description">
<paragraph id="8BE8435DA9776A4FFD39FAA61C66C447" blockId="2.[677,1210,1293,1700]" box="[677,821,1293,1319]" pageId="2" pageNumber="77">DESCRIPTION</paragraph>
<paragraph id="8BE8435DA9776A4FFD39FA861B48C487" blockId="2.[677,1210,1293,1700]" pageId="2" pageNumber="77">
Length
<emphasis id="B9239F4FA9776A4FFD64FA861C5AC424" box="[760,777,1325,1350]" italics="true" pageId="2" pageNumber="77">c.</emphasis>
<quantity id="4CAFEEB8A9776A4FFC93FA861C33C425" box="[783,864,1325,1351]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="77" unit="mm" value="32.0">32 mm</quantity>
, maximum width
<quantity id="4CAFEEB8A9776A4FFBB9FA861B2CC425" box="[1061,1151,1325,1351]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="77" unit="mm" value="2.8">2.8 mm</quantity>
. Coloration rather pale brown, largely marbled; head, antennae (except for a pallid tip), dorsum and metazona below paraterga being slightly darker brown, pattern becoming somewhat annulate. Legs usually paler. Eye patches blackish.
</paragraph>
<paragraph id="8BE8435DA9776A4FFD27FA401B6FC726" blockId="2.[677,1210,1293,1700]" pageId="2" pageNumber="77">Head width = segment 4 &gt;&gt; collum&gt; segment 2&gt; 3 &lt;5 &lt;6 = 20 &lt;7; after segment 20 or 21, body very gradually tapering toward telson.</paragraph>
<paragraph id="8BE8435DA9776A4EFD27F9E11D2BC7C1" blockId="2.[677,1210,1293,1700]" lastBlockId="3.[113,647,1546,1700]" lastPageId="3" lastPageNumber="78" pageId="2" pageNumber="77">
Body with 30 segments (29 pleurotergites, or rings, in terms of
<bibRefCitation id="EFC63EACA9776A4FFCBDF9C21CA0C7E6" author="ENGHOFF H. &amp; DOHLE W. &amp; BLOWER J. G." box="[801,1011,1641,1668]" pageId="2" pageNumber="77" pagination="103 - 234" refId="ref7873" refString="ENGHOFF H., DOHLE W. &amp; BLOWER J. G. 1993. - Anamorphosis in millipedes (Diplopoda) - the present state of knowledge with some developmental and phylogenetic considerations. Zoological Journal of the Linnean Society 109: 103 - 234." type="journal article" year="1993">
Enghoff
<emphasis id="B9239F4FA9776A4FFCE2F9C11CE2C7E1" box="[894,945,1641,1667]" italics="true" pageId="2" pageNumber="77">et al.</emphasis>
1993
</bibRefCitation>
). Head densely setose, clypeolabral region slightly convex.Eye patches triangular, each composed of 15-17 convex ocelli. Antennae very long and slender (
<figureCitation id="136C5FD8A9766A4EFE27F9811D5FC726" box="[443,524,1578,1604]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="3" pageNumber="78">Figs 1A</figureCitation>
;
<figureCitation id="136C5FD8A9766A4EFD89F9811D65C726" box="[533,566,1578,1604]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[197,1118,218,1566]" captionTargetId="figure-88@4.[168,1155,193,1628]" captionTargetPageId="4" captionText="FIG. 2. — Nepalella marmorata n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 4, front view; D, leg 5, caudal view; E, leg 7, front view; F, coxa 7, caudal view; G, leg 10, front view; H, leg 11, front view; I, anterior gonopods (leg-pair 8), front view. Scale bar: 0.8 mm." figureDoi="http://doi.org/10.5281/zenodo.5393506" httpUri="https://zenodo.org/record/5393506/files/figure.png" pageId="3" pageNumber="78">2A</figureCitation>
), reaching beyond body segment 5 dorsally; antennomere 7 with a peculiar flattened seta dorsally in distal third. Gnathochilarium without promentum (
<figureCitation id="136C5FD8A9766A4EFD85F9221D38C7C1" box="[537,619,1673,1699]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[197,1118,218,1566]" captionTargetId="figure-88@4.[168,1155,193,1628]" captionTargetPageId="4" captionText="FIG. 2. — Nepalella marmorata n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 4, front view; D, leg 5, caudal view; E, leg 7, front view; F, coxa 7, caudal view; G, leg 10, front view; H, leg 11, front view; I, anterior gonopods (leg-pair 8), front view. Scale bar: 0.8 mm." figureDoi="http://doi.org/10.5281/zenodo.5393506" httpUri="https://zenodo.org/record/5393506/files/figure.png" pageId="3" pageNumber="78">Fig. 2B</figureCitation>
).
</paragraph>
<caption id="DF2813D5A9766A4EFFECFA281E4BC4A3" ID-DOI="http://doi.org/10.5281/zenodo.5393504" ID-Zenodo-Dep="5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="3" pageNumber="78" startId="3.[112,123,1411,1428]" targetBox="[127,1195,217,1368]" targetPageId="3">
<paragraph id="8BE8435DA9766A4EFFECFA281E4BC4A3" blockId="3.[112,1210,1411,1473]" pageId="3" pageNumber="78">
FIG. 1. —
<emphasis id="B9239F4FA9766A4EFF5FFA281FB6C4F6" bold="true" box="[195,229,1411,1428]" pageId="3" pageNumber="78">A -C</emphasis>
,
<taxonomicName id="4C5738DEA9766A4EFF6CFA281ECFC4F6" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[240,412,1411,1428]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="marmorata" status="sp. nov.">
<emphasis id="B9239F4FA9766A4EFF6CFA281ECFC4F6" box="[240,412,1411,1428]" italics="true" pageId="3" pageNumber="78">Nepalella marmorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9766A4EFE3FFA281E85C4F6" box="[419,470,1411,1428]" pageId="3" pageNumber="78" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♂;
<emphasis id="B9239F4FA9766A4EFDDBFA281D3BC4F6" bold="true" box="[583,616,1411,1428]" pageId="3" pageNumber="78">D -F</emphasis>
,
<taxonomicName id="4C5738DEA9766A4EFDEFFA281D9AC4F6" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[627,713,1411,1428]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B9239F4FA9766A4EFDEFFA281D9AC4F6" box="[627,713,1411,1428]" italics="true" pageId="3" pageNumber="78">N. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9766A4EFD53FA281C50C4F6" box="[719,771,1411,1428]" pageId="3" pageNumber="78" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♂;
<emphasis id="B9239F4FA9766A4EFCE8FA281CD1C4F6" bold="true" box="[884,898,1411,1428]" pageId="3" pageNumber="78">G</emphasis>
,
<taxonomicName id="4C5738DEA9766A4EFC12FA281CB7C4F6" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[910,996,1411,1428]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B9239F4FA9766A4EFC12FA281CB7C4F6" box="[910,996,1411,1428]" italics="true" pageId="3" pageNumber="78">N. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9766A4EFC76FA281B4EC4F6" box="[1002,1053,1411,1428]" pageId="3" pageNumber="78" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♀;
<emphasis id="B9239F4FA9766A4EFB11FA281BC9C4F6" bold="true" box="[1165,1178,1411,1428]" pageId="3" pageNumber="78">A</emphasis>
,
<emphasis id="B9239F4FA9766A4EFB39FA281BE0C4F6" bold="true" box="[1189,1203,1411,1428]" pageId="3" pageNumber="78">D</emphasis>
, anterior body end, dorsal view;
<emphasis id="B9239F4FA9766A4EFEE5FA311ED5C4C9" bold="true" box="[377,390,1434,1451]" pageId="3" pageNumber="78">B</emphasis>
,
<emphasis id="B9239F4FA9766A4EFE0DFA311ECEC4C9" bold="true" box="[401,413,1434,1451]" pageId="3" pageNumber="78">E</emphasis>
, midbody segments, dorsal view;
<emphasis id="B9239F4FA9766A4EFD20FA311D99C4C9" bold="true" box="[700,714,1434,1451]" pageId="3" pageNumber="78">C</emphasis>
,
<emphasis id="B9239F4FA9766A4EFD49FA311DB3C4C9" bold="true" box="[725,736,1434,1451]" pageId="3" pageNumber="78">F</emphasis>
, same, lateral view;
<emphasis id="B9239F4FA9766A4EFC17FA311CCAC4C9" bold="true" box="[907,921,1434,1451]" pageId="3" pageNumber="78">G</emphasis>
, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm.
</paragraph>
</caption>
<paragraph id="8BE8435DA9766A48FD27F9A11EA7C070" blockId="3.[677,1210,1546,1700]" lastBlockId="5.[113,647,216,1700]" lastPageId="5" lastPageNumber="80" pageId="3" pageNumber="78">
Collum usual, obcordate in shape, with rudimentary paraterga (
<figureCitation id="136C5FD8A9766A4EFCA2F9811CDEC726" box="[830,909,1578,1604]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="3" pageNumber="78">Fig. 1A</figureCitation>
).Tegument smooth, shining, only prozona distinctly and densely striolate transversely. Metatergal setation 3 + 3, typical, macrochaetae short, rather thick but pointed, positioned on minute knobs; stricture between pro- and metazona shallow, inconspicuous (
<figureCitation id="136C5FD8A9706A48FEEBFF5C1EE2C070" box="[375,433,247,274]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="5" pageNumber="80">Fig. 1</figureCitation>
A-C).
</paragraph>
<caption id="DF2813D5A9716A49FFEDF9C81E6CC7C3" ID-DOI="http://doi.org/10.5281/zenodo.5393506" ID-Zenodo-Dep="5393506" httpUri="https://zenodo.org/record/5393506/files/figure.png" pageId="4" pageNumber="79" startId="4.[113,124,1635,1652]" targetBox="[197,1118,218,1566]" targetPageId="4">
<paragraph id="8BE8435DA9716A49FFEDF9C81E6CC7C3" blockId="4.[113,1210,1635,1697]" pageId="4" pageNumber="79">
FIG. 2. —
<taxonomicName id="4C5738DEA9716A49FF5EF9C81E3DC716" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[194,366,1635,1652]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="marmorata" status="sp. nov.">
<emphasis id="B9239F4FA9716A49FF5EF9C81E3DC716" box="[194,366,1635,1652]" italics="true" pageId="4" pageNumber="79">Nepalella marmorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9716A49FEEFF9C81EF6C716" box="[371,421,1635,1652]" pageId="4" pageNumber="79" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♂:
<emphasis id="B9239F4FA9716A49FD88F9C81D72C716" bold="true" box="[532,545,1635,1652]" pageId="4" pageNumber="79">A</emphasis>
, antenna;
<emphasis id="B9239F4FA9716A49FDE5F9C81DD5C716" bold="true" box="[633,646,1635,1652]" pageId="4" pageNumber="79">B</emphasis>
, gnathochilarium, ventral view;
<emphasis id="B9239F4FA9716A49FC13F9C81CCEC716" bold="true" box="[911,925,1635,1652]" pageId="4" pageNumber="79">C</emphasis>
, leg 4, front view;
<emphasis id="B9239F4FA9716A49FBAAF9C81B17C716" bold="true" box="[1078,1092,1635,1652]" pageId="4" pageNumber="79">D</emphasis>
, leg 5, caudal view;
<emphasis id="B9239F4FA9716A49FF3EF9D21FFDC7E8" bold="true" box="[162,174,1657,1674]" pageId="4" pageNumber="79">E</emphasis>
, leg 7, front view;
<emphasis id="B9239F4FA9716A49FED7F9D21E05C7E8" bold="true" box="[331,342,1657,1674]" pageId="4" pageNumber="79">F</emphasis>
, coxa 7, caudal view;
<emphasis id="B9239F4FA9716A49FD8FF9D21D72C7E8" bold="true" box="[531,545,1657,1674]" pageId="4" pageNumber="79">G</emphasis>
, leg 10, front view;
<emphasis id="B9239F4FA9716A49FD55F9D21D84C7E8" bold="true" box="[713,727,1657,1674]" pageId="4" pageNumber="79">H</emphasis>
, leg 11, front view;
<emphasis id="B9239F4FA9716A49FCE1F9D21CD0C7E8" bold="true" box="[893,899,1657,1674]" pageId="4" pageNumber="79">I</emphasis>
, anterior gonopods (leg-pair 8), front view. Scale bar: 0.8 mm.
</paragraph>
</caption>
<paragraph id="8BE8435DA9706A48FF14FEB31EC0C013" blockId="5.[113,647,216,1700]" pageId="5" pageNumber="80">
Paraterga small dorsolateral swellings/shoulders, not set off by peritremata, rather regularly rounded in dorsal view (
<figureCitation id="136C5FD8A9706A48FE89FEFD1E1CC013" box="[277,335,342,369]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="5" pageNumber="80">Fig. 1</figureCitation>
A-C).
</paragraph>
<paragraph id="8BE8435DA9706A48FF14FEDD1EFEC0B2" blockId="5.[113,647,216,1700]" pageId="5" pageNumber="80">
CIX (15) = 0.58; MIX (15) = 1.26; MA (15)
<emphasis id="B9239F4FA9706A48FDEBFEDC1DD4C0F2" box="[631,647,375,400]" italics="true" pageId="5" pageNumber="80"></emphasis>
160°; PIX (15) impossible to evaluate due to insufficiently developed paraterga.
</paragraph>
<paragraph id="8BE8435DA9706A48FF14FE7E1E59C3CF" blockId="5.[113,647,216,1700]" pageId="5" pageNumber="80">Axial suture distinct, pallid, usual; segment 7 only very slightly broader than adjacent ones, conspicuous due to slightly upturned, ridge-like pleural arches, without ventral apophyses. Limbus regularly microdentate, quite distinct ventrolaterally; similar minute denticles covering entire surface of pleural ridges.</paragraph>
<paragraph id="8BE8435DA9706A48FF14FD1F1E04C445" blockId="5.[113,647,216,1700]" pageId="5" pageNumber="80">
Legs long and slender, slightly longer in male,
<emphasis id="B9239F4FA9706A48FDEAFD1F1DD4C3AF" box="[630,647,692,717]" italics="true" pageId="5" pageNumber="80">c.</emphasis>
1.4 times as long as midbody height. Legs 1 and 2 slightly reduced as usual, tarsi with usual ventral brushes but without papillae; coxa 2 perforated by gonopore orifice. All subsequent legs conspicuously papillate over caudal face ofventral half (
<figureCitation id="136C5FD8A9706A48FDB7FCFA1D39C20E" box="[555,618,849,876]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[197,1118,218,1566]" captionTargetId="figure-88@4.[168,1155,193,1628]" captionTargetPageId="4" captionText="FIG. 2. — Nepalella marmorata n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 4, front view; D, leg 5, caudal view; E, leg 7, front view; F, coxa 7, caudal view; G, leg 10, front view; H, leg 11, front view; I, anterior gonopods (leg-pair 8), front view. Scale bar: 0.8 mm." figureDoi="http://doi.org/10.5281/zenodo.5393506" httpUri="https://zenodo.org/record/5393506/files/figure.png" pageId="5" pageNumber="80">Fig. 2</figureCitation>
C- E, G, H). Legs 3-7 distinctly incrassate, pairs 3 and 4 particularly so. Femora 3 and 4 each with a small but evident mushroom-like protuberance at midway ventrally (
<figureCitation id="136C5FD8A9706A48FEC5FC7B1EE3C288" box="[345,432,976,1002]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[197,1118,218,1566]" captionTargetId="figure-88@4.[168,1155,193,1628]" captionTargetPageId="4" captionText="FIG. 2. — Nepalella marmorata n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 4, front view; D, leg 5, caudal view; E, leg 7, front view; F, coxa 7, caudal view; G, leg 10, front view; H, leg 11, front view; I, anterior gonopods (leg-pair 8), front view. Scale bar: 0.8 mm." figureDoi="http://doi.org/10.5281/zenodo.5393506" httpUri="https://zenodo.org/record/5393506/files/figure.png" pageId="5" pageNumber="80">Fig. 2C</figureCitation>
). Coxae 5-7 microgranulate ventrally, prefemora 5-7 each with a microgranulate bulge dorso-parabasally (
<figureCitation id="136C5FD8A9706A48FDBDFBA41D2AC548" box="[545,633,1039,1066]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[197,1118,218,1566]" captionTargetId="figure-88@4.[168,1155,193,1628]" captionTargetPageId="4" captionText="FIG. 2. — Nepalella marmorata n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 4, front view; D, leg 5, caudal view; E, leg 7, front view; F, coxa 7, caudal view; G, leg 10, front view; H, leg 11, front view; I, anterior gonopods (leg-pair 8), front view. Scale bar: 0.8 mm." figureDoi="http://doi.org/10.5281/zenodo.5393506" httpUri="https://zenodo.org/record/5393506/files/figure.png" pageId="5" pageNumber="80">Fig. 2D</figureCitation>
). Coxa 7 and trochanter 7 with a distoventral digitiform outgrowth (
<figureCitation id="136C5FD8A9706A48FEB7FBE41EC9C50B" box="[299,410,1103,1129]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[197,1118,218,1566]" captionTargetId="figure-88@4.[168,1155,193,1628]" captionTargetPageId="4" captionText="FIG. 2. — Nepalella marmorata n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 4, front view; D, leg 5, caudal view; E, leg 7, front view; F, coxa 7, caudal view; G, leg 10, front view; H, leg 11, front view; I, anterior gonopods (leg-pair 8), front view. Scale bar: 0.8 mm." figureDoi="http://doi.org/10.5281/zenodo.5393506" httpUri="https://zenodo.org/record/5393506/files/figure.png" pageId="5" pageNumber="80">Fig. 2E, F</figureCitation>
). Legs 10 and 11 with coxal glands; coxae 10 conspicuously enlarged and setose (
<figureCitation id="136C5FD8A9706A48FF21FB251E41C5CA" box="[189,274,1166,1192]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[197,1118,218,1566]" captionTargetId="figure-88@4.[168,1155,193,1628]" captionTargetPageId="4" captionText="FIG. 2. — Nepalella marmorata n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 4, front view; D, leg 5, caudal view; E, leg 7, front view; F, coxa 7, caudal view; G, leg 10, front view; H, leg 11, front view; I, anterior gonopods (leg-pair 8), front view. Scale bar: 0.8 mm." figureDoi="http://doi.org/10.5281/zenodo.5393506" httpUri="https://zenodo.org/record/5393506/files/figure.png" pageId="5" pageNumber="80">Fig. 2G</figureCitation>
); each coxa 11 with a large, apically pilose, coniform funnel with a long parabasal seta on front face; prefemur 11 with a strong parabasal process ventrally (
<figureCitation id="136C5FD8A9706A48FEB3FB461ED6C465" box="[303,389,1261,1287]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="5" pageNumber="80">Fig. 1H</figureCitation>
). Claws invariably long, sabre-shaped, simple.
</paragraph>
<paragraph id="8BE8435DA9706A48FF14FA861D7BC4E4" blockId="5.[113,647,216,1700]" pageId="5" pageNumber="80">
Anterior gonopods (leg-pair 8) as usual very strongly reduced, sternum with a median and a lateral lobule, coxites spike-like (
<figureCitation id="136C5FD8A9706A48FE4CFAC71D4FC4E4" box="[464,540,1388,1414]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[197,1118,218,1566]" captionTargetId="figure-88@4.[168,1155,193,1628]" captionTargetPageId="4" captionText="FIG. 2. — Nepalella marmorata n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 4, front view; D, leg 5, caudal view; E, leg 7, front view; F, coxa 7, caudal view; G, leg 10, front view; H, leg 11, front view; I, anterior gonopods (leg-pair 8), front view. Scale bar: 0.8 mm." figureDoi="http://doi.org/10.5281/zenodo.5393506" httpUri="https://zenodo.org/record/5393506/files/figure.png" pageId="5" pageNumber="80">Fig. 2I</figureCitation>
).
</paragraph>
<paragraph id="8BE8435DA9706A48FF14FA271C0CC073" blockId="5.[113,647,216,1700]" lastBlockId="5.[677,1210,216,274]" pageId="5" pageNumber="80">
Posterior gonopods (leg-pair 9) hypertophied (
<figureCitation id="136C5FD8A9706A48FFE5FA001FA1C4A4" box="[121,242,1451,1478]" captionStart="FIG" captionStartId="6.[113,124,1484,1501]" captionTargetBox="[176,1108,220,1435]" captionTargetId="figure-51@6.[142,1182,202,1490]" captionTargetPageId="6" captionText="FIG. 3. — Nepalella marmorata n. sp., paratype ♂, posterior gonopods (leg-pair 9), front (A) and caudal (B) views, respectively. Scale bar: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393508" httpUri="https://zenodo.org/record/5393508/files/figure.png" pageId="5" pageNumber="80">Fig. 3A, B</figureCitation>
), with two ear-shaped folds at base on front face of colpocoxites and two highly conspicuous, extremely ramified, plumose, fimbriate, bush-like structures at base on caudal face; colpocoxites prominent, about as high as telopodites, with three longitudinal lamellae on caudal face, median lamella micropilose; telopoditomere 1 particularly strongly setose on posterior face, telopoditomere 2 infuscate and with a vestigial segment apically.
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9706A48FD39FE9C1C5AC02D" box="[677,777,311,337]" pageId="5" pageNumber="80" type="discussion">
<paragraph id="8BE8435DA9706A48FD39FE9C1C5AC02D" blockId="5.[677,1210,311,685]" box="[677,777,311,337]" pageId="5" pageNumber="80">REMARKS</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9706A48FD39FEFD1DB1C3CF" pageId="5" pageNumber="80" type="reference_group">
<paragraph id="8BE8435DA9706A48FD39FEFD1CADC32D" blockId="5.[677,1210,311,685]" pageId="5" pageNumber="80">
<taxonomicName id="4C5738DEA9706A48FD39FEFD1C2AC012" ID-CoL="46WVZ" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[677,889,342,368]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="marmorata" status="sp. nov.">
<emphasis id="B9239F4FA9706A48FD39FEFD1C2AC012" box="[677,889,342,368]" italics="true" pageId="5" pageNumber="80">Nepalella marmorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9706A48FCE2FEFC1CE4C013" box="[894,951,343,369]" pageId="5" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
differs from the similarly large and brown
<taxonomicName id="4C5738DEA9706A48FCC5FEDC1B15C0F2" ID-CoL="46WVY" authority="Shear, 2002" authorityName="Shear" authorityYear="2002" box="[857,1094,374,400]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="B9239F4FA9706A48FCC5FEDC1C92C0F2" box="[857,961,375,400]" italics="true" pageId="5" pageNumber="80">N. magna</emphasis>
<bibRefCitation id="EFC63EACA9706A48FC5BFEDD1B15C0F2" author="SHEAR W. A." box="[967,1094,374,400]" pageId="5" pageNumber="80" pagination="63 - 72" refId="ref8455" refString="SHEAR W. A. 2002. - Five new chordeumatidan millipeds from China: new species of Vieteuma (Kashmireumatidae) and Nepalella (Megalotylidae). Proceedings of the California Academy of Sciences 53 (6): 63 - 72." type="journal article" year="2002">Shear, 2002</bibRefCitation>
</taxonomicName>
the only congener based on females (
<bibRefCitation id="EFC63EACA9706A48FC53FE3D1B1CC0D2" author="SHEAR W. A." box="[975,1103,406,432]" pageId="5" pageNumber="80" pagination="63 - 72" refId="ref8455" refString="SHEAR W. A. 2002. - Five new chordeumatidan millipeds from China: new species of Vieteuma (Kashmireumatidae) and Nepalella (Megalotylidae). Proceedings of the California Academy of Sciences 53 (6): 63 - 72." type="journal article" year="2002">Shear 2002</bibRefCitation>
), and also coming from southern
<collectingCountry id="F34003CDA9706A48FC08FE1E1C85C0AD" box="[916,982,437,463]" name="China" pageId="5" pageNumber="80">China</collectingCountry>
(
<collectingRegion id="49938DBFA9706A48FC7EFE1D1B6BC0B2" box="[994,1080,438,464]" country="China" name="Yunnan" pageId="5" pageNumber="80">Yunnan</collectingRegion>
) primarily in the lesser number of ocelli per eye patch (15-17 versus 25). Regrettably, no material of the opposite sex of either species is available to reveal further differences (see also Key below).
</paragraph>
<paragraph id="8BE8435DA9706A48FD27FDFF1DB1C3CF" blockId="5.[677,1210,311,685]" pageId="5" pageNumber="80">
Based on the clearly pigmented body and ocelli,
<taxonomicName id="4C5738DEA9706A48FD39FDDF1C13C3EF" ID-CoL="46WVZ" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[677,832,628,653]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="marmorata" status="sp. nov.">
<emphasis id="B9239F4FA9706A48FD39FDDF1C13C3EF" box="[677,832,628,653]" italics="true" pageId="5" pageNumber="80">N. marmorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9706A48FCD6FDDF1CD4C3EC" box="[842,903,628,654]" pageId="5" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
seems to be only a troglophile.
</paragraph>
</subSubSection>
</treatment>
</document>