225 lines
29 KiB
XML
225 lines
29 KiB
XML
<document id="6B03A2B51601952FD0AD8C2AC9114EA6" ID="10.11646/zootaxa.4139.2.1" ID-DOI="10.11646/zootaxa.4139.2.1" ID-GBIF-Dataset="aeecbbb4-4764-41dd-9562-b25f91f9f70e" ID-ISSN="1175-5326" ID-Zenodo-Dep="265016" ID-ZooBank="2EAFF1B9-1FA2-44B1-89A3-476359314C5E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468994861279" checkinUser="plazi" docAuthor="Nguyen, Anh D. & Golovatch, Sergei I." docDate="2016" docId="03FECD21FFA7FFF8FF4F38D8FE8EFD74" docLanguage="en" docName="zootaxa.4139.2.1.pdf" docOrigin="Zootaxa 4139 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enghoffosoma digitatum Nguyen & Golovatch, 2016, sp. nov." docType="treatment" docVersion="11" lastPageNumber="165" masterDocId="FFC7B559FFACFFF6FFD83D24FFAEFF84" masterDocTitle="The millipede genus Enghoffosoma Golovatch, 1993 recorded in Vietnam for the first time, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="166" masterPageNumber="151" pageNumber="162" updateTime="1698365308467" updateUser="plazi">
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<mods:mods id="E92B2D96D067A3A25F5D4A60CC3AA48A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5ED5682654CAB9387416B5EAC7792776">
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<mods:title id="BDF80735AA037E6443FC019B1FB15DD6">The millipede genus Enghoffosoma Golovatch, 1993 recorded in Vietnam for the first time, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:name id="FA4298181305FBD411DE3A48760CA8A0" type="personal">
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<mods:roleTerm id="9079E7BD8DF45707434382069188B272">Author</mods:roleTerm>
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<mods:namePart id="DB9905A71F52C4D6D8620464D5846DBA">Nguyen, Anh D.</mods:namePart>
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</mods:name>
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<mods:name id="7D9756F8FB67E080F64C440F0016C877" type="personal">
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<mods:role id="D4AACDF0829CE323148141969439972F">
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<mods:roleTerm id="4187A9AD38DEBBA946B7984CF5B756B3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A600574E50AD5275E7C9814F019E65BA">Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4DF01898B5CEF8E5A735B64753261256">text</mods:typeOfResource>
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<mods:title id="DCF7CDC67F45D98CAAE3418329D2A09D">Zootaxa</mods:title>
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<mods:part id="20C5A7CE0A1CFB2D67BAFDCF62879FF9">
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<mods:date id="95C6AF88A2BCBE3E801C607C35F7F9B4">2016</mods:date>
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<mods:detail id="CFD4B680F7DF66B8BBB780D6E1EF8382" type="volume">
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<mods:number id="22D300B8E7F07B58A7057D0EFD319B4B">4139</mods:number>
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<mods:detail id="6A13B47994F34BBCDC7BC0298B02C890" type="issue">
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<mods:number id="9FC8E2A49BE6421B68BCE8C682A9BC9A">2</mods:number>
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<mods:classification id="F460085133659061E6DF137E65ABD742">journal article</mods:classification>
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<mods:identifier id="9693C2E1F08C99C705E3905E27E4E20A" type="DOI">10.11646/zootaxa.4139.2.1</mods:identifier>
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<mods:identifier id="57835A6FF8DBA1EB5D46D11601D858ED" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="96C83970F249B18FBC035DF191AF6BA1" type="Zenodo-Dep">265016</mods:identifier>
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<treatment id="03FECD21FFA7FFF8FF4F38D8FE8EFD74" ID-DOI="http://doi.org/10.5281/zenodo.6067537" ID-GBIF-Taxon="123198414" ID-Zenodo-Dep="6067537" LSID="urn:lsid:plazi:treatment:03FECD21FFA7FFF8FF4F38D8FE8EFD74" httpUri="http://treatment.plazi.org/id/03FECD21FFA7FFF8FF4F38D8FE8EFD74" lastPageId="14" lastPageNumber="165" pageId="11" pageNumber="162">
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<subSubSection id="C34D2FBCFFA7FFFDFF4F38D8FE89F9BC" pageId="11" pageNumber="162" type="nomenclature">
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<paragraph id="8BE87C37FFA7FFFDFF4F38D8FD87F993" blockId="11.[151,553,1532,1592]" box="[151,553,1532,1559]" pageId="11" pageNumber="162">
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<heading id="D0A0CB5BFFA7FFFDFF4F38D8FD87F993" bold="true" box="[151,553,1532,1559]" fontSize="11" level="1" pageId="11" pageNumber="162" reason="1">
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<emphasis id="B923A025FFA7FFFDFF4F38D8FD87F993" bold="true" box="[151,553,1532,1559]" pageId="11" pageNumber="162">
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<taxonomicName id="4C5707B4FFA7FFFDFF4F38D8FE6AF992" ID-CoL="39T4B" box="[151,452,1532,1558]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="digitatum" status="sp. nov.">
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<emphasis id="B923A025FFA7FFFDFF4F38D8FE6AF992" bold="true" box="[151,452,1532,1558]" italics="true" pageId="11" pageNumber="162">Enghoffosoma digitatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2101D5EFFA7FFFDFE1338D9FD87F993" box="[459,553,1533,1559]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE87C37FFA7FFFDFF4F3B3BFE89F9BC" blockId="11.[151,553,1532,1592]" box="[151,295,1567,1592]" pageId="11" pageNumber="162">
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(
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<figureCitation id="136C60B2FFA7FFFDFF473B3BFEB1F9BC" box="[159,287,1567,1592]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStartId-0="12.[151,250,1256,1278]" captionStartId-1="13.[151,250,1037,1059]" captionStartId-2="13.[151,250,1969,1991]" captionTargetBox-0="[164,1421,209,1220]" captionTargetBox-1="[165,1421,202,1004]" captionTargetBox-2="[151,1436,1108,1947]" captionTargetId-0="figure@12.[150,1436,193,1235]" captionTargetId-1="figure@13.[151,1436,193,1015]" captionTargetId-2="figure@13.[151,1436,1108,1947]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="13" captionText-0="FIGURE 12. Enghoffosoma digitatum sp. nov. Holotype, anterior segments, dorsal view (A), ventral view (B); midbody segments, dorsal view (D), lateral view (E); posterior segments, dorsal view (F), lateral view (G), ventral view (H); sternum between coxae 4, ventral view (C); hypoproct and paraprocts, ventral view (I). Scale = 1 mm." captionText-1="FIGURE 13. Enghoffosoma digitatum sp. nov. Holotype, right gonopod, mesal view (A); lateral view (B). Scale = 1 mm." captionText-2="FIGURE 14. Enghoffosoma digitatum sp. nov. Holotype, right gonopod, mesal view (A); postfemoral region, mesal view (B)." httpUri-0="https://zenodo.org/record/265028/files/figure.png" httpUri-1="https://zenodo.org/record/265029/files/figure.png" httpUri-2="https://zenodo.org/record/265030/files/figure.png" pageId="11" pageNumber="162">Figs 12–14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2FBCFFA7FFFDFF4F3B43FEAEF94C" pageId="11" pageNumber="162" type="materials_examined">
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<paragraph id="8BE87C37FFA7FFFDFF4F3B43FEAEF94C" blockId="11.[151,1436,1639,2024]" pageId="11" pageNumber="162">
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<emphasis id="B923A025FFA7FFFDFF4F3B43FE2EF904" bold="true" box="[151,384,1639,1664]" pageId="11" pageNumber="162">Material examined.</emphasis>
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<emphasis id="B923A025FFA7FFFDFE5F3B43FE40F9FB" box="[391,494,1639,1663]" italics="true" pageId="11" pageNumber="162">
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<typeStatus id="54ECC295FFA7FFFDFE5F3B43FE40F9FB" box="[391,494,1639,1663]" pageId="11" pageNumber="162" type="holotype">Holotype</typeStatus>
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</emphasis>
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male (
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<emphasis id="B923A025FFA7FFFDFDE33B43FCABF904" bold="true" box="[571,773,1639,1664]" pageId="11" pageNumber="162">IEBR-Myr-107H</emphasis>
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),
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<collectingCountry id="F3403CA7FFA7FFFDFCC53B43FCD1F904" box="[797,895,1639,1664]" name="Vietnam" pageId="11" pageNumber="162">Vietnam</collectingCountry>
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,
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<collectingRegion id="4993B2D5FFA7FFFDFC513B4CFC59F904" box="[905,1015,1639,1664]" country="Vietnam" name="Dong Nai" pageId="11" pageNumber="162">Dong Nai</collectingRegion>
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Prov., Cat Tien National Park, forest,
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<date id="FFE95AF7FFA7FFFDFF4F3BAFFE0AF920" box="[151,420,1675,1700]" pageId="11" pageNumber="162" value="2004-09-23" valueMax="2004-09-26" valueMin="2004-09-23">23–26 September 2004</date>
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, leg. Anh Duc Nguyen.
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<emphasis id="B923A025FFA7FFFDFD1A3BA8FC9AF927" box="[706,820,1676,1699]" italics="true" pageId="11" pageNumber="162">
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<typeStatus id="54ECC295FFA7FFFDFD1A3BA8FC9AF927" box="[706,820,1676,1699]" pageId="11" pageNumber="162" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount id="9D51B7BEFFA7FFFDFC9D3BA8FC0FF920" box="[837,929,1675,1700]" pageId="11" pageNumber="162" type="male">3 males</specimenCount>
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,
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<specimenCount id="9D51B7BEFFA7FFFDFC683BA8FBBAF920" box="[944,1044,1675,1700]" pageId="11" pageNumber="162" type="female">1 female</specimenCount>
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(
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<emphasis id="B923A025FFA7FFFDFBF03BAFFB40F920" bold="true" box="[1064,1262,1675,1700]" pageId="11" pageNumber="162">IEBR-Myr-107P</emphasis>
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), same data as
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<typeStatus id="54ECC295FFA7FFFDFF4F3B8BFF53F94C" box="[151,253,1711,1736]" pageId="11" pageNumber="162" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2FBCFFA7FFFDFF1F3BF7FB4AF8DC" pageId="11" pageNumber="162" type="diagnosis">
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<paragraph id="8BE87C37FFA7FFFDFF1F3BF7FEABF8B7" blockId="11.[151,1436,1639,2024]" pageId="11" pageNumber="162">
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<emphasis id="B923A025FFA7FFFDFF1F3BF7FEEFF968" bold="true" box="[199,321,1747,1772]" pageId="11" pageNumber="162">Diagnosis.</emphasis>
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The species is recognized by the paraterga being well developed; the gonopod postfemoral region consisting only of two processes: a thick, finger-shaped solenomere and a large, short, obscure, tuberculiform process
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<emphasis id="B923A025FFA7FFFDFF2A3A3FFF50F8B0" bold="true" box="[242,254,1819,1844]" pageId="11" pageNumber="162">e</emphasis>
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.
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</paragraph>
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<paragraph id="8BE87C37FFA7FFFDFF1F3A1BFB4AF8DC" blockId="11.[151,1436,1639,2024]" box="[199,1252,1855,1880]" pageId="11" pageNumber="162">
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<emphasis id="B923A025FFA7FFFDFF1F3A1BFEBAF8DC" bold="true" box="[199,276,1855,1880]" pageId="11" pageNumber="162">Name.</emphasis>
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“
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<taxonomicName id="4C5707B4FFA7FFFDFEFF3A1BFE3DF8D3" box="[295,403,1855,1879]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="digitatum">
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<emphasis id="B923A025FFA7FFFDFEFF3A1BFE3DF8D3" box="[295,403,1855,1879]" italics="true" pageId="11" pageNumber="162">digitatum</emphasis>
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</taxonomicName>
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”, an adjective in neuter gender to emphasize the finger-shaped solenomere.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D2FBCFFA7FFF8FF1F3A47FC6CFDB9" lastPageId="14" lastPageNumber="165" pageId="11" pageNumber="162" type="description">
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<paragraph id="8BE87C37FFA7FFFDFF1F3A47FDE4F863" blockId="11.[151,1436,1639,2024]" pageId="11" pageNumber="162">
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<emphasis id="B923A025FFA7FFFDFF1F3A47FEF7F8F8" bold="true" box="[199,345,1891,1916]" pageId="11" pageNumber="162">Description.</emphasis>
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Body length about
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<quantity id="4CAFD1D2FFA7FFFDFDEA3A47FD7BF8FF" box="[562,725,1891,1916]" metricMagnitude="-2" metricUnit="m" metricValue="3.615" metricValueMax="3.71" metricValueMin="3.52" pageId="11" pageNumber="162" unit="mm" value="36.15" valueMax="37.1" valueMin="35.2">35.2–37.1 mm</quantity>
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(male),
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<quantity id="4CAFD1D2FFA7FFFDFCE93A40FCD1F8FF" box="[817,895,1892,1916]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="11" pageNumber="162" unit="mm" value="41.0">41 mm</quantity>
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(female); width of midbody pro- and metazonae 2.9–3.0 mm (male),
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<quantity id="4CAFD1D2FFA7FFFDFE5F3AACFE4FF81B" box="[391,481,1928,1952]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="11" pageNumber="162" unit="mm" value="3.7">3.7 mm</quantity>
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(female) and 3.9–4.0 mm (male),
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<quantity id="4CAFD1D2FFA7FFFDFCAF3AA3FC7FF81B" box="[887,977,1927,1952]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="162" unit="mm" value="4.5">4.5 mm</quantity>
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(female), respectively.
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<typeStatus id="54ECC295FFA7FFFDFB3B3AA3FAE5F824" box="[1251,1355,1927,1952]" pageId="11" pageNumber="162" type="holotype">Holotype</typeStatus>
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length about
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<quantity id="4CAFD1D2FFA7FFFDFF043A88FEEAF847" box="[220,324,1964,1988]" metricMagnitude="-2" metricUnit="m" metricValue="3.71" pageId="11" pageNumber="162" unit="mm" value="37.1">37.1 mm</quantity>
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; width of pro- and metazonae ca. 3.0 and 4.0 mm, respectively. Body uniformly castaneous brown except for yellow paraterga and sterna.
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</paragraph>
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<caption id="DF282CBFFFA0FFFAFF4F39CCFBE4FABF" httpUri="https://zenodo.org/record/265028/files/figure.png" pageId="12" pageNumber="163" targetBox="[164,1421,209,1220]" targetPageId="12">
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<paragraph id="8BE87C37FFA0FFFAFF4F39CCFBE4FABF" blockId="12.[151,1436,1256,1339]" pageId="12" pageNumber="163">
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<emphasis id="B923A025FFA0FFFAFF4F39CCFE8DFB7A" bold="true" box="[151,291,1256,1278]" pageId="12" pageNumber="163">FIGURE 12.</emphasis>
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<taxonomicName id="4C5707B4FFA0FFFAFEF539CCFD85FB79" box="[301,555,1256,1277]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="digitatum" status="sp. nov.">
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<emphasis id="B923A025FFA0FFFAFEF539CCFD85FB79" box="[301,555,1256,1277]" italics="true" pageId="12" pageNumber="163">Enghoffosoma digitatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2101D5EFFA0FFFAFDED39CCFD24FB7A" box="[565,650,1256,1278]" pageId="12" pageNumber="163" rank="species">
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<emphasis id="B923A025FFA0FFFAFDED39CCFD24FB7A" bold="true" box="[565,650,1256,1278]" pageId="12" pageNumber="163">sp. nov.</emphasis>
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</taxonomicNameLabel>
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Holotype, anterior segments, dorsal view (A), ventral view (B); midbody segments, dorsal view (D), lateral view (E); posterior segments, dorsal view (F), lateral view (G), ventral view (H); sternum between coxae 4, ventral view (C); hypoproct and paraprocts, ventral view (I). Scale = 1 mm.
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</paragraph>
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</caption>
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<paragraph id="8BE87C37FFA0FFFAFF1F3846FF14FA47" blockId="12.[151,1437,1378,2016]" pageId="12" pageNumber="163">
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Clypeolabral region densely setose. Epicranial suture thin, line-shaped, distinct. Antennae short, claviform, nearly reaching to the end of segment 3 if stretched laterally. In length, antennomere 2=3=4=5=6>7>1 (
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<figureCitation id="136C60B2FFA0FFFAFAFD38A2FAD6FA1B" box="[1317,1400,1414,1439]" captionStart="FIGURE 12" captionStartId="12.[151,250,1256,1278]" captionTargetBox="[164,1421,209,1220]" captionTargetId="figure@12.[150,1436,193,1235]" captionTargetPageId="12" captionText="FIGURE 12. Enghoffosoma digitatum sp. nov. Holotype, anterior segments, dorsal view (A), ventral view (B); midbody segments, dorsal view (D), lateral view (E); posterior segments, dorsal view (F), lateral view (G), ventral view (H); sternum between coxae 4, ventral view (C); hypoproct and paraprocts, ventral view (I). Scale = 1 mm." httpUri="https://zenodo.org/record/265028/files/figure.png" pageId="12" pageNumber="163">Figs 12</figureCitation>
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A– B).
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</paragraph>
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<paragraph id="8BE87C37FFA0FFFAFF1F38EAFD5CF98F" blockId="12.[151,1437,1378,2016]" pageId="12" pageNumber="163">Collum nearly semi-circular; surface shining and smooth, with at least two rows of 3+3 and 3+3 setae. Paraterga broadly triangular, without lateral incisions.</paragraph>
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<paragraph id="8BE87C37FFA0FFFAFF1F3B3CFBA6F967" blockId="12.[151,1437,1378,2016]" pageId="12" pageNumber="163">In width, segment 3<4<head<collum=2=5–17, thereafter body tapering towards telson. Prozonae and sides below pataterga finely shagreened. Metazonae shining, with a row of 3+3 on fore halves. Metatergal transverse sulcus thin, line-shaped, starting with segment 4, but completely developed, reaching to base of paraterga on segments 5–19. Pleurosternal carinae well-developed, ridge-shaped, with a caudal acute denticle on segments 2– 10, reduced to a caudal acute denticle on segments 11–17, thereafter completely missing. Stricture between pro- and metazonae shallow, clearly striolate at bottom. Axial line thin and distinct.</paragraph>
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<paragraph id="8BE87C37FFA0FFFAFF1F3BD4FEACF813" blockId="12.[151,1437,1378,2016]" pageId="12" pageNumber="163">
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Paraterga (
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<figureCitation id="136C60B2FFA0FFFAFE983BCAFE38F883" box="[320,406,1774,1799]" captionStart="FIGURE 12" captionStartId="12.[151,250,1256,1278]" captionTargetBox="[164,1421,209,1220]" captionTargetId="figure@12.[150,1436,193,1235]" captionTargetPageId="12" captionText="FIGURE 12. Enghoffosoma digitatum sp. nov. Holotype, anterior segments, dorsal view (A), ventral view (B); midbody segments, dorsal view (D), lateral view (E); posterior segments, dorsal view (F), lateral view (G), ventral view (H); sternum between coxae 4, ventral view (C); hypoproct and paraprocts, ventral view (I). Scale = 1 mm." httpUri="https://zenodo.org/record/265028/files/figure.png" pageId="12" pageNumber="163">Figs 12</figureCitation>
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D–F) well developed, largely horizonal, only caudolateral corners somewhat upturned, but still lying below dorsum. Paraterga of segment 2 flabelliform, other paraterga subtriangular with a broadly rounded fore corner. Caudolateral corner acute, produced behind rear tergal margin into a small pointed tubercle in caudalmost segments (
|
||
<figureCitation id="136C60B2FFA0FFFAFE7A3A7EFE54F8F7" box="[418,506,1882,1907]" captionStart="FIGURE 12" captionStartId="12.[151,250,1256,1278]" captionTargetBox="[164,1421,209,1220]" captionTargetId="figure@12.[150,1436,193,1235]" captionTargetPageId="12" captionText="FIGURE 12. Enghoffosoma digitatum sp. nov. Holotype, anterior segments, dorsal view (A), ventral view (B); midbody segments, dorsal view (D), lateral view (E); posterior segments, dorsal view (F), lateral view (G), ventral view (H); sternum between coxae 4, ventral view (C); hypoproct and paraprocts, ventral view (I). Scale = 1 mm." httpUri="https://zenodo.org/record/265028/files/figure.png" pageId="12" pageNumber="163">Fig. 12</figureCitation>
|
||
F). Calluses thin, but more obvious on pore-bearing paraterga, without lateral incisions.
|
||
</paragraph>
|
||
<paragraph id="8BE87C37FFA0FFFAFF1F3A86FAAFF85B" blockId="12.[151,1437,1378,2016]" pageId="12" pageNumber="163">
|
||
Epiproct (
|
||
<figureCitation id="136C60B2FFA0FFFAFEE63A86FE37F83F" box="[318,409,1954,1979]" captionStart="FIGURE 12" captionStartId="12.[151,250,1256,1278]" captionTargetBox="[164,1421,209,1220]" captionTargetId="figure@12.[150,1436,193,1235]" captionTargetPageId="12" captionText="FIGURE 12. Enghoffosoma digitatum sp. nov. Holotype, anterior segments, dorsal view (A), ventral view (B); midbody segments, dorsal view (D), lateral view (E); posterior segments, dorsal view (F), lateral view (G), ventral view (H); sternum between coxae 4, ventral view (C); hypoproct and paraprocts, ventral view (I). Scale = 1 mm." httpUri="https://zenodo.org/record/265028/files/figure.png" pageId="12" pageNumber="163">Figs 12</figureCitation>
|
||
G–I) long, broadly truncated, without terminal tubercles. Tip not concave, with four spinnerets. Hypoproct broadly triangular, with two small, distolateral, setiferous knobs (
|
||
<figureCitation id="136C60B2FFA0FFFAFBA83AE2FB6AF85B" box="[1136,1220,1990,2015]" captionStart="FIGURE 12" captionStartId="12.[151,250,1256,1278]" captionTargetBox="[164,1421,209,1220]" captionTargetId="figure@12.[150,1436,193,1235]" captionTargetPageId="12" captionText="FIGURE 12. Enghoffosoma digitatum sp. nov. Holotype, anterior segments, dorsal view (A), ventral view (B); midbody segments, dorsal view (D), lateral view (E); posterior segments, dorsal view (F), lateral view (G), ventral view (H); sternum between coxae 4, ventral view (C); hypoproct and paraprocts, ventral view (I). Scale = 1 mm." httpUri="https://zenodo.org/record/265028/files/figure.png" pageId="12" pageNumber="163">Figs 12</figureCitation>
|
||
H–I).
|
||
</paragraph>
|
||
<caption id="DF282CBFFFA1FFFBFF4F3929FAC5FBA7" box="[151,1387,1037,1059]" httpUri="https://zenodo.org/record/265029/files/figure.png" pageId="13" pageNumber="164" targetBox="[165,1421,202,1004]" targetPageId="13">
|
||
<paragraph id="8BE87C37FFA1FFFBFF4F3929FAC5FBA7" blockId="13.[151,1387,1037,1059]" box="[151,1387,1037,1059]" pageId="13" pageNumber="164">
|
||
<emphasis id="B923A025FFA1FFFBFF4F3929FEB1FBA7" bold="true" box="[151,287,1037,1059]" pageId="13" pageNumber="164">FIGURE 13.</emphasis>
|
||
<taxonomicName id="4C5707B4FFA1FFFBFEFD3929FDB1FBA6" box="[293,543,1037,1058]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="digitatum" status="sp. nov.">
|
||
<emphasis id="B923A025FFA1FFFBFEFD3929FDB1FBA6" box="[293,543,1037,1058]" italics="true" pageId="13" pageNumber="164">Enghoffosoma digitatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923A025FFA1FFFBFDFD3929FDD8FBA7" bold="true" box="[549,630,1037,1059]" pageId="13" pageNumber="164">
|
||
<taxonomicNameLabel id="A2101D5EFFA1FFFBFDFD3929FDD8FBA7" box="[549,630,1037,1059]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Holotype, right gonopod, mesal view (A); lateral view (B). Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF282CBFFFA1FFFBFF4F3A95FA38F843" box="[151,1430,1969,1991]" httpUri="https://zenodo.org/record/265030/files/figure.png" pageId="13" pageNumber="164" targetBox="[151,1436,1108,1947]" targetPageId="13">
|
||
<paragraph id="8BE87C37FFA1FFFBFF4F3A95FA38F843" blockId="13.[151,1430,1969,1991]" box="[151,1430,1969,1991]" pageId="13" pageNumber="164">
|
||
<emphasis id="B923A025FFA1FFFBFF4F3A95FEB1F843" bold="true" box="[151,287,1969,1991]" pageId="13" pageNumber="164">FIGURE 14.</emphasis>
|
||
<taxonomicName id="4C5707B4FFA1FFFBFEFD3A95FDB0F842" box="[293,542,1969,1990]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="digitatum" status="sp. nov.">
|
||
<emphasis id="B923A025FFA1FFFBFEFD3A95FDB0F842" box="[293,542,1969,1990]" italics="true" pageId="13" pageNumber="164">Enghoffosoma digitatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2101D5EFFA1FFFBFDFC3A95FDDBF843" box="[548,629,1969,1991]" pageId="13" pageNumber="164" rank="species">
|
||
<emphasis id="B923A025FFA1FFFBFDFC3A95FDDBF843" bold="true" box="[548,629,1969,1991]" pageId="13" pageNumber="164">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Holotype, right gonopod, mesal view (A); postfemoral region, mesal view (B).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE87C37FFA2FFF8FF1F3DBCFEBDFF7C" blockId="14.[151,1437,151,752]" pageId="14" pageNumber="165">
|
||
Sterna densely setose; cross-impression distinct, with a small cone caudally near each coxa, rear cones being somewhat larger than fore ones. Male sternum 5 with a large, densely setose, trapeziform process between coxae 4 (
|
||
<figureCitation id="136C60B2FFA2FFF8FF473DFBFF5CFF7C" box="[159,242,223,248]" captionStart="FIGURE 12" captionStartId="12.[151,250,1256,1278]" captionTargetBox="[164,1421,209,1220]" captionTargetId="figure@12.[150,1436,193,1235]" captionTargetPageId="12" captionText="FIGURE 12. Enghoffosoma digitatum sp. nov. Holotype, anterior segments, dorsal view (A), ventral view (B); midbody segments, dorsal view (D), lateral view (E); posterior segments, dorsal view (F), lateral view (G), ventral view (H); sternum between coxae 4, ventral view (C); hypoproct and paraprocts, ventral view (I). Scale = 1 mm." httpUri="https://zenodo.org/record/265028/files/figure.png" pageId="14" pageNumber="165">Fig. 12</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
<paragraph id="8BE87C37FFA2FFF8FF1F3C21FA9BFEBB" blockId="14.[151,1437,151,752]" pageId="14" pageNumber="165">
|
||
Legs (
|
||
<figureCitation id="136C60B2FFA2FFF8FEC93C20FECBFE98" box="[273,357,260,285]" captionStart="FIGURE 12" captionStartId="12.[151,250,1256,1278]" captionTargetBox="[164,1421,209,1220]" captionTargetId="figure@12.[150,1436,193,1235]" captionTargetPageId="12" captionText="FIGURE 12. Enghoffosoma digitatum sp. nov. Holotype, anterior segments, dorsal view (A), ventral view (B); midbody segments, dorsal view (D), lateral view (E); posterior segments, dorsal view (F), lateral view (G), ventral view (H); sternum between coxae 4, ventral view (C); hypoproct and paraprocts, ventral view (I). Scale = 1 mm." httpUri="https://zenodo.org/record/265028/files/figure.png" pageId="14" pageNumber="165">Fig. 12</figureCitation>
|
||
E) about 1.4–1.5 times as long as midbody height (male). Male prefemora swollen dorsally. Femora without modifications. Tarsal brushes present until male leg 10, thereafter gradually thinning out.
|
||
</paragraph>
|
||
<paragraph id="8BE87C37FFA2FFF8FF1F3C68FC6CFDB9" blockId="14.[151,1437,151,752]" pageId="14" pageNumber="165">
|
||
Gonopod (
|
||
<figureCitation id="136C60B2FFA2FFF8FE983C68FE6CFEE0" box="[320,450,332,357]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="13.[151,250,1037,1059]" captionStartId-1="13.[151,250,1969,1991]" captionTargetBox-0="[165,1421,202,1004]" captionTargetBox-1="[151,1436,1108,1947]" captionTargetId-0="figure@13.[151,1436,193,1015]" captionTargetId-1="figure@13.[151,1436,1108,1947]" captionTargetPageId-0="13" captionTargetPageId-1="13" captionText-0="FIGURE 13. Enghoffosoma digitatum sp. nov. Holotype, right gonopod, mesal view (A); lateral view (B). Scale = 1 mm." captionText-1="FIGURE 14. Enghoffosoma digitatum sp. nov. Holotype, right gonopod, mesal view (A); postfemoral region, mesal view (B)." httpUri-0="https://zenodo.org/record/265029/files/figure.png" httpUri-1="https://zenodo.org/record/265030/files/figure.png" pageId="14" pageNumber="165">Figs 13–14</figureCitation>
|
||
) relatively simple. Coxite large and stout, about half as long as femorite; distoventral part sparsely setose. Prefemorite densely setose, separated laterally from femorite by a subtransverse sulcus. Femorite suberect, subcylindrical, long, weakly constricted at 2/3 of its extent, with neither processes nor other modifications. Postfemoral region consisting only of a solenomere and a process
|
||
<emphasis id="B923A025FFA2FFF8FBFA3C93FB80FE54" bold="true" box="[1058,1070,439,464]" pageId="14" pageNumber="165">e</emphasis>
|
||
, clearly separated from femorite by a cingulum. Solenomere very thick, oblique, finger-shaped, pointed at tip. Process
|
||
<emphasis id="B923A025FFA2FFF8FB053CF8FB47FE71" bold="true" box="[1245,1257,476,501]" pageId="14" pageNumber="165">e</emphasis>
|
||
large, obtuse, tuberculiform, slightly shorter than solenomere, somewhat curved ventrad. Seminal groove running distodorsad entirely mesally along femorite before entering onto a thick solenomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D2FBCFFA2FFF8FF1F3F63FE8EFD74" pageId="14" pageNumber="165" type="discussion">
|
||
<paragraph id="8BE87C37FFA2FFF8FF1F3F63FE8EFD74" blockId="14.[151,1437,151,752]" pageId="14" pageNumber="165">
|
||
<emphasis id="B923A025FFA2FFF8FF1F3F63FE95FDE4" bold="true" box="[199,315,583,608]" pageId="14" pageNumber="165">Remarks.</emphasis>
|
||
The new species is fairly similar to
|
||
<taxonomicName id="4C5707B4FFA2FFF8FD293F6DFAF9FDE4" authority="Likhitrakarn, Golovatch & Panha, 2014" authorityName="Likhitrakarn, Golovatch & Panha" authorityYear="2014" box="[753,1367,583,608]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="bispinum">
|
||
<emphasis id="B923A025FFA2FFF8FD293F6DFCD5FDE4" box="[753,891,584,608]" italics="true" pageId="14" pageNumber="165">E. bispinum</emphasis>
|
||
<bibRefCitation id="EFC601C6FFA2FFF8FC5E3F63FAF9FDE4" author="Likhitrakarn" box="[902,1367,583,608]" pageId="14" pageNumber="165" refString="Likhitrakarn, K., Golovatch, S. I. & Panha, S. (2014) Review of the Southeast Asian millipede genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species. Zootaxa, 3811 (4), 491 - 514. http: // dx. doi. org / 10.11646 / zootaxa. 3811.4.4" type="journal article" year="2014">Likhitrakarn, Golovatch & Panha, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingRegion id="4993B2D5FFA2FFF8FF4F3F49FEF0FD01" box="[151,350,620,645]" country="Thailand" name="Rayong" pageId="14" pageNumber="165">Rayong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3403CA7FFA2FFF8FEB23F48FE7EFD01" box="[362,464,620,645]" name="Thailand" pageId="14" pageNumber="165">Thailand</collectingCountry>
|
||
, in the gonopod postfemoral region consisting only of a solenomere and a process
|
||
<emphasis id="B923A025FFA2FFF8FA503F48FA3AFD01" bold="true" box="[1416,1428,620,645]" pageId="14" pageNumber="165">e</emphasis>
|
||
. However,
|
||
<taxonomicName id="4C5707B4FFA2FFF8FED33FB5FE3DFD2C" box="[267,403,656,680]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="bispinum">
|
||
<emphasis id="B923A025FFA2FFF8FED33FB5FE3DFD2C" box="[267,403,656,680]" italics="true" pageId="14" pageNumber="165">E. bispinum</emphasis>
|
||
</taxonomicName>
|
||
differs in the gonopod femorite being slightly curved, the solenomere with a small process terminating the seminal groove in the distal part, and process
|
||
<emphasis id="B923A025FFA2FFF8FCAD3F90FC2FFD49" bold="true" box="[885,897,692,717]" pageId="14" pageNumber="165">e</emphasis>
|
||
being pointed, erect, slightly shorter than the solenomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |