417 lines
40 KiB
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417 lines
40 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/afrinvertebr.63.76309" ID-GBIF-Dataset="4b66695a-c3d4-4711-8c87-00edd3951bf4" ID-Pensoft-Pub="2305-2562-1-19" ID-Pensoft-UUID="B0655A9F7BBC582691EC0F00E156BA38" ID-ZooBank="F849C700225A4923AE1962882F933E83" ModsDocID="2305-2562-63-1-19" checkinTime="1648229925038" checkinUser="pensoft" docAuthor="Boschert, Claire & Dikow, Torsten" docDate="2022" docId="3287E5085EDA5A858B991ED0E405D7FA" docLanguage="en" docName="AfrInvert 63(1): 19-75" docOrigin="African Invertebrates 63 (1)" docPubDate="2022-03-25" docSource="http://dx.doi.org/10.3897/afrinvertebr.63.76309" docTitle="Eremohaplomydas gobabebensis Boschert & Dikow 2022, sp. nov." docType="treatment" docUuid="745D49C1-62B8-4884-9F7F-2B82523373D3" docUuidSource="ZooBank" docVersion="8" id="B0655A9F7BBC582691EC0F00E156BA38" lastPageNumber="19" masterDocId="B0655A9F7BBC582691EC0F00E156BA38" masterDocTitle="Taxonomic revision of the mydas-fly genera Eremohaplomydas Bequaert, 1959, Haplomydas Bezzi, 1924, and Lachnocorynus Hesse, 1969 (Insecta, Diptera, Mydidae)" masterLastPageNumber="75" masterPageNumber="19" pageNumber="19" updateTime="1669128124301" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the mydas-fly genera Eremohaplomydas Bequaert, 1959, Haplomydas Bezzi, 1924, and Lachnocorynus Hesse, 1969 (Insecta, Diptera, Mydidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Boschert, Claire</mods:namePart>
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<mods:affiliation>Rice University, Houston, TX, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dikow, Torsten</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4816-2909</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, National Museum of Natural History, Smithsonian Institution, 10 th Street and Constitution Avenue NW, Washington, DC 20560, USA</mods:affiliation>
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<mods:nameIdentifier type="email">dikowt@si.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-03-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>63</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>75</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/afrinvertebr.63.76309</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/afrinvertebr.63.76309</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2305-2562-1-19</mods:identifier>
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<mods:identifier type="ZooBank">F849C700225A4923AE1962882F933E83</mods:identifier>
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<mods:identifier type="Pensoft-UUID">B0655A9F7BBC582691EC0F00E156BA38</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="194257702" LSID="urn:lsid:zoobank.org:act:745D49C1-62B8-4884-9F7F-2B82523373D3" httpUri="http://treatment.plazi.org/id/3287E5085EDA5A858B991ED0E405D7FA" lastPageNumber="19" pageId="0" pageNumber="19">
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<subSubSection pageId="0" pageNumber="19" type="nomenclature">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName LSID="http://zoobank.org/745D49C1-62B8-4884-9F7F-2B82523373D3" authority="Boschert & Dikow, 2022" authorityName="Boschert & Dikow" authorityYear="2022" class="Insecta" family="Mydidae" genus="Eremohaplomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremohaplomydas gobabebensis" order="Diptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="gobabebensis" status="sp. nov.">Eremohaplomydas gobabebensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="19">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">
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<figureCitation captionStart="Figures 10–12" captionStartId="F4" captionText="Figures 10 - 12. Eremohaplomydas gobabebensis sp. nov. (♂ paratype, USNMENT 01518012, Zenodo https: // doi. org / 10.5281 / zenodo. 6083928): 10 dorsal (Zenodo https: // doi. org / 10.5281 / zenodo. 6083969) 11 lateral (https: // doi. org / 10.5281 / zenodo. 6083971) 12 head anterior (https: // doi. org / 10.5281 / zenodo. 6083979). Scale bars: 5 mm." figureDoi="10.3897/afrinvertebr.63.76309.figures10-12" httpUri="https://binary.pensoft.net/fig/663582" pageId="0" pageNumber="19">Figs 10-12</figureCitation>
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<figureCitation captionStart="Figures 13–15" captionStartId="F5" captionText="Figures 13 - 15. Eremohaplomydas gobabebensis sp. nov. ♂ terminalia (paratype, USNMENT 01518012): 13 dorsal (Zenodo https: // doi. org / 10.5281 / zenodo. 6083973) 14 ventral (https: // doi. org / 10.5281 / zenodo. 6083977) 15 lateral (https: // doi. org / 10.5281 / zenodo. 6083975). Magnification = 75 x." figureDoi="10.3897/afrinvertebr.63.76309.figures13-15" httpUri="https://binary.pensoft.net/fig/663583" pageId="0" pageNumber="19">, 13-15</figureCitation>
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<figureCitation captionStart="Figures 31–34" captionStartId="F9" captionText="Figures 31 - 34. Heads of Eremohaplomydas species in ventro-lateral view: 31 E. desertorum ♂ paratype (MZLU- 2143: 2, 60 x magnification, Zenodo https: // doi. org / 10.5281 / zenodo. 6083967) 32 E. gobabebensis sp. nov. ♂ holotype (USNMENT 01518012, 75 x, https: // doi. org / 10.5281 / zenodo. 6083981) 33 E. stomachoris sp. nov. ♀ holotype (AAM- 003035, 75 x, https: // doi. org / 10.5281 / zenodo. 6083989) 34 E. whartoni sp. nov. ♂ holotype (NMSA-DIP- 92011, 75 x, https: // doi. org / 10.5281 / zenodo. 6084015)." figureDoi="10.3897/afrinvertebr.63.76309.figures31-34" httpUri="https://binary.pensoft.net/fig/663587" pageId="0" pageNumber="19">, 32</figureCitation>
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<figureCitation captionStart="Figures 53–55" captionStartId="F14" captionText="Figures 53 - 55. Metathoracic coxa in lateral view: 53 Eremohaplomydas gobabebensis sp. nov. (USNMENT 01518012, crop of Fig. 11) 54 Haplomydas crassipes (NMSA-DIP- 77049, crop Fig. 36) 55 Lachnocorynus chobeensis (NMSA-Dip- 43314, crop of Fig. 42)." figureDoi="10.3897/afrinvertebr.63.76309.figures53-55" httpUri="https://binary.pensoft.net/fig/663592" pageId="0" pageNumber="19">, 53</figureCitation>
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<figureCitation captionStart="Figure 56" captionStartId="F15" captionText="Figure 56. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of Eremohaplomydas desertorum, Eremohaplomydas gobabebensis sp. nov., Lachnocorynus chobeensis, and Lachnocorynus stenocephalus sp. nov. (SimpleMappr https: // www. simplemappr. net / map / 14089). Distribution and occurrence data available in Google Earth KML file https: // www. simplemappr. net / map / 14089. kml and also through GBIF (data-set https: // www. gbif. org / dataset / 993875 DD- 5915 - 4107 - 8707 - 835 D 5 A 8 D 1022, DOI https: // doi. org / 10.15468 / awpjz 9)." figureDoi="10.3897/afrinvertebr.63.76309.figure56" httpUri="https://binary.pensoft.net/fig/663593" pageId="0" pageNumber="19">, 56</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="diagnosis">
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<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="19">The species is distinguished from congeners by the densely arranged dorso-ventrally flattened setae on legs, the absence of the base of vein M3+M4, the overall golden pubescence, and the restricted distribution in the central Namib Desert.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="etymology">
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<paragraph pageId="0" pageNumber="19">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="19">This species is named after the Gobabeb Namib Research Institute (www.gobabeb.org) where it was collected for the first time in November 2018. The specific epithet is to be treated as a noun in apposition.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">Description.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">Female.</emphasis>
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unknown.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">
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Male.
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<emphasis italics="true" pageId="0" pageNumber="19">Head</emphasis>
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</emphasis>
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: black, facial gibbosity brown, in general golden pubescent, ventrally and posteriorly white pubescent, white setose, laterally compressed setae; width distinctly greater than thorax (at postpronotal lobe), interocular distance on vertex larger than at ventral eye margin; vertex between compound eyes
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<normalizedToken originalValue="±">+/-</normalizedToken>
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horizontally straight, medially only slightly below dorsal eye margin, vertex golden pubescent, white setose; ocellar triangle apubescent; facial gibbosity distinct, well-developed and discernible in lateral view, mystax covering entire facial gibbosity, white; parafacial area approximately as wide as
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<normalizedToken originalValue="½">1/2</normalizedToken>
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width of central facial gibbosity (at same level); frons not elevated, golden pubescent, white setose; occiput predominantly white pubescent, dorsally golden pubescent, white setose, median occipital sclerite white setose, laterally compressed setae; pocl macrosetae absent; postgena apubescent, long, sparsely white setose; clypeus comprised of inverted U-shaped sclerite, dorsal
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<normalizedToken originalValue="½">1/2</normalizedToken>
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sclerotized medially to form plate, recessed (concave), ventrally simple, posterior to proboscis, laterally connected to face by sclerotized cuticle; proboscis very short, vestigial, knob-like, yellow; labellum small, as wide as prementum, length indiscernible, sclerotization indiscernible; maxillary palpus laterally compressed (triangular), light brown, slightly longer than proboscis.
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</paragraph>
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<caption doi="10.3897/afrinvertebr.63.76309.figures10-12" httpUri="https://binary.pensoft.net/fig/663582" pageId="0" pageNumber="19" start="Figures 10–12" startId="F4">
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">Figures 10-12.</emphasis>
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<taxonomicName authorityName="Boschert & Dikow" authorityYear="2022" class="Insecta" family="Mydidae" genus="Eremohaplomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremohaplomydas gobabebensis" order="Diptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="gobabebensis">
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<emphasis italics="true" pageId="0" pageNumber="19">Eremohaplomydas gobabebensis</emphasis>
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</taxonomicName>
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sp. nov. (♂ paratype, USNMENT01518012, Zenodo https://doi.org/10.5281/zenodo.6083928):
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<emphasis bold="true" pageId="0" pageNumber="19">10</emphasis>
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dorsal (Zenodo https://doi.org/10.5281/zenodo.6083969)
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<emphasis bold="true" pageId="0" pageNumber="19">11</emphasis>
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lateral (https://doi.org/10.5281/zenodo.6083971)
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<emphasis bold="true" pageId="0" pageNumber="19">12</emphasis>
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head anterior (https://doi.org/10.5281/zenodo.6083979). Scale bars: 5 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Antenna</emphasis>
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: light brown to brown; scape white setose dorsally, asetose ventrally; pedicel white setose dorsally and ventrally; postpedicel cylindrical in proximal 1/5, symmetrically bulbous in distal 4/5, ≥ 5.0 times as long as combined length of scape and pedicel, asetose; apical seta-like sensory element situated apically in cavity on postpedicel.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Thorax</emphasis>
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: brown, scutum golden pubescent, pleura white pubescent; scutum uniformly black, surface entirely smooth, golden pubescent, scutal setation comprised of long white setae with distinct rows of long dorsocentral setae and dense lateral scutal setae; dc setae pre- and postsuturally white, acr setae absent, lateral scutal setae white, npl setae 0, spal setae 0, pal setae 0; proepisternum apubescent, long white setose; proepimeron grey pubescent, asetose; antepronotum antero-medially smooth (without any indentation); lateral postpronotum long white setose; postpronotal lobe light brown, golden to light brown pubescent, long white setose; scutellum golden pubescent, discal scutellar setae absent, apical scutellar setae absent; mesopostnotum golden pubescent, asetose; anatergite golden pubescent, asetose; katatergite white pubescent, long white setose, slightly elevated, smoothly convex; anepisternum white pubescent, anteriorly white setose, posteriorly densely long white setose, scattered long white setose centrally; katepisternum dorsally white pubescent, ventrally apubescent, asetose; anepimeron white pubescent, long white setose; katepimeron white pubescent, asetose; meron white pubescent dorsally, sparsely white pubescent ventrally, asetose; metakatepisternum large; metanepisternum white pubescent, asetose; metepimeron yellow (same color as T1), white pubescent, long white setose,
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<normalizedToken originalValue="±">+/-</normalizedToken>
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flat, infra-halter sclerite absent.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Legs</emphasis>
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: light brown to brown, setation comprised of white laterally compressed setae predominantly covering surface; pro coxa apubescent, sparse white laterally compressed setae; mes coxa apubescent, asetose anteriorly, short white laterally compressed setae posteriorly; met coxa laterally unsclerotized (membrane between coxa and metakatepisternum clearly visible), apubescent, asetose anteriorly, short white laterally compressed setae posteriorly; met trochanter setose medially; pro + mes femur light brown to brown, met femur light brown to brown, distinctly clubbed for nearly entire length, macrosetose, 1 antero-ventral and 1 postero-ventral row of macrosetae, postero-ventrally long white, appressed setose; pro tibia laterally arched; mes tibia laterally arched; met tibia laterally arched, met tibia cylindrical with ventral keel terminating into distinct spur, spur not projecting beyond tip of tibia, postero-laterally long white, appressed setose; pro + mes tarsomere 1 approximately as long as individual tarsomeres 2, 3, or 4, met tarsomere 1 as long as individual tarsomeres 2, 3, or 4; pulvillus well-developed, as long as well-developed claw, and as wide as base of claw; setiform empodium absent.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Wing</emphasis>
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: length = 4.2-5.7 mm; hyaline throughout, veins light yellow, microtrichia absent; cells r1, r4, m3, + cua closed, r5 open; C terminating at junction with R1; Sc long, terminating in C proximal to r-m; R4 terminates in R1; R5 terminates in R1 and R4 simultaneously; auxiliary vein (R3) at base of R4 absent; R4 and R5 widest apart medially; r-m distinct, R4+5 and M1 apart, connected by crossvein; M1 curves slightly anteriorly at r-m, M1 (or M1+M2) terminates in C (not reaching wing margin); base of M3+M4 absent, M3+M4 not terminating together in C (not reaching wing margin), M4 and CuA split proximally to m-cu (cell m3 narrow proximally); CuP straight, cell cup wide, CuP and wing margin further apart proximally than distally; alula well-developed; halter light yellow, apubescent, asetose.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Abdomen</emphasis>
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: light brown to brown, setation comprised of dense long white setae, T2-4 parallel-sided and not constricted waist-like, T surface entirely smooth; T1-4 light brown, T5-7 brown; T entirely golden pubescent; T1-7 long white setose; S1-5 brown with white posterior margin, S6-7 dark brown; S apubescent; S1 asetose, S2-7 long white setose; bullae on T2 oval, brown, surface entirely smooth, T2 surface anterior to bullae smooth.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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♂
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">abdomen and terminalia</emphasis>
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: T1-8 well-developed; T7-8 anteriorly with 2 lateral apodemes; S6 regular, without any special setation postero-medially; S8 simple plate, entire (undivided) ventro-medially, not fused to T8 dorso-laterally; epandrium formed by single sclerite (fused medially
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<normalizedToken originalValue="±">+/-</normalizedToken>
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entirely), distally in dorsal view pointed postero-laterally; subepandrial sclerite without lateral or median protuberances; hypandrium
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<normalizedToken originalValue="±">+/-</normalizedToken>
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flat, divided ventro-medially by unsclerotized area into 2 sclerotized halves, entirely fused with gonocoxite, forming a gonocoxite-hypandrial complex, supra-hypandrial sclerite absent; gonocoxite simple, short, hook-like, without median or lateral protuberance, gonocoxal apodeme absent; 2 functional phallic prongs, short with broad lateral flange, medio-distally connected, parallel or diverging laterally, distally straight or only diverging slightly laterally; phallic epimere absent; lateral ejaculatory process absent; ejaculatory apodeme formed by single dorso-ventrally oriented plate; ventro-median margin of parameral sheath heavily sclerotized (appearing entirely closed); parameral sheath long, sperm sac entirely covered; sperm sac appearing
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<normalizedToken originalValue="±">+/-</normalizedToken>
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heavily sclerotized.
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</paragraph>
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<caption doi="10.3897/afrinvertebr.63.76309.figures13-15" httpUri="https://binary.pensoft.net/fig/663583" pageId="0" pageNumber="19" start="Figures 13–15" startId="F5">
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">Figures 13-15.</emphasis>
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<taxonomicName authorityName="Boschert & Dikow" authorityYear="2022" class="Insecta" family="Mydidae" genus="Eremohaplomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremohaplomydas gobabebensis" order="Diptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="gobabebensis">
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<emphasis italics="true" pageId="0" pageNumber="19">Eremohaplomydas gobabebensis</emphasis>
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</taxonomicName>
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sp. nov. ♂ terminalia (paratype, USNMENT01518012):
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<emphasis bold="true" pageId="0" pageNumber="19">13</emphasis>
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dorsal (Zenodo https://doi.org/10.5281/zenodo.6083973)
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<emphasis bold="true" pageId="0" pageNumber="19">14</emphasis>
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ventral (https://doi.org/10.5281/zenodo.6083977)
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<emphasis bold="true" pageId="0" pageNumber="19">15</emphasis>
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lateral (https://doi.org/10.5281/zenodo.6083975). Magnification = 75
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<normalizedToken originalValue="×">x</normalizedToken>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="type locality">
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<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Namibia: Erongo: Namib-Naukluft National Park, Gobabeb 20 km NW on D1983, Kuiseb riverbed (
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<geoCoordinate degrees="23" direction="south" minutes="24" orientation="latitude" precision="15" seconds="56" value="-23.415556">23°24'56"S</geoCoordinate>
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,
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<geoCoordinate degrees="014" direction="east" minutes="54" orientation="longitude" precision="15" seconds="43" value="14.911944">014°54'43"E</geoCoordinate>
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, -23.41556, 14.91194).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="materials_examined">
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<paragraph pageId="0" pageNumber="19">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<materialsCitation ID-GBIF-Occurrence="3738022347" collectedFrom="dry, sandy, partly wooded riverbed, resting on sand" collectingDate="2018-11-24" collectionCode="NMNW" collectorName="Dikow, T." country="Namibia" county="Namib-Naukluft National Park" elevation="317" latitude="-23.415556" location="Kuiseb riverbed" longLatPrecision="21" longitude="14.911944" municipality="Gobabeb" specimenCode="USNMENT01518262" specimenCount="1" specimenCount-male="1" stateProvince="Erongo" typeStatus="holotype">
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<collectingCountry name="Namibia">Namibia</collectingCountry>
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:
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<collectingRegion country="Namibia" name="Erongo">Erongo</collectingRegion>
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:
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<specimenCount type="male">1♂</specimenCount>
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<collectingCounty>Namib-Naukluft National Park</collectingCounty>
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,
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<collectingMunicipality>Gobabeb</collectingMunicipality>
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<locationDeviation>
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
|
||
NW on D1983
|
||
</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:3287E5085EDA5A858B991ED0E405D7FA:A8644FB316AFFC9B214C1AD85865BE84" country="Namibia" county="Namib-Naukluft National Park" latitude="-23.415556" longLatPrecision="21" longitude="14.911944" municipality="Gobabeb" name="Kuiseb riverbed" stateProvince="Erongo">Kuiseb riverbed</location>
|
||
,
|
||
<geoCoordinate degrees="23" direction="south" minutes="24" orientation="latitude" precision="15" seconds="56" value="-23.415556">23°24'56"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="014" direction="east" minutes="54" orientation="longitude" precision="15" seconds="43" value="14.911944">014°54'43"E</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.17" unit="m" value="317.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.17" unit="m" value="317.0">317 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate value="2018-11-24">2018-11-24</collectingDate>
|
||
collected a.m. (9:00-noon),
|
||
<collectedFrom>dry, sandy, partly wooded riverbed, resting on sand</collectedFrom>
|
||
,
|
||
<collectorName>Dikow, T.</collectorName>
|
||
(
|
||
<specimenCode>USNMENT01518262</specimenCode>
|
||
,
|
||
<typeStatus>Holotype</typeStatus>
|
||
,
|
||
<collectionCode>NMNW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3738022353" collectedFrom="dry, sandy, partly wooded riverbed, resting on sand" collectingDate="2018-11-24" collectionCode="NMNW" collectorName="Dikow, T." country="Namibia" county="Namib-Naukluft National Park" elevation="317" latitude="-23.415556" location="Kuiseb riverbed" longLatPrecision="21" longitude="14.911944" municipality="Gobabeb" specimenCode="USNMENT01518263" specimenCount="1" specimenCount-male="1" stateProvince="Erongo" typeStatus="paratype">
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
<collectingCounty>Namib-Naukluft National Park</collectingCounty>
|
||
,
|
||
<collectingMunicipality>Gobabeb</collectingMunicipality>
|
||
<locationDeviation>
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
|
||
NW on D1983
|
||
</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:3287E5085EDA5A858B991ED0E405D7FA:BF8ED9D44F927D698BE49FF8B66B0272" country="Namibia" county="Namib-Naukluft National Park" latitude="-23.415556" longLatPrecision="21" longitude="14.911944" municipality="Gobabeb" name="Kuiseb riverbed" stateProvince="Erongo">Kuiseb riverbed</location>
|
||
,
|
||
<geoCoordinate degrees="23" direction="south" minutes="24" orientation="latitude" precision="15" seconds="56" value="-23.415556">23°24'56"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="014" direction="east" minutes="54" orientation="longitude" precision="15" seconds="43" value="14.911944">014°54'43"E</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.17" unit="m" value="317.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.17" unit="m" value="317.0">317 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate value="2018-11-24">2018-11-24</collectingDate>
|
||
collected a.m. (9:00-noon),
|
||
<collectedFrom>dry, sandy, partly wooded riverbed, resting on sand</collectedFrom>
|
||
,
|
||
<collectorName>Dikow, T.</collectorName>
|
||
(
|
||
<specimenCode>USNMENT01518263</specimenCode>
|
||
,
|
||
<typeStatus>Paratype</typeStatus>
|
||
,
|
||
<collectionCode>NMNW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3738022331" collectedFrom="dry, sandy, partly wooded riverbed, resting on sand" collectingDate="2018-11-24" collectionCode="USNM" collectorName="Dikow, T." country="Namibia" county="Namib-Naukluft National Park" elevation="317" latitude="-23.415556" location="Kuiseb riverbed" longLatPrecision="21" longitude="14.911944" municipality="Gobabeb" specimenCode="USNMENT01518261" specimenCount="1" specimenCount-male="1" stateProvince="Erongo" typeStatus="paratype">
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
<collectingCounty>Namib-Naukluft National Park</collectingCounty>
|
||
,
|
||
<collectingMunicipality>Gobabeb</collectingMunicipality>
|
||
<locationDeviation>
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
|
||
NW on D1983
|
||
</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:3287E5085EDA5A858B991ED0E405D7FA:0F7C12368E155AE051EF4E4706456E56" country="Namibia" county="Namib-Naukluft National Park" latitude="-23.415556" longLatPrecision="21" longitude="14.911944" municipality="Gobabeb" name="Kuiseb riverbed" stateProvince="Erongo">Kuiseb riverbed</location>
|
||
,
|
||
<geoCoordinate degrees="23" direction="south" minutes="24" orientation="latitude" precision="15" seconds="56" value="-23.415556">23°24'56"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="014" direction="east" minutes="54" orientation="longitude" precision="15" seconds="43" value="14.911944">014°54'43"E</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.17" unit="m" value="317.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.17" unit="m" value="317.0">317 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate value="2018-11-24">2018-11-24</collectingDate>
|
||
collected a.m. (9:00-noon),
|
||
<collectedFrom>dry, sandy, partly wooded riverbed, resting on sand</collectedFrom>
|
||
,
|
||
<collectorName>Dikow, T.</collectorName>
|
||
(
|
||
<specimenCode>USNMENT01518261</specimenCode>
|
||
,
|
||
<typeStatus>Paratype</typeStatus>
|
||
,
|
||
<collectionCode>USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3738022307" collectedFrom="dry, sandy, partly wooded riverbed, resting on sand" collectingDate="2018-11-24" collectionCode="USNM" collectorName="Dikow, T." country="Namibia" county="Namib-Naukluft National Park" elevation="317" latitude="-23.415556" location="Kuiseb riverbed" longLatPrecision="21" longitude="14.911944" municipality="Gobabeb" specimenCode="USNMENT01518260" specimenCount="1" specimenCount-male="1" stateProvince="Erongo" typeStatus="paratype">
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
<collectingCounty>Namib-Naukluft National Park</collectingCounty>
|
||
,
|
||
<collectingMunicipality>Gobabeb</collectingMunicipality>
|
||
<locationDeviation>
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
|
||
NW on D1983
|
||
</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:3287E5085EDA5A858B991ED0E405D7FA:10DDE115AFEEEAE66EAC79E2C0CAB020" country="Namibia" county="Namib-Naukluft National Park" latitude="-23.415556" longLatPrecision="21" longitude="14.911944" municipality="Gobabeb" name="Kuiseb riverbed" stateProvince="Erongo">Kuiseb riverbed</location>
|
||
,
|
||
<geoCoordinate degrees="23" direction="south" minutes="24" orientation="latitude" precision="15" seconds="56" value="-23.415556">23°24'56"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="014" direction="east" minutes="54" orientation="longitude" precision="15" seconds="43" value="14.911944">014°54'43"E</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.17" unit="m" value="317.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.17" unit="m" value="317.0">317 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate value="2018-11-24">2018-11-24</collectingDate>
|
||
collected a.m. (9:00-noon),
|
||
<collectedFrom>dry, sandy, partly wooded riverbed, resting on sand</collectedFrom>
|
||
,
|
||
<collectorName>Dikow, T.</collectorName>
|
||
(
|
||
<specimenCode>USNMENT01518260</specimenCode>
|
||
,
|
||
<typeStatus>Paratype</typeStatus>
|
||
,
|
||
<collectionCode>USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3738022314" collectedFrom="small vegetated dunes, resting on sand" collectingDate="2018-11-21" collectionCode="USNM" collectorName="Dikow, T." country="Namibia" elevation="401" httpUri="https://doi.org/10.5281/zenodo.6083928" latitude="-23.563334" location="Gobabeb" longLatPrecision="21" longitude="15.032778" municipality="Namib-Naukluft National Park" specimenCode="USNMENT01518012" specimenCount="1" specimenCount-male="1" stateProvince="Erongo" typeStatus="paratype">
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
<collectingMunicipality>Namib-Naukluft National Park</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:3287E5085EDA5A858B991ED0E405D7FA:E0EB35387AB65EFE9249B268F19CB057" country="Namibia" latitude="-23.563334" longLatPrecision="21" longitude="15.032778" municipality="Namib-Naukluft National Park" name="Gobabeb" stateProvince="Erongo">Gobabeb</location>
|
||
,
|
||
<locationDeviation>dunes W of Kuiseb riverbed</locationDeviation>
|
||
,
|
||
<geoCoordinate degrees="23" direction="south" minutes="33" orientation="latitude" precision="15" seconds="48" value="-23.563334">23°33'48"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="015" direction="east" minutes="01" orientation="longitude" precision="15" seconds="58" value="15.032778">015°01'58"E</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.01" unit="m" value="401.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.01" unit="m" value="401.0">401 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate value="2018-11-21">2018-11-21</collectingDate>
|
||
collected a.m. (9:00-noon),
|
||
<collectedFrom>small vegetated dunes, resting on sand</collectedFrom>
|
||
,
|
||
<collectorName>Dikow, T.</collectorName>
|
||
(
|
||
<specimenCode>USNMENT01518012</specimenCode>
|
||
,
|
||
<typeStatus>Paratype</typeStatus>
|
||
,
|
||
<collectionCode>USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3738022362" collectedFrom="partly vegetated dune, resting on sand" collectingDate="2018-11-23" collectionCode="USNM" collectorName="Dikow, T." country="Namibia" elevation="398" latitude="-23.563889" location="Gobabeb" longLatPrecision="21" longitude="15.033055" municipality="Namib-Naukluft National Park" specimenCode="USNMENT01518339" specimenCount="1" specimenCount-male="1" stateProvince="Erongo" typeStatus="paratype">
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
<collectingMunicipality>Namib-Naukluft National Park</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:3287E5085EDA5A858B991ED0E405D7FA:92E83AB0F0785790D992EC1CA4ECCFE5" country="Namibia" latitude="-23.563889" longLatPrecision="21" longitude="15.033055" municipality="Namib-Naukluft National Park" name="Gobabeb" stateProvince="Erongo">Gobabeb</location>
|
||
,
|
||
<locationDeviation>small dunes W of Kuiseb River</locationDeviation>
|
||
,
|
||
<geoCoordinate degrees="23" direction="south" minutes="33" orientation="latitude" precision="15" seconds="50" value="-23.563889">23°33'50"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="015" direction="east" minutes="01" orientation="longitude" precision="15" seconds="59" value="15.033055">015°01'59"E</geoCoordinate>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.9799999999999995" unit="m" value="398.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.9799999999999995" unit="m" value="398.0">398 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate value="2018-11-23">2018-11-23</collectingDate>
|
||
collected p.m. (noon-15:00),
|
||
<collectedFrom>partly vegetated dune, resting on sand</collectedFrom>
|
||
,
|
||
<collectorName>Dikow, T.</collectorName>
|
||
(
|
||
<specimenCode>USNMENT01518339</specimenCode>
|
||
,
|
||
<typeStatus>Paratype</typeStatus>
|
||
,
|
||
<collectionCode>USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="19" type="distribution">
|
||
<paragraph pageId="0" pageNumber="19">Distribution, biodiversity hotspots, phenology, and biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
Known only from two localities in the central Namib Desert in Namibia (Fig.
|
||
<figureCitation captionStart="Figure 56" captionStartId="F15" captionText="Figure 56. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of Eremohaplomydas desertorum, Eremohaplomydas gobabebensis sp. nov., Lachnocorynus chobeensis, and Lachnocorynus stenocephalus sp. nov. (SimpleMappr https: // www. simplemappr. net / map / 14089). Distribution and occurrence data available in Google Earth KML file https: // www. simplemappr. net / map / 14089. kml and also through GBIF (data-set https: // www. gbif. org / dataset / 993875 DD- 5915 - 4107 - 8707 - 835 D 5 A 8 D 1022, DOI https: // doi. org / 10.15468 / awpjz 9)." figureDoi="10.3897/afrinvertebr.63.76309.figure56" httpUri="https://binary.pensoft.net/fig/663593" pageId="0" pageNumber="19">56</figureCitation>
|
||
). A rarely collected species known only from seven specimens from three collecting events in 2018 (Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Collecting event summary for Eremohaplomydas, Haplomydas, and Lachnocorynus species." httpUri="http://table.plazi.org/id/8DF9F799C695B7BAA951697DD1292320" pageId="0" pageNumber="19" tableUuid="8DF9F799C695B7BAA951697DD1292320">1</tableCitation>
|
||
). The species is not known to occur in any currently recognized biodiversity hotspot. Adult flies are active in November in late spring (Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Seasonal imago flight activity of Eremohaplomydas, Haplomydas, and Lachnocorynus species through number of specimens collected and unique collecting events in each month (data given as specimens / collecting events). Months abbreviated starting with July. * = iNaturalist observation." httpUri="http://table.plazi.org/id/FD020CB799315F8304F2901FC17DB00E" pageId="0" pageNumber="19" tableUuid="FD020CB799315F8304F2901FC17DB00E">2</tableCitation>
|
||
), which is a time at the beginning of a moister period and rising temperatures in this part of the Namib Desert (data for Gobabeb, Namibia, see https://www.worldweatheronline.com/gobabeb-weather/erongo/na.aspx). So far, only males have been collected and they were observed to fly very low above the ground and appeared to inspect the base of single grass plants and dart at high speed across the sand to the next plant. At the Gobabeb locality, the flies were collected flying among
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Centropodia" kingdom="Plantae" lsidName="Centropodia glauca" order="Poales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="glauca">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Centropodia glauca</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="H.G.L.Reichenbach" authorityYear="1828" baseAuthorityName="R.Brown" class="Liliopsida" family="Poaceae" kingdom="Plantae" lsidName="" order="Poales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
|
||
, Gha Grass, https://www.gbif.org/species/5680035) and at the 20 km N Gobabeb locality the flies darted among
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Cladoraphis" kingdom="Plantae" lsidName="Cladoraphis spinosa" order="Poales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="spinosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Cladoraphis spinosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="A.R.Franchet" authorityYear="1887" class="Liliopsida" family="Poaceae" kingdom="Plantae" lsidName="" order="Poales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
|
||
, Spiny Love Grass, https://www.gbif.org/species/4152290, see habitat photographs with the grasses in the foreground in Figs
|
||
<figureCitation captionStart="Figures 1–2" captionStartId="F1" captionText="Figures 1 - 2. Habitat photographs where Eremohaplomydas gobabebensis sp. nov. was observed and collected: 1 sparsely vegetated small sand dune West of Kuiseb riverbed at Gobabeb, Namibia (23 ° 33 ' 50 " S, 015 ° 01 ' 59 " E, note grass Centropodia glauca in foreground), taken on 23 Nov 2018 (Zenodo https: // doi. org / 10.5281 / zenodo. 6263467) 2 margin of dry Kuiseb riverbed, 20 km NW on D 1983 of Gobabeb, Namibia (23 ° 24 ' 56 " S, 014 ° 54 ' 43 " E, note grass Cladoraphis spinosa in foreground), taken on 24 Nov 2018 (https: // doi. org / 10.5281 / zenodo. 6263266). Photographs by T. Dikow." figureDoi="10.3897/afrinvertebr.63.76309.figures1-2" httpUri="https://binary.pensoft.net/fig/663579" pageId="0" pageNumber="19">1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 1–2" captionStartId="F1" captionText="Figures 1 - 2. Habitat photographs where Eremohaplomydas gobabebensis sp. nov. was observed and collected: 1 sparsely vegetated small sand dune West of Kuiseb riverbed at Gobabeb, Namibia (23 ° 33 ' 50 " S, 015 ° 01 ' 59 " E, note grass Centropodia glauca in foreground), taken on 23 Nov 2018 (Zenodo https: // doi. org / 10.5281 / zenodo. 6263467) 2 margin of dry Kuiseb riverbed, 20 km NW on D 1983 of Gobabeb, Namibia (23 ° 24 ' 56 " S, 014 ° 54 ' 43 " E, note grass Cladoraphis spinosa in foreground), taken on 24 Nov 2018 (https: // doi. org / 10.5281 / zenodo. 6263266). Photographs by T. Dikow." figureDoi="10.3897/afrinvertebr.63.76309.figures1-2" httpUri="https://binary.pensoft.net/fig/663579" pageId="0" pageNumber="19">2</figureCitation>
|
||
). Both grass species are native and widely distributed in the western parts of southern Africa including the Namib Desert (
|
||
<bibRefCitation author="van Oudtshoorn, F" journalOrPublisher="Briza, Pretoria, 3 rd revised edn." pageId="0" pageNumber="19" refId="B23" refString="van Oudtshoorn, F, 2012. Guide to grasses of southern Africa. Briza, Pretoria, 3rd revised edn." title="Guide to grasses of southern Africa." year="2012">van Oudtshoorn 2012</bibRefCitation>
|
||
). The male flies possibly inspected the grasses in search for females resting in the shade although the junior author was not able to observe or collect any females. In general, the flies were very difficult to observe and collect due to their high flight speed, light colouration, and small size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="19" type="remarks">
|
||
<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
<bibRefCitation author="Wharton, RA" journalOrPublisher="Annals of the Transvaal Museum" pageId="0" pageNumber="19" pagination="145 - 151" publicationUrl="https://hdl.handle.net/10520/AJA00411752_165" refId="B24" refString="Wharton, RA, 1982. Observations on the behaviour, phenology and habitat preferences of mydas flies in the central Namib Desert (Diptera: Mydidae). Annals of the Transvaal Museum 33 (9): 145 - 151, https://hdl.handle.net/10520/AJA00411752_165" title="Observations on the behaviour, phenology and habitat preferences of mydas flies in the central Namib Desert (Diptera: Mydidae)." url="https://hdl.handle.net/10520/AJA00411752_165" volume="33" year="1982">Wharton (1982)</bibRefCitation>
|
||
did not collect this species in his seminal year-long study of
|
||
<taxonomicName authorityName="Latreille" authorityYear="1806" class="Insecta" family="Mydidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="family">Mydidae</taxonomicName>
|
||
at Gobabeb.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |