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<mods:title id="76A031268E7A335D6662BE303C44318C">The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent: taxonomy with notes on habits and distributional patterns</mods:title>
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<mods:namePart id="A0A8393084F119245F4A3C8CEF31FD63">Raj, Shiju T.</mods:namePart>
<mods:affiliation id="8FAE77B33E707C281E896907F3FE4140">Post Graduate and Research Department of Zoology The New College, Royapettah Chennai- 670014, Tamil Nadu, INDIA</mods:affiliation>
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<mods:namePart id="808DCA1706CF085C3A81FCA89C8B6EC3">Sabu, Thomas K.</mods:namePart>
<mods:affiliation id="08BD5B45B2DD80206D21F2F21E66D788">Litter Entomology Research Unit Post Graduate and Research Department of Zoology St. Josephs College, Devagiri Calicut- 673008, Kerala, INDIA</mods:affiliation>
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<mods:affiliation id="A6CD301D01A5B91CA44A89A7F8C5CFA6">Zhao Danyang Department of Entomology College of Natural Resources and Environment South China Agricultural University, Wushan Guangzhou- 510640, CHINA</mods:affiliation>
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<treatment id="03B5EA1BD4673D26FF2E330ECF1B00DB" ID-DOI="http://doi.org/10.5281/zenodo.4670958" ID-GBIF-Taxon="180879990" ID-Zenodo-Dep="4670958" LSID="urn:lsid:plazi:treatment:03B5EA1BD4673D26FF2E330ECF1B00DB" httpUri="http://treatment.plazi.org/id/03B5EA1BD4673D26FF2E330ECF1B00DB" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">
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<taxonomicName id="4C1C208ED4673D27FF2E330EC80E073D" ID-CoL="6SKY4" authority="(Klug, 1834)" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[192,574,1381,1405]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B968871FD4673D27FF2E330EC80E073D" bold="true" box="[192,574,1381,1405]" pageId="10" pageNumber="9">
<emphasis id="B968871FD4673D27FF2E330ECBA3073C" bold="true" box="[192,403,1381,1405]" italics="true" pageId="10" pageNumber="9">Omphra atrata</emphasis>
(
<bibRefCitation id="EF8D26FCD4673D27FE4D330EC805073D" author="Klug" box="[419,565,1381,1405]" firstAuthor="Klug" pageId="10" pageNumber="9" refId="ref8313" refString="Klug, J. C. F. 1834. Bebersicht der Carabici der Sammlung. Jahrbucher der Insectenkunde, mit besonderer Rucksicht auf die Sammlung im Konigi. Museum zu Berlin; Berlin. 396 p." type="book" year="1834">Klug, 1834</bibRefCitation>
)
</emphasis>
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<subSubSection id="C3060886D4673D27FF2E33EECBBE07DD" box="[192,398,1413,1437]" pageId="10" pageNumber="9" type="description">
<paragraph id="8BA35B0DD4673D27FF2E33EECBBE07DD" blockId="10.[192,574,1381,1437]" box="[192,398,1413,1437]" pageId="10" pageNumber="9">
<figureCitation id="13274788D4673D27FF2E33EECB0707DD" box="[192,311,1413,1437]" captionStart="Figures 28" captionStartId="6.[192,288,1381,1403]" captionTargetBox="[221,1396,248,1339]" captionTargetId="figure-138@6.[192,1440,213,1407]" captionTargetPageId="6" captionText="Figures 28. Habitus of Omphra species. 2) O. hirta syntype. 3) O. pilosa syntype. 4) O. rufipes syntype. 5) O. atrata syntype. 6) O. complanata holotype. 7) O. rotundicollis holotype. 8) O. drumonti n. sp. holotype." figureDoi="http://doi.org/10.5281/zenodo.4670938" httpUri="https://zenodo.org/record/4670938/files/figure.png" pageId="10" pageNumber="9">Figures 5</figureCitation>
,
<figureCitation id="13274788D4673D27FEAD33EECB5307DD" box="[323,355,1413,1437]" captionStart="Figures 912" captionStartId="7.[192,288,1008,1030]" captionTargetBox="[192,1440,192,991]" captionTargetId="figure-241@7.[192,1440,192,991]" captionTargetPageId="7" captionText="Figures 912. Elytral apex. 9) Pointed and not emarginate. 10) Straight and not emarginate. 11) Oblique and weakly emarginate. 12) Straight and deeply emarginate." figureDoi="http://doi.org/10.5281/zenodo.4670940" httpUri="https://zenodo.org/record/4670940/files/figure.png" pageId="10" pageNumber="9">10</figureCitation>
,
<figureCitation id="13274788D4673D27FE9E33EECBBE07DD" box="[368,398,1413,1437]" captionStart="Figures 1314" captionStartId="8.[192,288,1178,1200]" captionTargetBox="[192,1440,192,1161]" captionTargetId="figure-273@8.[192,1440,192,1161]" captionTargetPageId="8" captionText="Figures 1314. Last antennomere 13) oval and 14) elongate oval." figureDoi="http://doi.org/10.5281/zenodo.4670944" httpUri="https://zenodo.org/record/4670944/files/figure.png" pageId="10" pageNumber="9">14</figureCitation>
</paragraph>
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<taxonomicName id="4C1C208ED4673D27FF2E33AECBE5079D" ID-CoL="6SKY4" authority="(Klug)" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[192,469,1477,1501]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B968871FD4673D27FF2E33AECB4D079D" box="[192,381,1477,1501]" italics="true" pageId="10" pageNumber="9">Omphra atrata</emphasis>
(Klug)
</taxonomicName>
;
<treatmentCitation id="0ABD7D1CD4673D27FE0C33AEC887079D" author="Reiche, M." box="[482,695,1477,1501]" page="330" pageId="10" pageNumber="9" year="1843">
<bibRefCitation id="EF8D26FCD4673D27FE0C33AEC887079D" author="Reiche" box="[482,695,1477,1501]" firstAuthor="Reiche" journalOrPublisher="Annales de la Societe Entomologique de France" page="330" pageId="10" pageNumber="9" pagination="323 - 344" part="11" refId="ref8878" refString="Reiche, M. 1843. Recherches sur les Helluonides, ou Revision du genre Helluo, Bonelli et Dejean. Annales de la Societe Entomologique de France 11 [1842]: 323 - 344." title="Recherches sur les Helluonides, ou Revision du genre Helluo, Bonelli et Dejean" type="journal article" year="1843">Reiche 1843: 330</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="EF8D26FCD4673D27FD2A33AEC9B5079D" author="Andrewes" box="[708,901,1477,1501]" firstAuthor="Andrewes" pageId="10" pageNumber="9" pagination="145 - 195" refId="ref7697" refString="Andrewes, H. E. 1921 a. Notes on synonymy and on some types of Oriental Carabidae in various foreign collections. The Transactions of the Entomological Society of London 3: 145 - 195." type="journal article" year="1921">Andrewes 1921</bibRefCitation>
(b): 346;
<treatmentCitation id="0ABD7D1CD4673D27FC0133AECEFD079D" author="Chopra, B." box="[1007,1229,1477,1501]" page="145" pageId="10" pageNumber="9" year="1924">
<bibRefCitation id="EF8D26FCD4673D27FC0133AECEFD079D" author="Chopra" box="[1007,1229,1477,1501]" firstAuthor="Chopra" page="145" pageId="10" pageNumber="9" pagination="145 - 151" refId="ref8035" refString="Chopra, B. 1924. The larva of an Indian carabid beetle. Records of Indian Museum 26: 145 - 151." type="journal article" year="1924">Chopra 1924: 145</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD7D1CD4673D27FB3433AECB2F07BD" author="Andrewes, H. E." page="101" pageId="10" pageNumber="9" year="1927">
<bibRefCitation id="EF8D26FCD4673D27FB3433AECB2F07BD" author="Andrewes" firstAuthor="Andrewes" page="101" pageId="10" pageNumber="9" pagination="97 - 111" refId="ref7800" refString="Andrewes, H. E. 1927. Papers on Oriental Carabidae-XIX. The Annals and Magazine of Natural History 9 (19): 97 - 111." type="journal article" year="1927">Andrewes 1927: 101</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD7D1CD4673D27FEC5338ECB9407BD" author="Andrewes, H. E." box="[299,420,1509,1533]" page="236" pageId="10" pageNumber="9" year="1930">
<bibRefCitation id="EF8D26FCD4673D27FEC5338ECB9407BD" author="Andrewes" box="[299,420,1509,1533]" firstAuthor="Andrewes" page="236" pageId="10" pageNumber="9" refId="ref7830" refString="Andrewes, H. E. 1930. Catalogue of Indian insects (Part 18: Carabidae). Government of India Central Publication Calcutta; Calcutta 389 p." type="book" year="1930">1930: 236</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD7D1CD4673D27FE5F338EC8B107BD" author="Csiki, E." box="[433,641,1509,1533]" page="1577" pageId="10" pageNumber="9" year="1932">
<bibRefCitation id="EF8D26FCD4673D27FE5F338EC8B107BD" author="Csiki" box="[433,641,1509,1533]" firstAuthor="Csiki" page="1577" pageId="10" pageNumber="9" pagination="1279 - 1598" refId="ref8059" refString="Csiki, E. 1932. Carabidae: Harpalinae VII (Pars 124). Coleopterorum Catalogus 3 (3): 1279 - 1598." type="journal article" year="1932">Csiki 1932: 1577</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD7D1CD4673D27FD63338DC95307BD" author="Lorenz, W." box="[653,867,1509,1533]" page="511" pageId="10" pageNumber="9" year="2005">
<bibRefCitation id="EF8D26FCD4673D27FD63338DC95307BD" author="Lorenz" box="[653,867,1509,1533]" firstAuthor="Lorenz" page="511" pageId="10" pageNumber="9" refId="ref8478" refString="Lorenz, W. 2005. Systematic list of extant ground beetles of the world. Tutzing: Published by the author; Tutzing. 530 p." type="book" year="2005">Lorenz 2005: 511</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD7D1CD4673D27FC81338ECE4207BD" author="Zhao, D. &amp; T. Deuve &amp; M. Tian" box="[879,1138,1509,1533]" page="371" pageId="10" pageNumber="9" year="2008">
<bibRefCitation id="EF8D26FCD4673D27FC81338ECE4207BD" author="Zhao" box="[879,1138,1509,1533]" etAl="et al." firstAuthor="Zhao" page="371" pageId="10" pageNumber="9" pagination="367 - 378" refId="ref9061" refString="Zhao, D., T. Deuve, and M. Tian. 2008. A review of the genus Omphra Reiche (Coleoptera: Carabidae: Helluonini). Oriental Insects 42: 367 - 378." type="journal article" year="2008">Zhao et al. 2008: 371</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BA35B0DD4673D27FF2E306EC804045D" blockId="10.[192,1440,1477,1565]" box="[192,564,1541,1565]" pageId="10" pageNumber="9">
<treatmentCitationGroup id="AB0C7C23D4673D27FF2E306EC804045D" box="[192,564,1541,1565]" pageId="10" pageNumber="9">
<treatmentCitation id="0ABD7D1CD4673D27FF2E306EC81F045D" author="Klug, J. C. F." box="[192,559,1541,1565]" page="72" pageId="10" pageNumber="9" year="1834">
<taxonomicName id="4C1C208ED4673D27FF2E306EC81F045D" authority="Klug, 1834: 72" authorityName="Klug" authorityPageNumber="72" authorityYear="1834" box="[192,559,1541,1565]" class="Insecta" family="Carabidae" genus="Helluo" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="B968871FD4673D27FF2E306ECB42045D" box="[192,370,1541,1565]" italics="true" pageId="10" pageNumber="9">Helluo atratus</emphasis>
<bibRefCitation id="EF8D26FCD4673D27FE94306EC81F045D" author="Klug" box="[378,559,1541,1565]" firstAuthor="Klug" page="72" pageId="10" pageNumber="9" refId="ref8313" refString="Klug, J. C. F. 1834. Bebersicht der Carabici der Sammlung. Jahrbucher der Insectenkunde, mit besonderer Rucksicht auf die Sammlung im Konigi. Museum zu Berlin; Berlin. 396 p." type="book" year="1834">Klug, 1834: 72</bibRefCitation>
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.
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<paragraph id="8BA35B0DD4673D27FF2E302ECB5A04DD" blockId="10.[192,1440,1605,1885]" pageId="10" pageNumber="9">
<emphasis id="B968871FD4673D27FF2E302ECBFA041D" bold="true" box="[192,458,1605,1629]" pageId="10" pageNumber="9">Description. Color.</emphasis>
Dorsal side black, ventral side brownish red. Labrum and mentum brownish red, coxae and trochanter black or brownish red, legs black. Dorsal and ventral body setae black or brownish red.
</paragraph>
<paragraph id="8BA35B0DD4673D27FF1E30CEC82504BD" blockId="10.[192,1440,1605,1885]" pageId="10" pageNumber="9">
<emphasis id="B968871FD4673D27FF1E30CECB7004FD" bold="true" box="[240,320,1701,1725]" pageId="10" pageNumber="9">Head.</emphasis>
Labrum transversely not extended. Clypeus with 6-8 setae. Mentum rectangular; median tooth with 3 setae, base of mentum with 4 setae. 4
<superScript id="7C69F645D4673D27FCAE30AFC9600492" attach="left" box="[832,848,1732,1746]" fontSize="6" pageId="10" pageNumber="9">th</superScript>
palpomere of labium and maxilla dilated. Last antennomere elongate oval.
</paragraph>
<paragraph id="8BA35B0DD4673D27FF1E316ECACF057D" blockId="10.[192,1440,1605,1885]" pageId="10" pageNumber="9">
<emphasis id="B968871FD4673D27FF1E316ECBB1055D" bold="true" box="[240,385,1797,1821]" pageId="10" pageNumber="9">Pronotum.</emphasis>
Slightly convex, widest before or behind middle. Lateral margin weakly sinuate towards base.
</paragraph>
<paragraph id="8BA35B0DD4673D27FF1E312EC805051D" blockId="10.[192,1440,1605,1885]" box="[240,565,1861,1885]" pageId="10" pageNumber="9">
<emphasis id="B968871FD4673D27FF1E312ECBB4051D" bold="true" box="[240,388,1861,1885]" pageId="10" pageNumber="9">Scutellum.</emphasis>
Not punctate.
</paragraph>
<paragraph id="8BA35B0DD4663D26FF1E36A8C87D02BB" blockId="11.[192,1440,195,347]" pageId="11" pageNumber="10">
<emphasis id="B968871FD4663D26FF1E36A8CB7F029B" bold="true" box="[240,335,195,219]" pageId="11" pageNumber="10">Elytra.</emphasis>
Elongate oval, widest behind middle. Humerus pronounced. Intervals weakly convex. Apex truncate, without emargination.
</paragraph>
<paragraph id="8BA35B0DD4663D26FF1E3768CF90035B" blockId="11.[192,1440,195,347]" box="[240,1440,259,283]" pageId="11" pageNumber="10">
<emphasis id="B968871FD4663D26FF1E3768CB45035B" bold="true" box="[240,373,259,283]" pageId="11" pageNumber="10">Genitalia.</emphasis>
Male: Apical lamella of aedeagus narrow and pointed at apex. Female genitalia unknown.
</paragraph>
<paragraph id="8BA35B0DD4663D26FF1E3748CBCD031B" blockId="11.[192,1440,195,347]" pageId="11" pageNumber="10">
<emphasis id="B968871FD4663D26FF1E3748C854037B" bold="true" box="[240,612,291,315]" pageId="11" pageNumber="10">Measurements. Male (n=5),</emphasis>
TL = 14.0 and 15.0, TW = 4.9 and 5.3, PL = 3.1 and 3.3, PW = 3.8 and 4.1, EL = 8.0 and 9.0.
</paragraph>
<paragraph id="8BA35B0DD4663D26FF2E37E8CFA0039B" blockId="11.[192,1441,387,475]" pageId="11" pageNumber="10">
<emphasis id="B968871FD4663D26FF2E37E8C80003DB" bold="true" box="[192,560,387,411]" pageId="11" pageNumber="10">Specimens examined (n=5).</emphasis>
<materialsCitation id="3B745150D4663D26FDDB37E8CEE803DB" ID-GBIF-Occurrence="3069154323" box="[565,1240,387,412]" collectionCode="ZMHB" collectorName="No." country="India" location="West Bengal" pageId="11" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="syntype">
<typeStatus id="54A7E5AFD4663D26FDDB37E8C89803DB" box="[565,680,387,411]" pageId="11" pageNumber="10" type="syntype">Syntypes</typeStatus>
.
<collectingCountry id="F30B1B9DD4663D26FD5F37EFC93703DC" box="[689,775,388,412]" name="India" pageId="11" pageNumber="10">INDIA</collectingCountry>
:
<collectingRegion id="49D895EFD4663D26FCFE37EFC99B03DB" box="[784,939,387,411]" country="India" name="West Bengal" pageId="11" pageNumber="10">West Bengal</collectingRegion>
,
<collectorName id="26E93EDBD4663D26FC5B37EFC9EF03DB" box="[949,991,388,411]" pageId="11" pageNumber="10">No.</collectorName>
576,
<specimenCount id="9D1A9084D4663D26FBF337E8CE5D03DB" box="[1053,1133,387,411]" pageId="11" pageNumber="10" type="male">1 male</specimenCount>
(
<collectionCode id="ED0DC3C8D4663D26FB9437E8CEE103DB" box="[1146,1233,387,411]" pageId="11" pageNumber="10">ZMHB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B745150D4663D26FB0C37EFC8DA03FB" ID-GBIF-Occurrence="3069154315" collectionCode="MNHN" collectorName="Deccan &amp; Coll. Chaudoir &amp; Reiche" country="India" location="INDIA" pageId="11" pageNumber="10" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<collectingCountry id="F30B1B9DD4663D26FB0C37EFCF0803DC" box="[1250,1336,388,412]" name="India" pageId="11" pageNumber="10">INDIA</collectingCountry>
:
<collectorName id="26E93EDBD4663D26FAAF37EFCFAC03DB" box="[1345,1436,388,411]" pageId="11" pageNumber="10">Deccan</collectorName>
,
<collectorName id="26E93EDBD4663D26FF2E37C8CB5F03FB" box="[192,367,419,443]" pageId="11" pageNumber="10">Coll. Chaudoir</collectorName>
ex coll.
<collectorName id="26E93EDBD4663D26FE2437C8C82E03FB" box="[458,542,419,443]" pageId="11" pageNumber="10">Reiche</collectorName>
,
<specimenCount id="9D1A9084D4663D26FDC937C8C84603FB" box="[551,630,419,443]" pageId="11" pageNumber="10" type="male">1 male</specimenCount>
(
<collectionCode id="ED0DC3C8D4663D26FD6A37C8C8D203FB" box="[644,738,419,443]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="10">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B745150D4663D26FD1D37CFCFBC039B" ID-GBIF-Occurrence="3069154325" collectionCode="MNHN, ZMHB, NHMW" collectorName="Eastern &amp; Chaudoir" country="India" location="West Bengal" pageId="11" pageNumber="10" specimenCount="3" specimenCount-male="3" stateProvince="West Bengal" typeStatus="syntype">
<collectorName id="26E93EDBD4663D26FD1D37CFC96403FB" box="[755,852,420,443]" pageId="11" pageNumber="10">Eastern</collectorName>
India, ex coll.
<collectorName id="26E93EDBD4663D26FC1237C8CE4003FB" box="[1020,1136,419,443]" pageId="11" pageNumber="10">Chaudoir</collectorName>
,
<specimenCount id="9D1A9084D4663D26FB9437C8CEF903FB" box="[1146,1225,419,443]" pageId="11" pageNumber="10" type="male">1 male</specimenCount>
(
<collectionCode id="ED0DC3C8D4663D26FB3937C8CF0603FB" box="[1239,1334,419,443]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="10">MNHN</collectionCode>
).
<collectingCountry id="F30B1B9DD4663D26FAA937CFCFAD03FC" box="[1351,1437,420,444]" name="India" pageId="11" pageNumber="10">INDIA</collectingCountry>
:
<collectingRegion id="49D895EFD4663D26FF2E37AFCB6F039B" box="[192,351,451,475]" country="India" name="West Bengal" pageId="11" pageNumber="10">West Bengal</collectingRegion>
, the number 576,
<specimenCount id="9D1A9084D4663D26FDD337A8C8A0039B" box="[573,656,451,475]" pageId="11" pageNumber="10" type="male">1 male</specimenCount>
(
<collectionCode id="ED0DC3C8D4663D26FD7137A8C8C7039B" box="[671,759,451,475]" pageId="11" pageNumber="10">ZMHB</collectionCode>
). Locality unknown, Collect. Plason.
<specimenCount id="9D1A9084D4663D26FB5337A8CF3F039B" box="[1213,1295,451,475]" pageId="11" pageNumber="10" type="male">1 male</specimenCount>
(
<collectionCode id="ED0DC3C8D4663D26FAF137A8CFB4039B" box="[1311,1412,451,475]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="10" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3060886D4663D26FF2E3468CBF4005B" box="[192,452,515,539]" pageId="11" pageNumber="10" type="distribution">
<paragraph id="8BA35B0DD4663D26FF2E3468CBF4005B" blockId="11.[192,452,515,539]" box="[192,452,515,539]" pageId="11" pageNumber="10">
<emphasis id="B968871FD4663D26FF2E3468CB44005B" bold="true" box="[192,372,515,539]" pageId="11" pageNumber="10">Distribution.</emphasis>
<collectingCountry id="F30B1B9DD4663D26FE953468CBF0005B" box="[379,448,515,539]" name="India" pageId="11" pageNumber="10">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3060886D4663D26FF2E3428CF1B00DB" pageId="11" pageNumber="10" type="discussion">
<paragraph id="8BA35B0DD4663D26FF2E3428CF1B00DB" blockId="11.[192,1440,579,667]" pageId="11" pageNumber="10">
<emphasis id="B968871FD4663D26FF2E3428CB74001B" bold="true" box="[192,324,579,603]" pageId="11" pageNumber="10">Remarks.</emphasis>
<taxonomicName id="4C1C208ED4663D26FEA53428C836001B" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[331,518,579,603]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B968871FD4663D26FEA53428C836001B" box="[331,518,579,603]" italics="true" pageId="11" pageNumber="10">Omphra atrata</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C1C208ED4663D26FD04342FC9B2001B" authority="Raj &amp; Sabu &amp; Danyang, 2012" authorityName="Raj &amp; Sabu &amp; Danyang" authorityYear="2012" box="[746,898,579,603]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="drumonti" status="sp. nov.">
<emphasis id="B968871FD4663D26FD04342FC9B2001B" box="[746,898,579,603]" italics="true" pageId="11" pageNumber="10">O. drumonti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B3A64D4663D26FC67342FC9FB001B" box="[905,971,580,603]" pageId="11" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
having truncate and not emarginated elytral apex. It is distinguished from
<taxonomicName id="4C1C208ED4663D26FD68340FC92D003B" authority="Raj &amp; Sabu &amp; Danyang, 2012" authorityName="Raj &amp; Sabu &amp; Danyang" authorityYear="2012" box="[646,797,611,635]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="drumonti" status="sp. nov.">
<emphasis id="B968871FD4663D26FD68340FC92D003B" box="[646,797,611,635]" italics="true" pageId="11" pageNumber="10">O. drumonti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B3A64D4663D26FCCD340FC954003B" box="[803,868,612,635]" pageId="11" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
by the presence of a slightly convex pronotum, pronounced elytral humerus and reddish brown anterior margin the of abdominal sternites.
</paragraph>
</subSubSection>
</treatment>
</document>