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<document id="58978004423A4A073043D3F34D453821" ID-CLB-Dataset="7452" ID-DOI="10.5281/zenodo.4670934" ID-GBIF-Dataset="0c820dc3-bd71-4eba-902e-59bd37283060" ID-ISSN="1942-1354" ID-Zenodo-Dep="4670934" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612980874391" checkinUser="felipe" docAuthor="Raj, Shiju T., Sabu, Thomas K. &amp; Danyang, Zhao" docDate="2012" docId="03B5EA1BD4613D20FF2E32AAC93107DB" docLanguage="en" docName="InsectaMundi.2012.0206.1-15.pdf" docOrigin="Insecta Mundi 2012 (206)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Omphra drumonti Raj &amp; Sabu &amp; Danyang 2012, n. sp." docType="treatment" docVersion="5" lastPageNumber="12" masterDocId="FF8C9263D46D3D2DFFEE366BCA300240" masterDocTitle="The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent: taxonomy with notes on habits and distributional patterns" masterLastPageNumber="15" masterPageNumber="1" pageNumber="11" updateTime="1698928787536" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="F710FBBF86A3C8DCED1EC0027D380427">The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent: taxonomy with notes on habits and distributional patterns</mods:title>
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<mods:name id="634AEFE667D30B03AAA1F018419571DD" type="personal">
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<mods:namePart id="74D5E9A71FD7B2948A6C1C90E42C6BDF">Raj, Shiju T.</mods:namePart>
<mods:affiliation id="DD5B4A8222274FBF95494F912C7DEC8B">Post Graduate and Research Department of Zoology The New College, Royapettah Chennai- 670014, Tamil Nadu, INDIA</mods:affiliation>
<mods:nameIdentifier id="60A8CB104CA2D24B1ED62886DFBFBCA9" type="email">shijutraj@gmail.com</mods:nameIdentifier>
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<mods:namePart id="275AE1F9C90B5F016871016AA072682C">Sabu, Thomas K.</mods:namePart>
<mods:affiliation id="8A884EEB419446CA7017C9229510A7FC">Litter Entomology Research Unit Post Graduate and Research Department of Zoology St. Josephs College, Devagiri Calicut- 673008, Kerala, INDIA</mods:affiliation>
<mods:nameIdentifier id="C454EC69117E5FD4F059D10619819033" type="email">sabukthomas@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B32AF3843121F2973C01BC6AA74AE9F9">Danyang, Zhao</mods:namePart>
<mods:affiliation id="9AD79294602FD5CCF9055586AB3BC68B">Zhao Danyang Department of Entomology College of Natural Resources and Environment South China Agricultural University, Wushan Guangzhou- 510640, CHINA</mods:affiliation>
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<mods:title id="5BD43748356E1B5E7C0561EFEEF46E59">Insecta Mundi</mods:title>
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<mods:date id="A56D52FA0DDC5FA582C6115358639AC9">2012</mods:date>
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<mods:number id="6E2D1E4029DC1896A22471A0F8436443">2012-02-29</mods:number>
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<mods:number id="DA36165060034803EC7984495477D1AC">2012</mods:number>
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<treatment id="03B5EA1BD4613D20FF2E32AAC93107DB" ID-DOI="http://doi.org/10.5281/zenodo.4670964" ID-GBIF-Taxon="180879994" ID-Zenodo-Dep="4670964" LSID="urn:lsid:plazi:treatment:03B5EA1BD4613D20FF2E32AAC93107DB" httpUri="http://treatment.plazi.org/id/03B5EA1BD4613D20FF2E32AAC93107DB" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11">
<subSubSection id="C3060886D4613D21FF2E32AAC83B069A" box="[192,523,1217,1242]" pageId="12" pageNumber="11" type="nomenclature">
<paragraph id="8BA35B0DD4613D21FF2E32AAC83B069A" blockId="12.[192,523,1217,1274]" box="[192,523,1217,1242]" pageId="12" pageNumber="11">
<heading id="D0EBEC61D4613D21FF2E32AAC83B069A" bold="true" box="[192,523,1217,1242]" fontSize="10" level="2" pageId="12" pageNumber="11" reason="3">
<emphasis id="B968871FD4613D21FF2E32AAC83B069A" bold="true" box="[192,523,1217,1242]" pageId="12" pageNumber="11">
<taxonomicName id="4C1C208ED4613D21FF2E32AACB8C0699" ID-CoL="74KY4" authority="Raj &amp; Sabu &amp; Danyang, 2012" authorityName="Raj &amp; Sabu &amp; Danyang" authorityYear="2012" box="[192,444,1217,1241]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="drumonti" status="sp. nov.">
<emphasis id="B968871FD4613D21FF2E32AACB8C0699" bold="true" box="[192,444,1217,1241]" italics="true" pageId="12" pageNumber="11">Omphra drumonti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B3A64D4613D21FE2A32A9C83B069A" box="[452,523,1218,1242]" pageId="12" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3060886D4613D20FF2E3289CF61011B" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11" type="description">
<paragraph id="8BA35B0DD4613D21FF2E3289CBBE06B9" blockId="12.[192,523,1217,1274]" box="[192,398,1249,1274]" pageId="12" pageNumber="11">
<figureCitation id="13274788D4613D21FF2E3289CB0606BA" box="[192,310,1250,1274]" captionStart="Figures 28" captionStartId="6.[192,288,1381,1403]" captionTargetBox="[221,1396,248,1339]" captionTargetId="figure-138@6.[192,1440,213,1407]" captionTargetPageId="6" captionText="Figures 28. Habitus of Omphra species. 2) O. hirta syntype. 3) O. pilosa syntype. 4) O. rufipes syntype. 5) O. atrata syntype. 6) O. complanata holotype. 7) O. rotundicollis holotype. 8) O. drumonti n. sp. holotype." figureDoi="http://doi.org/10.5281/zenodo.4670938" httpUri="https://zenodo.org/record/4670938/files/figure.png" pageId="12" pageNumber="11">Figures 8</figureCitation>
,
<figureCitation id="13274788D4613D21FEAD3289CB5306BA" box="[323,355,1250,1274]" captionStart="Figures 912" captionStartId="7.[192,288,1008,1030]" captionTargetBox="[192,1440,192,991]" captionTargetId="figure-241@7.[192,1440,192,991]" captionTargetPageId="7" captionText="Figures 912. Elytral apex. 9) Pointed and not emarginate. 10) Straight and not emarginate. 11) Oblique and weakly emarginate. 12) Straight and deeply emarginate." figureDoi="http://doi.org/10.5281/zenodo.4670940" httpUri="https://zenodo.org/record/4670940/files/figure.png" pageId="12" pageNumber="11">10</figureCitation>
,
<figureCitation id="13274788D4613D21FE9E328ACBBE06B9" box="[368,398,1249,1273]" captionStart="Figure 15" captionStartId="9.[816,898,1042,1064]" captionTargetBox="[816,1440,193,1025]" captionTargetId="figure-508@9.[816,1440,193,1025]" captionText="Figure 15. Color pattern of sternites of Omphra drumonti n. sp." figureDoi="http://doi.org/10.5281/zenodo.4670946" httpUri="https://zenodo.org/record/4670946/files/figure.png" pageId="12" pageNumber="11">15</figureCitation>
</paragraph>
<paragraph id="8BA35B0DD4613D21FF2E334AC9C4071A" blockId="12.[192,1441,1313,1914]" pageId="12" pageNumber="11">
<emphasis id="B968871FD4613D21FF2E334ACBF80779" bold="true" box="[192,456,1313,1337]" pageId="12" pageNumber="11">Description. Color.</emphasis>
Dorsal and ventral side of the body brownish red; anterior margin of sternites distinctly yellowish red, middle region and posterior margin black.
</paragraph>
<paragraph id="8BA35B0DD4613D21FF1E3309C81D0539" blockId="12.[192,1441,1313,1914]" pageId="12" pageNumber="11">
<emphasis id="B968871FD4613D21FF1E3309CB70073A" bold="true" box="[240,320,1378,1402]" pageId="12" pageNumber="11">Head.</emphasis>
Large, slightly narrower than pronotum, widest between eyes, moderately setose and punctate. Antenna elongate, filiform, setose, scape elongate, terminal antennomeres of the unique
<typeStatus id="54A7E5AFD4613D21FAD933E9CF9007DA" box="[1335,1440,1410,1434]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
is broken off. Clypeus transverse, anterior margin straight, frontoclypeal suture faintly rounded, with a depression at anterior side, 3 setae on each side, and elevated in the middle. Frons punctate, frontal foveae setose and depressed, middle region and sides elevated and glabrous. Vertex punctate and setose at the sides, glabrous at middle. Eyes big, semicircular, laterally well projected, supraorbital and suborbital region setose and punctate. Occiput strongly convex and glabrous. Labrum transverse and short, brownish red, apical margin emarginate, slightly round, 6 setae along apical margin, outer apical angles rounded with setae. Labial and maxillary palpomeres broken off. Mandibles simple, strongly arcuate, exposed, apex acuminate, slightly asymmetrical. Ligula wide, deeply depressed at sides, with a deep longitudinal central channel, apex rounded and pale red, two pairs of setae on each side of the channel close to apex. Lacinia, galea, glossa and paraglossa not visible. Mentum with one pair of setae on each side of base, median tooth strongly developed and stout, triangular, shorter than lateral lobes, base with one setae on each side, lateral lobes strongly developed and stout, pointed at apex. Submentum transverse, narrow, two setae present near middle. Gula narrow, glabrous, slightly convex, smooth and glossy. Two deep punctures present at the joining of submentum and gula. Genae punctate, smooth and convex.
</paragraph>
<paragraph id="8BA35B0DD4603D20FF1E36A8C9DB039B" blockId="13.[192,1440,195,859]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF1E36A8CBB4029B" bold="true" box="[240,388,195,219]" pageId="13" pageNumber="12">Pronotum.</emphasis>
Strongly convex, large and wide, almost round, widest in middle, apical and basal margin not straight, both apical and basal margins with a dense fringe of short setae, punctures dense on lateral area, lateral margin rounded anteriorly, sinuate close to base, without marginal border, median line well developed and a row of punctures present on both sides, apical and basal line faint and covered with dense punctures; central portion of pronotum glossy and with a few punctures, sides of basal margin emarginated, anterior angles rounded. Lateral margins bordered, sinuate near base, elongate setae present; base narrower than apex, hind angles nearly obtuse. Ventral side of pronotum brownish red, proepisternum densely punctate and pilose, proepimeron impunctate, prosternal process sparsely punctate and with a bunch of elongate setae on the apex.
</paragraph>
<paragraph id="8BA35B0DD4603D20FF1E3788C8EB03BB" blockId="13.[192,1440,195,859]" box="[240,731,483,508]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF1E3788CBB403BB" bold="true" box="[240,388,483,507]" pageId="13" pageNumber="12">Scutellum.</emphasis>
Asetose, sparsely punctate.
</paragraph>
<paragraph id="8BA35B0DD4603D20FF1E3468CBA300DB" blockId="13.[192,1440,195,859]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF1E3468CB7F005B" bold="true" box="[240,335,515,539]" pageId="13" pageNumber="12">Elytra.</emphasis>
Elongate, twice as long as pronotum, lateral margins subparallel; humerus pronounced, setose, rounded, without tooth. Surface glossy and densely punctate. Intervals gently convex, regularly and densely pilose with two rows of short black setae, interval 7 with 2 elongate setae. Lateral margins bordered, with elongate setae; apical margin truncate, without emargination, elongate setae present at apical margin.
</paragraph>
<paragraph id="8BA35B0DD4603D20FF1E34C8CBCB00FB" blockId="13.[192,1440,195,859]" box="[240,507,675,699]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF1E34C8CBA600FB" bold="true" box="[240,406,675,699]" pageId="13" pageNumber="12">Hind wings.</emphasis>
Absent.
</paragraph>
<paragraph id="8BA35B0DD4603D20FF1E34A8C9A5009B" blockId="13.[192,1440,195,859]" box="[240,917,707,731]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF1E34A8CB08009B" bold="true" box="[240,312,707,731]" pageId="13" pageNumber="12">Legs.</emphasis>
Stout and elongate and densely setose, dark red.
</paragraph>
<paragraph id="8BA35B0DD4603D20FF1E3488CF67015B" blockId="13.[192,1440,195,859]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF1E3488CB6600BB" bold="true" box="[240,342,739,763]" pageId="13" pageNumber="12">Venter.</emphasis>
Elongate setae present on lateral margins, anterior margin of sternites yellowish red and posterior margin dark red, 5
<superScript id="7C69F645D4603D20FDF03569C81E0150" attach="left" box="[542,558,770,784]" fontSize="6" pageId="13" pageNumber="12">th</superScript>
and 6
<superScript id="7C69F645D4603D20FD973569C8B90150" attach="left" box="[633,649,770,784]" fontSize="6" pageId="13" pageNumber="12">th</superScript>
abdominal sternite with two elongate setae in the middle.
</paragraph>
<paragraph id="8BA35B0DD4603D20FF1E3548C981017B" blockId="13.[192,1440,195,859]" box="[240,945,803,827]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF1E3548CB41017B" bold="true" box="[240,369,803,827]" pageId="13" pageNumber="12">Genitalia</emphasis>
. Male: unknown. Female: genitalia broken off.
</paragraph>
<paragraph id="8BA35B0DD4603D20FF1E3528CF61011B" blockId="13.[192,1440,195,859]" box="[240,1361,835,859]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF1E3528C8E0011B" bold="true" box="[240,720,835,859]" pageId="13" pageNumber="12">
Measurements. Female (
<typeStatus id="54A7E5AFD4603D20FDD13528C8F1011B" box="[575,705,835,859]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
),
</emphasis>
TL = 18.0, TW = 5.6, PL = 3.9, PW = 5.6, EL = 10.0.
</paragraph>
</subSubSection>
<subSubSection id="C3060886D4603D20FF2E35E8C8FC065B" pageId="13" pageNumber="12" type="materials_examined">
<paragraph id="8BA35B0DD4603D20FF2E35E8CE8B019B" blockId="13.[192,1440,899,987]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF2E35E8CBE801DB" bold="true" box="[192,472,899,923]" pageId="13" pageNumber="12">Type material (n=1).</emphasis>
<materialsCitation id="3B745150D4603D20FE3135E8CE8B019B" ID-GBIF-Occurrence="3069154317" collectionCode="ISNB" collectorName="R. Mus. Hist. Nat. Belg." country="India" location="Western Ghats" pageId="13" pageNumber="12" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<typeStatus id="54A7E5AFD4603D20FE3135E8C86101DB" box="[479,593,899,923]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
,
<collectingCountry id="F30B1B9DD4603D20FDB335EFC88401DC" box="[605,692,900,924]" name="India" pageId="13" pageNumber="12">INDIA</collectingCountry>
.
<collectingRegion id="49D895EFD4603D20FD5135E8C96801DB" box="[703,856,899,923]" country="India" name="Tamil Nadu" pageId="13" pageNumber="12">Tamil Nadu</collectingRegion>
: Kodicanal (=tropical montane evergreen forest ecoregion of the
<location id="8EC30DD6D4603D20FE6C35CFC80A01FB" LSID="urn:lsid:plazi:treatment:03B5EA1BD4613D20FF2E32AAC93107DB:8EC30DD6D4603D20FE6C35CFC80A01FB" box="[386,570,931,955]" country="India" name="Western Ghats" pageId="13" pageNumber="12" stateProvince="Tamil Nadu">Western Ghats</location>
at Kodaikanal),
<collectorName id="26E93EDBD4603D20FCEC35CFCE1201FB" box="[770,1058,931,955]" pageId="13" pageNumber="12">R. Mus. Hist. Nat. Belg.</collectorName>
I.G. 12.595, unknown collector,
<specimenCount id="9D1A9084D4603D20FF2E35A8CB18019B" box="[192,296,963,987]" pageId="13" pageNumber="12" type="female">1 female</specimenCount>
(
<collectionCode id="ED0DC3C8D4603D20FED735A8CBB2019B" box="[313,386,963,987]" pageId="13" pageNumber="12">ISNB</collectionCode>
). Labial and maxillary palpi and last antennomeres are broken off.
</materialsCitation>
</paragraph>
<paragraph id="8BA35B0DD4603D20FF2E3268C8FC065B" blockId="13.[192,716,1027,1051]" box="[192,716,1027,1051]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF2E3268C801065B" bold="true" box="[192,561,1027,1051]" pageId="13" pageNumber="12">Geographical distribution.</emphasis>
South
<collectingCountry id="F30B1B9DD4603D20FD643268C8FC065B" box="[650,716,1027,1051]" name="India" pageId="13" pageNumber="12">India</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C3060886D4603D20FF2E3228C874063B" pageId="13" pageNumber="12" type="etymology">
<paragraph id="8BA35B0DD4603D20FF2E3228C874063B" blockId="13.[192,1440,1091,1147]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF2E3228CB6C061B" bold="true" box="[192,348,1091,1115]" pageId="13" pageNumber="12">Etymology.</emphasis>
Species is named for Mr. Alain Drumont (ISNB,
<collectingRegion id="49D895EFD4603D20FC3C3228CE7A061B" box="[978,1098,1091,1115]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="13" pageNumber="12">Bruxelles</collectingRegion>
,
<collectingCountry id="F30B1B9DD4603D20FBB73228CEF4061B" box="[1113,1220,1091,1115]" name="Belgium" pageId="13" pageNumber="12">Belgium</collectingCountry>
), whose generous help enabled the present study.
</paragraph>
</subSubSection>
<subSubSection id="C3060886D4603D20FF2E32C8C93107DB" pageId="13" pageNumber="12" type="discussion">
<paragraph id="8BA35B0DD4603D20FF2E32C8C93107DB" blockId="13.[192,1440,1187,1435]" pageId="13" pageNumber="12">
<emphasis id="B968871FD4603D20FF2E32C8CB7406FB" bold="true" box="[192,324,1187,1211]" pageId="13" pageNumber="12">Remarks.</emphasis>
<taxonomicName id="4C1C208ED4603D20FEA532C8C81D06FB" authorityName="Raj &amp; Sabu &amp; Danyang" authorityYear="2012" box="[331,557,1187,1211]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="drumonti">
<emphasis id="B968871FD4603D20FEA532C8C81D06FB" box="[331,557,1187,1211]" italics="true" pageId="13" pageNumber="12">Omphra drumonti</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C1C208ED4603D20FCFF32CFC9B106FB" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[785,897,1188,1211]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B968871FD4603D20FCFF32CFC9B106FB" box="[785,897,1188,1211]" italics="true" pageId="13" pageNumber="12">O. atrata</emphasis>
</taxonomicName>
having truncate and in lacking an emarginated elytral apex. It is distinguishable from
<taxonomicName id="4C1C208ED4603D20FD0C32AFC963069B" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[738,851,1220,1243]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B968871FD4603D20FD0C32AFC963069B" box="[738,851,1220,1243]" italics="true" pageId="13" pageNumber="12">O. atrata</emphasis>
</taxonomicName>
by the presence of a strongly convex pronotum, less pronounced elytral humerus and distinct yellowish red anterior margin of the abdominal sternites.
<taxonomicName id="4C1C208ED4603D20FF2E3368CB92075B" authorityName="Raj &amp; Sabu &amp; Danyang" authorityYear="2012" box="[192,418,1283,1307]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="drumonti">
<emphasis id="B968871FD4603D20FF2E3368CB92075B" box="[192,418,1283,1307]" italics="true" pageId="13" pageNumber="12">Omphra drumonti</emphasis>
</taxonomicName>
is distinct from
<taxonomicName id="4C1C208ED4603D20FD84336FC92E075B" authorityName="Reiche" authorityYear="1843" box="[618,798,1283,1307]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="complanata">
<emphasis id="B968871FD4603D20FD84336FC92E075B" box="[618,798,1283,1307]" italics="true" pageId="13" pageNumber="12">O. complanata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C208ED4603D20FCB4336FC9E0075B" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[858,976,1283,1307]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B968871FD4603D20FCB4336FC95F075B" box="[858,879,1284,1307]" italics="true" pageId="13" pageNumber="12">O</emphasis>
.
<emphasis id="B968871FD4603D20FC933368C9E0075B" box="[893,976,1283,1307]" italics="true" pageId="13" pageNumber="12">rufipes</emphasis>
</taxonomicName>
in the absence of emarginated elytral apex; further it differs from
<taxonomicName id="4C1C208ED4603D20FDFC334FC8F5077B" authorityName="Reiche" authorityYear="1843" box="[530,709,1315,1339]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="complanata">
<emphasis id="B968871FD4603D20FDFC334FC8F5077B" box="[530,709,1315,1339]" italics="true" pageId="13" pageNumber="12">O. complanata</emphasis>
</taxonomicName>
in the possession of an elongated oval last antennomere and dilated 4
<superScript id="7C69F645D4603D20FEC33329CB0D0710" attach="left" box="[301,317,1346,1360]" fontSize="6" pageId="13" pageNumber="12">th</superScript>
palpomere in contrast to the oval last antennomere and undilated 4
<superScript id="7C69F645D4603D20FB7E3329CE900710" attach="left" box="[1168,1184,1346,1360]" fontSize="6" pageId="13" pageNumber="12">th</superScript>
palpomere.
<taxonomicName id="4C1C208ED4603D20FAD73328CB04073B" authorityName="Raj &amp; Sabu &amp; Danyang" authorityYear="2012" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="drumonti">
<emphasis id="B968871FD4603D20FAD73328CB04073B" italics="true" pageId="13" pageNumber="12">Omphra drumonti</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1C208ED4603D20FE3F330FC877073B" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[465,583,1379,1403]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B968871FD4603D20FE3F330FC877073B" box="[465,583,1379,1403]" italics="true" pageId="13" pageNumber="12">O. rufipes</emphasis>
</taxonomicName>
in having a convex pronotum and brownish red legs in contrast to the flat pronotum and bright red legs in
<emphasis id="B968871FD4603D20FD6C33EFC93107DB" box="[642,769,1411,1435]" italics="true" pageId="13" pageNumber="12">
<taxonomicName id="4C1C208ED4603D20FD6C33EFC8CC07DB" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[642,764,1411,1435]" class="Insecta" family="Carabidae" genus="Omphra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="rufipes">O. rufipes</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>