198 lines
20 KiB
XML
198 lines
20 KiB
XML
<document ID-DOI="10.11646/zootaxa.5263.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7804268" ID-ZooBank="20CB3CA0-BEF9-474C-8931-6A7948B9CA61" approvalRequired="5" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="4" checkinTime="1680760231104" checkinUser="plazi" docAuthor="Řezáč, Milan, Cardoso, Pedro & Řezáčová, Veronika" docDate="2023" docId="039CE028E872FFC0FF12FD83FE46629E" docLanguage="en" docName="zootaxa.5263.3.2.pdf" docOrigin="Zootaxa 5263 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpactea Bristowe 1939" docType="treatment" docVersion="2" lastPageNumber="337" masterDocId="FFA59850E870FFC2FF85FFA1FFD2667D" masterDocTitle="Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal" masterLastPageNumber="364" masterPageNumber="335" pageNumber="337" updateTime="1680760785065" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Řezáč, Milan</mods:namePart>
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<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cardoso, Pedro</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-8119-9960</mods:nameIdentifier>
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<mods:affiliation>Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History Luomus, University of Helsinki, P. O. Box 17, FI- 00014 Helsinki, Finland. pedro. cardoso @ helsinki. fi; https: // orcid. org / 0000 - 0001 - 8119 - 9960</mods:affiliation>
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<mods:nameIdentifier type="email">pedro.cardoso@helsinki.fi</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Řezáčová, Veronika</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-1749-0355</mods:nameIdentifier>
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<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia & strver @ seznam. cz; https: // orcid. org / 0000 - 0002 - 1749 - 0355</mods:affiliation>
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<mods:nameIdentifier type="email">strver@seznam.cz</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5263</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>335</mods:start>
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<mods:end>364</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.3.2</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5263.3.2</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7804268</mods:identifier>
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<mods:identifier type="ZooBank">20CB3CA0-BEF9-474C-8931-6A7948B9CA61</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7804272" ID-Zenodo-Dep="7804272" LSID="urn:lsid:plazi:treatment:039CE028E872FFC0FF12FD83FE46629E" httpUri="http://treatment.plazi.org/id/039CE028E872FFC0FF12FD83FE46629E" lastPageNumber="337" pageId="2" pageNumber="337">
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<subSubSection box="[151,557,546,573]" pageId="2" pageNumber="337" type="nomenclature">
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<paragraph blockId="2.[151,557,546,573]" box="[151,557,546,573]" pageId="2" pageNumber="337">
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<heading bold="true" box="[151,557,546,573]" fontSize="11" level="1" pageId="2" pageNumber="337" reason="1">
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<emphasis bold="true" box="[151,557,546,573]" pageId="2" pageNumber="337">
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Genus
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<taxonomicName ID-CoL="4TS5" ID-ENA="94013" authority="Bristowe, 1939" authorityName="Bristowe" authorityYear="1939" box="[238,557,546,573]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[238,365,547,573]" italics="true" pageId="2" pageNumber="337">Harpactea</emphasis>
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<bibRefCitation author="Bristowe, W. S." box="[373,557,546,572]" pageId="2" pageNumber="337" refId="ref19005" refString="Bristowe, W. S. (1939) The Comity of Spiders. Ray Society, London, 228 pp." type="book" year="1939">Bristowe, 1939</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,888,615,639]" pageId="2" pageNumber="337" type="materials_examined">
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<paragraph blockId="2.[151,888,615,639]" box="[151,888,615,639]" pageId="2" pageNumber="337">
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<typeStatus box="[151,201,615,639]" pageId="2" pageNumber="337">Type</typeStatus>
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species
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<taxonomicName authority="Blackwall, 1832" authorityName="Blackwall" authorityYear="1832" box="[287,636,615,639]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="latreillii">
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<emphasis box="[287,461,615,639]" italics="true" pageId="2" pageNumber="337">Dysdera latreillii</emphasis>
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<bibRefCitation author="Blackwall, J." box="[468,636,615,639]" pageId="2" pageNumber="337" pagination="190 - 191" refId="ref19048" refString="Blackwall, J. (1832) Description of a species of Arachnida, hitherto uncharacterized, belonging to the family Araneidae. London and Edinburgh Philosophical Magazine and Journal of Science, Series 3, 1, 190 - 191. https: // doi. org / 10.1080 / 14786443208647870" type="journal article" year="1832">Blackwall, 1832</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="337" type="discussion">
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<paragraph blockId="2.[151,1437,685,1251]" pageId="2" pageNumber="337">
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<emphasis bold="true" box="[151,267,685,711]" pageId="2" pageNumber="337">Remarks.</emphasis>
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<taxonomicName baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[278,523,685,711]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="hombergii">
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<emphasis box="[278,523,685,711]" italics="true" pageId="2" pageNumber="337">Harpactea hombergii</emphasis>
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</taxonomicName>
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is currently stated as the
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<typeStatus box="[834,882,685,711]" pageId="2" pageNumber="337">type</typeStatus>
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species of the genus
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<taxonomicName authorityName="Bristowe" authorityYear="1939" box="[1142,1262,686,711]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis box="[1142,1262,686,711]" italics="true" pageId="2" pageNumber="337">Harpactea</emphasis>
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</taxonomicName>
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(
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<bibRefCitation author="World Spider Catalog" pageId="2" pageNumber="337" refId="ref21764" refString="World Spider Catalog (2021) World Spider Catalog, Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 2 February 2021)" type="url" year="2021">World Spider Catalog 2021</bibRefCitation>
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). It is through synonymy of
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<taxonomicName authorityName="Scopoli" authorityYear="1763" box="[629,834,721,747]" class="Arachnida" family="Agelenidae" genus="Aranea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="hombergii">
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<emphasis box="[629,834,721,747]" italics="true" pageId="2" pageNumber="337">Aranea hombergii</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Blackwall" authorityYear="1832" box="[890,1084,721,747]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="latreillii">
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<emphasis box="[890,1084,721,747]" italics="true" pageId="2" pageNumber="337">Dysdera latreillii</emphasis>
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</taxonomicName>
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.
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<bibRefCitation author="Templeton, R." box="[1099,1300,721,748]" pageId="2" pageNumber="337" pagination="400 - 408" refId="ref21598" refString="Templeton, R. (1835) On the spiders of the genus Dysdera Latr. with the description of a new allied genus. The Zoological Journal, 5, 400 - 408." type="journal article" year="1835">Templeton (1835)</bibRefCitation>
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described a new dysderid genus
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<taxonomicName authorityName="Swainson" authorityYear="1833" box="[379,496,758,783]" class="Insecta" family="Carabidae" genus="Harpactes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis box="[379,496,758,783]" italics="true" pageId="2" pageNumber="337">Harpactes</emphasis>
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</taxonomicName>
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for
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<taxonomicName authorityName="Blackwall" authorityYear="1832" box="[542,734,757,783]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="latreillii">
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<emphasis box="[542,734,757,783]" italics="true" pageId="2" pageNumber="337">Dysdera latreillii</emphasis>
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</taxonomicName>
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. He was not sure whether his specimens were conspecific with
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<bibRefCitation author="Blackwall, J." box="[151,366,793,819]" pageId="2" pageNumber="337" pagination="190 - 191" refId="ref19048" refString="Blackwall, J. (1832) Description of a species of Arachnida, hitherto uncharacterized, belonging to the family Araneidae. London and Edinburgh Philosophical Magazine and Journal of Science, Series 3, 1, 190 - 191. https: // doi. org / 10.1080 / 14786443208647870" type="journal article" year="1832">Blackwall’s (1832)</bibRefCitation>
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<emphasis box="[375,575,793,819]" italics="true" pageId="2" pageNumber="337">
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<taxonomicName authorityName="Blackwall" authorityYear="1832" box="[375,571,793,819]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="latreillii">Dysdera latreillii</taxonomicName>
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,
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</emphasis>
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therefore he called it
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<taxonomicName authorityName="Blackwall" authorityYear="1832" box="[827,953,793,819]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="latreillii">
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<emphasis box="[827,953,793,819]" italics="true" pageId="2" pageNumber="337">D. latreillii</emphasis>
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</taxonomicName>
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with question mark. Vigors (in
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<bibRefCitation author="Templeton, R." pageId="2" pageNumber="337" pagination="400 - 408" refId="ref21598" refString="Templeton, R. (1835) On the spiders of the genus Dysdera Latr. with the description of a new allied genus. The Zoological Journal, 5, 400 - 408." type="journal article" year="1835">Templeton 1835</bibRefCitation>
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), the editor of the paper, emended the name to
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<taxonomicName box="[754,908,829,855]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="templetoni">
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<emphasis box="[754,908,829,855]" italics="true" pageId="2" pageNumber="337">D. templetoni</emphasis>
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</taxonomicName>
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. Later,
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<bibRefCitation author="Bristowe, W. S." box="[997,1182,829,855]" pageId="2" pageNumber="337" refId="ref19005" refString="Bristowe, W. S. (1939) The Comity of Spiders. Ray Society, London, 228 pp." type="book" year="1939">Bristowe (1939)</bibRefCitation>
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noticed that the name
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<taxonomicName authorityName="Swainson" authorityYear="1833" box="[151,268,866,891]" class="Insecta" family="Carabidae" genus="Harpactes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,268,866,891]" italics="true" pageId="2" pageNumber="337">Harpactes</emphasis>
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</taxonomicName>
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was already preoccupied, so he replaced it with
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<taxonomicName authorityName="Bristowe" authorityYear="1939" box="[824,944,866,891]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis box="[824,944,866,891]" italics="true" pageId="2" pageNumber="337">Harpactea</emphasis>
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</taxonomicName>
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. In summary, the
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<typeStatus box="[1151,1199,865,891]" pageId="2" pageNumber="337">type</typeStatus>
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species of the genus
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<taxonomicName authorityName="Bristowe" authorityYear="1939" box="[151,271,902,927]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,271,902,927]" italics="true" pageId="2" pageNumber="337">Harpactea</emphasis>
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</taxonomicName>
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is undoubtedly
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<taxonomicName authorityName="Blackwall" authorityYear="1832" box="[451,644,901,927]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="latreillii">
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<emphasis box="[451,644,901,927]" italics="true" pageId="2" pageNumber="337">Dysdera latreillii</emphasis>
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</taxonomicName>
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, not
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<taxonomicName authorityName="Scopoli" authorityYear="1763" box="[700,904,901,927]" class="Arachnida" family="Agelenidae" genus="Aranea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="hombergii">
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<emphasis box="[700,904,901,927]" italics="true" pageId="2" pageNumber="337">Aranea hombergii</emphasis>
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</taxonomicName>
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(see also
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<bibRefCitation author="Thaler, K. & Knoflach, B." box="[1015,1294,901,927]" pageId="2" pageNumber="337" pagination="413 - 444" refId="ref21633" refString="Thaler, K. & Knoflach, B. (2002) Zur Faunistik der Spinnen (Araneae) von Osterreich: Atypidae, Haplogynae, Eresidae, Zodariidae, Mimetidae. Linzer biologische Beitrage, 34, 413 - 444." type="journal article" year="2002">Thaler & Knoflach 2002</bibRefCitation>
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,
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<bibRefCitation author="Rezac, M. & Gasparo, F. & Kral, J. & Heneberg, P." pageId="2" pageNumber="337" pagination="451 - 474" refId="ref21095" refString="Rezac, M., Gasparo, F., Kral, J. & Heneberg, P. (2014) Integrative taxonomy and evolutionary history of a newly revealed Dysdera ninnii complex (Araneae: Dysderidae). Zoological Journal of the Linnean Society, 172, 451 - 474. https: // doi. org / 10.1111 / zoj. 12177" type="journal article" year="2014">
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Řezáč
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<emphasis box="[1380,1437,901,927]" italics="true" pageId="2" pageNumber="337">et al.</emphasis>
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2014
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</bibRefCitation>
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).
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</paragraph>
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<paragraph blockId="2.[151,1437,685,1251]" pageId="2" pageNumber="337">
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<emphasis bold="true" box="[199,469,973,999]" pageId="2" pageNumber="337">Diagnostic characters.</emphasis>
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Diagnostic characters of
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<taxonomicName authorityName="Bristowe" authorityYear="1939" box="[772,892,974,999]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis box="[772,892,974,999]" italics="true" pageId="2" pageNumber="337">Harpactea</emphasis>
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</taxonomicName>
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include the body size, colour of prosoma, leg spination and the shape of male chelicerae and copulatory organs.
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<taxonomicName authorityName="Bristowe" authorityYear="1939" box="[924,1044,1010,1035]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis box="[924,1044,1010,1035]" italics="true" pageId="2" pageNumber="337">Harpactea</emphasis>
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</taxonomicName>
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are usually small to medium size spiders (carapace length
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<quantity box="[426,562,1045,1071]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="2.3" metricValueMin="1.3" pageId="2" pageNumber="337" unit="mm" value="1.7999999999999998" valueMax="2.3" valueMin="1.3">1.3–2.3 mm</quantity>
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, although members of the group
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<emphasis box="[927,1042,1045,1071]" italics="true" pageId="2" pageNumber="337">rubicunda</emphasis>
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from the eastern Meditteranean are usuay larger), with a homogeneous body shape (
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<figureCitation box="[683,747,1081,1107]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="2" pageNumber="337">Fig. 1</figureCitation>
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), although some species have elongated body and appendages (
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<emphasis box="[161,201,1118,1143]" italics="true" pageId="2" pageNumber="337">e.g.</emphasis>
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,
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<figureCitation box="[216,303,1117,1143]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="2" pageNumber="337">Fig. 1C</figureCitation>
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). The prosoma is brown or yellow, usually darker in the anterior part. The abdomen shows no color patterns, its cuticle is hardly pigmented, so that grey or brownish midgut is visible. The shape of the chelicerae and the arrangement of the cheliceral teeth are uniform and characteristic for the majority of species of the genus (see the
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1980" box="[192,256,1225,1251]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="fageli">
|
||
<emphasis box="[192,256,1225,1251]" italics="true" pageId="2" pageNumber="337">fageli</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus box="[263,311,1225,1251]" pageId="2" pageNumber="337">type</typeStatus>
|
||
below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |