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<document id="388A66AA9A2921A6C63640F059EA8800" ID-CLB-Dataset="297149" ID-DOI="10.1093/zoolinnean/zlad118" ID-GBIF-Dataset="657c53fd-7c2f-4a59-ade4-152de4cf8d03" ID-ISSN="0024-4082" ID-Zenodo-Dep="11241048" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1716363684985" checkinUser="plazi" docAuthor="Sánchez-García, Alba, Sendra, Alberto, Davis, Steven R. &amp; Grimaldi, David A." docDate="2024" docId="039787F42D1FFFACEB61FE41FD6BFEDD" docLanguage="en" docName="zlad118.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (1)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad118" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Rostricampa Sanchez-Garcia, Sendra and Grimaldi" docType="treatment" docVersion="6" lastPageNumber="154" masterDocId="FFAEFF8C2D12FFBEE826FFA0FFF8F917" masterDocTitle=" Dawn hexapods in Cenozoic ambers (Diplura: Campodeoidea)" masterLastPageNumber="158" masterPageNumber="136" pageNumber="149" updateTime="1732901998317" updateUser="ExternalLinkService">
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<mods:titleInfo id="2D18AFCD5A4F3F051BE8CE856E7EBC74">
<mods:title id="5E3641177C4E355C8E82093332E40394"> Dawn hexapods in Cenozoic ambers (Diplura: Campodeoidea)</mods:title>
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<mods:name id="D82F55CAE35974D6CA57C75FCB2CA512" type="personal">
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<mods:namePart id="D8192B25E612C981FF061B8FDFACAFB9">Sánchez-García, Alba</mods:namePart>
<mods:affiliation id="6B0E16BF988AABCED8D9F8ABAD698181">Instituto Geológico y Minero de España (IGME), CSIC, València, Spain</mods:affiliation>
<mods:nameIdentifier id="543F79E810D0059FF76FAEB262D51BB4" type="email">alba.sanchez@csic.es</mods:nameIdentifier>
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<mods:namePart id="B960F3D68322F59DE02B3350DA925651">Sendra, Alberto</mods:namePart>
<mods:affiliation id="FD7C69603E1074DB87B67C21B84C6A5D">Coleccions Torres-Sala i Siro de Fez, Servei de Patrimoni Històric, Ajuntament de València, València, Spain &amp; Departament de Didàctica de les Ciències Experimentals i Socials, Facultat de Magisteri, Universitat de València, València, Spain</mods:affiliation>
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<mods:namePart id="8E6F645B9BDF961F306EFF9AF23888CB">Davis, Steven R.</mods:namePart>
<mods:affiliation id="EC9B4A0DFC5115252E871A8F4517CD60">Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA</mods:affiliation>
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</mods:role>
<mods:namePart id="FD21DAE2664B5117A6423CD2F84E672D">Grimaldi, David A.</mods:namePart>
<mods:affiliation id="43ED4F851981605FFF22EB6E10C45535">Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA</mods:affiliation>
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<mods:date id="EF68C653F50257FF748B49ECED1C81DD">2024</mods:date>
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<treatment id="039787F42D1FFFACEB61FE41FD6BFEDD" ID-DOI="http://doi.org/10.5281/zenodo.11354292" ID-GBIF-Taxon="231564027" ID-Zenodo-Dep="11354292" LSID="urn:lsid:plazi:treatment:039787F42D1FFFACEB61FE41FD6BFEDD" httpUri="http://treatment.plazi.org/id/039787F42D1FFFACEB61FE41FD6BFEDD" lastPageId="18" lastPageNumber="154" pageId="13" pageNumber="149" scope_class="Entognatha" scope_family="Campodeidae" scope_order="Diplura" scope_subFamily="Campodeinae" scope_superFamily="Campodeoidea">
<subSubSection id="C32465692D1FFFB3EB61FE41FC21FB4E" pageId="13" pageNumber="149" type="nomenclature">
<paragraph id="8B8136E22D1FFFB3EB61FE41FA4CF8EB" blockId="13.[839,1460,481,508]" box="[839,1460,481,508]" pageId="13" pageNumber="149">
<heading id="D0C9818E2D1FFFB3EB61FE41FA4CF8EB" bold="true" box="[839,1460,481,508]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="149" reason="2">
<emphasis id="B94AEAF02D1FFFB3EB61FE41FA4CF8EB" bold="true" box="[839,1460,481,508]" pageId="13" pageNumber="149">
<taxonomicName id="4C3E4D612D1FFFB3EB61FE41FAA1F8EC" authority="Sanchez-Garcia, Sendra and Grimaldi" authorityName="Sanchez-Garcia, Sendra and Grimaldi" box="[839,1369,481,508]" class="Diplura" family="Campodeidae" genus="Rostricampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAF02D1FFFB3EB61FE41FC30F8EC" bold="true" box="[839,968,481,507]" italics="true" pageId="13" pageNumber="149">Rostricampa</emphasis>
Sánchez-García, Sendra and Grimaldi
</taxonomicName>
gen nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8136E22D1FFFB3EB1FFD82FC21FB4E" blockId="13.[825,1474,546,601]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3EB1FFD82FC80FB2E" box="[825,888,546,569]" italics="true" pageId="13" pageNumber="149">LSID:</emphasis>
<uri id="FFAF3AE02D1FFFB3EBA6FD82FC21FB4E" pageId="13" pageNumber="149">
urn:lsid:zoobank.org:act:
<uuid id="FF980C372D1FFFB3EC5CFD82FC21FB4E" pageId="13" pageNumber="149">A4331BAB-13E2-494A-AF61- 1226546147A1</uuid>
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</paragraph>
</subSubSection>
<subSubSection id="C32465692D1FFFB3EB1FFDDBFB2EFC75" pageId="13" pageNumber="149" type="etymology">
<paragraph id="8B8136E22D1FFFB3EB1FFDDBFAABFBC6" blockId="13.[825,1474,635,721]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3EB1FFDDBFC50FB84" box="[825,936,635,659]" italics="true" pageId="13" pageNumber="149">Etymology:</emphasis>
The generic name is a combination of the Latin
<emphasis id="B94AEAF02D1FFFB3EB1FFD3BFC7DFBA5" box="[825,901,667,690]" italics="true" pageId="13" pageNumber="149">rostrum</emphasis>
(bill, beak, snout, muzzle) and the typical suffix for campodeids,
<emphasis id="B94AEAF02D1FFFB3EBE6FD1AFBF9FBC6" box="[960,1025,698,721]" italics="true" pageId="13" pageNumber="149">campa</emphasis>
. Gender of the name is feminine.
</paragraph>
<paragraph id="8B8136E22D1FFFB3EB1FFD53FB81FA3D" blockId="13.[825,1474,755,810]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3EB1FFD53FC44FA1D" box="[825,956,755,778]" italics="true" pageId="13" pageNumber="149">
<typeStatus id="548588402D1FFFB3EB1FFD53FC91FA1D" box="[825,873,755,778]" pageId="13" pageNumber="149">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C3E4D612D1FFFB3EBECFD53FC6EFA3D" authority="Sanchez-Garcia, Sendra and Grimaldi" authorityName="Sanchez-Garcia, Sendra and Grimaldi" authorityYear="2024" class="Diplura" family="Campodeidae" genus="Rostricampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="149" phylum="Arthropoda" rank="species" species="engeli" status="sp. nov.">
<emphasis id="B94AEAF02D1FFFB3EBECFD53FB70FA1C" box="[970,1160,755,779]" italics="true" pageId="13" pageNumber="149">Rostricampa engeli</emphasis>
Sánchez-García, Sendra and Grimaldi
</taxonomicName>
<taxonomicNameLabel id="A279578B2D1FFFB3EBBDFCB2FC1FFA3E" box="[923,999,786,809]" pageId="13" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
by monotypy.
</paragraph>
<paragraph id="8B8136E22D1FFFB3EB1FFCECFB2EFDED" blockId="13.[825,1475,843,1274]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3EB1FFCECFC67FA74" box="[825,927,844,867]" italics="true" pageId="13" pageNumber="149">Diagnosis:</emphasis>
Male. Frontal process, clypeus and labium forming a sclerotized beak protruding from head. Antenna with at least 20 antennomeres, moniliform. Distribution of thoracic macrosetae: pronotum with 1 + 1
<emphasis id="B94AEAF02D1FFFB3EC90FC09FB30FAD6" box="[1206,1224,937,961]" italics="true" pageId="13" pageNumber="149">la</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1FFFB3ED39FC09FACAFAD6" box="[1311,1330,937,961]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
; mesonotum with 1 + 1
<emphasis id="B94AEAF02D1FFFB3EB92FC69FC2BFAF7" box="[948,979,969,992]" italics="true" pageId="13" pageNumber="149">ma</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1FFFB3EC00FC69FBC0FAF6" box="[1062,1080,969,993]" italics="true" pageId="13" pageNumber="149">la</emphasis>
1 + 1
<emphasis id="B94AEAF02D1FFFB3ECA3FC69FB60FAF6" box="[1157,1176,969,993]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
; metanotum with 1 + 1
<emphasis id="B94AEAF02D1FFFB3ED8EFC69FA43FAF6" box="[1448,1467,969,993]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
; marginal setae thin. Metathoracic leg reaching abdominal segment IV; femur without dorsal macrosetae and with one ventral macroseta; tibia with one ventral macroseta; claws subequal, with small lateral crests; pretarsal lateral processes laminar and curved, smooth. Styli with a ventromedial macroseta bi- or trifurcate. Cerci slightly shorter than body, with 25 articles, with whorls of setae and macrosetae both barbed and smooth. Male urosternite I with enlarged subtrapezoidal subcoxal appendages with glandular setae in a large distal area.
</paragraph>
<paragraph id="8B8136E22D1FFFB3EBAFFA88FB2EFC75" blockId="13.[905,1395,1319,1378]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3EBAFFA88FB2EFC75" bold="true" pageId="13" pageNumber="149">
<taxonomicName id="4C3E4D612D1FFFB3EBAFFA88FB7DFC76" authority="Sanchez-Garcia, Sendra and Grimaldi" authorityName="Sanchez-Garcia, Sendra and Grimaldi" authorityYear="2024" class="Diplura" family="Campodeidae" genus="Rostricampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="149" phylum="Arthropoda" rank="species" species="engeli" status="sp. nov.">
<emphasis id="B94AEAF02D1FFFB3EBAFFA88FBBFFC54" bold="true" box="[905,1095,1320,1347]" italics="true" pageId="13" pageNumber="149">Rostricampa engeli</emphasis>
Sánchez-García, Sendra and Grimaldi
</taxonomicName>
<taxonomicNameLabel id="A279578B2D1FFFB3ECACFAE8FB2EFC75" box="[1162,1238,1352,1378]" pageId="13" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32465692D1FFFACEC1EFA26FD6BFEDD" lastPageId="18" lastPageNumber="154" pageId="13" pageNumber="149" type="description">
<paragraph id="8B8136E22D1FFFB3EC1EFA26FB3BFCB7" blockId="13.[1080,1219,1414,1440]" box="[1080,1219,1414,1440]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3EC1EFA26FB3BFCB7" bold="true" box="[1080,1219,1414,1440]" pageId="13" pageNumber="149">
(
<figureCitation id="13052A672D1FFFB3EC65FA26FB41FCB7" box="[1091,1209,1414,1440]" captionStart-0="Figure 11" captionStart-1="Figure 12" captionStart-2="Figure 13" captionStart-3="Figure 14" captionStartId-0="14.[113,178,1470,1494]" captionStartId-1="15.[129,194,1836,1860]" captionStartId-2="16.[113,178,1809,1833]" captionStartId-3="17.[129,194,1833,1857]" captionTargetBox-0="[117,1455,147,1439]" captionTargetBox-1="[220,1382,147,1805]" captionTargetBox-2="[117,1455,147,1779]" captionTargetBox-3="[133,1469,150,1802]" captionTargetId-0="figure-230@14.[114,1458,144,1442]" captionTargetId-1="figure-8@15.[217,1385,144,1808]" captionTargetId-2="figure-7@16.[114,1458,144,1782]" captionTargetId-3="figure-9@17.[129,1473,144,1805]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionText-0="Figure 11. Dorsal photomicrograph of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber.A, dorsolateral habitus. B, detail of head in lateral view.C, detail of anterior part of the head." captionText-1="Figure 12. Camera lucida drawings of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber. A, detail of head. B, dorsolateral habitus. C, detail of stylus and eversible vesicle of urosternite II. D, detail of stylus and eversible vesicle of urosternite V. E, detail of stylus and eversible vesicle of urosternite VI." captionText-2="Figure 13. Photomicrographs of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber. A, detail of left antennomeres IVVI.B, detail of ventral femoral macroseta of right leg I. C, detail of pretarsus; asterisks indicate the lateral processes. D, detail of subcoxal appendage of urosternite I. E, detail of stylus and eversible vesicle of urosternite V.F, detail of stylus and eversible vesicle of urosternite VI. Abbreviations: T, trichobothria." captionText-3="Figure 14. Computed tomography scan images of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber. A, lateral habitus of anterior part of body. B, dorsal habitus of anterior part of body. C, head in dorsal view. D, head in ventral view. E, head in lateral view.AB and CE are to the same scale." figureDoi-0="http://doi.org/10.5281/zenodo.11241072" figureDoi-1="http://doi.org/10.5281/zenodo.11241074" figureDoi-2="http://doi.org/10.5281/zenodo.11241076" figureDoi-3="http://doi.org/10.5281/zenodo.11241078" httpUri-0="https://zenodo.org/record/11241072/files/figure.png" httpUri-1="https://zenodo.org/record/11241074/files/figure.png" httpUri-2="https://zenodo.org/record/11241076/files/figure.png" httpUri-3="https://zenodo.org/record/11241078/files/figure.png" pageId="13" pageNumber="149">Figs 1114</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B8136E22D1FFFB3EB1FFA67FC13FCE9" blockId="13.[825,1474,1478,1534]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3EB1FFA67FC80FCC9" box="[825,888,1479,1502]" italics="true" pageId="13" pageNumber="149">LSID:</emphasis>
<uri id="FFAF3AE02D1FFFB3EB5FFA66FC13FCE9" pageId="13" pageNumber="149">
urn:lsid:zoobank.org:act:
<uuid id="FF980C372D1FFFB3EC51FA66FC13FCE9" pageId="13" pageNumber="149">C9561FC1-BAC2-44F4-8FAA-7D7E2CD53F06</uuid>
</uri>
</paragraph>
<paragraph id="8B8136E22D1FFFB3EB1FF9BFFA43FF40" blockId="13.[825,1474,1567,1623]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3EB1FF9BFFC50FF20" box="[825,936,1567,1591]" italics="true" pageId="13" pageNumber="149">Etymology:</emphasis>
The specific epithet is a patronym in honour of a great mentor, colleague and entomologist, Dr Michael S. Engel.
</paragraph>
<paragraph id="8B8136E22D1FFFB3EB1FF9D8FAF3FEBD" blockId="13.[825,1474,1656,1962]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3EB1FF9D8FC6DFF87" box="[825,917,1656,1680]" italics="true" pageId="13" pageNumber="149">Material:</emphasis>
<materialsCitation id="3B563CBF2D1FFFB3EBBDF9D8FAF3FEBD" ID-GBIF-Occurrence="4867528301" collectionCode="AMNH" collectorName="K. Luzzi &amp; EpoTek" pageId="13" pageNumber="149" specimenCount="1" typeStatus="holotype">
<typeStatus id="548588402D1FFFB3EBBDF9D8FC06FF87" box="[923,1022,1656,1680]" pageId="13" pageNumber="149" type="holotype">Holotype</typeStatus>
<emphasis id="B94AEAF02D1FFFB3EC22F9D8FB1DFF87" box="[1028,1253,1656,1680]" italics="true" pageId="13" pageNumber="149">
<collectionCode id="ED2FAE272D1FFFB3EC22F9D8FBABFF98" box="[1028,1107,1656,1679]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="149">AMNH</collectionCode>
DR-KLCa002
</emphasis>
, male, adult; donated by
<collectorName id="26CB53342D1FFFB3EB7DF938FC4BFFB8" box="[859,947,1687,1711]" pageId="13" pageNumber="149">K. Luzzi</collectorName>
and held in the
<collectionCode id="ED2FAE272D1FFFB3EC43F938FB41FFA7" box="[1125,1209,1688,1712]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="149">AMNH</collectionCode>
; specimen virtually complete, with tips of both antennae cut off at the amber surface; a reddish halo is around most of the specimen; there are bubbles in the anterior and posterior part of the body, and all abdominal vesicles are extruded, probably as a stress behaviour attributable to being trapped in resin; observable laterally (both sides) and dorsally; the piece is very transparent, orange in colour, embedded in a small block of
<collectorName id="26CB53342D1FFFB3EC64F8D3FB68FE9C" box="[1090,1168,1907,1931]" pageId="13" pageNumber="149">EpoTek</collectorName>
resin trimmed to 1.0 cm × 1.0 cm ×
<quantity id="4CC69B072D1FFFB3EB50F832FC45FEBE" box="[886,957,1938,1962]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="149" unit="cm" value="0.4">0.4 cm</quantity>
; syninclusions include one acari.
</materialsCitation>
</paragraph>
<caption id="DF41666A2D1CFFB0E857FA1EFCC6FCE5" ID-DOI="http://doi.org/10.5281/zenodo.11241072" ID-Zenodo-Dep="11241072" httpUri="https://zenodo.org/record/11241072/files/figure.png" pageId="14" pageNumber="150" startId="14.[113,178,1470,1494]" targetBox="[117,1455,147,1439]" targetPageId="14" targetType="figure">
<paragraph id="8B8136E22D1CFFB0E857FA1EFCC6FCE5" blockId="14.[113,1442,1470,1522]" pageId="14" pageNumber="150">
<emphasis id="B94AEAF02D1CFFB0E857FA1EFF2EFCC1" bold="true" box="[113,214,1470,1494]" pageId="14" pageNumber="150">Figure 11.</emphasis>
Dorsal photomicrograph of holotype of
<taxonomicName id="4C3E4D612D1CFFB0EA72FA1EFC98FCC2" authority=", AMNH" authorityName="AMNH" box="[596,864,1470,1494]" class="Diplura" family="Campodeidae" genus="Rostricampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="150" phylum="Arthropoda" rank="species" species="engeli">
<emphasis id="B94AEAF02D1CFFB0EA72FA1EFCF6FCC1" bold="true" box="[596,782,1470,1494]" italics="true" pageId="14" pageNumber="150">Rostricampa engeli</emphasis>
,
<emphasis id="B94AEAF02D1CFFB0EB31FA1EFC98FCC2" box="[791,864,1470,1493]" italics="true" pageId="14" pageNumber="150">AMNH</emphasis>
</taxonomicName>
<emphasis id="B94AEAF02D1CFFB0EB43FA1EFC10FCC1" box="[869,1000,1470,1494]" italics="true" pageId="14" pageNumber="150">DR-KLCa002</emphasis>
, in Miocene Dominican amber. A, dorsolateral habitus. B, detail of head in lateral view. C, detail of anterior part of the head.
</paragraph>
</caption>
<paragraph id="8B8136E22D1CFFB0E857F982FDD3FF2D" blockId="14.[113,555,1570,1594]" box="[113,555,1570,1594]" pageId="14" pageNumber="150">
<emphasis id="B94AEAF02D1CFFB0E857F982FF1DFF2E" box="[113,229,1570,1593]" italics="true" pageId="14" pageNumber="150">Occurrence:</emphasis>
Miocene,
<collectingCountry id="F32976722D1CFFB0E969F982FDDFFF2D" box="[335,551,1570,1594]" name="Dominican Republic" pageId="14" pageNumber="150">Dominican Republic</collectingCountry>
.
</paragraph>
<paragraph id="8B8136E22D1CFFB0E857F9FCFE0CFF63" blockId="14.[113,500,1627,1652]" box="[113,500,1627,1652]" pageId="14" pageNumber="150">
<emphasis id="B94AEAF02D1CFFB0E857F9FCFF2FFF64" box="[113,215,1628,1651]" italics="true" pageId="14" pageNumber="150">Diagnosis:</emphasis>
As for genus, by monotypy.
</paragraph>
<paragraph id="8B8136E22D1CFFB0E857F931FE9EFFBF" blockId="14.[113,358,1680,1704]" box="[113,358,1680,1704]" pageId="14" pageNumber="150">
<emphasis id="B94AEAF02D1CFFB0E857F931FF10FFBF" box="[113,232,1681,1704]" italics="true" pageId="14" pageNumber="150">Description:</emphasis>
Male, adult.
</paragraph>
<paragraph id="8B8136E22D1CFFB0E857F965FD0BFE4D" blockId="14.[113,762,1733,1882]" pageId="14" pageNumber="150">
<emphasis id="B94AEAF02D1CFFB0E857F965FF53FFCA" box="[113,171,1733,1757]" italics="true" pageId="14" pageNumber="150">Body:</emphasis>
Length
<quantity id="4CC69B072D1CFFB0E921F965FE9DFFCB" box="[263,357,1733,1757]" metricMagnitude="-3" metricUnit="m" metricValue="4.66" pageId="14" pageNumber="150" unit="mm" value="4.66">4.66 mm</quantity>
(
<figureCitation id="13052A672D1CFFB0E95FF965FE2AFFCA" box="[377,466,1733,1757]" captionStart="Figure 11" captionStartId="14.[113,178,1470,1494]" captionTargetBox="[117,1455,147,1439]" captionTargetId="figure-230@14.[114,1458,144,1442]" captionTargetPageId="14" captionText="Figure 11. Dorsal photomicrograph of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber.A, dorsolateral habitus. B, detail of head in lateral view.C, detail of anterior part of the head." figureDoi="http://doi.org/10.5281/zenodo.11241072" httpUri="https://zenodo.org/record/11241072/files/figure.png" pageId="14" pageNumber="150">Fig. 11A</figureCitation>
). Cuticle very irregular and webby, probably owing to slightly pyritized nature of the specimen (there is a reddish halo around most of the specimen), with no distinct microstructures, probably smooth; body with short, thin, clothing setae, smooth or with few thin distal barbs.
</paragraph>
<paragraph id="8B8136E22D1CFFACE857F8DBFF56F9D0" blockId="14.[113,763,1915,1971]" lastBlockId="18.[113,763,144,199]" lastPageId="18" lastPageNumber="154" pageId="14" pageNumber="150">
<emphasis id="B94AEAF02D1CFFB0E857F8DBFF48FE84" box="[113,176,1915,1939]" italics="true" pageId="14" pageNumber="150">Head:</emphasis>
Length
<quantity id="4CC69B072D1CFFB0E928F8DBFE89FE84" box="[270,369,1915,1939]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="14" pageNumber="150" unit="mm" value="0.73">0.73 mm</quantity>
, 0.16 × length of body. Frontal process well developed, together with an elongated clypeus and probably the labium that form a tapered, apparently sclerotized beak protruding from head (
<figureCitation id="13052A672D1CFFB0EC8CF9E1FAFDFF4E" box="[1194,1285,1601,1625]" captionStart="Figure 11" captionStartId="14.[113,178,1470,1494]" captionTargetBox="[117,1455,147,1439]" captionTargetId="figure-230@14.[114,1458,144,1442]" captionTargetPageId="14" captionText="Figure 11. Dorsal photomicrograph of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber.A, dorsolateral habitus. B, detail of head in lateral view.C, detail of anterior part of the head." figureDoi="http://doi.org/10.5281/zenodo.11241072" httpUri="https://zenodo.org/record/11241072/files/figure.png" pageId="14" pageNumber="150">Fig. 11B</figureCitation>
); on the lateral side is a group of ~20 very fine setae, three of these close to the base of the antenna raised into low tubercles, which might be membranous (
<figureCitation id="13052A672D1CFFB0EBE0F93FFBD8FFA0" box="[966,1056,1695,1719]" captionStart="Figure 11" captionStartId="14.[113,178,1470,1494]" captionTargetBox="[117,1455,147,1439]" captionTargetId="figure-230@14.[114,1458,144,1442]" captionTargetPageId="14" captionText="Figure 11. Dorsal photomicrograph of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber.A, dorsolateral habitus. B, detail of head in lateral view.C, detail of anterior part of the head." figureDoi="http://doi.org/10.5281/zenodo.11241072" httpUri="https://zenodo.org/record/11241072/files/figure.png" pageId="14" pageNumber="150">Fig. 11C</figureCitation>
); apical part of head showing a higher grade of sclerotization. Labial palp suboval and small, setose. Both antennae incomplete, maximum length as preserved
<quantity id="4CC69B072D1CFFB0EB0CF95DFC74FE02" box="[810,908,1789,1813]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="14" pageNumber="150" unit="mm" value="2.1">2.10 mm</quantity>
, with ≥ 20 antennomeres; base of the antenna with a rather large recessed area, the margins of which form a steep edge; antennomeres I and II subrectangular; antennomere III onwards moniliform, twice as long as wide (length
<quantity id="4CC69B072D1CFFB0ED69F8FBFA56FE64" box="[1359,1454,1883,1907]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="150" unit="mm" value="0.12">0.12 mm</quantity>
, width
<quantity id="4CC69B072D1CFFB0EB56F8DAFC2EFE84" box="[880,982,1914,1939]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="150" unit="mm" value="0.06">0.06 mm</quantity>
); trichobothria visible in antennomeres III VI (
<figureCitation id="13052A672D1CFFB0EB7FF83AFC4CFEA5" box="[857,948,1946,1970]" captionStart="Figure 13" captionStartId="16.[113,178,1809,1833]" captionTargetBox="[117,1455,147,1779]" captionTargetId="figure-7@16.[114,1458,144,1782]" captionTargetPageId="16" captionText="Figure 13. Photomicrographs of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber. A, detail of left antennomeres IVVI.B, detail of ventral femoral macroseta of right leg I. C, detail of pretarsus; asterisks indicate the lateral processes. D, detail of subcoxal appendage of urosternite I. E, detail of stylus and eversible vesicle of urosternite V.F, detail of stylus and eversible vesicle of urosternite VI. Abbreviations: T, trichobothria." figureDoi="http://doi.org/10.5281/zenodo.11241076" httpUri="https://zenodo.org/record/11241076/files/figure.png" pageId="14" pageNumber="150">Fig. 13A</figureCitation>
); medial and distal antennomeres with whorls of barbed setae combined with whorls of smooth setae (
<figureCitation id="13052A672D00FFACEAF5FF30FF65F9D0" captionStart="Figure 13" captionStartId="16.[113,178,1809,1833]" captionTargetBox="[117,1455,147,1779]" captionTargetId="figure-7@16.[114,1458,144,1782]" captionTargetPageId="16" captionText="Figure 13. Photomicrographs of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber. A, detail of left antennomeres IVVI.B, detail of ventral femoral macroseta of right leg I. C, detail of pretarsus; asterisks indicate the lateral processes. D, detail of subcoxal appendage of urosternite I. E, detail of stylus and eversible vesicle of urosternite V.F, detail of stylus and eversible vesicle of urosternite VI. Abbreviations: T, trichobothria." figureDoi="http://doi.org/10.5281/zenodo.11241076" httpUri="https://zenodo.org/record/11241076/files/figure.png" pageId="18" pageNumber="154">Fig. 13A</figureCitation>
).
</paragraph>
<caption id="DF41666A2D1DFFB1E8A7F88CFDC2FE6B" ID-DOI="http://doi.org/10.5281/zenodo.11241074" ID-Zenodo-Dep="11241074" httpUri="https://zenodo.org/record/11241074/files/figure.png" pageId="15" pageNumber="151" startId="15.[129,194,1836,1860]" targetBox="[220,1382,147,1805]" targetPageId="15" targetType="figure">
<paragraph id="8B8136E22D1DFFB1E8A7F88CFDC2FE6B" blockId="15.[129,1471,1836,1916]" pageId="15" pageNumber="151">
<emphasis id="B94AEAF02D1DFFB1E8A7F88CFF1DFE53" bold="true" box="[129,229,1836,1860]" pageId="15" pageNumber="151">Figure 12.</emphasis>
Camera lucida drawings of holotype of
<taxonomicName id="4C3E4D612D1DFFB1EA7AF88CFC90FE53" authority=", AMNH" authorityName="AMNH" box="[604,872,1836,1860]" class="Diplura" family="Campodeidae" genus="Rostricampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="15" pageNumber="151" phylum="Arthropoda" rank="species" species="engeli">
<emphasis id="B94AEAF02D1DFFB1EA7AF88CFCEDFE53" bold="true" box="[604,789,1836,1860]" italics="true" pageId="15" pageNumber="151">Rostricampa engeli</emphasis>
,
<emphasis id="B94AEAF02D1DFFB1EB39F88DFC90FE53" box="[799,872,1837,1860]" italics="true" pageId="15" pageNumber="151">AMNH</emphasis>
</taxonomicName>
<emphasis id="B94AEAF02D1DFFB1EB4BF88CFC08FE53" box="[877,1008,1836,1860]" italics="true" pageId="15" pageNumber="151">DR-KLCa002</emphasis>
, in Miocene Dominican amber. A, detail of head. B, dorsolateral habitus. C, detail of stylus and eversible vesicle of urosternite II. D, detail of stylus and eversible vesicle of urosternite V. E, detail of stylus and eversible vesicle of urosternite VI.
</paragraph>
</caption>
<caption id="DF41666A2D02FFAEE857F8B1FDD6FE69" ID-DOI="http://doi.org/10.5281/zenodo.11241076" ID-Zenodo-Dep="11241076" httpUri="https://zenodo.org/record/11241076/files/figure.png" pageId="16" pageNumber="152" startId="16.[113,178,1809,1833]" targetBox="[117,1455,147,1779]" targetPageId="16" targetType="figure">
<paragraph id="8B8136E22D02FFAEE857F8B1FDD6FE69" blockId="16.[113,1434,1809,1918]" pageId="16" pageNumber="152">
<emphasis id="B94AEAF02D02FFAEE857F8B1FF2EFE3D" bold="true" box="[113,214,1809,1834]" pageId="16" pageNumber="152">Figure 13.</emphasis>
Photomicrographs of holotype of
<taxonomicName id="4C3E4D612D02FFAEEA3CF8B2FCDEFE3E" authority=", AMNH" authorityName="AMNH" box="[538,806,1809,1834]" class="Diplura" family="Campodeidae" genus="Rostricampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="16" pageNumber="152" phylum="Arthropoda" rank="species" species="engeli">
<emphasis id="B94AEAF02D02FFAEEA3CF8B2FD2CFE3E" bold="true" box="[538,724,1809,1834]" italics="true" pageId="16" pageNumber="152">Rostricampa engeli</emphasis>
,
<emphasis id="B94AEAF02D02FFAEEAFBF8B2FCDEFE3E" box="[733,806,1810,1833]" italics="true" pageId="16" pageNumber="152">AMNH</emphasis>
</taxonomicName>
<emphasis id="B94AEAF02D02FFAEEB0DF8B2FC56FE3D" box="[811,942,1810,1834]" italics="true" pageId="16" pageNumber="152">DR-KLCa002</emphasis>
, in Miocene Dominican amber. A, detail of left antennomeres IVVI. B, detail of ventral femoral macroseta of right leg I. C, detail of pretarsus; asterisks indicate the lateral processes. D, detail of subcoxal appendage of urosternite I. E, detail of stylus and eversible vesicle of urosternite V. F, detail of stylus and eversible vesicle of urosternite VI. Abbreviations: T, trichobothria.
</paragraph>
</caption>
<caption id="DF41666A2D03FFAFE8A7F889FDB9FE6E" ID-DOI="http://doi.org/10.5281/zenodo.11241078" ID-Zenodo-Dep="11241078" httpUri="https://zenodo.org/record/11241078/files/figure.png" pageId="17" pageNumber="153" startId="17.[129,194,1833,1857]" targetBox="[133,1469,150,1802]" targetPageId="17" targetType="figure">
<paragraph id="8B8136E22D03FFAFE8A7F889FA74FE56" blockId="17.[129,1441,1832,1913]" box="[129,1420,1832,1857]" pageId="17" pageNumber="153">
<emphasis id="B94AEAF02D03FFAFE8A7F889FF1DFE56" bold="true" box="[129,229,1833,1857]" pageId="17" pageNumber="153">Figure 14.</emphasis>
Computed tomography scan images of holotype of
<taxonomicName id="4C3E4D612D03FFAFEAEAF889FC21FE57" authority=", AMNH" authorityName="AMNH" box="[716,985,1832,1857]" class="Diplura" family="Campodeidae" genus="Rostricampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="17" pageNumber="153" phylum="Arthropoda" rank="species" species="engeli">
<emphasis id="B94AEAF02D03FFAFEAEAF889FC7EFE57" bold="true" box="[716,902,1832,1857]" italics="true" pageId="17" pageNumber="153">Rostricampa engeli</emphasis>
,
<emphasis id="B94AEAF02D03FFAFEBB6F889FC21FE57" box="[912,985,1833,1856]" italics="true" pageId="17" pageNumber="153">AMNH</emphasis>
</taxonomicName>
<emphasis id="B94AEAF02D03FFAFEBFBF889FB99FE56" box="[989,1121,1833,1857]" italics="true" pageId="17" pageNumber="153">DR-KLCa002</emphasis>
, in Miocene Dominican amber.
</paragraph>
<paragraph id="8B8136E22D03FFAFE8A7F8E5FDB9FE6E" blockId="17.[129,1441,1832,1913]" pageId="17" pageNumber="153">A, lateral habitus of anterior part of body. B, dorsal habitus of anterior part of body. C, head in dorsal view. D, head in ventral view. E, head in lateral view. AB and CE are to the same scale.</paragraph>
</caption>
<paragraph id="8B8136E22D00FFACE857FF49FDECFBC0" blockId="18.[113,763,233,727]" pageId="18" pageNumber="154">
<emphasis id="B94AEAF02D00FFACE857FF49FF47F816" box="[113,191,233,257]" italics="true" pageId="18" pageNumber="154">Thorax:</emphasis>
Length
<quantity id="4CC69B072D00FFACE93AFF49FE84F817" box="[284,380,233,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="18" pageNumber="154" unit="mm" value="1.43">1.43 mm</quantity>
, 0.31 × length of body. Distribution of macrosetae: pronotum with 1 + 1
<emphasis id="B94AEAF02D00FFACE9CDFEA8FE05F837" box="[491,509,264,288]" italics="true" pageId="18" pageNumber="154">la</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D00FFACEA63FEA8FDA0F837" box="[581,600,264,288]" italics="true" pageId="18" pageNumber="154">lp</emphasis>
,
<emphasis id="B94AEAF02D00FFACEA45FEA9FD7AF837" box="[611,642,265,288]" italics="true" pageId="18" pageNumber="154">ma</emphasis>
not visible; mesonotum with 1 + 1
<emphasis id="B94AEAF02D00FFACE941FE88FE7EF828" box="[359,390,296,319]" italics="true" pageId="18" pageNumber="154">ma</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D00FFACE9F6FE87FE1AF828" box="[464,482,295,319]" italics="true" pageId="18" pageNumber="154">la</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D00FFACEA0AFE87FDC7F828" box="[556,575,295,319]" italics="true" pageId="18" pageNumber="154">lp</emphasis>
; metanotum with 1 + 1
<emphasis id="B94AEAF02D00FFACE889FEE7FF3AF848" box="[175,194,327,351]" italics="true" pageId="18" pageNumber="154">lp</emphasis>
,
<emphasis id="B94AEAF02D00FFACE8E8FEE7FF15F849" box="[206,237,327,350]" italics="true" pageId="18" pageNumber="154">ma</emphasis>
not visible; macrosetae very long and thin, barbed on one side along distal half or third; marginal setae longer than clothing setae and thin, with a few thin distal barbs. Legs slightly elongated; metathoracic leg reaching abdominal segment IV; length of metathoracic leg segments: femur
<quantity id="4CC69B072D00FFACEA14FE64FD68F8CC" box="[562,656,452,476]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="18" pageNumber="154" unit="mm" value="0.36">0.36 mm</quantity>
, tibia
<quantity id="4CC69B072D00FFACEAE8FE64FF64F8EC" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="18" pageNumber="154" unit="mm" value="0.37">0.37 mm</quantity>
, tarsus including pretarsus
<quantity id="4CC69B072D00FFACE993FE43FDEDF8EC" box="[437,533,483,507]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="18" pageNumber="154" unit="mm" value="0.29">0.29 mm</quantity>
; femur with one long, thin ventral macroseta, with short barbs along distal half or third (
<figureCitation id="13052A672D00FFACE892FD82FEF0FB2D" box="[180,264,546,570]" captionStart="Figure 13" captionStartId="16.[113,178,1809,1833]" captionTargetBox="[117,1455,147,1779]" captionTargetId="figure-7@16.[114,1458,144,1782]" captionTargetPageId="16" captionText="Figure 13. Photomicrographs of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber. A, detail of left antennomeres IVVI.B, detail of ventral femoral macroseta of right leg I. C, detail of pretarsus; asterisks indicate the lateral processes. D, detail of subcoxal appendage of urosternite I. E, detail of stylus and eversible vesicle of urosternite V.F, detail of stylus and eversible vesicle of urosternite VI. Abbreviations: T, trichobothria." figureDoi="http://doi.org/10.5281/zenodo.11241076" httpUri="https://zenodo.org/record/11241076/files/figure.png" pageId="18" pageNumber="154">Fig. 13B</figureCitation>
); tibia with one long ventral macroseta, and two thin calcars with short barbs distally; femur and tibia without dorsal macrosetae; tarsus with typical setae along ventral side; subapical tarsal setae longer than other tarsal setae; pretarsus with subequal claws, with small lateral crests; pretarsal lateral processes laminar and curved (
<figureCitation id="13052A672D00FFACE98AFD1EFDFBFBC0" box="[428,515,702,727]" captionStart="Figure 13" captionStartId="16.[113,178,1809,1833]" captionTargetBox="[117,1455,147,1779]" captionTargetId="figure-7@16.[114,1458,144,1782]" captionTargetPageId="16" captionText="Figure 13. Photomicrographs of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber. A, detail of left antennomeres IVVI.B, detail of ventral femoral macroseta of right leg I. C, detail of pretarsus; asterisks indicate the lateral processes. D, detail of subcoxal appendage of urosternite I. E, detail of stylus and eversible vesicle of urosternite V.F, detail of stylus and eversible vesicle of urosternite VI. Abbreviations: T, trichobothria." figureDoi="http://doi.org/10.5281/zenodo.11241076" httpUri="https://zenodo.org/record/11241076/files/figure.png" pageId="18" pageNumber="154">Fig. 13C</figureCitation>
).
</paragraph>
<paragraph id="8B8136E22D00FFACE857FD58FE36FC54" blockId="18.[113,763,760,1348]" pageId="18" pageNumber="154">
<emphasis id="B94AEAF02D00FFACE857FD58FF2EFA07" box="[113,214,760,784]" italics="true" pageId="18" pageNumber="154">Abdomen:</emphasis>
Length
<quantity id="4CC69B072D00FFACE91AFD58FE5AFA18" box="[316,418,760,784]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="154" unit="mm" value="2.5">2.50 mm</quantity>
, 0.54 × length of body. Visible macrosetae on tergites: VVII with 2 macrosetae; VIIIIX with 3 + 3
<emphasis id="B94AEAF02D00FFACE8C0FC96FF01FA59" box="[230,249,822,846]" italics="true" pageId="18" pageNumber="154">lp</emphasis>
; X with 2 macrosetae; macrosetae very long and thin, barbed along distal half or third. Pairs of styli and eversible vesicles present from sternites II to VII (
<figureCitation id="13052A672D00FFACEA1FFCD5FD5FFA9A" box="[569,679,885,909]" captionStart="Figure 13" captionStartId="16.[113,178,1809,1833]" captionTargetBox="[117,1455,147,1779]" captionTargetId="figure-7@16.[114,1458,144,1782]" captionTargetPageId="16" captionText="Figure 13. Photomicrographs of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber. A, detail of left antennomeres IVVI.B, detail of ventral femoral macroseta of right leg I. C, detail of pretarsus; asterisks indicate the lateral processes. D, detail of subcoxal appendage of urosternite I. E, detail of stylus and eversible vesicle of urosternite V.F, detail of stylus and eversible vesicle of urosternite VI. Abbreviations: T, trichobothria." figureDoi="http://doi.org/10.5281/zenodo.11241076" httpUri="https://zenodo.org/record/11241076/files/figure.png" pageId="18" pageNumber="154">Fig. 13E, F</figureCitation>
); stylus length
<quantity id="4CC69B072D00FFACE89AFC35FEE5FABB" box="[188,285,917,941]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="18" pageNumber="154" unit="mm" value="0.07">0.07 mm</quantity>
, width
<quantity id="4CC69B072D00FFACE956FC35FE2AFABB" box="[368,466,917,941]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="154" unit="mm" value="0.02">0.02 mm</quantity>
, with fine transverse wrinkling; stylus with smooth apical and subapical macrosetae plus a ventromedial macroseta always bifurcate (in one it is trifurcate); overall, there are six to seven setae on the stylus. On the side of the stylus opposite the eversible vesicle is a pair of long macrosetae, barbedalongdistalhalforthird, visiblefromsternites I to VII; the macroseta closest to the stylus is shortest; all eversible vesicles extruded, ovoid, of variable dimensions. Both cerci complete (non-regenerated), length
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, 0.91 × length of body; with 25 articles, length of basal articles
<quantity id="4CC69B072D00FFACEA69FB0FFD56FDD1" box="[591,686,1199,1223]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="154" unit="mm" value="0.1">0.10 mm</quantity>
, length of medial and distal articles
<quantity id="4CC69B072D00FFACE9ACFB6EFE10FDF2" box="[394,488,1230,1254]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="154" unit="mm" value="0.25">0.25 mm</quantity>
; with two medial whorls of long, thin macrosetae with thin barbs along distal half, accompanied by two whorls of long, thin, smooth setae, plus a distal whorl of thin, short, smooth setae.
</paragraph>
<paragraph id="8B8136E22D00FFACE857FAC5FE8EFCCC" blockId="18.[113,763,1381,1499]" pageId="18" pageNumber="154">
<emphasis id="B94AEAF02D00FFACE857FAC5FE70FC6A" box="[113,392,1381,1405]" italics="true" pageId="18" pageNumber="154">Secondary sexual characters:</emphasis>
Male urosternite I with thick, large, subtrapezoidal subcoxal appendages (
<figureCitation id="13052A672D00FFACEA20FA24FD9CFC8B" box="[518,612,1412,1436]" captionStart="Figure 13" captionStartId="16.[113,178,1809,1833]" captionTargetBox="[117,1455,147,1779]" captionTargetId="figure-7@16.[114,1458,144,1782]" captionTargetPageId="16" captionText="Figure 13. Photomicrographs of holotype of Rostricampa engeli, AMNH DR-KLCa002, in Miocene Dominican amber. A, detail of left antennomeres IVVI.B, detail of ventral femoral macroseta of right leg I. C, detail of pretarsus; asterisks indicate the lateral processes. D, detail of subcoxal appendage of urosternite I. E, detail of stylus and eversible vesicle of urosternite V.F, detail of stylus and eversible vesicle of urosternite VI. Abbreviations: T, trichobothria." figureDoi="http://doi.org/10.5281/zenodo.11241076" httpUri="https://zenodo.org/record/11241076/files/figure.png" pageId="18" pageNumber="154">Fig. 13D</figureCitation>
), length
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, basal width
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, distal width
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, with glandular setae in a large distal area.
</paragraph>
<paragraph id="8B8136E22D00FFACE857FA5CFD6BFEDD" blockId="18.[113,763,1532,1995]" pageId="18" pageNumber="154">
<emphasis id="B94AEAF02D00FFACE857FA5CFE4AFF03" box="[113,434,1532,1556]" italics="true" pageId="18" pageNumber="154">
Remarks:
<taxonomicName id="4C3E4D612D00FFACE8CEFA5DFE4AFF03" box="[232,434,1532,1556]" class="Diplura" family="Campodeidae" genus="Rostricampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="18" pageNumber="154" phylum="Arthropoda" rank="species" species="engeli">Rostricampa engeli</taxonomicName>
</emphasis>
fits within the subfamily
<taxonomicName id="4C3E4D612D00FFACE857F9BCFEF9FF23" authorityName="Conde" authorityYear="1956" box="[113,257,1564,1588]" class="Diplura" family="Campodeidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="18" pageNumber="154" phylum="Arthropoda" rank="subFamily" subFamily="Campodeinae">Campodeinae</taxonomicName>
owing to the presence of a maximum of 3 + 3 macrosetae on the pronotum (
<bibRefCitation id="EFAF4B132D00FFACE9F2F99BFDA6FF44" author="Conde B" box="[468,606,1595,1619]" pageId="18" pageNumber="154" pagination="1 - 202" refId="ref10405" refString="Conde B. Materiaux pour une Monographie des Diploures Campodeides. Memoires du Museum National d'Histoire naturelle, Serie A. Zoologie 1956; 12: 1 - 202." type="journal article" year="1956">Condé 1956</bibRefCitation>
, Paclt 1957). However, the morphology of the new taxon is remarkably different from any other
<taxonomicName id="4C3E4D612D00FFACE943F9DAFE4EFF85" authorityName="Borner" authorityYear="1904" box="[357,438,1658,1682]" class="Diplura" kingdom="Animalia" pageId="18" pageNumber="154" phylum="Arthropoda" rank="class">Diplura</taxonomicName>
. The excellent preservation of the fossil and the use of various techniques for its examination has allowed us to describe the extraordinary morphology of the head (uniquely modified among all
<taxonomicName id="4C3E4D612D00FFACE9CCF978FDC5FFE7" authorityName="Borner" authorityYear="1904" box="[490,573,1752,1776]" class="Diplura" kingdom="Animalia" pageId="18" pageNumber="154" phylum="Arthropoda" rank="class">Diplura</taxonomicName>
), which is characterized by a sclerotized beak protruding from head. Among
<taxonomicName id="4C3E4D612D00FFACE857F8B6FEFAFE39" authorityName="Conde" authorityYear="1956" box="[113,258,1814,1838]" class="Diplura" family="Campodeidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="18" pageNumber="155" phylum="Arthropoda" rank="subFamily" subFamily="Campodeinae">Campodeinae</taxonomicName>
,
<taxonomicName id="4C3E4D612D00FFACE92BF8B7FE92FE39" box="[269,362,1814,1838]" class="Diplura" family="Campodeidae" genus="Rostricampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="18" pageNumber="154" phylum="Arthropoda" rank="species" species="engeli">
<emphasis id="B94AEAF02D00FFACE92BF8B7FE92FE39" box="[269,362,1814,1838]" italics="true" pageId="18" pageNumber="154">Ro. engeli</emphasis>
</taxonomicName>
resembles the genus
<taxonomicName id="4C3E4D612D00FFACEA6EF8B6FD4AFE39" authorityName="Westwood" authorityYear="1842" box="[584,690,1814,1838]" class="Diplura" family="Campodeidae" genus="Campodea" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="18" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAF02D00FFACEA6EF8B6FD4AFE39" box="[584,690,1814,1838]" italics="true" pageId="18" pageNumber="154">Campodea</emphasis>
</taxonomicName>
by the presence of 3 + 3 M (
<emphasis id="B94AEAF02D00FFACE96DF896FE92FE5A" box="[331,362,1846,1869]" italics="true" pageId="18" pageNumber="154">ma</emphasis>
,
<emphasis id="B94AEAF02D00FFACE952F896FE7EFE59" box="[372,390,1846,1870]" italics="true" pageId="18" pageNumber="154">la</emphasis>
,
<emphasis id="B94AEAF02D00FFACE9B7F896FE5CFE59" box="[401,420,1846,1870]" italics="true" pageId="18" pageNumber="154">lp</emphasis>
) on the pro- and mesonotum, the metanotum with
<emphasis id="B94AEAF02D00FFACE902F8F5FECFFE7A" box="[292,311,1877,1901]" italics="true" pageId="18" pageNumber="154">lp</emphasis>
macrosetae, and the presence of non-setose lateral processes of the pretarsus (
<bibRefCitation id="EFAF4B132D00FFACE9EEF8D4FDBFFE9B" author="Conde B" box="[456,583,1908,1932]" pageId="18" pageNumber="154" pagination="1 - 202" refId="ref10405" refString="Conde B. Materiaux pour une Monographie des Diploures Campodeides. Memoires du Museum National d'Histoire naturelle, Serie A. Zoologie 1956; 12: 1 - 202." type="journal article" year="1956">Condé 1956</bibRefCitation>
, Paclt 1957). The specimen is identified as an adult male owing to the presence of the enlarged subcoxal appendages of the urosternite I.
</paragraph>
</subSubSection>
</treatment>
</document>