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<document id="A5F5FA3966D7FB8841F06EFBD17E21DB" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FF8884718CFBFF07DF13FDB0" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes sinuosus Mariano &amp; Flowers (in Mariano, Flowers &amp; Froehlich 2011" docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="22" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="81864E1964BB78A0E40F75CBDD25B01D">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<treatment id="039787A6FF8884718CFBFF07DF13FDB0" ID-DOI="http://doi.org/10.5281/zenodo.3811757" ID-GBIF-Taxon="163420794" ID-Zenodo-Dep="3811757" LSID="urn:lsid:plazi:treatment:039787A6FF8884718CFBFF07DF13FDB0" httpUri="http://treatment.plazi.org/id/039787A6FF8884718CFBFF07DF13FDB0" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection id="C324653BFF8884718CFBFF07DA65FF59" box="[151,347,225,251]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8B8136B0FF8884718CFBFF07DA65FF59" blockId="21.[151,347,225,251]" box="[151,347,225,251]" pageId="21" pageNumber="22">
<heading id="D0C981DCFF8884718CFBFF07DA65FF59" bold="true" box="[151,347,225,251]" fontSize="10" level="2" pageId="21" pageNumber="22" reason="2">
<emphasis id="B94AEAA2FF8884718CFBFF07DA65FF59" bold="true" box="[151,347,225,251]" pageId="21" pageNumber="22">
Group
<taxonomicName id="4C3E4D33FF8884718C9DFF07DA65FF59" ID-CoL="56LL4" authorityName="Mariano &amp; Flowers (in Mariano, Flowers &amp; Froehlich" authorityYear="2011" box="[241,347,225,251]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sinuosus">
<emphasis id="B94AEAA2FF8884718C9DFF07DA65FF59" bold="true" box="[241,347,225,251]" italics="true" pageId="21" pageNumber="22">sinuosus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFF8884718CFBFEC1DF13FDB0" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8B8136B0FF8884718CFBFEC1DF34FE52" blockId="21.[151,1437,295,530]" pageId="21" pageNumber="22">
<emphasis id="B94AEAA2FF8884718CFBFEC1DA28FEE0" bold="true" box="[151,278,295,322]" pageId="21" pageNumber="22">Definition.</emphasis>
Penis lobes small and extremely simplified, without apical widening; telopenes unusually short, unrolled and bent ventrally-proximally (
<figureCitation id="13052A35FF8884718E52FEADD9ABFEC7" box="[574,661,331,357]" captionStart="FIGURES 2534" captionStartId="27.[151,264,188,213]" captionTargetId="figure@26.[151,1436,181,1962]" captionTargetPageId="26" captionText="FIGURES 2534. Thraulodes ludmilae sp. n.: 25, 27, male imago; 26, apex of fore wing; 2830, fore, middle and hind legs of male imago; 31, subimaginal exuviae of hind leg; 32, costal margin of fore wing (arrows show proximal costal cross veins); 33, genitalia of male imago; 34, left half of subimaginal exuviae of mesonotum (2526, 2834, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736905" httpUri="https://zenodo.org/record/3736905/files/figure.png" pageId="21" pageNumber="22">Figs 33</figureCitation>
,
<figureCitation id="13052A35FF8884718ECEFEADD9D5FEC7" box="[674,747,331,357]" captionStart="FIGURES 3541" captionStartId="28.[151,264,1854,1879]" captionTargetBox="[151,1436,185,1773]" captionTargetId="figure@28.[151,1436,181,1782]" captionTargetPageId="28" captionText="FIGURES 3541. Thraulodes ludmilae sp. n.: 35, genitalia of male imago in ventral view; 36, subimaginal exuviae of styliger and gonostyli at the same magnification; 37, larval exuviae of protogonostyli and protopenis at the same magnification; 38, imaginal styliger and penis enlarged; 39, subimaginal exuviae of penis at the same magnification; 40, genitalia of male imago in lateral view; 41, hind wing (3539, 41, holotype). Abbreviations: bl, basal lobe; gr, groove of telopenis; dp, dorsal projection of styliger; m-v.r, medio-ventral ridge (recurved flap); tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736907" httpUri="https://zenodo.org/record/3736907/files/figure.png" pageId="21" pageNumber="22">38, 40</figureCitation>
,
<figureCitation id="13052A35FF8884718E9AFEACD801FEC7" box="[758,831,330,357]" captionStart="FIGURES 5259" captionStartId="30.[151,264,1809,1834]" captionTargetBox="[151,1436,166,1789]" captionTargetId="figure@30.[151,1436,166,1789]" captionTargetPageId="30" captionText="FIGURES 5259. Thraulodes sinuosus. 5253, male imago in dorsal and ventral view; 54, genitalia of male imago with focus on gonostyli; 55, the same, with focus on penis; 56, right half of subimaginal exuviae of mesonotum; 5758, female imago in ventral and lateral view; 59, pterostigma of fore wing." figureDoi="http://doi.org/10.5281/zenodo.3736911" httpUri="https://zenodo.org/record/3736911/files/figure.png" pageId="21" pageNumber="22">5455</figureCitation>
,
<figureCitation id="13052A35FF8884718F27FEADD855FEC4" box="[843,875,331,358]" captionStart="FIGURES 6063" captionStartId="31.[151,264,1219,1244]" captionTargetBox="[154,1432,171,1193]" captionTargetId="figure@31.[151,1437,166,1199]" captionTargetPageId="31" captionText="FIGURES 6063. Thraulodes sinuosus. 60, genitalia of male imago in ventral view; 61, their subimaginal exuviae at the same magnification; 62, larval exuviae of protogonostyli and protopenis at the same magnification; 63, hind wing." figureDoi="http://doi.org/10.5281/zenodo.3736913" httpUri="https://zenodo.org/record/3736913/files/figure.png" pageId="21" pageNumber="22">60</figureCitation>
). In imago of both sexes, ventral side of pterothorax mostly ocher, with transverse orange band crossing anterior part of mesothoracic episterna and basisternum; abdominal sternum I brown, contrastingly darker than thoracic sterna and remainder abdominal sterna (
<figureCitation id="13052A35FF888471897BFE76DE50FE08" box="[1303,1390,400,426]" captionStart="FIGURES 2534" captionStartId="27.[151,264,188,213]" captionTargetId="figure@26.[151,1436,181,1962]" captionTargetPageId="26" captionText="FIGURES 2534. Thraulodes ludmilae sp. n.: 25, 27, male imago; 26, apex of fore wing; 2830, fore, middle and hind legs of male imago; 31, subimaginal exuviae of hind leg; 32, costal margin of fore wing (arrows show proximal costal cross veins); 33, genitalia of male imago; 34, left half of subimaginal exuviae of mesonotum (2526, 2834, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736905" httpUri="https://zenodo.org/record/3736905/files/figure.png" pageId="21" pageNumber="22">Figs 27</figureCitation>
,
<figureCitation id="13052A35FF8884718915FE69DB89FE6F" captionStart="FIGURES 5259" captionStartId="30.[151,264,1809,1834]" captionTargetBox="[151,1436,166,1789]" captionTargetId="figure@30.[151,1436,166,1789]" captionTargetPageId="30" captionText="FIGURES 5259. Thraulodes sinuosus. 5253, male imago in dorsal and ventral view; 54, genitalia of male imago with focus on gonostyli; 55, the same, with focus on penis; 56, right half of subimaginal exuviae of mesonotum; 5758, female imago in ventral and lateral view; 59, pterostigma of fore wing." figureDoi="http://doi.org/10.5281/zenodo.3736911" httpUri="https://zenodo.org/record/3736911/files/figure.png" pageId="21" pageNumber="22">53, 57</figureCitation>
); femora with one multicolored apical band; in male, abdominal terga IIVI whitish, terga VIIX orange (
<figureCitation id="13052A35FF8884718907FE55DB8BFE52" captionStart="FIGURES 2534" captionStartId="27.[151,264,188,213]" captionTargetId="figure@26.[151,1436,181,1962]" captionTargetPageId="26" captionText="FIGURES 2534. Thraulodes ludmilae sp. n.: 25, 27, male imago; 26, apex of fore wing; 2830, fore, middle and hind legs of male imago; 31, subimaginal exuviae of hind leg; 32, costal margin of fore wing (arrows show proximal costal cross veins); 33, genitalia of male imago; 34, left half of subimaginal exuviae of mesonotum (2526, 2834, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736905" httpUri="https://zenodo.org/record/3736905/files/figure.png" pageId="21" pageNumber="22">Figs 25</figureCitation>
,
<figureCitation id="13052A35FF8884718CADFE33DBE0FE52" box="[193,222,469,496]" captionStart="FIGURES 5259" captionStartId="30.[151,264,1809,1834]" captionTargetBox="[151,1436,166,1789]" captionTargetId="figure@30.[151,1436,166,1789]" captionTargetPageId="30" captionText="FIGURES 5259. Thraulodes sinuosus. 5253, male imago in dorsal and ventral view; 54, genitalia of male imago with focus on gonostyli; 55, the same, with focus on penis; 56, right half of subimaginal exuviae of mesonotum; 5758, female imago in ventral and lateral view; 59, pterostigma of fore wing." figureDoi="http://doi.org/10.5281/zenodo.3736911" httpUri="https://zenodo.org/record/3736911/files/figure.png" pageId="21" pageNumber="22">52</figureCitation>
) (the same coloration of legs and male abdomen in many other species).
</paragraph>
<paragraph id="8B8136B0FF8884718CABFE11DF13FDB0" blockId="21.[151,1437,295,530]" box="[199,1069,503,530]" pageId="21" pageNumber="22">
<emphasis id="B94AEAA2FF8884718CABFE11DA59FDB0" bold="true" box="[199,359,503,530]" pageId="21" pageNumber="22">Composition.</emphasis>
<taxonomicName id="4C3E4D33FF8884718D02FE1EDAC5FDB0" box="[366,507,504,530]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
<emphasis id="B94AEAA2FF8884718D02FE1EDAC5FDB0" box="[366,507,504,530]" italics="true" pageId="21" pageNumber="22">Th. ludmilae</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FF8884718E6EFE1ED97CFDB0" bold="true" box="[514,578,504,530]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="A27957D9FF8884718E6EFE1ED97CFDB0" box="[514,578,504,530]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3E4D33FF8884718E14FE1EDF17FDB0" authority="Mariano &amp; Flowers 2011" authorityName="Mariano &amp; Flowers (in Mariano, Flowers &amp; Froehlich" authorityYear="2011" box="[632,1065,504,530]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sinuosus">
<emphasis id="B94AEAA2FF8884718E14FE1ED83DFDB0" box="[632,771,504,530]" italics="true" pageId="21" pageNumber="22">Th. sinuosus</emphasis>
<bibRefCitation id="EFAF4B41FF8884718F67FE1EDF17FDB0" author="Mariano, R. &amp; Flowers, R. W. &amp; Froehlich, C. G." box="[779,1065,504,530]" pageId="21" pageNumber="22" pagination="387 - 393" refId="ref71295" refString="Mariano, R., Flowers, R. W. &amp; Froehlich, C. G. (2011) Four new species, a new synonymy and new record for Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae). Annales de Limnologie, 47, 387 - 393. https: // doi. org / 10.1051 / limn / 2011044" type="journal article" year="2011">Mariano &amp; Flowers 2011</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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