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<document id="CEAACC0098E3B8415A04F497B41E60CA" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E59FFD494FD4D2CED75ED27" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) suensoni Kataev 1997" docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="21" updateTime="1698562432667" updateUser="plazi">
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<mods:title id="D5F0AD0D642B6C217699093A7A545E81">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="C6D5361A79AA052243A2D42F74DE8031">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="852D42377EEC04020632FCE770B735EA">Liang, Hongbin</mods:namePart>
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<treatment id="039787812E59FFD494FD4D2CED75ED27" ID-DOI="http://doi.org/10.5281/zenodo.6114272" ID-GBIF-Taxon="127671646" ID-Zenodo-Dep="6114272" LSID="urn:lsid:plazi:treatment:039787812E59FFD494FD4D2CED75ED27" httpUri="http://treatment.plazi.org/id/039787812E59FFD494FD4D2CED75ED27" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<subSubSection id="C324651C2E59FFD594FD4D2CE878E858" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8B8136972E59FFD594FD4D2CEA27E83B" blockId="20.[151,768,1799,1858]" box="[151,768,1799,1825]" pageId="20" pageNumber="21">
<heading id="D0C981FB2E59FFD594FD4D2CEA27E83B" bold="true" box="[151,768,1799,1825]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<taxonomicName id="4C3E4D142E59FFD594FD4D2CEA27E83B" ID-CoL="6LDVB" authority="Kataev, 1997" authorityName="Kataev" authorityYear="1997" box="[151,768,1799,1825]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="suensoni" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E59FFD594FD4D2CEA27E83B" bold="true" box="[151,768,1799,1825]" pageId="20" pageNumber="21">
<emphasis id="B94AEA852E59FFD594FD4D2CE82EE83B" bold="true" box="[151,265,1799,1825]" italics="true" pageId="20" pageNumber="21">Harpalus</emphasis>
(
<emphasis id="B94AEA852E59FFD595704D2CE8FDE83B" bold="true" box="[282,474,1799,1825]" italics="true" pageId="20" pageNumber="21">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E59FFD595814D2CEB70E83B" bold="true" box="[491,599,1799,1825]" italics="true" pageId="20" pageNumber="21">suensoni</emphasis>
<bibRefCitation id="EFAF4B662E59FFD596344D2CEA27E83B" author="Kataev" box="[606,768,1799,1825]" pageId="20" pageNumber="21" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev, 1997</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E59FFD594FD4D02E878E858" blockId="20.[151,768,1799,1858]" box="[151,351,1833,1858]" pageId="20" pageNumber="21">(Figs 67, 4243)</paragraph>
</subSubSection>
<subSubSection id="C324651C2E59FFD594FD4D59EDAFE892" box="[151,1160,1906,1928]" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="8B8136972E59FFD594FD4D59EDAFE892" blockId="20.[151,1160,1906,1928]" box="[151,1160,1906,1928]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="AB2E11B92E59FFD594FD4D59EDAFE892" box="[151,1160,1906,1928]" pageId="20" pageNumber="21">
<taxonomicName id="4C3E4D142E59FFD594FD4D59EB30E892" ID-CoL="6LDVB" authority="Kataev, 1997: 131" authorityName="Kataev" authorityPageNumber="131" authorityYear="1997" box="[151,535,1906,1928]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="suensoni">
<emphasis id="B94AEA852E59FFD594FD4D59E871E892" box="[151,342,1906,1928]" italics="true" pageId="20" pageNumber="21">Harpalus suensoni</emphasis>
<treatmentCitation id="0A9F10862E59FFD595314D58EB30E892" author="Kataev" box="[347,535,1906,1928]" page="131" pageId="20" pageNumber="21" year="1997">
<bibRefCitation id="EFAF4B662E59FFD595314D58E8C3E892" author="Kataev" box="[347,484,1906,1928]" pageId="20" pageNumber="21" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev, 1997</bibRefCitation>
: 131
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E59FFD5964B4D58EB74E892" box="[545,595,1907,1928]" pageId="20" pageNumber="21">Type</typeStatus>
locality: Soochow (
<geoCoordinate id="EE0A50502E59FFD597754D59EA54E892" box="[799,883,1906,1928]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="31.35">31°21'N</geoCoordinate>
<geoCoordinate id="EE0A50502E59FFD597134D59EAFDE892" box="[889,986,1906,1928]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="120.666664">120°40'E</geoCoordinate>
), Jiangsu,
<collectingCountry id="F32976072E59FFD5902E4D59EDA3E892" box="[1092,1156,1906,1928]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E59FFD494FD4D9FEB1FEE06" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8B8136972E59FFD594FD4D9FECA2E8EB" blockId="20.[151,1437,1972,2033]" pageId="20" pageNumber="21">
<emphasis id="B94AEA852E59FFD594FD4D9FE819E8D7" bold="true" box="[151,318,1972,1997]" pageId="20" pageNumber="21">
<typeStatus id="548588352E59FFD594FD4D9FE9F5E8D7" box="[151,210,1972,1997]" pageId="20" pageNumber="21">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E59FFD595264D9FE89EE8D7" box="[332,441,1972,1997]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
: ♂, labeled &quot;
<collectingCountry id="F32976072E59FFD596224D9FEBB7E8D7" box="[584,656,1972,1997]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
, Jiangsu, Soochow,
<geoCoordinate id="EE0A50502E59FFD597194D9EEAE8E8D7" box="[883,975,1973,1997]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="31.35">31°21'N</geoCoordinate>
<geoCoordinate id="EE0A50502E59FFD597BD4D9EED66E8D7" box="[983,1089,1973,1997]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="120.666664">120°40'E</geoCoordinate>
,
<date id="FF8010572E59FFD590264D9FEDE9E8D7" box="[1100,1230,1972,1997]" pageId="20" pageNumber="21" value="1921-09-25">25.IX.1921</date>
, coll. E. Suenson&quot; (
<collectionCode id="ED2FAE522E59FFD594F54DF2E9D1E8EB" box="[159,246,2009,2033]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="21">ZMUC</collectionCode>
).
<typeStatus id="548588352E59FFD595624DF2E85EE8EB" box="[264,377,2009,2033]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D38FD1E2E59FFD595EE4DF2E899E8EA" box="[388,446,2009,2033]" pageId="20" pageNumber="21" type="generic">13 ♂</specimenCount>
, 10 ♀ from
<collectingCountry id="F32976072E59FFD5962C4DF3EBAEE8EB" box="[582,649,2008,2033]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
(Jiangsu and Zhejiang) (
<collectionCode id="ED2FAE522E59FFD597CA4DF2EAD0E8EB" box="[928,1015,2009,2033]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="21">ZMUC</collectionCode>
); and
<specimenCount id="9D38FD1E2E59FFD590504DF2ED43E8EA" box="[1082,1124,2009,2033]" pageId="20" pageNumber="21" type="generic">1 ♂</specimenCount>
from
<collectingCountry id="F32976072E59FFD590C24DF3EC12E8EA" box="[1192,1333,2008,2033]" name="South Korea" pageId="20" pageNumber="21">South Korea</collectingCountry>
(
<collectionCode id="ED2FAE522E59FFD5912E4DF2EC5FE8EB" box="[1348,1400,2009,2033]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="20" pageNumber="21">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E58FFD494AD4ABCEB1FEE06" blockId="21.[151,1437,151,573]" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD494AD4ABCEB0CEFAA" bold="true" box="[199,555,151,176]" pageId="21" pageNumber="22">Additional material examined</emphasis>
.
<specimenCount id="9D38FD1E2E58FFD496534ABCEBF4EFAA" box="[569,723,151,176]" pageId="21" pageNumber="22" type="generic">36 specimens</specimenCount>
, including the following ones not published before.
<collectingCountry id="F32976072E58FFD4917F4ABCEC79EFAA" box="[1301,1374,151,176]" name="China" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD4917F4ABCEC79EFAA" bold="true" box="[1301,1374,151,176]" pageId="21" pageNumber="22">China</emphasis>
</collectingCountry>
.
<specimenCount id="9D38FD1E2E58FFD491064AB3ECB0EFB5" box="[1388,1431,152,176]" pageId="21" pageNumber="22" type="generic">3 ♂</specimenCount>
, 1 ♀, &quot;
<emphasis id="B94AEA852E58FFD494844A97E844EFCE" box="[238,355,188,212]" italics="true" pageId="21" pageNumber="22">crenulatus</emphasis>
Chaudoir [this name has never been published],
<collectingCountry id="F32976072E58FFD497BF4A97ED3FEFCF" box="[981,1048,188,213]" name="China" pageId="21" pageNumber="22">Chine</collectingCountry>
boreale, S. Stevens&quot; (
<collectionCode id="ED2FAE522E58FFD4915F4A97ECB6EFCE" LSID="urn:lsid:biocol.org:col:34988" box="[1333,1425,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="22">MNHN</collectionCode>
). ZHEJIANG: 1 ♀, &quot;Chekiang [= Zhejiang], Hangcheou, A. Pichon 1925&quot; (
<collectionCode id="ED2FAE522E58FFD4978E4ACBED67EFE2" LSID="urn:lsid:biocol.org:col:34988" box="[996,1088,224,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="22">MNHN</collectionCode>
). YUNNAN:
<specimenCount id="9D38FD1E2E58FFD490B14ACBEC2FEFED" box="[1243,1288,224,248]" pageId="21" pageNumber="22" type="generic">1 ♂</specimenCount>
, Dengchuan,
<date id="FF8010572E58FFD494FD4B2FE80CEE07" box="[151,299,260,285]" pageId="21" pageNumber="22" value="1937-08-25">25.VIII.1937</date>
, C.W. Wang leg. (
<collectionCode id="ED2FAE522E58FFD4959D4B2FEB0BEE06" box="[503,556,260,284]" pageId="21" pageNumber="22">IOZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E58FFD494AD4B0CEAA1EED5" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8B8136972E58FFD494AD4B0CEAA1EED5" blockId="21.[151,1437,151,573]" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD494AD4B0CE81DEE5A" bold="true" box="[199,314,295,320]" pageId="21" pageNumber="22">Diagnosis</emphasis>
. This species is very similar in external characters, including protibial apical spur dentate at margins (Fig. 6), and male genitalia to
<taxonomicName id="4C3E4D142E58FFD4958C4B66EB72EE7E" box="[486,597,332,356]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E58FFD4958C4B66EB72EE7E" box="[486,597,332,356]" italics="true" pageId="21" pageNumber="22">H. tridens</emphasis>
</taxonomicName>
, but is distinguished from the latter by having elytral striae finely punctate, elytral disc without microsculpture, only two lateral intervals densely punctate apically, intervals 6 and 7 with few punctures before apex, subapical sinuation of elytra markedly deeper (Fig. 7), and internal sac of aedeagus (
<figureCitation id="13052A122E58FFD491004BBFE9F8EECA" captionStart="FIGURES 40 43" captionStartId="20.[151,263,1670,1693]" captionTargetBox="[319,1267,654,1642]" captionTargetId="figure@20.[309,1278,645,1649]" captionTargetPageId="20" captionText="FIGURES 40 43. Harpalus, median lobe of aedeagus. 40 41, H. tridens (Shaanxi); 42 43, H. suensoni (Zhejiang). 40, 42, dorsal view; 41, 43, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287894/files/figure.png" pageId="21" pageNumber="22">Figs 4243</figureCitation>
) with a large medial spiny patch. Body length 9.713.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E58FFD494AD4BF7ED75ED27" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8B8136972E58FFD494AD4BF7EB54ED0D" blockId="21.[151,1437,151,573]" pageId="21" pageNumber="22">
<emphasis id="B94AEA852E58FFD494AD4BF7E87EEEEF" bold="true" box="[199,345,476,501]" pageId="21" pageNumber="22">Distribution</emphasis>
.
<collectingCountry id="F32976072E58FFD495074BF7E892EEEF" box="[365,437,476,501]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
: Shaanxi, Sichuan, Jiangsu, Zhejiang, Hubei (
<bibRefCitation id="EFAF4B662E58FFD497B14BF7ED98EEEE" author="Kataev" box="[987,1215,476,500]" pageId="21" pageNumber="22" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E58FFD490534BF6ED5FEEEE" box="[1081,1144,476,500]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2003
</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E58FFD490BB4BF6EC78EEEF" author="Wrase" box="[1233,1375,476,501]" pageId="21" pageNumber="22" refString="Wrase, D. W. (2005) Nomenclatorial, taxonomic and faunistic notes on some Palaearctic genera and species of ground-beetles (Coleoptera, Carabidae: Apotomini, Chlaeniini, Cyclosomini, Harpalini, Lebiini, Licinini, Platynini, Siagonini, Sphodrini). Linzer biologische Beitrage, 37 (1), 815 - 874." type="journal article" year="2005">Wrase 2005</bibRefCitation>
) and Yunnan. It is the first record from Yunnan.
</paragraph>
<paragraph id="8B8136972E58FFD494AD480FED75ED27" blockId="21.[151,1437,151,573]" box="[199,1106,548,573]" pageId="21" pageNumber="22">
The species is also known from
<collectingCountry id="F32976072E58FFD4965A480FEB9BED26" box="[560,700,548,573]" name="South Korea" pageId="21" pageNumber="22">South Korea</collectingCountry>
(
<bibRefCitation id="EFAF4B662E58FFD496A1480FEA79ED26" author="Kataev" box="[715,862,548,572]" pageId="21" pageNumber="22" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E58FFD49700480FED61ED27" author="Paik" box="[874,1094,548,573]" pageId="21" pageNumber="22" refString="Paik, J. - Ch. &amp; Moon, Ch. - S. (2005) Some ground bettles (Coleoptera, Carabidae) from Korea (15). Korean Journal of Soil Zoology, 10 (1 - 2), 42 - 54." type="journal article" year="2005">Paik &amp; Moon 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>