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<document id="3084843EB3F41E5A712072A64CEF6038" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E52FFE094FD4FC2E9F0EE07" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) singularis Tschitscherine 1906" docType="treatment" docVersion="7" lastPageNumber="33" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="32" updateTime="1698562432667" updateUser="plazi">
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<mods:title id="ACAB1115E2415CA224A5BCD58AA4145F">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="FDAE208AF6C707A6ABE0C539BCF47D78">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="C493FD0E79178F6C6365FB5820D732F5">Liang, Hongbin</mods:namePart>
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<mods:date id="5257D3A84D1B6166AB4659E1365DD5BD">2015</mods:date>
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<treatment id="039787812E52FFE094FD4FC2E9F0EE07" ID-DOI="http://doi.org/10.5281/zenodo.6114296" ID-GBIF-Taxon="127671649" ID-Zenodo-Dep="6114296" LSID="urn:lsid:plazi:treatment:039787812E52FFE094FD4FC2E9F0EE07" httpUri="http://treatment.plazi.org/id/039787812E52FFE094FD4FC2E9F0EE07" lastPageId="33" lastPageNumber="33" pageId="31" pageNumber="32">
<subSubSection id="C324651C2E52FFDE94FD4FC2E800E93E" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8B8136972E52FFDE94FD4FC2EA4FE919" blockId="31.[151,872,1513,1573]" box="[151,872,1513,1539]" pageId="31" pageNumber="32">
<heading id="D0C981FB2E52FFDE94FD4FC2EA4FE919" bold="true" box="[151,872,1513,1539]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<taxonomicName id="4C3E4D142E52FFDE94FD4FC2EA4FE919" ID-CoL="6LDVF" authority="Tschitscherine, 1906" authorityName="Tschitscherine" authorityYear="1906" box="[151,872,1513,1539]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="singularis" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E52FFDE94FD4FC2EA4FE919" bold="true" box="[151,872,1513,1539]" pageId="31" pageNumber="32">
<emphasis id="B94AEA852E52FFDE94FD4FC2E82EE919" bold="true" box="[151,265,1513,1539]" italics="true" pageId="31" pageNumber="32">Harpalus</emphasis>
(
<emphasis id="B94AEA852E52FFDE95704FC2E8FDE919" bold="true" box="[282,474,1513,1539]" italics="true" pageId="31" pageNumber="32">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E52FFDE95814FC2EB43E919" bold="true" box="[491,612,1513,1539]" italics="true" pageId="31" pageNumber="32">singularis</emphasis>
<bibRefCitation id="EFAF4B662E52FFDE96014FC2EA4FE919" author="Tschitscherine" box="[619,872,1513,1539]" pageId="31" pageNumber="32" refString="Tschitscherine, T. (1906) Notes detachees sur les Harpalini de l'Asie orientale. Horae Societatis Entomologicae Rossicae, 37 (3 / 4), 247 - 292." type="journal article" year="1906">Tschitschérine, 1906</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E52FFDE94FD4C27E800E93E" blockId="31.[151,872,1513,1573]" box="[151,295,1548,1573]" pageId="31" pageNumber="32">
(
<figureCitation id="13052A122E52FFDE94F54C27E838E93E" box="[159,287,1548,1573]" captionStart="FIGURES 82 85" captionStartId="33.[151,263,1137,1160]" captionTargetBox="[259,1328,342,1109]" captionTargetId="figure@33.[252,1334,339,1116]" captionTargetPageId="33" captionText="FIGURES 82 85. Harpalus, median lobe of aedeagus. 82 83, H. singularis (Sichuan, lectotype); 84 85, H. azumai (Shima Island). 82, 84, view from the left side; 83, 85, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287901/files/figure.png" pageId="31" pageNumber="32">Figs 8283</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E52FFDE94FD4C7FE8AFE9DC" pageId="31" pageNumber="32" type="reference_group">
<paragraph id="8B8136972E52FFDE94FD4C7FECBBE9BD" blockId="31.[151,1436,1620,1734]" pageId="31" pageNumber="32">
<taxonomicName id="4C3E4D142E52FFDE94FD4C7FE892E973" ID-CoL="6LDV8" box="[151,437,1620,1642]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tridens" subGenus="Ophonus">
<emphasis id="B94AEA852E52FFDE94FD4C7FE9D0E973" box="[151,247,1620,1641]" italics="true" pageId="31" pageNumber="32">Harpalus</emphasis>
(
<emphasis id="B94AEA852E52FFDE956F4C7FE846E973" box="[261,353,1620,1641]" italics="true" pageId="31" pageNumber="32">Ophonus</emphasis>
)
<emphasis id="B94AEA852E52FFDE95054C7FE892E973" box="[367,437,1620,1641]" italics="true" pageId="31" pageNumber="32">tridens</emphasis>
</taxonomicName>
<taxonomicName id="4C3E4D142E52FFDE95D64C7EEA7CE970" authority="Tschitscherine, 1906: 250" authorityName="Tschitscherine" authorityPageNumber="250" authorityYear="1906" box="[444,859,1620,1642]" pageId="31" pageNumber="32" rank="variety" variety="singularis">
var.
<emphasis id="B94AEA852E52FFDE958D4C7FEB6BE973" box="[487,588,1620,1641]" italics="true" pageId="31" pageNumber="32">singularis</emphasis>
<bibRefCitation id="EFAF4B662E52FFDE96394C7FEA0FE970" author="Tschitscherine" box="[595,808,1620,1642]" pageId="31" pageNumber="32" refString="Tschitscherine, T. (1906) Notes detachees sur les Harpalini de l'Asie orientale. Horae Societatis Entomologicae Rossicae, 37 (3 / 4), 247 - 292." type="journal article" year="1906">Tschitschérine, 1906</bibRefCitation>
: 250
</taxonomicName>
.
<typeStatus id="548588352E52FFDE970E4C7EEAB1E970" box="[868,918,1621,1642]" pageId="31" pageNumber="32">Type</typeStatus>
locality: &quot;vallee du Ja-ho&quot;, Sichuan,
<collectingCountry id="F32976072E52FFDE91654C7FEC68E970" box="[1295,1359,1620,1642]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
.
<taxonomicName id="4C3E4D142E52FFDE94FD4C58EBD4E992" ID-CoL="6LDNZ" authority="Schauberger, 1930: 173" authorityName="Schauberger" authorityPageNumber="173" authorityYear="1930" box="[151,755,1650,1672]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="szetschuanus" subGenus="Pardileus">
<emphasis id="B94AEA852E52FFDE94FD4C58E9D0E992" box="[151,247,1651,1672]" italics="true" pageId="31" pageNumber="32">Harpalus</emphasis>
(
<emphasis id="B94AEA852E52FFDE956F4C58E840E992" box="[261,359,1651,1672]" italics="true" pageId="31" pageNumber="32">Pardileus</emphasis>
)
<emphasis id="B94AEA852E52FFDE951F4C58E8DEE992" box="[373,505,1651,1672]" italics="true" pageId="31" pageNumber="32">szetschuanus</emphasis>
<bibRefCitation id="EFAF4B662E52FFDE966A4C59EBE7E992" author="Schauberger" box="[512,704,1650,1672]" pageId="31" pageNumber="32" refString="Schauberger, E. (1930) Zur Kenntnis der palaarktischen Harpalinen (VII. Beitrag). Coleopterologisches Centralblatt, 4, 169 - 218." type="journal article" year="1930">Schauberger, 1930</bibRefCitation>
: 173
</taxonomicName>
.
<typeStatus id="548588352E52FFDE96964C58EA09E992" box="[764,814,1651,1672]" pageId="31" pageNumber="32">Type</typeStatus>
locality: &quot;Kwanshien&quot;, Sichuan,
<collectingCountry id="F32976072E52FFDE90154C59ED98E992" box="[1151,1215,1650,1672]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
.
<taxonomicName id="4C3E4D142E52FFDE94FD4CBAEB48E9BD" ID-CoL="8LFZW" authority="Huang, 1993: 451" authorityName="Huang" authorityPageNumber="451" authorityYear="1993" box="[151,623,1681,1703]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="chengjiangensis">
<emphasis id="B94AEA852E52FFDE94FD4CBAE883E9BC" box="[151,420,1681,1702]" italics="true" pageId="31" pageNumber="32">Harpalus chengjiangensis</emphasis>
<bibRefCitation id="EFAF4B662E52FFDE95C44CB9EB1EE9BD" author="Huang" box="[430,569,1681,1703]" pageId="31" pageNumber="32" refString="Huang, T. (1993) Three new species of the genus Harpalus from China (Coleoptera: Carabidae). Acta Zootaxonomica Sinica, 18 (4), 451 - 455." type="journal article" year="1993">Huang, 1993</bibRefCitation>
: 451
</taxonomicName>
, 454,
<emphasis id="B94AEA852E52FFDE96DE4CBAEBD9E9BC" bold="true" box="[692,766,1681,1702]" pageId="31" pageNumber="32">syn. n.</emphasis>
<typeStatus id="548588352E52FFDE97624CB9EA1DE9BD" box="[776,826,1682,1703]" pageId="31" pageNumber="32">Type</typeStatus>
locality: Chengjiang, Beibei (
<geoCoordinate id="EE0A50502E52FFDE90164CBAEDF7E9BD" box="[1148,1232,1681,1703]" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="925" value="29.85">29°51'N</geoCoordinate>
<geoCoordinate id="EE0A50502E52FFDE90B14CBAEC1CE9BD" box="[1243,1339,1681,1703]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="106.45">106°27'E</geoCoordinate>
),
<quantity id="4CC69B722E52FFDE91384CBAECBFE9BD" box="[1362,1432,1681,1703]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="31" pageNumber="32" unit="m" value="260.0">260 m</quantity>
,
</paragraph>
<paragraph id="8B8136972E52FFDE94AD4C9BE8AFE9DC" blockId="31.[151,1436,1620,1734]" box="[199,392,1712,1734]" pageId="31" pageNumber="32">
Chongqing,
<collectingCountry id="F32976072E52FFDE952E4C9BE8A3E9DC" box="[324,388,1712,1734]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E52FFE194FD4CD9ED59ECD2" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8B8136972E52FFDE94FD4CD9EC7BE835" blockId="31.[151,1436,1778,2019]" pageId="31" pageNumber="32">
<emphasis id="B94AEA852E52FFDE94FD4CD9E866E811" bold="true" box="[151,321,1778,1803]" pageId="31" pageNumber="32">
<typeStatus id="548588352E52FFDE94FD4CD9E9F5E811" box="[151,210,1778,1803]" pageId="31" pageNumber="32">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E52FFDE95394CD8E8E4E811" box="[339,451,1779,1803]" pageId="31" pageNumber="32" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C3E4D142E52FFDE959A4CD8EB43E811" box="[496,612,1779,1803]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E52FFDE959A4CD8EB43E811" box="[496,612,1779,1803]" italics="true" pageId="31" pageNumber="32">H. tridens</emphasis>
</taxonomicName>
<taxonomicName id="4C3E4D142E52FFDE96054CD8EA4DE811" authority="Tschit." authorityName="Tschit." box="[623,874,1778,1803]" pageId="31" pageNumber="32" rank="variety" variety="singularis">
var.
<emphasis id="B94AEA852E52FFDE96C94CD8EA34E811" box="[675,787,1779,1803]" italics="true" pageId="31" pageNumber="32">singularis</emphasis>
Tschit.
</taxonomicName>
[designated by
<bibRefCitation id="EFAF4B662E52FFDE904D4CD8EC29E811" author="Kataev" box="[1063,1294,1779,1803]" pageId="31" pageNumber="32" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev (1997: 134)</bibRefCitation>
]: ♂, labeled &quot;
<taxonomicName id="4C3E4D142E52FFDE94C84D3DE9D7E834" box="[162,240,1814,1838]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B94AEA852E52FFDE94C84D3DE9D7E834" box="[162,240,1814,1838]" italics="true" pageId="31" pageNumber="32">tridens</emphasis>
</taxonomicName>
<taxonomicName id="4C3E4D142E52FFDE949D4D3CE8B0E834" box="[247,407,1814,1838]" pageId="31" pageNumber="32" rank="variety" variety="singularis">
var.
<emphasis id="B94AEA852E52FFDE954D4D3DE8B0E834" box="[295,407,1814,1838]" italics="true" pageId="31" pageNumber="32">singularis</emphasis>
</taxonomicName>
m., Typ, Tschitscherin det.&quot;, &quot;Val. Ya R.,
<date id="FF8010572E52FFDE97064D3DED33E835" box="[876,1044,1814,1839]" pageId="31" pageNumber="32" value="1993-03-29" valueMax="1993-04-05" valueMin="1993-03-29">29.III.5.IV.93</date>
, Potanin&quot; [in Cyrillic] (
<collectionCode id="ED2FAE522E52FFDE91764D3CEC77E835" box="[1308,1360,1815,1839]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="31" pageNumber="32">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E52FFE194AD4D11ED59ECD2" blockId="31.[151,1436,1778,2019]" lastBlockId="32.[151,1437,151,2013]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<emphasis id="B94AEA852E52FFDE94AD4D11EB09E849" bold="true" box="[199,558,1850,1875]" pageId="31" pageNumber="32">Additional material examined</emphasis>
[in addition to the specimens listed by
<bibRefCitation id="EFAF4B662E52FFDE97854D10EDB0E849" author="Kataev" box="[1007,1175,1851,1875]" pageId="31" pageNumber="32" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev (1997)</bibRefCitation>
].
<collectingCountry id="F32976072E52FFDE90C34D11EDD5E849" box="[1193,1266,1850,1875]" name="China" pageId="31" pageNumber="32">
<emphasis id="B94AEA852E52FFDE90C34D11EDD5E849" bold="true" box="[1193,1266,1850,1875]" pageId="31" pageNumber="32">China</emphasis>
</collectingCountry>
. SICHUAN:
<specimenCount id="9D38FD1E2E52FFDE91E44D10E989E86C" pageId="31" pageNumber="32" type="generic">1 ♂</specimenCount>
, &quot;Beipei near Chungking, 1956&quot;, Hwang Tien-yun leg. (
<collectionCode id="ED2FAE522E52FFDE97244D74EAA5E86D" box="[846,898,1887,1911]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="31" pageNumber="32">ZIN</collectionCode>
); 1 ♀, S of Ying-kuan-chiai,
<date id="FF8010572E52FFDE908F4D74EC57E86D" box="[1253,1392,1887,1911]" pageId="31" pageNumber="32" value="1929-07-14">14.VII.1929</date>
, H. Stevens leg. (
<collectionCode id="ED2FAE522E52FFDE95454DA8E8A1E881" LSID="urn:lsid:biocol.org:col:34795" box="[303,390,1923,1947]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="31" pageNumber="32">FMNH</collectionCode>
).
<collectionCode id="ED2FAE522E52FFDE95F04DA8E8D8E881" box="[410,511,1923,1947]" pageId="31" pageNumber="32">HUNAN</collectionCode>
: 2 f, Baojing Co.,
<quantity id="4CC69B722E52FFDE96BC4DA8EA07E880" box="[726,800,1923,1947]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="m" value="300.0">300 m</quantity>
,
<date id="FF8010572E52FFDE97474DA8EA93E881" box="[813,948,1923,1947]" pageId="31" pageNumber="32" value="1987-08-01">1.VIII.1987</date>
, Liao Subai leg. (
<collectionCode id="ED2FAE522E52FFDE90144DA8ED95E881" box="[1150,1202,1923,1947]" pageId="31" pageNumber="32">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E52FFDE90AD4DA8EDD3E880" box="[1223,1268,1923,1947]" pageId="31" pageNumber="32" type="generic">1 ♂</specimenCount>
, Jianghua Co.,
<date id="FF8010572E52FFDE94FD4D8CE9D1E8A5" box="[151,246,1959,1983]" pageId="31" pageNumber="32" value="1978-06">VI.1978</date>
, Peng Jianwen &amp; Yin Shicai leg. (
<collectionCode id="ED2FAE522E52FFDE96194D8CEB80E8A5" box="[627,679,1959,1983]" pageId="31" pageNumber="32">IOZ</collectionCode>
); ZHEJIANG:
<specimenCount id="9D38FD1E2E52FFDE973B4D8CEA5AE8A4" box="[849,893,1959,1983]" pageId="31" pageNumber="32" type="generic">1 ♂</specimenCount>
, Xian Ju Forest Station, 1977, Gao Zhaowei leg. (
<collectionCode id="ED2FAE522E52FFDE94F54DE0E9F4E8F9" box="[159,211,1995,2019]" pageId="31" pageNumber="32">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E52FFDE94834DE0E830E8F8" box="[233,279,1995,2019]" pageId="31" pageNumber="32" type="generic">1 ♂</specimenCount>
, &quot;Mokan Shan,
<date id="FF8010572E52FFDE95BB4DE1EB74E8F9" box="[465,595,1994,2019]" pageId="31" pageNumber="32" value="1936-06-20">20.VI.1936</date>
, O. Piel leg.&quot; (
<collectionCode id="ED2FAE522E52FFDE976A4DE0EA13E8F9" box="[768,820,1995,2019]" pageId="31" pageNumber="32">IOZ</collectionCode>
). GUIZHOU: 1 ♀, &quot;Kouy-Tcheou [= Guizhou], Abbe Largeteau&quot; (
<collectionCode id="ED2FAE522E6DFFE1954B4AB3E85AEFAA" LSID="urn:lsid:biocol.org:col:34988" box="[289,381,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="32" pageNumber="33">MNHN</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE195FA4AB3E89AEFB5" box="[400,445,152,176]" pageId="32" pageNumber="33" type="generic">3 ♂</specimenCount>
, 1 ♀, &quot;Kouy-Tcheou [=Guizhou], Reg. de Pin-Fa, Pere Cavalerie, 1910&quot; (
<collectionCode id="ED2FAE522E6DFFE190914AB3EC70EFAA" LSID="urn:lsid:biocol.org:col:34988" box="[1275,1367,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="32" pageNumber="33">MNHN</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE191014AB3ECB0EFB5" box="[1387,1431,152,176]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, 1 ♀, Xishui County, Dabaitang,
<quantity id="4CC69B722E6DFFE1966E4A97EB69EFCE" box="[516,590,188,213]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="32" pageNumber="33" unit="m" value="550.0">550 m</quantity>
,
<date id="FF8010572E6DFFE196304A97EBFBEFCE" box="[602,732,188,212]" pageId="32" pageNumber="33" value="2000-09-24">24.IX.2000</date>
, Liang H.B.,
<collectionCode id="ED2FAE522E6DFFE1971F4A97EA83EFCE" box="[885,932,188,212]" pageId="32" pageNumber="33">IOZ</collectionCode>
&amp; Guizhou Univ. Joint. Expedition (
<collectionCode id="ED2FAE522E6DFFE1912C4A97EC5DEFCE" box="[1350,1402,188,212]" pageId="32" pageNumber="33">IOZ</collectionCode>
); 1 ♀, Guiyang, Huaxi,
<date id="FF8010572E6DFFE195E14AF4EB2EEFE2" box="[395,521,223,248]" pageId="32" pageNumber="33" value="2007-04-15">15.IV.2007</date>
, Y. Liu leg. (
<collectionCode id="ED2FAE522E6DFFE196C54ACBEBC4EFE2" box="[687,739,224,248]" pageId="32" pageNumber="33">IOZ</collectionCode>
). GUANGXI:
<specimenCount id="9D38FD1E2E6DFFE197F84ACBEAE4EFED" box="[914,963,224,248]" pageId="32" pageNumber="33" type="generic">3 ♂</specimenCount>
, 1 ♀, Jinxiu,
<quantity id="4CC69B722E6DFFE190054ACBED9AEFED" box="[1135,1213,224,248]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="m" value="200.0">200 m</quantity>
,
<date id="FF8010572E6DFFE190A44AF4EC64EFE2" box="[1230,1347,223,248]" pageId="32" pageNumber="33" value="1999-05-15">15.V.1999</date>
, Zhang Xuezhong leg. (
<collectionCode id="ED2FAE522E6DFFE1953A4B2FE8A3EE06" box="[336,388,260,284]" pageId="32" pageNumber="33">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE195F14B2FE8EAEE06" box="[411,461,260,284]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
,1 ♀, Jinxiu,
<date id="FF8010572E6DFFE1960E4B2FEBBDEE07" box="[612,666,260,285]" pageId="32" pageNumber="33">15.X</date>
,1981 (
<collectionCode id="ED2FAE522E6DFFE196874B2FEA06EE06" box="[749,801,260,284]" pageId="32" pageNumber="33">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE197524B2FEA4FEE06" box="[824,872,260,284]" pageId="32" pageNumber="33" type="generic">3 ♂</specimenCount>
, Jinxiu,
<quantity id="4CC69B722E6DFFE197A74B2FED3EEE06" box="[973,1049,260,284]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="32" pageNumber="33" unit="m" value="900.0">900 m</quantity>
,
<date id="FF8010572E6DFFE190424B2FEDB9EE07" box="[1064,1182,260,285]" pageId="32" pageNumber="33" value="1999-05-16">16.V.1999</date>
, Gao Mingyuan &amp; Li Wenzhu leg. (
<collectionCode id="ED2FAE522E6DFFE195524B03E84BEE5A" box="[312,364,296,320]" pageId="32" pageNumber="33">IOZ</collectionCode>
). FUJIAN: 1 ♀, W Fujian, ca
<quantity id="4CC69B722E6DFFE196BF4B03EA36EE5A" box="[725,785,295,320]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="km" value="2.0">2 km</quantity>
SE Xinqiao,
<geoCoordinate id="EE0A50502E6DFFE197D84B0CED36EE5A" box="[946,1041,295,320]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="555" value="27.05">27.05°N</geoCoordinate>
<geoCoordinate id="EE0A50502E6DFFE190714B03ED5FEE5A" box="[1051,1144,296,320]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="5555" value="117.1">117.1°E</geoCoordinate>
,
<date id="FF8010572E6DFFE190EC4B0CEDDBEE5A" box="[1158,1276,295,320]" pageId="32" pageNumber="33" value="2005-05-10">10.V.2005</date>
, J. Turna leg. (cWR);
<specimenCount id="9D38FD1E2E6DFFE1949B4B67E83AEE7E" box="[241,285,332,356]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, 2 ♀, W Fujian, Taining E env., pitfall traps,
<geoCoordinate id="EE0A50502E6DFFE1977A4B67EA78EE7F" box="[784,863,332,357]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="26.15">26°9'N</geoCoordinate>
<geoCoordinate id="EE0A50502E6DFFE1970F4B67EAE9EE7F" box="[869,974,332,357]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="117.25">117°15'E</geoCoordinate>
,
<date id="FF8010572E6DFFE197B34B67ED4DEE7F" box="[985,1130,332,357]" pageId="32" pageNumber="33" value="2005-05-09" valueMax="2005-05-28" valueMin="2005-05-09">928.V.2005</date>
, J. Turna leg. (cZRS,
<collectionCode id="ED2FAE522E6DFFE191364B67ECA8EE7E" box="[1372,1423,332,356]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="32" pageNumber="33">ZIN</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE194FD4B5BE9E0EE9D" box="[151,199,368,392]" pageId="32" pageNumber="33" type="generic">2 ♂</specimenCount>
, 3 ♀, Sangang / light trap, 1982.
<date id="FF8010572E6DFFE1963C4B5BEBBFEE92" box="[598,664,368,392]" pageId="32" pageNumber="33" value="1920-09">IX.20</date>
, Liao Subai leg. [in
<taxonomicName id="4C3E4D142E6DFFE197F94B44EAD4EE92" box="[915,1011,367,392]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="32" pageNumber="37" rank="genus">Chinese</taxonomicName>
], &quot;
<taxonomicName id="4C3E4D142E6DFFE1907E4B5BECB0EE92" authority="Huang" authorityName="Huang" box="[1044,1431,368,392]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="chengjiangensis">
<emphasis id="B94AEA852E6DFFE1907E4B5BEC19EE92" box="[1044,1342,368,392]" italics="true" pageId="32" pageNumber="33">Harpalus chengjiangensis</emphasis>
Huang
</taxonomicName>
, 1993.6.23, det. Huang Tongling&quot; (
<collectionCode id="ED2FAE522E6DFFE196714BBFEB68EEB6" box="[539,591,404,428]" pageId="32" pageNumber="33">IOZ</collectionCode>
); 1 ♀, Sangang,
<quantity id="4CC69B722E6DFFE197674BBFEA77EEB6" box="[781,848,404,428]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="32" pageNumber="33" unit="m" value="740.0">740m</quantity>
,
<date id="FF8010572E6DFFE197364BBFEAFAEEB6" box="[860,989,404,428]" pageId="32" pageNumber="33" value="1982-09-17">17.IX.1982</date>
, Liao Subai [in
<taxonomicName id="4C3E4D142E6DFFE190FF4BBFEDD5EEB7" box="[1173,1266,404,429]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="32" pageNumber="37" rank="genus">Chinese</taxonomicName>
] (
<collectionCode id="ED2FAE522E6DFFE191624BBFEC1DEEB6" box="[1288,1338,404,428]" pageId="32" pageNumber="33">IOZ</collectionCode>
) [det. as
<taxonomicName id="4C3E4D142E6DFFE194FD4B92E848EECA" box="[151,367,440,464]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="chengjiangensis">
<emphasis id="B94AEA852E6DFFE194FD4B92E848EECA" box="[151,367,440,464]" italics="true" pageId="32" pageNumber="33">H. chengjiangensis</emphasis>
</taxonomicName>
by Huang Tongling];
<specimenCount id="9D38FD1E2E6DFFE196034B93EBB2EED5" box="[617,661,440,464]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, Sangang,
<quantity id="4CC69B722E6DFFE1977B4B93EA73EECA" box="[785,852,440,464]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="32" pageNumber="33" unit="m" value="740.0">740m</quantity>
,
<date id="FF8010572E6DFFE197354B93EAC6EECA" box="[863,993,440,464]" pageId="32" pageNumber="33" value="1982-09-18">18.IX.1982</date>
, Liao Subai [in
<taxonomicName id="4C3E4D142E6DFFE190FD4B9CEDD3EECA" box="[1175,1268,439,464]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="32" pageNumber="37" rank="genus">Chinese</taxonomicName>
] (
<collectionCode id="ED2FAE522E6DFFE191604B93EC1BEECA" box="[1290,1340,440,464]" pageId="32" pageNumber="33">IOZ</collectionCode>
) [det. as
<taxonomicName id="4C3E4D142E6DFFE194FD4BF6E855EEEE" box="[151,370,476,500]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="chengjiangensis">
<emphasis id="B94AEA852E6DFFE194FD4BF6E855EEEE" box="[151,370,476,500]" italics="true" pageId="32" pageNumber="33">H. chengjiangensis</emphasis>
</taxonomicName>
by Huang Tongling]; 1 ♀, Sangang (Wuyishan),
<date id="FF8010572E6DFFE197AA4BF7ED65EEEE" box="[960,1090,476,500]" pageId="32" pageNumber="33" value="1982-09-20">20.IX.1982</date>
, Liao Subai leg. (
<collectionCode id="ED2FAE522E6DFFE191704BF7EC69EEEE" box="[1306,1358,476,500]" pageId="32" pageNumber="33">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE1910C4BF7ECB0EEEE" box="[1382,1431,476,501]" pageId="32" pageNumber="33" type="generic">5 ♂</specimenCount>
, Sangang,
<date id="FF8010572E6DFFE1956F4BD4E8A0ED02" box="[261,391,511,536]" pageId="32" pageNumber="33" value="1982-09-16">16.IX.1982</date>
, Liao Subai leg. (
<collectionCode id="ED2FAE522E6DFFE19620482BEB59ED02" box="[586,638,512,536]" pageId="32" pageNumber="33">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE196FB482BEB9AED0D" box="[657,701,512,536]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, Sangang,
<date id="FF8010572E6DFFE1975D482BEA9EED02" box="[823,953,512,536]" pageId="32" pageNumber="33" value="1979-09-28">28.IX.1979</date>
, Xu Jianfei leg. (
<collectionCode id="ED2FAE522E6DFFE1901F482BED8EED02" box="[1141,1193,512,536]" pageId="32" pageNumber="33">IOZ</collectionCode>
); 1 ♀, Wuyishan,
<quantity id="4CC69B722E6DFFE19118482BE988ED26" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="32" pageNumber="33" unit="m" value="740.0">740 m</quantity>
,
<date id="FF8010572E6DFFE194D1480FE81BED27" box="[187,316,548,573]" pageId="32" pageNumber="33" value="1982-09-16">16.IX.1982</date>
, Zhao Angang leg. (
<collectionCode id="ED2FAE522E6DFFE1964B480FEB72ED26" box="[545,597,548,572]" pageId="32" pageNumber="33">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE19602480FEBB2ED26" box="[616,661,548,572]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, Sanming Co.,
<date id="FF8010572E6DFFE1972B480FEA9DED26" box="[833,954,548,572]" pageId="32" pageNumber="33" value="1981-10-17">17.X.1981</date>
, no collector (
<collectionCode id="ED2FAE522E6DFFE19033480FEDAAED26" box="[1113,1165,548,572]" pageId="32" pageNumber="33">IOZ</collectionCode>
).
<collectingCountry id="F32976072E6DFFE190CA480FEC2AED26" box="[1184,1293,548,572]" name="Taiwan" pageId="32" pageNumber="33">TAIWAN</collectingCountry>
:
<specimenCount id="9D38FD1E2E6DFFE19174480FEC61ED26" box="[1310,1350,548,572]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, Taipei county, Neitong Forest Recreation Area,
<quantity id="4CC69B722E6DFFE1960F486CEBB9ED7A" box="[613,670,583,608]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="32" pageNumber="33" unit="km" value="6.0">6 km</quantity>
S Wulai, swept from vegetation,
<date id="FF8010572E6DFFE190734863EDAEED7A" box="[1049,1161,584,608]" pageId="32" pageNumber="33" value="2002-04-07">7.IV.2002</date>
, Gy. Fábián &amp; O. Merkl leg. (
<collectionCode id="ED2FAE522E6DFFE194B84847E833ED9E" box="[210,276,620,644]" pageId="32" pageNumber="33">TMB</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE195414847E87DED9E" box="[299,346,620,644]" pageId="32" pageNumber="33" type="generic">2 ♂</specimenCount>
, 3 ♀, Taipei county, Haeng-Lu Dyi, around lights,
<date id="FF8010572E6DFFE197C54847ED6EED9E" box="[943,1097,620,644]" pageId="32" pageNumber="33" value="2002-04-02" valueMax="2002-04-21" valueMin="2002-04-02">221.IV.2002</date>
, Gy. Fábián &amp; O. Merkl leg. (
<collectionCode id="ED2FAE522E6DFFE194F548BBE9C6EDB2" box="[159,225,656,680]" pageId="32" pageNumber="33">TMB</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE1949F48BBE805EDBD" box="[245,290,656,680]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, Ilan county, Fu-Shan Botanical Garden,
<quantity id="4CC69B722E6DFFE1969C48BBEA67EDBD" box="[758,832,656,680]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="32" pageNumber="33" unit="m" value="700.0">700 m</quantity>
, at light,
<date id="FF8010572E6DFFE197C048A4ED70EDB2" box="[938,1111,655,680]" pageId="32" pageNumber="33" value="2000-09-25" valueMax="2000-09-27" valueMin="2000-09-25">2527.IX.2000</date>
, L. Papp, L. Peregovits &amp; L. Ronkay leg. (
<collectionCode id="ED2FAE522E6DFFE1955E489FE851EDD6" box="[308,374,692,716]" pageId="32" pageNumber="33">TMB</collectionCode>
); 1 ♀, Taipei Co., Beitou (NW Taipei City), &quot;Yangmingshan cemetetery&quot;,
<quantity id="4CC69B722E6DFFE190B1489FEC0FEDD6" box="[1243,1320,692,716]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="33" unit="m" value="300.0">300 m</quantity>
, decaying trunc of Pinus,
<date id="FF8010572E6DFFE1952948F3E89BEDEA" box="[323,444,728,752]" pageId="32" pageNumber="33" value="2007-10-22">22.X.2007</date>
, S. Vit leg. (cWR).
<collectingCountry id="F32976072E6DFFE196F148FCEBD8EDEA" box="[667,767,727,752]" name="Vietnam" pageId="32" pageNumber="33">
<emphasis id="B94AEA852E6DFFE196F148FCEBD8EDEA" bold="true" box="[667,767,727,752]" pageId="32" pageNumber="33">Vietnam</emphasis>
</collectingCountry>
.
<specimenCount id="9D38FD1E2E6DFFE1976548F3EA1BEDF5" box="[783,828,728,752]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, W Binly,
<quantity id="4CC69B722E6DFFE197DF48F3ED2AEDF5" box="[949,1037,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="33" unit="m" value="1200.0">1200 m</quantity>
,
<date id="FF8010572E6DFFE1907348FCEDA9EDEA" box="[1049,1166,727,752]" pageId="32" pageNumber="33" value="1963-05-28">28.V.1963</date>
, Kabakov leg. (cAK);
<specimenCount id="9D38FD1E2E6DFFE191E448F3E989EC0E" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, N
<collectingCountry id="F32976072E6DFFE194BD48D7E81EEC0F" box="[215,313,764,789]" name="Vietnam" pageId="32" pageNumber="33">Vietnam</collectingCountry>
, Lang Son Prov., Huu Lien,
<date id="FF8010572E6DFFE1961648D7EBFFEC0E" box="[636,728,764,788]" pageId="32" pageNumber="33" value="2010-04">IV.2010</date>
, Spiridonov leg. (
<collectionCode id="ED2FAE522E6DFFE197CB48D7EAF2EC0E" box="[929,981,764,788]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="32" pageNumber="33">ZIN</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE1978048D7ED30EC0E" box="[1002,1047,764,788]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, Lao Cai Prov., near Cat Cat Vill.,
<quantity id="4CC69B722E6DFFE194FD4934E9C2EC22" box="[151,229,799,824]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="32" pageNumber="33" unit="km" value="2.5">2.5 km</quantity>
SW of Sa Pa,
<geoCoordinate id="EE0A50502E6DFFE195ED490BEB2BEC22" box="[391,524,800,824]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="22.326946">22°19'37&quot;N</geoCoordinate>
<geoCoordinate id="EE0A50502E6DFFE19678490BEB84EC22" box="[530,675,800,824]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="103.8225">103°49'21&quot;E</geoCoordinate>
,
<quantity id="4CC69B722E6DFFE196C54934EA6BEC2D" box="[687,844,799,824]" metricMagnitude="3" metricUnit="m" metricValue="1.425" metricValueMax="1.45" metricValueMin="1.4" pageId="32" pageNumber="33" unit="m" value="1425.0" valueMax="1450.0" valueMin="1400.0">14001450 m</quantity>
,
<date id="FF8010572E6DFFE1973D490BEA8EEC22" box="[855,937,800,824]" pageId="32" pageNumber="33" value="2010-05">V.2010</date>
, A.V. Abramov leg. (
<collectionCode id="ED2FAE522E6DFFE190F2490BEDEBEC22" box="[1176,1228,800,824]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="32" pageNumber="33">ZIN</collectionCode>
); 1 ♀, Hoang Lien Son Prov., Sa Pa,
<date id="FF8010572E6DFFE19506496FEB2EEC47" box="[364,521,836,861]" pageId="32" pageNumber="33" value="1990-05-11" valueMax="1990-05-15" valueMin="1990-05-11">1115.V.1990</date>
, V. Kubaň leg. (
<collectionCode id="ED2FAE522E6DFFE196A1496FEA02EC46" box="[715,805,836,860]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="32" pageNumber="33">NHMB</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE19751496FEA4CEC46" box="[827,875,836,860]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, &quot;
<collectingCountry id="F32976072E6DFFE197E8496FEAF3EC47" box="[898,980,836,861]" name="Vietnam" pageId="32" pageNumber="33">Tonkin</collectingCountry>
sept., frontiere de
<collectingCountry id="F32976072E6DFFE190D9496FEDDFEC47" box="[1203,1272,836,861]" name="China" pageId="32" pageNumber="33">Chine</collectingCountry>
, ha-Giang, A. Weiss, 1901&quot; (
<collectionCode id="ED2FAE522E6DFFE195504943E8B1EC9A" LSID="urn:lsid:biocol.org:col:34988" box="[314,406,872,896]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="32" pageNumber="33">MNHN</collectionCode>
); 1 ♀,
<collectingCountry id="F32976072E6DFFE19588494CEB13EC9A" box="[482,564,871,896]" name="Vietnam" pageId="32" pageNumber="33">Tonkin</collectingCountry>
, Vinhphu Prov., Tamdao,
<date id="FF8010572E6DFFE1973D494CEAD3EC9A" box="[855,1012,871,896]" pageId="32" pageNumber="33" value="1985-06-02" valueMax="1985-06-11" valueMin="1985-06-02">211.VI.1985</date>
, Kuban leg. (
<collectionCode id="ED2FAE522E6DFFE190E04943EDC3EC9A" box="[1162,1252,872,896]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="32" pageNumber="33">NHMB</collectionCode>
).
<collectingCountry id="F32976072E6DFFE1909C494CEC08EC9A" box="[1270,1327,871,896]" name="Laos" pageId="32" pageNumber="33">
<emphasis id="B94AEA852E6DFFE1909C494CEC08EC9A" bold="true" box="[1270,1327,871,896]" pageId="32" pageNumber="33">Laos</emphasis>
</collectingCountry>
. 1 ♀, NE
<collectingCountry id="F32976072E6DFFE194FD49A6E9F7ECBE" box="[151,208,909,932]" name="Laos" pageId="32" pageNumber="33">Laos</collectingCountry>
, Pr. Hua Phan Ban Saleui, Phou Pan (Mt.). ca
<geoCoordinate id="EE0A50502E6DFFE1977849A7EA49ECBE" box="[786,878,908,932]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="20.2">20°12'N</geoCoordinate>
<geoCoordinate id="EE0A50502E6DFFE1971149A7EAC2ECBE" box="[891,997,908,932]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="104.01667">104°01'E</geoCoordinate>
,
<quantity id="4CC69B722E6DFFE1979D49A7EDBEECBE" box="[1015,1177,908,932]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="32" pageNumber="33" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">13001900 m</quantity>
,
<date id="FF8010572E6DFFE190C149A7EC56ECBF" box="[1195,1393,908,933]" pageId="32" pageNumber="33" value="2012-04-11" valueMax="2012-05-15" valueMin="2012-04-11">11.IV.15.V.2012</date>
, C. Holzschuh leg. (
<collectionCode id="ED2FAE522E6DFFE19524499BE8B5ECD2" box="[334,402,944,968]" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="32" pageNumber="33">NME</collectionCode>
);
<specimenCount id="9D38FD1E2E6DFFE195CF499BE8F6ECDD" box="[421,465,944,968]" pageId="32" pageNumber="33" type="generic">1 ♂</specimenCount>
, same as preceding, but
<date id="FF8010572E6DFFE1968E499BEA53ECD2" box="[740,884,944,968]" pageId="32" pageNumber="33" value="2011-05-01" valueMax="2011-05-31" valueMin="2011-05-01">131.V.2011</date>
, local collector (
<collectionCode id="ED2FAE522E6DFFE19044499BED55ECD2" box="[1070,1138,944,968]" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="32" pageNumber="33">NME</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E6DFFE194AD49FFEB5DEBDE" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8B8136972E6DFFE194AD49FFEB5DEBDE" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="33">
<emphasis id="B94AEA852E6DFFE194AD49FFE81DECF7" bold="true" box="[199,314,980,1005]" pageId="32" pageNumber="33">Diagnosis</emphasis>
. This species is somewhat similar to
<taxonomicName id="4C3E4D142E6DFFE1968E49FEEA7FECF6" box="[740,856,980,1004]" class="Insecta" family="Carabidae" genus="Pardileus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94AEA852E6DFFE1968E49FEEA7FECF6" box="[740,856,980,1004]" italics="true" pageId="32" pageNumber="33">H. indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3E4D142E6DFFE197FA49FEED1AECF6" box="[912,1085,980,1004]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hauserianus">
<emphasis id="B94AEA852E6DFFE197FA49FEED1AECF6" box="[912,1085,980,1004]" italics="true" pageId="32" pageNumber="33">H. hauserianus</emphasis>
</taxonomicName>
in having elytra with only two lateral intervals punctate and with microsculpture on disc in baso-medial portion consisting of narrow transverse meshes, but distinctly differs from two these species in having more prominent humeral denticle, visible dorsally, and in shape of pronotum with rather deep lateral depressions, separated from somewhat deep and narrow basal foveae by convex areas. Besides, one or two lateral intervals of elytra are very finely pubescent. Metafemur has three setigerous pores along posterior margin. The median lobe of aedeagus (
<figureCitation id="13052A122E6DFFE197844EACED4AEBBA" box="[1006,1133,1159,1184]" captionStart="FIGURES 82 85" captionStartId="33.[151,263,1137,1160]" captionTargetBox="[259,1328,342,1109]" captionTargetId="figure@33.[252,1334,339,1116]" captionTargetPageId="33" captionText="FIGURES 82 85. Harpalus, median lobe of aedeagus. 82 83, H. singularis (Sichuan, lectotype); 84 85, H. azumai (Shima Island). 82, 84, view from the left side; 83, 85, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287901/files/figure.png" pageId="32" pageNumber="33">Figs 8283</figureCitation>
) has two separate spines in the internal sac. Body length
<quantity id="4CC69B722E6DFFE195B54E87EB51EBDE" box="[479,630,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" metricValueMax="1.1199999999999999" metricValueMin="0.9199999999999999" pageId="32" pageNumber="33" unit="mm" value="10.2" valueMax="11.2" valueMin="9.2">9.211.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E6DFFE194AD4EE4EB5FEADA" pageId="32" pageNumber="33" type="distribution">
<paragraph id="8B8136972E6DFFE194AD4EE4EB28EA62" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="33">
<emphasis id="B94AEA852E6DFFE194AD4EE4E87EEBF2" bold="true" box="[199,345,1231,1256]" pageId="32" pageNumber="33">Distribution</emphasis>
.
<collectingCountry id="F32976072E6DFFE195004EE4E896EBF2" box="[362,433,1231,1256]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
: mainly in the central and southern parts. In
<collectingCountry id="F32976072E6DFFE197D54EE4ED22EBF2" box="[959,1029,1231,1256]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
, the species was previously known from Zhejiang, Hubei, Hunan, Sichuan, Guizhou, Guangdong, Guangxi, Yunnan and Fujian (
<bibRefCitation id="EFAF4B662E6DFFE190D54EDFECBFEA16" author="Kataev" box="[1215,1432,1268,1292]" pageId="32" pageNumber="33" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E6DFFE191734EDEEC73EA16" box="[1305,1364,1268,1292]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2003
</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E6DFFE194FD4F3CE85AEA2A" author="Liang" box="[151,381,1303,1328]" pageId="32" pageNumber="33" refString="Liang, H. - B. &amp; Liu, Y. (2007) Carabids of Leigongshan Reserve in Guizhou Province. In: Li, Z. Zh., Yang, M. - F. &amp; Jin, D. - Ch. (Eds.), Insects from Leigongshan Landscape. Guizhou Science and Technology Publishing House, Guiyang, pp. 236 - 247. [in Chinese with English summary]" type="book chapter" year="2007">Liang &amp; Liu 2007</bibRefCitation>
). We examined additionally specimens from
<collectingCountry id="F32976072E6DFFE197C84F3CEADFEA2A" box="[930,1016,1303,1328]" name="Taiwan" pageId="32" pageNumber="33">Taiwan</collectingCountry>
. The records by
<bibRefCitation id="EFAF4B662E6DFFE190A54F32EC70EA2A" author="Hua" box="[1231,1367,1304,1328]" pageId="32" pageNumber="33" refString="Hua, L. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp." type="book" year="2002">Hua (2002)</bibRefCitation>
from Heilongjiang, Jilin, Nei Mongol, Hebei, Henan, Shaanxi, Gansu, Anhui, Jiangsu, Jiangxi, Hunan and Xizang seem to be based on misidentifications.
</paragraph>
<paragraph id="8B8136972E6DFFE194AD4FAFEB5FEADA" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="33">
The species is also distributed in
<collectingCountry id="F32976072E6DFFE196544FAFEBBAEA87" box="[574,669,1412,1437]" name="Vietnam" pageId="32" pageNumber="33">Vietnam</collectingCountry>
(
<bibRefCitation id="EFAF4B662E6DFFE196C64FAFEA19EA86" author="Kataev" box="[684,830,1412,1436]" pageId="32" pageNumber="33" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
) and
<collectingCountry id="F32976072E6DFFE197144FAEEA9FEA86" box="[894,952,1413,1436]" name="Laos" pageId="32" pageNumber="33">Laos</collectingCountry>
. The species is reported from
<collectingCountry id="F32976072E6DFFE191664FAEEC65EA86" box="[1292,1346,1413,1436]" name="Laos" pageId="32" pageNumber="33">Laos</collectingCountry>
and the
<taxonomicName id="4C3E4D142E6DFFE194FD4F8CE9D6EADA" box="[151,241,1447,1472]" family="Nimaviridae" genus="Chinese" higherTaxonomySource="GBIF" kingdom="Viruses" pageId="32" pageNumber="37" rank="genus">Chinese</taxonomicName>
province
<collectingCountry id="F32976072E6DFFE1950B4F8CE894EADA" box="[353,435,1447,1472]" name="Taiwan" pageId="32" pageNumber="33">Taiwan</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E6DFFE194AD4FE7E87AE857" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8B8136972E6DFFE194AD4FE7E87AE857" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="33">
<emphasis id="B94AEA852E6DFFE194AD4FE7E813EAFF" bold="true" box="[199,308,1484,1509]" pageId="32" pageNumber="33">Remarks</emphasis>
. As stated by
<bibRefCitation id="EFAF4B662E6DFFE195884FE6EBAEEAFE" author="Kataev" box="[482,649,1484,1508]" pageId="32" pageNumber="33" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev (1997)</bibRefCitation>
,
<taxonomicName id="4C3E4D142E6DFFE196F34FE6EA08EAFE" box="[665,815,1484,1508]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B94AEA852E6DFFE196F34FE6EA08EAFE" box="[665,815,1484,1508]" italics="true" pageId="32" pageNumber="33">H. singularis</emphasis>
</taxonomicName>
is very similar and apparently closely related to two allopatric Japanese species:
<taxonomicName id="4C3E4D142E6DFFE195BE4FDAEBABE912" authority="Habu" authorityName="Habu" box="[468,652,1519,1544]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="azumai">
<emphasis id="B94AEA852E6DFFE195BE4FDAEB60E91D" box="[468,583,1520,1544]" italics="true" pageId="32" pageNumber="33">H. azumai</emphasis>
Habu
</taxonomicName>
from Ryukyu Islands and
<taxonomicName id="4C3E4D142E6DFFE197D24FDAEDF0E912" authority="Habu" authorityName="Habu" box="[952,1239,1519,1544]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="aogashimensis">
<emphasis id="B94AEA852E6DFFE197D24FDAEDA7E912" box="[952,1152,1520,1544]" italics="true" pageId="32" pageNumber="33">H. aogashimensis</emphasis>
(Habu)
</taxonomicName>
from Aogashima Island. All three species seem to form a monophyletic subgroup based on unique combination of characters: shape of pronotum with distinct obtuse basal angles, blunt at apex, microsculpture on elytral disc in baso-medial portion consisting of clearly transverse meshes, presence of three (rarely four) setigerous pores along posterior margin of metafemur and aedeagus with two large spines in internal sac.
<taxonomicName id="4C3E4D142E6DFFE1973A4CABED36E98D" box="[848,1041,1664,1688]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="azumai">
<emphasis id="B94AEA852E6DFFE1973A4CABED36E98D" box="[848,1041,1664,1688]" italics="true" pageId="32" pageNumber="33">Harpalus azumai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C3E4D142E6DFFE191604CAAECBBE982" box="[1290,1436,1664,1688]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B94AEA852E6DFFE191604CAAECBBE982" box="[1290,1436,1664,1688]" italics="true" pageId="32" pageNumber="33">H. singularis</emphasis>
</taxonomicName>
by elytra punctate on all intervals and pronotum with flatter latero-basal areas (its median lobe is illustrated in
<figureCitation id="13052A122E6DFFE1910F4C8FE9C5E9FA" captionStart="FIGURES 82 85" captionStartId="33.[151,263,1137,1160]" captionTargetBox="[259,1328,342,1109]" captionTargetId="figure@33.[252,1334,339,1116]" captionTargetPageId="33" captionText="FIGURES 82 85. Harpalus, median lobe of aedeagus. 82 83, H. singularis (Sichuan, lectotype); 84 85, H. azumai (Shima Island). 82, 84, view from the left side; 83, 85, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287901/files/figure.png" pageId="32" pageNumber="33">Figs. 8485</figureCitation>
).
<taxonomicName id="4C3E4D142E6DFFE1949E4CE3EB2EE9FA" box="[244,521,1736,1760]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="aogashimensis">
<emphasis id="B94AEA852E6DFFE1949E4CE3EB2EE9FA" box="[244,521,1736,1760]" italics="true" pageId="32" pageNumber="33">Harpalus aogashimensis</emphasis>
</taxonomicName>
, which is known to us only from Habus descriptions (
<bibRefCitation id="EFAF4B662E6DFFE190004CECEDCDE9FA" author="Habu" box="[1130,1258,1735,1760]" pageId="32" pageNumber="33" refString="Habu, A. (1957) On some species of the Carabidae from Aogashima taken by Miss. S. Kobayashi. Kontyu, 25, 66 - 72." type="journal article" year="1957">Habu 1957</bibRefCitation>
,
<bibRefCitation id="EFAF4B662E6DFFE1909F4CECEC17E9FA" author="Habu" box="[1269,1328,1735,1760]" pageId="32" pageNumber="33" refString="Habu, A. (1968) Revision of the Japanese species belonging to the subgenus-group Ophonus of the genus Harpalus (Coleoptera, Carabidae). Bulletin of the National Institute of Agricultural Sciences (Japan), Series C, 22, 283 - 327." type="journal article" year="1968">1968</bibRefCitation>
,
<bibRefCitation id="EFAF4B662E6DFFE191504CE3EC52E9FA" author="Habu" box="[1338,1397,1736,1760]" pageId="32" pageNumber="33" refString="Habu, A. (1973) Carabidae: Harpalini (Insecta, Coleoptera). Fauna Japonica. Keigaku Publishing Co., Tokio, xiii + 430 pp." type="book" year="1973">1973</bibRefCitation>
), is recognizable by pronotum with impunctate baso-medial area; like
<taxonomicName id="4C3E4D142E6DFFE197ED4CC6ED3CE81E" box="[903,1051,1772,1796]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B94AEA852E6DFFE197ED4CC6ED3CE81E" box="[903,1051,1772,1796]" italics="true" pageId="32" pageNumber="33">H. singularis</emphasis>
</taxonomicName>
, its elytra are impunctate on disc. The records of
<taxonomicName id="4C3E4D142E6DFFE195334D3AEB0EE832" box="[345,553,1808,1832]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="aogashimensis">
<emphasis id="B94AEA852E6DFFE195334D3AEB0EE832" box="[345,553,1808,1832]" italics="true" pageId="32" pageNumber="33">H. aogashimensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F32976072E6DFFE196174D24EBE7E832" box="[637,704,1807,1832]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
(Guangdong and Sichuan) (
<bibRefCitation id="EFAF4B662E6DFFE190764D3BEDB1E832" author="Hua" box="[1052,1174,1808,1832]" pageId="32" pageNumber="33" refString="Hua, L. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp." type="book" year="2002">Hua 2002</bibRefCitation>
) are surely based on misidentification.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E6DFFE094AD4D73E9F0EE07" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8B8136972E6DFFE094AD4D73E9F0EE07" blockId="32.[151,1437,151,2013]" lastBlockId="33.[151,1436,151,285]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<taxonomicName id="4C3E4D142E6DFFE194AD4D73E8D6E86A" box="[199,497,1880,1904]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="chengjiangensis">
<emphasis id="B94AEA852E6DFFE194AD4D73E8D6E86A" box="[199,497,1880,1904]" italics="true" pageId="32" pageNumber="33">Harpalus chengjiangensis</emphasis>
</taxonomicName>
was described from two specimens (male and female) collected in Chengjiang, Beibei, Chongqing,
<collectingCountry id="F32976072E6DFFE195104D57E8E6E88F" box="[378,449,1916,1941]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
. According to the original description (
<bibRefCitation id="EFAF4B662E6DFFE197144D57ED2AE88E" author="Huang" box="[894,1037,1916,1940]" pageId="32" pageNumber="33" refString="Huang, T. (1993) Three new species of the genus Harpalus from China (Coleoptera: Carabidae). Acta Zootaxonomica Sinica, 18 (4), 451 - 455." type="journal article" year="1993">Huang 1993</bibRefCitation>
), this species is &quot;close to
<taxonomicName id="4C3E4D142E6DFFE191584D56ECB2E88E" box="[1330,1429,1916,1940]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B94AEA852E6DFFE191584D56ECB2E88E" box="[1330,1429,1916,1940]" italics="true" pageId="32" pageNumber="33">H. babai</emphasis>
</taxonomicName>
, but differs in dorsal surface of head glabrous, interval 7 without additional pores, hind femur with 3 setae near hind margin, sternite 5 glabrous&quot;. We had no opportunity to study the
<typeStatus id="548588352E6DFFE197EF4DEEEA92E8C7" box="[901,949,1989,2013]" pageId="32" pageNumber="33">type</typeStatus>
specimens, but several males and females examined by us from Fujian (see material above), which were determined by Huang Tongling as
<taxonomicName id="4C3E4D142E6CFFE091EB4AB2E86CEFCE" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="chengjiangensis">
<emphasis id="B94AEA852E6CFFE091EB4AB2E86CEFCE" italics="true" pageId="33" pageNumber="34">H. chengjiangensis</emphasis>
</taxonomicName>
, belong to
<taxonomicName id="4C3E4D142E6CFFE0958E4A96EB58EFCE" box="[484,639,188,212]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B94AEA852E6CFFE0958E4A96EB58EFCE" box="[484,639,188,212]" italics="true" pageId="33" pageNumber="34">H. singularis</emphasis>
</taxonomicName>
. Because all the distinctive characters mentioned in the original description of
<taxonomicName id="4C3E4D142E6CFFE095514ACAEB33EFE2" box="[315,532,224,248]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="chengjiangensis">
<emphasis id="B94AEA852E6CFFE095514ACAEB33EFE2" box="[315,532,224,248]" italics="true" pageId="33" pageNumber="34">H. chengjiangensis</emphasis>
</taxonomicName>
fit those of
<taxonomicName id="4C3E4D142E6CFFE096CA4ACAEA13EFE2" box="[672,820,224,248]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B94AEA852E6CFFE096CA4ACAEA13EFE2" box="[672,820,224,248]" italics="true" pageId="33" pageNumber="34">H. singularis</emphasis>
</taxonomicName>
, we treat the former name as a junior synonym of the latter.
</paragraph>
</subSubSection>
</treatment>
</document>