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<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.3.37464" ID-GBIF-Dataset="8ac0c1d8-2ebc-40bd-bcf6-7216ce5d8d66" ID-GBIF-Taxon="160689241" ID-Pensoft-Pub="2535-0730-2-109" ID-Pensoft-UUID="48BB2ADCDFB750ACA7D9EC306EC80801" ID-ZooBank="87602625AF8D4A3FBAE5F35C09FB6C00" ModsDocID="2535-0730-2-109" checkinTime="1575997079287" checkinUser="pensoft" docAuthor="Monod, Lionel, Duperre, Nadine &amp; Harms, Danilo" docDate="2019" docId="67AEC7BDF56B59658EE548A9C7606EF5" docLanguage="en" docName="EvolutSyst 3(2): 109-200" docOrigin="Evolutionary Systematics 3 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.3.37464" docTitle="Cercophonius granulosus" docType="treatment" docVersion="4" id="48BB2ADCDFB750ACA7D9EC306EC80801" lastPageNumber="109" masterDocId="48BB2ADCDFB750ACA7D9EC306EC80801" masterDocTitle="An annotated catalogue of the scorpion types (Arachnida, Scorpiones) held in the Zoological Museum Hamburg. Part I: Parvorder Iurida Soleglad &amp; Fet, 2003" masterLastPageNumber="200" masterPageNumber="109" pageId="0" pageNumber="109" updateTime="1643462502119" updateUser="ExternalLinkService">
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<mods:title>An annotated catalogue of the scorpion types (Arachnida, Scorpiones) held in the Zoological Museum Hamburg. Part I: Parvorder Iurida Soleglad &amp; Fet, 2003</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Monod, Lionel</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Duperre, Nadine</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Harms, Danilo</mods:namePart>
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<mods:title>Evolutionary Systematics</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>3</mods:number>
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<mods:number>2</mods:number>
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<mods:start>109</mods:start>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.3.37464</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.3.37464</mods:identifier>
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<treatment ID-GBIF-Taxon="160689241" LSID="urn:lsid:plazi:treatment:67AEC7BDF56B59658EE548A9C7606EF5" httpUri="http://treatment.plazi.org/id/67AEC7BDF56B59658EE548A9C7606EF5" lastPageNumber="109" pageId="0" pageNumber="109">
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<taxonomicName LSID="67AEC7BD-F56B-5965-8EE5-48A9C7606EF5" class="Arachnida" family="Bothriuridae" genus="Cercophonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercophonius granulosus" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="granulosus">Cercophonius granulosus</taxonomicName>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cercophonius granulosus Kraepelin, 1908, female lectotype, habitus A dorsal aspect B ventral aspect. Scale bars: 10 mm." httpUri="https://binary.pensoft.net/fig/355022" pageId="0" pageNumber="109">Fig. 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="reference_group">
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Arachnida" family="Bothriuridae" genus="Cercophonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercophonius granulosus" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="granulosus">Cercophonius granulosus</taxonomicName>
Kraepelin, 1908c: 102-103
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="current combination">
<paragraph pageId="0" pageNumber="109">Current combination.</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Arachnida" family="Bothriuridae" genus="Cercophonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercophonius granulosus" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis italics="true" pageId="0" pageNumber="109">Cercophonius granulosus</emphasis>
</taxonomicName>
Kraepelin, 1908
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="lectotype">
<paragraph pageId="0" pageNumber="109">Lectotype.</paragraph>
<paragraph pageId="0" pageNumber="109">
♀ (
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Bothriurus titschacki Werner, 1939 [= Centromachetes titschacki (Werner, 1939)], male holotype (A-C), female paratype (D-F): A, D dorsal aspect of habitus B, E ventral aspect of habitus C, F pro-lateral aspect of chela. Scale bars: 10 mm (A-B, D-E), 0.5 mm (C, F)." httpUri="https://binary.pensoft.net/fig/355021" pageId="0" pageNumber="109">
Fig. 9
<normalizedToken originalValue="AB">A-B</normalizedToken>
</figureCitation>
, ZMH-A0000909), [Western Australia], Moonyoonooka (station 82) [
<geoCoordinate degrees="28" direction="south" minutes="46" orientation="latitude" precision="15" seconds="51" value="-28.780832">
28°46
<normalizedToken originalValue="51”">'51&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate degrees="114" direction="east" minutes="43" orientation="longitude" precision="15" seconds="37" value="114.726944">
114°43
<normalizedToken originalValue="37”">'37&quot;</normalizedToken>
E
</geoCoordinate>
], 13.07.1905, Wilhelm Michaelsen leg.,
<normalizedToken originalValue="Hamburgs">Hamburg's</normalizedToken>
South-West Australia Expedition 1905, ded. 06.1908.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="paralectotype">
<paragraph pageId="0" pageNumber="109">Paralectotype.</paragraph>
<paragraph pageId="0" pageNumber="109">
♀ (ZMH-A0002236), Western Australia, Geraldton (station 75) [
<geoCoordinate degrees="28" direction="south" minutes="46" orientation="latitude" precision="15" seconds="44" value="-28.778889">
28°46
<normalizedToken originalValue="44”">'44&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate degrees="114" direction="east" minutes="36" orientation="longitude" precision="15" seconds="52" value="114.61444">
114°36
<normalizedToken originalValue="52”">'52&quot;</normalizedToken>
E
</geoCoordinate>
], 16.07.1905, Wilhelm Michaelsen leg.,
<normalizedToken originalValue="Hamburgs">Hamburg's</normalizedToken>
South-West Australia Expedition 1905, ded. 06.1908.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="remarks">
<paragraph pageId="0" pageNumber="109">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="109">
<bibRefCitation author="Kraepelin, K" editor="Michaelsen, W" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pageId="0" pageNumber="109" refId="B78" refString="Kraepelin, K, Michaelsen, W, Hartmeyer, R, 1908c. Scorpiones. In: MichaelsenWHartmeyerR (Eds) Die Fauna Suedwest-Australiens, Ergebnisse der Hamburger Suedwest-Australischen Forschungsreise 1905.[Vol. 2.] Verlag Gustav Fischer, Jena, 87-104." title="Scorpiones. In: MichaelsenWHartmeyerR (Eds) Die Fauna Suedwest-Australiens, Ergebnisse der Hamburger Suedwest-Australischen Forschungsreise 1905. [Vol. 2.] Verlag Gustav Fischer, Jena, 87 - 104." year="1908 c">Kraepelin (1908c)</bibRefCitation>
listed 2 females from Moonyoonooka and 1 juvenile from Geraldton.
<bibRefCitation author="Acosta, LE" journalOrPublisher="Boletin de la Sociedad de Biologia de Concepcion" pageId="0" pageNumber="109" pagination="7 - 27" refId="B1" refString="Acosta, LE, 1990. El genero Cercophonius Peters, 1861 (Scorpiones, Bothriuridae). Boletin de la Sociedad de Biologia de Concepcion 61: 7 - 27" title="El genero Cercophonius Peters, 1861 (Scorpiones, Bothriuridae)." volume="61" year="1990">Acosta (1990)</bibRefCitation>
noted that the material was only composed of 1 female from Moonyoonooka and 1 female from Geraldton, with no trace of a juvenile specimen. The female from Moonyoonooka erroneously considered as holotype by
<bibRefCitation author="Weidner, F" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="0" pageNumber="109" pagination="89 - 142" refId="B175" refString="Weidner, F, 1959. Die Entomologischen Sammlungen des Zoologischen Staatsinstituts und Zoologischen Museums Hamburg, I, Teil, Parathropoda und Chelicerata I. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 54: 89 - 142" title="Die Entomologischen Sammlungen des Zoologischen Staatsinstituts und Zoologischen Museums Hamburg, I, Teil, Parathropoda und Chelicerata I." volume="54" year="1959">Weidner (1959)</bibRefCitation>
was designated as lectotype and the remaining specimen as paralectotype by
<bibRefCitation author="Acosta, LE" journalOrPublisher="Boletin de la Sociedad de Biologia de Concepcion" pageId="0" pageNumber="109" pagination="7 - 27" refId="B1" refString="Acosta, LE, 1990. El genero Cercophonius Peters, 1861 (Scorpiones, Bothriuridae). Boletin de la Sociedad de Biologia de Concepcion 61: 7 - 27" title="El genero Cercophonius Peters, 1861 (Scorpiones, Bothriuridae)." volume="61" year="1990">Acosta (1990)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="remarks">
<paragraph pageId="0" pageNumber="109">Remarks on collector.</paragraph>
<paragraph pageId="0" pageNumber="109">
J. Wilhelm Michaelsen (1860-1937) was a curator at the ZMH and a specialist of
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Oligochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oligochaeta" order="Asterales" pageId="0" pageNumber="109" phylum="Tracheophyta" rank="subClass" subClass="Oligochaeta">Oligochaeta</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1038/140308a0" author="Monro, CCA" journalOrPublisher="Nature" pageId="0" pageNumber="109" pagination="308 - 309" refId="B120" refString="Monro, CCA, 1937. Prof. J. W. Michaelsen. Nature 140: 308 - 309" title="Prof. J. W. Michaelsen." url="https://doi.org/10.1038/140308a0" volume="140" year="1937">Monro 1937</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1080/09397140.2012.10649001" author="Sherlock, E" journalOrPublisher="Zoology in the Middle East, Supplementum" pageId="0" pageNumber="109" pagination="177 - 187" refId="B156" refString="Sherlock, E, Berridge, L, 2012. History of the earthworm collections at the Natural History Museum, London (Oligochaeta). Advances of the 5th International Oligochaeta Taxonomy Meeting. Zoology in the Middle East, Supplementum 4: 177 - 187" title="History of the earthworm collections at the Natural History Museum, London (Oligochaeta). Advances of the 5 th International Oligochaeta Taxonomy Meeting." url="https://doi.org/10.1080/09397140.2012.10649001" volume="4" year="2012">Sherlock and Berridge 2012</bibRefCitation>
). He undertook three major expeditions to Chile, South Africa and Western Australia (see
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4375.1.1" author="Harms, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="109" pagination="1 - 58" refId="B33" refString="Harms, D, Duperre, N, 2018. An annotated type catalogue of the camel spiders (Arachnida: Solifugae) held in the Zoological Museum Hamburg. Zootaxa 4375: 1 - 58" title="An annotated type catalogue of the camel spiders (Arachnida: Solifugae) held in the Zoological Museum Hamburg." url="https://doi.org/10.11646/zootaxa.4375.1.1" volume="4375" year="2018">
Harms and
<normalizedToken originalValue="Dupérré">Duperre</normalizedToken>
2018
</bibRefCitation>
for details). In 1905, the Berlin and Hamburg Museums organized a major expedition to southwestern Australia that was led by Michaelsen and Robert Hartmeyer (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.7416" author="Michaelsen, W" editor="Michaelsen, W" journalOrPublisher="Verlag von Gustav Fischer, Jena" pageId="0" pageNumber="109" refId="B118" refString="Michaelsen, W, Hartmeyer, R, , 1907. Reisebericht. In: Michaelsen, W, Hartmeyer, R, Eds., Die Fauna Suedwest-Australiens. Ergebnisse der Hamburger Suedwest-Australischen Forschungsreise 1905, 1(1). . Verlag von Gustav Fischer, Jena" title="Reisebericht" url="https://doi.org/10.5962/bhl.title.7416" volumeTitle="Die Fauna Suedwest-Australiens. Ergebnisse der Hamburger Suedwest-Australischen Forschungsreise 1905, 1 (1)." year="1907">Michaelsen and Hartmeyer 1907</bibRefCitation>
[see travel map for detail of collecting localities]). Numerous spiders (
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Oligochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oligochaeta" order="Araneae" pageId="0" pageNumber="109" phylum="Tracheophyta" rank="order">Araneae</taxonomicName>
) species were described by
<bibRefCitation author="Simon, E" editor="Michaelsen, W" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="109" refId="B168" refString="Simon, E, Michaelsen, W, Hartmeyer, R, 1908. Araneae, Iere partie. In: MichaelsenWHartmeyerR (Eds) Die Fauna Sudwest-Australiens, 1.Verlag von Gustav Fischer, Jena, 359-446." title="Araneae, Iere partie. In: MichaelsenWHartmeyerR (Eds) Die Fauna Sudwest-Australiens, 1. Verlag von Gustav Fischer, Jena, 359 - 446." year="1908">Simon (1908</bibRefCitation>
,
<bibRefCitation author="Simon, E" editor="Michaelsen, W" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="109" refId="B169" refString="Simon, E, Michaelsen, W, Hartmeyer, R, 1909. Spiders, 2eme partie. In: MichaelsenWHartmeyerR (Eds) Die Fauna Sudwest-Australiens, 2.Verlag von Gustav Fischer, Jena, 155-212." title="Spiders, 2 eme partie. In: MichaelsenWHartmeyerR (Eds) Die Fauna Sudwest-Australiens, 2. Verlag von Gustav Fischer, Jena, 155 - 212." year="1909">1909</bibRefCitation>
) based on the material collected during this expedition. The type specimens of these species are lodged in the Western Australian Museum (
<bibRefCitation author="Main, BY" journalOrPublisher="Record of the Western Australian Museum" pageId="0" pageNumber="109" pagination="703 - 712" refId="B112" refString="Main, BY, Harvey, MS, 1992. Type material of spiders (Chelicerata: Araneae) collected on the Michaelsen and Hartmeyer Expedition of 1905 and lodged in the Western Australian Museum. Record of the Western Australian Museum 15: 703 - 712" title="Type material of spiders (Chelicerata: Araneae) collected on the Michaelsen and Hartmeyer Expedition of 1905 and lodged in the Western Australian Museum." volume="15" year="1992">Main and Harvey 1992</bibRefCitation>
). Interestingly, the type specimens of Australian scorpion species described by Kraepelin based on material collected during the same expedition were not sent back to Australia but remained in the collections of the Hamburg Museum.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/355022" pageId="0" pageNumber="109" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 10.</emphasis>
<taxonomicName class="Arachnida" family="Bothriuridae" genus="Cercophonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercophonius granulosus" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis italics="true" pageId="0" pageNumber="109">Cercophonius granulosus</emphasis>
</taxonomicName>
Kraepelin, 1908, female lectotype, habitus
<emphasis bold="true" pageId="0" pageNumber="109">A</emphasis>
dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="109">B</emphasis>
ventral aspect. Scale bars: 10 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>